344 lines
43 KiB
XML
344 lines
43 KiB
XML
<document id="7729B457989583FC0B53504993EFE313" ID-CLB-Dataset="55716" ID-DOI="10.5852/ejt.2022.817.1763" ID-GBIF-Dataset="daf1b8a9-0042-4b39-8f9c-df87cb1603f5" ID-ISSN="2118-9773" ID-Zenodo-Dep="6518289" ID-ZooBank="22C5B528-F7D7-40F2-8877-C30FF6057157" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1651674041341" checkinUser="felipe" docAuthor="Tshernyshev, Sergei E." docDate="2022" docId="1E7A87E1505A5C1AFD17955D35F06BBB" docLanguage="en" docName="ejt-817_58-77.pdf" docOrigin="European Journal of Taxonomy 817" docSource="http://dx.doi.org/10.5852/ejt.2022.817.1763" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Tonyattalus Tshernyshev 2022, gen. nov." docType="treatment" docUuid="DF2416DD-2780-4C55-805F-C4000309D01E" docUuidSource="ZooBank" docVersion="9" lastPageNumber="63" masterDocId="E243FF9950595C1FFF8D9279370A6F24" masterDocTitle="Soft-winged flower beetles (Coleoptera: Malachiidae) of the United Arab Emirates" masterLastPageNumber="77" masterPageNumber="58" pageNumber="61" updateTime="1699329578614" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="D193809AA4DF49B02EC120AB7632B44C">
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<mods:title id="D5B3FD1CB2D6F880BBDED36F618F5CBA">Soft-winged flower beetles (Coleoptera: Malachiidae) of the United Arab Emirates</mods:title>
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<mods:name id="5E54DC132EBEFD94F8C4A5A0ECB6EAA7" type="personal">
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<mods:namePart id="E6D009A6F121D33050B3B71386FB5493">Tshernyshev, Sergei E.</mods:namePart>
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<mods:affiliation id="F59F76BA669DF298DAD4AD1B4A143FB1">Institute of Systematics and Ecology of Animals, Siberian Branch of the Russian Academy of Sciences, Frunze Street 11, 630091 Novosibirsk, Russia. Tomsk State University, Lenina prospekt 36, Tomsk 634050, Russia. Email: sch-sch @ mail. ru</mods:affiliation>
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<mods:nameIdentifier id="7FCFB17454D7EC2C65E423B4DD6FE267" type="email">sch-sch@mail.ru</mods:nameIdentifier>
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<mods:title id="F4F2FF89C8589BAEAB742367ABAAF345">European Journal of Taxonomy</mods:title>
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<mods:part id="ED6041C9E52C44683047B49EEE1BE6BE">
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<mods:date id="1762A389AB66287EAA20E631414B59AE">2022</mods:date>
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<mods:number id="36CCC65E312A4A28EFC07704EA0D8B37">2022-04-29</mods:number>
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<mods:number id="F384088FB75F6960D31D8B703662371C">817</mods:number>
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<mods:url id="65593F20DDCF22B9AE414548C26EF889">http://dx.doi.org/10.5852/ejt.2022.817.1763</mods:url>
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<treatment id="1E7A87E1505A5C1AFD17955D35F06BBB" ID-DOI="http://doi.org/10.5281/zenodo.6521859" ID-GBIF-Taxon="195191934" ID-Zenodo-Dep="6521859" LSID="urn:lsid:zoobank.org:act:DF2416DD-2780-4C55-805F-C4000309D01E" httpUri="http://treatment.plazi.org/id/1E7A87E1505A5C1AFD17955D35F06BBB" lastPageId="5" lastPageNumber="63" pageId="3" pageNumber="61">
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<subSubSection id="DEC9657C505A5C1CFD17955D3490681B" box="[666,922,1828,1855]" pageId="3" pageNumber="61" type="nomenclature">
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<paragraph id="966C36F7505A5C1CFD17955D3490681B" blockId="3.[372,1215,1828,1891]" box="[666,922,1828,1855]" pageId="3" pageNumber="61">
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<heading id="CD24819B505A5C1CFD17955D3490681B" box="[666,922,1828,1855]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="61" reason="2">
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<taxonomicName id="51D34D74505A5C1CFD17955D3422681B" authority="Tshernyshev, 2022" authorityName="Tshernyshev" authorityYear="2022" box="[666,808,1828,1855]" class="Insecta" family="Melyridae" genus="Tonyattalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="61" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="A4A7EAE5505A5C1CFD17955D3422681B" bold="true" box="[666,808,1828,1855]" italics="true" pageId="3" pageNumber="61">Tonyattalus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="BF94579E505A5C1CFCA2955C3490681B" box="[815,922,1829,1855]" pageId="3" pageNumber="61" rank="genus">gen. nov.</taxonomicNameLabel>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="DEC9657C505A5C1CFEF9953133B56847" box="[372,1215,1864,1891]" pageId="3" pageNumber="61" type="description">
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<paragraph id="966C36F7505A5C1CFEF9953133B56847" blockId="3.[372,1215,1828,1891]" box="[372,1215,1864,1891]" pageId="3" pageNumber="61">
