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<document ID-CLB-Dataset="294695" ID-DOI="http://dx.doi.org/10.3897/zookeys.1198.119152" ID-GBIF-Dataset="fe1e56c4-cdbb-410d-bb4b-5efc6ba075a7" ID-Pensoft-Pub="1313-2970-1198-173" ID-Pensoft-UUID="2C1DBEEC3E0156D5AB788A2E51505363" ID-ZooBank="C840CCE9A6974AEB85D86E3DD33B1D7F" ModsDocID="1313-2970-1198-173" checkinTime="1713961486013" checkinUser="pensoft" docAuthor="Hlavac, Peter, Banar, Petr &amp; Stoces, Dominik" docDate="2024" docId="1FC3B260550E555CAAA06144B160A53E" docLanguage="en" docName="ZooKeys 1198: 173-184" docOrigin="ZooKeys 1198" docPubDate="2024-04-24" docSource="http://dx.doi.org/10.3897/zookeys.1198.119152" docTitle="Commatocerus concinnus Besuchet &amp; Cuccodoro 2011" docType="treatment" docVersion="2" id="2C1DBEEC3E0156D5AB788A2E51505363" lastPageNumber="173" masterDocId="2C1DBEEC3E0156D5AB788A2E51505363" masterDocTitle="Clavigeritae (Coleoptera, Staphylinidae, Pselaphinae) of the Arabian Peninsula with the description of a new species of Corynotopsis Jeannel, 1951 from Oman" masterLastPageNumber="184" masterPageNumber="173" pageNumber="173" updateTime="1713962098711" updateUser="ExternalLinkService">
<mods:mods id="8FF714100B6DFD60330855457E8A2D7B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7726D424D425AA1E958ACD865F66CF1A">
<mods:title id="972F90300BAE1A9A085302FF04948654">Clavigeritae (Coleoptera, Staphylinidae, Pselaphinae) of the Arabian Peninsula with the description of a new species of Corynotopsis Jeannel, 1951 from Oman</mods:title>
</mods:titleInfo>
<mods:name id="2E96F0AD0BEE6F1A245C47ADBFE03F7B" type="personal">
<mods:role id="7A491D9D8C47049BC6AF07BA8943604C">
<mods:roleTerm id="052FBA09F127CF626438EEE4FA608B5D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4D6B3D26DABFF13A2F22B62528860843">Hlavac, Peter</mods:namePart>
<mods:affiliation id="0A7332A717C0B256A8EB00BC9F4E1E22">Department of Zoology, Fisheries, Hydrobiology and Apiculture, Faculty of AgriSciences, Mendel University, Zemedelska 1, Brno, CZ- 613 00, Czech Republic</mods:affiliation>
</mods:name>
<mods:name id="E165E0F30D4A1B27A2B9B8E82E2CF98C" type="personal">
<mods:role id="DF36312ED99E2CAC9BBF4DF9737729D7">
<mods:roleTerm id="A2A07A67F3191C0DE3990E8B0FC55D5D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B9E9FE45C9900581E4FF9B0534ACC741">Banar, Petr</mods:namePart>
<mods:affiliation id="5AF77B0FFC2E41D9833915B8D23984B5">Department of Zoology, Fisheries, Hydrobiology and Apiculture, Faculty of AgriSciences, Mendel University, Zemedelska 1, Brno, CZ- 613 00, Czech Republic</mods:affiliation>
<mods:nameIdentifier id="463D33EC0A42E92283224738FEA46AD9" type="email">petrbanar@seznam.cz</mods:nameIdentifier>
</mods:name>
<mods:name id="577335A801BB14438EBD66C8542EE1B1" type="personal">
<mods:role id="01356B6F923C1E89809697F6E91CD3C0">
<mods:roleTerm id="B2EA4C3B17FC7F7F4A7877B436324557">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F29686EBA5E2D48CF93544A1B843D99F">Stoces, Dominik</mods:namePart>
<mods:nameIdentifier id="C6801847D297592A0D8C4743599C66D9" type="ORCID">https://orcid.org/0000-0002-1867-5327</mods:nameIdentifier>
<mods:affiliation id="E4BAA35F10745BC2A05B205CB866BE84">Department of Zoology, Fisheries, Hydrobiology and Apiculture, Faculty of AgriSciences, Mendel University, Zemedelska 1, Brno, CZ- 613 00, Czech Republic</mods:affiliation>
</mods:name>
<mods:typeOfResource id="010EBA06A486346A8EB88CA52F47F942">text</mods:typeOfResource>
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<mods:title id="CD5E6A54D0E98536AE2F2FBE82523058">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="37B9763E3ED4FAF930FAD32F10A1F7AB">