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<uri id="E2423AF5505A5C1CFEF9953133B56847" box="[372,1215,1864,1891]" pageId="3" pageNumber="61">
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urn:lsid:zoobank.org:act:
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<uuid id="E2750C22505A5C1CFD02953133B56847" box="[655,1215,1864,1891]" pageId="3" pageNumber="61">DF2416DD-2780-4C55-805F-C4000309D01E</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="DEC9657C505A5C1CFF3095F0330C68F6" pageId="3" pageNumber="61" type="type_taxon">
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<paragraph id="966C36F7505A5C1CFF3095F036536887" blockId="3.[189,345,1929,1955]" box="[189,345,1929,1955]" pageId="3" pageNumber="61">
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<heading id="CD24819B505A5C1CFF3095F036536887" bold="true" box="[189,345,1929,1955]" fontSize="11" level="3" pageId="3" pageNumber="61" reason="3">
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<emphasis id="A4A7EAE5505A5C1CFF3095F036536887" bold="true" box="[189,345,1929,1955]" pageId="3" pageNumber="61">
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<typeStatus id="49688855505A5C1CFF3095F037F16887" box="[189,251,1929,1955]" pageId="3" pageNumber="61">Type</typeStatus>
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species
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="966C36F7505A5C1CFF3095C1330C68F6" blockId="3.[189,1030,1976,2002]" box="[189,1030,1976,2002]" pageId="3" pageNumber="61">
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<taxonomicName id="51D34D74505A5C1CFF3095C136C468F6" authority="Tshernyshev, 2022" authorityName="Tshernyshev" authorityYear="2022" box="[189,462,1976,2002]" class="Insecta" family="Melyridae" genus="Tonyattalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="61" phylum="Arthropoda" rank="species" species="vanharteni" status="gen. et sp. nov.">
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<emphasis id="A4A7EAE5505A5C1CFF3095C136C468F6" box="[189,462,1976,2002]" italics="true" pageId="3" pageNumber="61">Tonyattalus vanharteni</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="BF94579E505A5C1CFE5B95C1358368F6" box="[470,649,1976,2002]" pageId="3" pageNumber="61" rank="species">gen. et sp. nov.</taxonomicNameLabel>
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, designated in the present paper.
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</paragraph>
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</subSubSection>
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<subSubSection id="DEC9657C505D5C1BFF30937136856A43" pageId="4" pageNumber="62" type="diagnosis">
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<paragraph id="966C36F7505D5C1BFF309371363D6E06" blockId="4.[189,311,264,290]" box="[189,311,264,290]" pageId="4" pageNumber="62">
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<heading id="CD24819B505D5C1BFF309371363D6E06" bold="true" box="[189,311,264,290]" fontSize="11" level="3" pageId="4" pageNumber="62" reason="3">
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<emphasis id="A4A7EAE5505D5C1BFF309371363D6E06" bold="true" box="[189,311,264,290]" pageId="4" pageNumber="62">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="966C36F7505D5C1BFF30934E36736D4F" blockId="4.[189,1399,311,619]" pageId="4" pageNumber="62">
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Moderately small-sized (ca 2.0–
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<quantity id="512B9B12505D5C1BFDCE934E35A86E76" box="[579,674,311,338]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="4" pageNumber="62" unit="mm" value="2.3">2.3 mm</quantity>
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) malachiid beetles with the body subparallel and oval with abdomen not completely covered by elytra (
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<figureCitation id="0EE82A72505D5C1BFD5A9323345E6E50" box="[727,852,346,372]" captionStart="Figs 1–10" captionStartId="6.[189,242,1889,1915]" captionTargetBox="[222,1352,267,1823]" captionTargetId="figure-75@6.[219,1368,265,1849]" captionTargetPageId="6" captionText="Figs 1–10. Tonyattalus vanharteni gen. et sp. nov., holotype (SCH_ISEA_000133), ♂ (1–8), ♀ (9– 10). 1, 9. External appearance, dorsal view. 2, 10. External appearance, lateral view. 3. Left antenna. 4. Head and pronotum, dorsal view. 5. Left anterior tarsus. 6. Pygidium. 7. Ultimate abdominal ventrite. 8. Aedeagus, and tegmen, lateral view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6518293" httpUri="https://zenodo.org/record/6518293/files/figure.png" pageId="4" pageNumber="62">Figs 1–10</figureCitation>
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). Colouration from yellow with brown bases and apices of elytra, to black with yellow spots on pronotum and elytra; lacking metallic lustre. Antennae dentate in females and flabellate in males, not long, reaching the base of the elytra; in males 1
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antennomere enlarged, sub-conic, 2
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antennomere oval, half as long as the 3
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<superScript id="61A69BBF505D5C1BFBE293BA33776EF5" attach="left" box="[1135,1149,451,465]" fontSize="6" pageId="4" pageNumber="62">rd</superScript>
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antennomere, the 3
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<superScript id="61A69BBF505D5C1BFAE593BA327C6EF5" attach="none" box="[1384,1398,451,465]" fontSize="6" pageId="4" pageNumber="62">rd</superScript>
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antennomere triangular, not wider than the 4
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<superScript id="61A69BBF505D5C1BFD47939F35DD6ED0" attach="none" box="[714,727,486,500]" fontSize="6" pageId="4" pageNumber="62">th</superScript>
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, 4
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<superScript id="61A69BBF505D5C1BFD78939F34086ED0" attach="left" box="[757,770,486,500]" fontSize="6" pageId="4" pageNumber="62">th</superScript>
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antennomere wide triangular, with somewhat dilated outer edge, 5
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with more strongly dilated outer edge, 6
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<superScript id="61A69BBF505D5C1BFCCB907034596D33" attach="none" box="[838,851,521,535]" fontSize="6" pageId="4" pageNumber="62">th</superScript>
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–10
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<superScript id="61A69BBF505D5C1BFCF2907034866D33" attach="left" box="[895,908,521,535]" fontSize="6" pageId="4" pageNumber="62">th</superScript>
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segments flabellate, cylindrical with long dilated outer edges, terminal antennomere distinctly the longest, curved at basal third, evenly rounded at apex (
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<figureCitation id="0EE82A72505D5C1BFEAF902836636D4F" box="[290,361,593,619]" captionStart="Figs 1–10" captionStartId="6.[189,242,1889,1915]" captionTargetBox="[222,1352,267,1823]" captionTargetId="figure-75@6.[219,1368,265,1849]" captionTargetPageId="6" captionText="Figs 1–10. Tonyattalus vanharteni gen. et sp. nov., holotype (SCH_ISEA_000133), ♂ (1–8), ♀ (9– 10). 1, 9. External appearance, dorsal view. 2, 10. External appearance, lateral view. 3. Left antenna. 4. Head and pronotum, dorsal view. 5. Left anterior tarsus. 6. Pygidium. 7. Ultimate abdominal ventrite. 8. Aedeagus, and tegmen, lateral view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6518293" httpUri="https://zenodo.org/record/6518293/files/figure.png" pageId="4" pageNumber="62">Fig. 3</figureCitation>
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).
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</paragraph>
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<paragraph id="966C36F7505D5C1BFF3090EB346E6C79" blockId="4.[187,1398,658,861]" pageId="4" pageNumber="62">
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Head narrower than pronotum, slightly elongate, eyes small, weakly protruding over the contour of head; frons and inter-ocular area flat and slightly impressed, glabrous, lacking depression or protuberances; clypeus slightly elongate, transverse, straight; labrum short and transverse; maxillary palpi short, ultimate palpomere thin, cylindrical, slightly narrowed and flattened apically. Pronotum (
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<figureCitation id="0EE82A72505D5C1BFB5A908532676C32" box="[1239,1389,764,790]" captionStart="Figs 1–10" captionStartId="6.[189,242,1889,1915]" captionTargetBox="[222,1352,267,1823]" captionTargetId="figure-75@6.[219,1368,265,1849]" captionTargetPageId="6" captionText="Figs 1–10. Tonyattalus vanharteni gen. et sp. nov., holotype (SCH_ISEA_000133), ♂ (1–8), ♀ (9– 10). 1, 9. External appearance, dorsal view. 2, 10. External appearance, lateral view. 3. Left antenna. 4. Head and pronotum, dorsal view. 5. Left anterior tarsus. 6. Pygidium. 7. Ultimate abdominal ventrite. 8. Aedeagus, and tegmen, lateral view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6518293" httpUri="https://zenodo.org/record/6518293/files/figure.png" pageId="4" pageNumber="62">Figs 2, 4, 10</figureCitation>
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) transverse, with rounded lateral sides, not sinuate or depressed at base; scutellum small, rectangular, narrow, slightly transverse, with slightly subparallel apex.