<mods:date id="5978F44746FDD048C4D831F658121155">2024</mods:date>
<mods:detail id="8531A3341E086E1C948C4941204E66B8" type="pubDate">
<mods:number id="85EF0E510F84FAE98AA774B76CF488D5">2024-04-24</mods:number>
</mods:detail>
<mods:detail id="AB67CAE8D29C95217CBA6C6445F12D5A" type="volume">
<mods:number id="FD34459702D67D7600334A6FD54060F4">1198</mods:number>
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<mods:start id="91833AD5F994D026A78F6650D7FF244F">173</mods:start>
<mods:end id="135E6B3E30AB66F850B5C2639F0875D5">184</mods:end>
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<mods:location id="85709C853AC969E3EAD882DEB57E1788">
<mods:url id="9C83164DCE5DD228E6C6C8EFFC2546BC">http://dx.doi.org/10.3897/zookeys.1198.119152</mods:url>
</mods:location>
<mods:classification id="5ADE3195F10553C88B6E0E58611CDEBE">journal article</mods:classification>
<mods:identifier id="835249A5C661FE40D0AA83F4CC63D187" type="DOI">http://dx.doi.org/10.3897/zookeys.1198.119152</mods:identifier>
<mods:identifier id="895841B56F8FE65E8ACBF9EA875AB400" type="Pensoft-Pub">1313-2970-1198-173</mods:identifier>
<mods:identifier id="9722C37F83FEF8F9EBE23F5D4106704B" type="ZooBank">C840CCE9A6974AEB85D86E3DD33B1D7F</mods:identifier>
<mods:identifier id="BCCEE836508B85863E0508C0793253A7" type="Pensoft-UUID">2C1DBEEC3E0156D5AB788A2E51505363</mods:identifier>
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<subSection id="4943C75CBC26FD3C0CE0D08147FB2F5F" pageId="0" pageNumber="173" type="taxonomy">
<treatment id="1FC3B260550E555CAAA06144B160A53E" LSID="urn:lsid:plazi:treatment:1FC3B260550E555CAAA06144B160A53E" httpUri="http://treatment.plazi.org/id/1FC3B260550E555CAAA06144B160A53E" lastPageNumber="173" pageId="0" pageNumber="173">
<subSubSection id="37BD94AB1E6AFBEDCF3B7987B7B4CFE4" pageId="0" pageNumber="173" type="nomenclature">
<paragraph id="9F8F837634463CA568541D7B4389DAAC" pageId="0" pageNumber="173">
<taxonomicName id="D1636C8504F8723E6465777A5B26C890" ID-CoL="XFCL" LSID="1FC3B260-550E-555C-AAA0-6144B160A53E" authority="Besuchet &amp; Cuccodoro, 2011" authorityName="Besuchet &amp; Cuccodoro" authorityYear="2011" class="Insecta" family="Staphylinidae" genus="Commatocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Commatocerus concinnus" order="Coleoptera" pageId="0" pageNumber="173" phylum="Arthropoda" rank="species" species="concinnus">Commatocerus concinnus Besuchet &amp; Cuccodoro, 2011</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="02458007D36EBC6201220D63FE7F6733" pageId="0" pageNumber="173" type="reference_group">
<paragraph id="EC401733F757CD8CF9FC42D8D1B1CCDD" pageId="0" pageNumber="173">
<taxonomicName id="2D785CBE64654305177BDCF4FC65FFD8" authorityName="Besuchet &amp; Cuccodoro" authorityYear="2011" class="Insecta" family="Staphylinidae" genus="Commatocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Commatocerus concinnus" order="Coleoptera" pageId="0" pageNumber="173" phylum="Arthropoda" rank="species" species="concinnus">Commatocerus concinnus</taxonomicName>
Besuchet &amp; Cuccodoro, 2011: 165, plate 14 (habitus), figs 22-24 (aedeagus); type locality: United Arab Emirates, Ras al Khaimah, Wadi Shawkah,
<geoCoordinate id="9DB550AB3BD20AF49ABC5F8F4C1E4406" degrees="25" direction="north" minutes="6" orientation="latitude" precision="1" seconds="15.30" value="25.10425">25°6'15.30&quot;N</geoCoordinate>
,
<geoCoordinate id="E47FBCF9D9B08E41DC07D46729C9D22D" degrees="56" direction="east" minutes="2" orientation="longitude" precision="1" seconds="47.55" value="56.04654">56°2'47.55&quot;E</geoCoordinate>
.
</paragraph>
</subSubSection>
<subSubSection id="B3E99F1D29213E0CFC691AE422D86499" pageId="0" pageNumber="173" type="materials_examined">
<paragraph id="11014E3869F912AAA2D68CB03B70C78B" pageId="0" pageNumber="173">Material examined.</paragraph>
<paragraph id="1D212B96CE40A31AF9A79C7B01336527" pageId="0" pageNumber="173">
<materialsCitation id="7951494CD3958428955FC327201B1612" collectingDate="2019-03-29" collectorName="Lgt. T. Kopecky" country="Oman" county="Subayb" determinerName="P. Hlavac" elevation="1442" latitude="23.235546" location="Oman" longLatPrecision="1" longitude="57.14944" municipality="Ad" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="DA61D2BF309F1A927E414457A2AB147D" count="1" type="male">1 ♂</specimenCount>
,
<specimenCount id="F7A84292E11EF17457B44D9165E117BA" count="1" type="female">1 ♀</specimenCount>
,
<emphasis id="7CB039A3D55C50548FB7FC798ADFD9D2" bold="true" pageId="0" pageNumber="173">
<collectingCountry id="46550954C385B022B241B964488DD776" name="Oman">Oman</collectingCountry>
</emphasis>
: two labels &quot;
<location id="CD3E264D83781771131CC74FBB69C852" LSID="urn:lsid:plazi:treatment:1FC3B260550E555CAAA06144B160A53E:CD3E264D83781771131CC74FBB69C852" country="Oman" county="Subayb" latitude="23.235546" longLatPrecision="1" longitude="57.14944" municipality="Ad" name="Oman">Oman</location>
, 28-
<collectingDate id="5916BB83390012384D25AB814670B698" value="2019-03-29">29.3.2019</collectingDate>
,
<collectingMunicipality id="5CF734C2EC75F6C0A8F37FDA1B30BA86">Ad</collectingMunicipality>
/ Dakhilyiah Gov., near
<collectingCounty id="7FAC1FF2325D9930C1D4DB107DF22F39">Subayb</collectingCounty>
/
<geoCoordinate id="19F54EDDDA4017D5387044F605636018" degrees="23" direction="north" minutes="14" orientation="latitude" precision="1" seconds="7.966" value="23.235546">23°14'7.966&quot;N</geoCoordinate>
,
<geoCoordinate id="8F736BFC76B64A5718E49DB618A677D5" degrees="57" direction="east" minutes="8" orientation="longitude" precision="1" seconds="57.977966" value="57.14944">57°8'57.977966&quot;E</geoCoordinate>
, / wadi,
<quantity id="B24FCDC2A8E4CBCD045A63DBE723A824" metricMagnitude="3" metricUnit="m" metricValue="1.4419999999999997" unit="m" value="1442.0">
<elevation id="A9CB3DAFCDB11C24EB44B9454D7AFF4B" metricMagnitude="3" metricUnit="m" metricValue="1.4419999999999997" unit="m" value="1442.0">1442 m</elevation>
</quantity>
,
<collectorName id="31A2A4D9DCDCEAFA0A2A2458214759BF">
Lgt. T.
<normalizedToken id="B14005C086147DDB2BFFC1C1A04653CF" originalValue="Kopecký“">Kopecky&quot;</normalizedToken>
</collectorName>
[white, printed], &quot;
<emphasis id="ED866D038560BBA7D47E653F480951E8" italics="true" pageId="0" pageNumber="173">COMMATOCERUS</emphasis>
/
<emphasis id="940EC265CB79AFD84E5766F1723CD274" italics="true" pageId="0" pageNumber="173">concinnus</emphasis>
Bes &amp; Cucc. /
<determinerName id="82839A985BA6097B797C7CFA43F3EA62">
P.