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</paragraph>
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<paragraph id="966C36F7505D5C1BFF3091FA36856A43" blockId="4.[188,1399,899,1383]" pageId="4" pageNumber="62">
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Elytra subparallel, weakly expanded posteriorly and not covering last segments of abdomen in both sexes; humeri indistinct and small; surface densely punctured, covered with light dense adpressed short hairs. Anterior tarsi (
|
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<figureCitation id="0EE82A72505D5C1BFD8D91B335406CC1" box="[512,586,970,997]" captionStart="Figs 1–10" captionStartId="6.[189,242,1889,1915]" captionTargetBox="[222,1352,267,1823]" captionTargetId="figure-75@6.[219,1368,265,1849]" captionTargetPageId="6" captionText="Figs 1–10. Tonyattalus vanharteni gen. et sp. nov., holotype (SCH_ISEA_000133), ♂ (1–8), ♀ (9– 10). 1, 9. External appearance, dorsal view. 2, 10. External appearance, lateral view. 3. Left antenna. 4. Head and pronotum, dorsal view. 5. Left anterior tarsus. 6. Pygidium. 7. Ultimate abdominal ventrite. 8. Aedeagus, and tegmen, lateral view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6518293" httpUri="https://zenodo.org/record/6518293/files/figure.png" pageId="4" pageNumber="62">Fig. 5</figureCitation>
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) five-segmented, compressed, with a comb above the 2
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segment, 2
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<superScript id="61A69BBF505D5C1BFF41919537D76CDE" attach="none" box="[204,221,1004,1018]" fontSize="6" pageId="4" pageNumber="62">nd</superScript>
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–3
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<superScript id="61A69BBF505D5C1BFF77919536026CDE" attach="left" box="[250,264,1004,1018]" fontSize="6" pageId="4" pageNumber="62">rd</superScript>
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tarsomeres equal in length, 1
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is 1.2 × as long as 2
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, the terminal tarsomere slightly narrower and depressed, the longest, as long as 1
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–4
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tarsomeres in anterior legs, 1
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<superScript id="61A69BBF505D5C1BFC60966934F26B3A" attach="none" box="[1005,1016,1040,1054]" fontSize="6" pageId="4" pageNumber="62">st</superScript>
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–3
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in intermediate legs and 1
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– 2
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<superScript id="61A69BBF505D5C1BFF41964A37D76B65" attach="left" box="[204,221,1075,1089]" fontSize="6" pageId="4" pageNumber="62">nd</superScript>
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in posterior legs; claws thin, weakly curved and sharp, with dentate small lamellae at base; femora and tibiae simple, not swollen or strongly curved, lacking indentation. Metathorax weakly swollen, narrow, transverse, lacking appendages or tuft of hairs, but with specific straight and edged distal side, which is slightly lamellate. Pygidium (
|
||
<figureCitation id="0EE82A72505D5C1BFD0396E435DC6B9C" box="[654,726,1181,1208]" captionStart="Figs 1–10" captionStartId="6.[189,242,1889,1915]" captionTargetBox="[222,1352,267,1823]" captionTargetId="figure-75@6.[219,1368,265,1849]" captionTargetPageId="6" captionText="Figs 1–10. Tonyattalus vanharteni gen. et sp. nov., holotype (SCH_ISEA_000133), ♂ (1–8), ♀ (9– 10). 1, 9. External appearance, dorsal view. 2, 10. External appearance, lateral view. 3. Left antenna. 4. Head and pronotum, dorsal view. 5. Left anterior tarsus. 6. Pygidium. 7. Ultimate abdominal ventrite. 8. Aedeagus, and tegmen, lateral view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6518293" httpUri="https://zenodo.org/record/6518293/files/figure.png" pageId="4" pageNumber="62">Fig. 6</figureCitation>