<normalizedToken id="7CB2A18A893E24395F4D9D8FF6AD5965" originalValue="Hlaváč">Hlavac</normalizedToken>
</determinerName>
det., 2023&quot; [white, printed] (PCTK, PCPH). New record for Oman
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="50554164245A7E25F7B2B3A435E2F1B1" pageId="0" pageNumber="173" type="natural history">
<paragraph id="EE3430211E2A16BA53B4573EC24C7661" pageId="0" pageNumber="173">Natural history.</paragraph>
<paragraph id="8F9937FE63B3F60F04749C3946C32F3D" pageId="0" pageNumber="173">
The two specimens were collected from two different ant nests under stones. One with an unknown species of
<taxonomicName id="B1FD4B7A626A5643D183AB5DD37D4EA5" class="Insecta" family="Formicidae" genus="Lasius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lasius" order="Hymenoptera" pageId="0" pageNumber="173" phylum="Arthropoda" rank="genus">
<emphasis id="6F53E52C9B1329B277F4BF00680B726F" italics="true" pageId="0" pageNumber="173">Lasius</emphasis>
</taxonomicName>
Fabricius, 1804 and the other with
<taxonomicName id="20775BA9943FA75AA30BC7C108D41B91" class="Insecta" family="Formicidae" genus="Lepisiota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lepisiota" order="Coleoptera" pageId="0" pageNumber="173" phylum="Arthropoda" rank="genus">
<emphasis id="D2C4460700AD2F954D724437FCC2F9B6" italics="true" pageId="0" pageNumber="173">Lepisiota</emphasis>
</taxonomicName>
(cf.)
<taxonomicName id="13EC95BE7BA9899259161DA727CD9CC8" baseAuthorityName="Kuznetsov-Ugamsky" baseAuthorityYear="1929" class="Insecta" family="Formicidae" genus="Lepisiota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lepisiota spinisquama" order="Coleoptera" pageId="0" pageNumber="173" phylum="Arthropoda" rank="species" species="spinisquama">
<emphasis id="8A4ABB4DC4188D9DF018DCC0F2C74E1F" italics="true" pageId="0" pageNumber="173">Lepisiota spinisquama</emphasis>
</taxonomicName>
(Kuznetsov-Ugamsky, 1929), M. Sharaf det., 2023, collected in the evening on an open, dry area in the mountains (Fig.
<figureCitation id="89637F6A48C1DA001B1622C25A92A49A" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A Oman, Ad Dakhilyiah Gov., near Subayb wadi, locality with the occurrence of Commatocerus concinnus B distribution map of all Clavigeritae species known from the Arabian Peninsula." figureDoi="10.3897/zookeys.1198.119152.figure6" httpUri="https://binary.pensoft.net/fig/1031833" pageId="0" pageNumber="173">6A</figureCitation>
).
</paragraph>
<caption id="55C4C70487E06CD5BFA74D3EE404DF51" doi="10.3897/zookeys.1198.119152.figure5" httpUri="https://binary.pensoft.net/fig/1031832" pageId="0" pageNumber="173" start="Figure 5" startId="F5">
<paragraph id="AD3C8C2A0EA59D78706B9D99AADFAEBF" pageId="0" pageNumber="173">
<emphasis id="3F52097D4CE1CA6573D473FAA25D8E87" bold="true" pageId="0" pageNumber="173">Figure 5.</emphasis>
Type locality of
<taxonomicName id="AF0EEC5C0A35624099737D760FC6FB4F" authorityName="Hlaváč &amp; Baňař &amp; Stočes" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Corynotopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Corynotopsis omanicus" order="Coleoptera" pageId="0" pageNumber="173" phylum="Arthropoda" rank="species" species="omanicus">
<emphasis id="A6E1A1D44077979CDBC800CCC5E67598" italics="true" pageId="0" pageNumber="173">Corynotopsis omanicus</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</caption>
<caption id="55D8480A5681B2E441B253C8DDBDBF2D" doi="10.3897/zookeys.1198.119152.figure6" httpUri="https://binary.pensoft.net/fig/1031833" pageId="0" pageNumber="173" start="Figure 6" startId="F6">
<paragraph id="44BC029B4F8FD7B2B90FC382E34F3BF9" pageId="0" pageNumber="173">
<emphasis id="CC8A2ED88D55569F4FD49B46F9C5A2AB" bold="true" pageId="0" pageNumber="173">Figure 6.</emphasis>
<emphasis id="5F45880FA1FB1A64EBA35D8FCB991A9D" bold="true" pageId="0" pageNumber="173">A</emphasis>
Oman, Ad Dakhilyiah Gov., near Subayb wadi, locality with the occurrence of
<taxonomicName id="BD126939F93EF5C8F5B5B5FE4505149A" authorityName="Besuchet &amp; Cuccodoro" authorityYear="2011" class="Insecta" family="Staphylinidae" genus="Commatocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Commatocerus concinnus" order="Coleoptera" pageId="0" pageNumber="173" phylum="Arthropoda" rank="species" species="concinnus">
<emphasis id="BE80295CA2F652FC9EBBAA1FAC807170" italics="true" pageId="0" pageNumber="173">Commatocerus concinnus</emphasis>
</taxonomicName>
<emphasis id="ADF53D9FBDE81DDD82D7C2FA15AC13D3" bold="true" pageId="0" pageNumber="173">B</emphasis>
distribution map of all
<taxonomicName id="4F2234DA20A76533A59F63269F7E10F8" authorityName="Raffray" authorityYear="1882" family="Staphylinidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="173" rank="superTribe" superTribe="Clavigeritae">Clavigeritae</taxonomicName>
species known from the Arabian Peninsula.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="5C8BA92013682136168D204BAB68755F" pageId="0" pageNumber="173" type="remarks">
<paragraph id="CCC5D83CACAB5D9A77F1310871E33B6D" pageId="0" pageNumber="173">Remarks.</paragraph>
<paragraph id="7244866F337878BEF6B1F7B84643135A" pageId="0" pageNumber="173">
<taxonomicName id="89ED104184153CA8A9A5D62C823204DE" authorityName="Besuchet &amp; Cuccodoro" authorityYear="2011" class="Insecta" family="Staphylinidae" genus="Commatocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Commatocerus concinnus" order="Coleoptera" pageId="0" pageNumber="173" phylum="Arthropoda" rank="species" species="concinnus">
<emphasis id="B0F1DA36965F802E1D9ED12D2B9F5AE7" italics="true" pageId="0" pageNumber="173">Commatocerus concinnus</emphasis>
</taxonomicName>
was described based on one male from United Arab Emirates, Ras al Khaimah, Wadi Shawkah,
<geoCoordinate id="DC1C83C118FEC07628E7E7BD3D46E6A9" degrees="25" direction="north" minutes="6" orientation="latitude" precision="1" seconds="15.30" value="25.10425">25°6'15.30&quot;N</geoCoordinate>
,
<geoCoordinate id="848743D5C61B86F35E35A0FF6AE00297" degrees="56" direction="east" minutes="2" orientation="longitude" precision="1" seconds="47.55" value="56.04654">56°2'47.55&quot;E</geoCoordinate>
, collected by a water-trap. The species is readily separated from its congeners by having the terminal antennomeres cylindrical, instead of distally expanded.
</paragraph>
<paragraph id="1ED15E9ECDF992F7B23E6E25C26CEE26" pageId="0" pageNumber="173">
The generic name
<taxonomicName id="A86CEF1353A6FCDADF2F917A2DDA456A" authorityName="Raffray" authorityYear="1882" class="Insecta" family="Staphylinidae" genus="Commatocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Commatocerus" order="Coleoptera" pageId="0" pageNumber="173" phylum="Arthropoda" rank="genus">
<emphasis id="B39D11BED7506B36D6B17FAF7718FCB3" italics="true" pageId="0" pageNumber="173">Commatocerus</emphasis>
</taxonomicName>
has been subjected to much instability. It was established by
<bibRefCitation id="6AE0CE1302C5A5EC637456AC2E936CD7" author="Raffray, A" journalOrPublisher="Revue d'Entomologie" pageId="0" pageNumber="173" pagination="1 - 16" refId="B11" refString="Raffray, A, 1882. Pselaphides nouveaux ou peu connus. 1er memoire. Revue d'Entomologie 1: 1 - 16" title="Pselaphides nouveaux ou peu connus. 1 er memoire." volume="1" year="1882">Raffray (1882</bibRefCitation>
: 1) for
<taxonomicName id="DD5EE4A51EC91239DE92F268456E14DE" authorityName="Raffray" authorityYear="1882" class="Insecta" family="Staphylinidae" genus="Commatocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Commatocerus elegantulus" order="Coleoptera" pageId="0" pageNumber="173" phylum="Arthropoda" rank="species" species="elegantulus">
<emphasis id="13715351277D44B08EA53CD70DAA8228" italics="true" pageId="0" pageNumber="173">Commatocerus elegantulus</emphasis>
</taxonomicName>
Raffray. In the same year,
<bibRefCitation id="E08CF25E9C861BA513C872522E210809" DOI="https://doi.org/10.1002/mmnd.48018820229" author="Reitter, E" journalOrPublisher="Verhandlungen des Naturforschenden Vereines in Bruenn" pageId="0" pageNumber="173" pagination="177 - 211" refId="B13" refString="Reitter, E, 1882. Versuch einer systematischen Eintheilung der Clavigeriden und Pselaphiden. Verhandlungen des Naturforschenden Vereines in Bruenn 20 (1881): 177 - 211, DOI: https://doi.org/10.1002/mmnd.48018820229" title="Versuch einer systematischen Eintheilung der Clavigeriden und Pselaphiden." url="https://doi.org/10.1002/mmnd.48018820229" volume="20" year="1882">Reitter (1882)</bibRefCitation>
synonymized it with the genus
<taxonomicName id="D491F5ED58656708F3F28B929D55E00F" class="Insecta" family="Staphylinidae" genus="Fustiger" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fustiger" order="Coleoptera" pageId="0" pageNumber="173" phylum="Arthropoda" rank="genus">
<emphasis id="EC0EA37493D5CF394D2318BA55051E0A" italics="true" pageId="0" pageNumber="173">Fustiger</emphasis>
</taxonomicName>
LeConte, 1866 under the name
<taxonomicName id="9EA7C14ED2B38EB9E7D43745D4E47E0D" genus="Comatocerus" lsidName="Comatocerus" pageId="0" pageNumber="173" rank="genus">
<emphasis id="2409AE08B7CA2A9A009717C6A61F4CBE" italics="true" pageId="0" pageNumber="173">Comatocerus</emphasis>
</taxonomicName>
[sic] (
<bibRefCitation id="955E71B80C7BAB083C46EC1FA43C63A4" DOI="https://doi.org/10.1002/mmnd.48018820229" author="Reitter, E" journalOrPublisher="Verhandlungen des Naturforschenden Vereines in Bruenn" pageId="0" pageNumber="173" pagination="177 - 211" refId="B13" refString="Reitter, E, 1882. Versuch einer systematischen Eintheilung der Clavigeriden und Pselaphiden. Verhandlungen des Naturforschenden Vereines in Bruenn 20 (1881): 177 - 211, DOI: https://doi.org/10.1002/mmnd.48018820229" title="Versuch einer systematischen Eintheilung der Clavigeriden und Pselaphiden." url="https://doi.org/10.1002/mmnd.48018820229" volume="20" year="1882">Reitter 1882</bibRefCitation>
: 200).
<bibRefCitation id="31E66C0AFD10111E6ACD7783D5391412" author="Jeannel, R" journalOrPublisher="Memoires du Museum National d'Histoire Naturelle, (Zoologie)" pageId="0" pageNumber="173" pagination="1 - 226" refId="B7" refString="Jeannel, R, 1949. Les Pselaphides de L'Afrique orientale. Memoires du Museum National d'Histoire Naturelle, (Zoologie) 29 (1): 1 - 226" title="Les Pselaphides de L'Afrique orientale." volume="29" year="1949">Jeannel (1949</bibRefCitation>
: 37) redescribed the genus and resurrected its generic status. He also pointed out that the American and Old World species of the genus
<taxonomicName id="45474EFC7BC1178EBB46357498D3D61E" class="Insecta" family="Staphylinidae" genus="Fustiger" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fustiger" order="Coleoptera" pageId="0" pageNumber="173" phylum="Arthropoda" rank="genus">
<emphasis id="C9A75D5CFCA4169B1AF2D1CA37227825" italics="true" pageId="0" pageNumber="173">Fustiger</emphasis>
</taxonomicName>
cannot be congeneric and should be placed in separate genera. Later,
<bibRefCitation id="D5ABF921E8408FFDF0B15302045A3483" author="Jeannel, R" journalOrPublisher="Annals &amp; Magazine of Natural History" pageId="0" pageNumber="173" refId="B9" refString="Jeannel, R, 1954. Pselaphides recueillis par N. Leleup au Congo Belge sous les auspices de l'Institut pour la Recherche Scientifique en Afrique Centrale (I.R.S.A.C.). VI-XI. Annales du Musee Royal du Congo Belge, Tervuren (Serie 8°: Sciences Zoologiques) 33: 1-175." title="Pselaphides recueillis par N. Leleup au Congo Belge sous les auspices de l'Institut pour la Recherche Scientifique en Afrique Centrale (I. R. S. A. C.). VI-XI. Annales du Musee Royal du Congo Belge, Tervuren (Serie 8 °: Sciences Zoologiques) 33: 1 - 175." year="1954">Jeannel (1954</bibRefCitation>
: 152) discussed the status of similar genera
<taxonomicName id="2268D8F38CD28F5FB25BCDD308852098" authorityName="Raffray" authorityYear="1882" class="Insecta" family="Staphylinidae" genus="Commatocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Commatocerus" order="Coleoptera" pageId="0" pageNumber="173" phylum="Arthropoda" rank="genus">
<emphasis id="A3CACAC36742E82EAEB12FC3104A1B40" italics="true" pageId="0" pageNumber="173">Commatocerus</emphasis>
</taxonomicName>
,
<taxonomicName id="5AF2344CD47AB1A580B1581B94EE188E" authorityName="Wasmann" authorityYear="1912" class="Insecta" family="Staphylinidae" genus="Fustigerinus" kingdom="Animalia" lsidName="Fustigerinus" order="Coleoptera" pageId="0" pageNumber="173" phylum="Arthropoda" rank="genus">
<emphasis id="1C16A264481B4670335D706D7215B596" italics="true" pageId="0" pageNumber="173">Fustigerinus</emphasis>
</taxonomicName>
Wasmann, 1912 and its actual synonym
<taxonomicName id="39C227AE7DF5F0513E09111994047F26" authorityName="Jeannel" authorityYear="1949" class="Insecta" family="Staphylinidae" genus="Neocommatocerus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Neocommatocerus" order="Coleoptera" pageId="0" pageNumber="173" phylum="Arthropoda" rank="genus">
<emphasis id="134CD9F1DE7DF2796D91BF5D20653471" italics="true" pageId="0" pageNumber="173">Neocommatocerus</emphasis>
</taxonomicName>
Jeannel, 1949.
<bibRefCitation id="98874AB563C9004A2C59774B7BB4F6D5" author="Celis, MJ" journalOrPublisher="Revue de Zoologie et de Botanique Africaines" pageId="0" pageNumber="173" pagination="440 - 446" refId="B4" refString="Celis, MJ, 1975. Contributions a l'etude des Clavigerines de l'Afrique (Coleoptera Pselaphidae). 10. Le genre Commatocerus Raffray (Trib. Fustigerini). Revue de Zoologie et de Botanique Africaines 89 (2): 440 - 446" title="Contributions a l'etude des Clavigerines de l'Afrique (Coleoptera Pselaphidae). 10. Le genre Commatocerus Raffray (Trib. Fustigerini)." volume="89" year="1975">
<normalizedToken id="D69D4CABEBD3601EDCAEE4278A2FC460" originalValue="Célis">Celis</normalizedToken>
(1975
</bibRefCitation>
: 441) confirmed the generic status of
<taxonomicName id="0ED586FE2274A54A5A59D10F0D2019BA" authorityName="Raffray" authorityYear="1882" class="Insecta" family="Staphylinidae" genus="Commatocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Commatocerus" order="Coleoptera" pageId="0" pageNumber="173" phylum="Arthropoda" rank="genus">
<emphasis id="AB2738AAA44D18784F0FDEE172012749" italics="true" pageId="0" pageNumber="173">Commatocerus</emphasis>
</taxonomicName>
.
<bibRefCitation id="0FFB3595636606C808D7509655FBF5F8" author="Besuchet, C" journalOrPublisher="Entomologica Basiliensia" pageId="0" pageNumber="173" pagination="261 - 267" refId="B1" refString="Besuchet, C, 1977. Ergebnisse der Bhutan-Expedition 1972 des Naturhistorischen Museums in Basel. Coleoptera: Fam. Pselaphidae Subf. Clavigerinae Gen. Fustiger BREND. mit Ein schluss der indischen Arten. Entomologica Basiliensia 2: 261 - 267" title="Ergebnisse der Bhutan-Expedition 1972 des Naturhistorischen Museums in Basel. Coleoptera: Fam. Pselaphidae Subf. Clavigerinae Gen. Fustiger BREND. mit Ein schluss der indischen Arten." volume="2" year="1977">Besuchet (1977</bibRefCitation>
: 261) again synonymized it with
<taxonomicName id="C7AA20D48F43990AD1A16F8CD91EFFD4" class="Insecta" family="Staphylinidae" genus="Fustiger" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fustiger" order="Coleoptera" pageId="0" pageNumber="173" phylum="Arthropoda" rank="genus">
<emphasis id="FF3BF5FB715A4BF9EB10E99B397638F5" italics="true" pageId="0" pageNumber="173">Fustiger</emphasis>
</taxonomicName>
and provided a key to species of India. When describing
<taxonomicName id="EAD96DE5B2A9F518A208881BD2C912BB" authorityName="Besuchet &amp; Cuccodoro" authorityYear="2011" class="Insecta" family="Staphylinidae" genus="Commatocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Commatocerus concinnus" order="Coleoptera" pageId="0" pageNumber="173" phylum="Arthropoda" rank="species" species="concinnus">
<emphasis id="B3933563DFB5C13A3D8ED51817CC7D30" italics="true" pageId="0" pageNumber="173">Commatocerus concinnus</emphasis>
</taxonomicName>
(
<bibRefCitation id="A5A350542A1A72E42B39297C962A4AA4" author="Besuchet, C" editor="Harten, A van" journalOrPublisher="Vol. 4. Dar Al Ummah Printing, Publishing, Distribution and Advertising, Abu Dhabi, UAE" pageId="0" pageNumber="173" pagination="151 - 167" refId="B3" refString="Besuchet, C, Cuccodoro, G, 2011. Order Coleoptera, family Staphylinidae, 151-157. In: Harten, A van, Ed., Arthropod fauna of the United Arab Emirates. Vol. 4. Dar Al Ummah Printing, Publishing, Distribution and Advertising, Abu Dhabi, UAE: 151 - 167" title="Order Coleoptera, family Staphylinidae, 151 - 157." volumeTitle="Arthropod fauna of the United Arab Emirates." year="2011">Besuchet and Cuccodoro 2011</bibRefCitation>
: 164), the authors discussed and supported the validity of the genus and they apparently overlooked the synonymy of Besuchet from 1977.
</paragraph>
</subSubSection>
<subSubSection id="5977C5C2635707B675A7D6B3931BCC18" pageId="0" pageNumber="173" type="species included">
<paragraph id="A1F5BC086B0A9FD7F79D07168B45317C" pageId="0" pageNumber="173">Species included.</paragraph>
<paragraph id="122BA466E5C6263965E095ABB724B9C5" pageId="0" pageNumber="173">
<taxonomicName id="DB1A596610111A6639E342B449C83ECA" authorityName="Besuchet &amp; Cuccodoro" authorityYear="2011" class="Insecta" family="Staphylinidae" genus="Commatocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Commatocerus concinnus" order="Coleoptera" pageId="0" pageNumber="173" phylum="Arthropoda" rank="species" species="concinnus">
<emphasis id="AD3CB772FF36F7E7EA5E6E497B115C22" italics="true" pageId="0" pageNumber="173">Commatocerus concinnus</emphasis>
</taxonomicName>
Besuchet &amp; Cuccodoro, 2011,
<taxonomicName id="B5BD61411F8CDE66130430B2567FCBE9" family="Staphylinidae" kingdom="Animalia" lsidName="C. elegantulus" order="Coleoptera" pageId="0" pageNumber="173" rank="species" species="elegantulus">
<emphasis id="6C549286621A38FA7A1827DE5A52ABF5" italics="true" pageId="0" pageNumber="173">C. elegantulus</emphasis>
</taxonomicName>
Raffray, 1882,
<taxonomicName id="F3C25184E0037889C0A9EBC8D45B11F7" family="Staphylinidae" kingdom="Animalia" lsidName="C. leleupi" order="Coleoptera" pageId="0" pageNumber="173" rank="species" species="leleupi">
<emphasis id="F2F464F89C8660161BE2CF773B6FBEB4" italics="true" pageId="0" pageNumber="173">C. leleupi</emphasis>
</taxonomicName>
Jeannel, 1953 and
<taxonomicName id="EF55723ED9AD3B3135D65DD559FF13F1" family="Staphylinidae" kingdom="Animalia" lsidName="C. turkmenicus" order="Coleoptera" pageId="0" pageNumber="173" rank="species" species="turkmenicus">
<emphasis id="2F5C8BACE4214E2D5C610D2E186F8215" italics="true" pageId="0" pageNumber="173">C. turkmenicus</emphasis>
</taxonomicName>
Kryzhanovski, 1957.
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>
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