|
||
) undivided, evenly rounded and narrowed distally, with oval emargination in middle; 8
|
||
<superScript id="61A69BBF505D5C1BFDA896B935386BEA" attach="left" box="[549,562,1216,1230]" fontSize="6" pageId="4" pageNumber="62">th</superScript>
|
||
ultimate abdominal ventrite (
|
||
<figureCitation id="0EE82A72505D5C1BFC0396B934D96BFF" box="[910,979,1216,1243]" captionStart="Figs 1–10" captionStartId="6.[189,242,1889,1915]" captionTargetBox="[222,1352,267,1823]" captionTargetId="figure-75@6.[219,1368,265,1849]" captionTargetPageId="6" captionText="Figs 1–10. Tonyattalus vanharteni gen. et sp. nov., holotype (SCH_ISEA_000133), ♂ (1–8), ♀ (9– 10). 1, 9. External appearance, dorsal view. 2, 10. External appearance, lateral view. 3. Left antenna. 4. Head and pronotum, dorsal view. 5. Left anterior tarsus. 6. Pygidium. 7. Ultimate abdominal ventrite. 8. Aedeagus, and tegmen, lateral view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6518293" httpUri="https://zenodo.org/record/6518293/files/figure.png" pageId="4" pageNumber="62">Fig. 7</figureCitation>
|
||
) strongly narrowed, bilaciniate, but both lobes narrowly adjoined, evenly rounded distally, with round-triangular emargination in middle and sharp lateral processes; aedeagus (
|
||
<figureCitation id="0EE82A72505D5C1BFD0F977E35C26A05" box="[642,712,1287,1313]" captionStart="Figs 1–10" captionStartId="6.[189,242,1889,1915]" captionTargetBox="[222,1352,267,1823]" captionTargetId="figure-75@6.[219,1368,265,1849]" captionTargetPageId="6" captionText="Figs 1–10. Tonyattalus vanharteni gen. et sp. nov., holotype (SCH_ISEA_000133), ♂ (1–8), ♀ (9– 10). 1, 9. External appearance, dorsal view. 2, 10. External appearance, lateral view. 3. Left antenna. 4. Head and pronotum, dorsal view. 5. Left anterior tarsus. 6. Pygidium. 7. Ultimate abdominal ventrite. 8. Aedeagus, and tegmen, lateral view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6518293" httpUri="https://zenodo.org/record/6518293/files/figure.png" pageId="4" pageNumber="62">Fig. 8</figureCitation>
|
||
) elongate and strongly curved dorsally, typical of
|
||
<taxonomicName id="51D34D74505D5C1BFA98977E32786A05" authorityName="Abeille de Perrin" authorityYear="1890" box="[1301,1394,1287,1313]" class="Insecta" family="Melyridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="62" phylum="Arthropoda" rank="tribe" tribe="Attalini">Attalini</taxonomicName>
|
||
, with a long sharpened bristle in the inner sac in middle of distal part, tegmen wide, transverse, with thin short appendages.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DEC9657C505D5C1BFF3097F635256939" pageId="4" pageNumber="62" type="etymology">
|
||
<paragraph id="966C36F7505D5C1BFF3097F6364F6A8D" blockId="4.[189,325,1423,1449]" box="[189,325,1423,1449]" pageId="4" pageNumber="62">
|
||
<heading id="CD24819B505D5C1BFF3097F6364F6A8D" bold="true" box="[189,325,1423,1449]" fontSize="11" level="3" pageId="4" pageNumber="62" reason="3">
|
||
<emphasis id="A4A7EAE5505D5C1BFF3097F6364F6A8D" bold="true" box="[189,325,1423,1449]" pageId="4" pageNumber="62">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="966C36F7505D5C1BFF3097C435256939" blockId="4.[189,1399,1469,1566]" pageId="4" pageNumber="62">
|
||
The new genus ‘
|
||
<taxonomicName id="51D34D74505D5C1BFE0997C435076AF3" authorityName="Tshernyshev" authorityYear="2022" box="[388,525,1469,1495]" class="Insecta" family="Melyridae" genus="Tonyattalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="62" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A4A7EAE5505D5C1BFE0997C435076AF3" box="[388,525,1469,1495]" italics="true" pageId="4" pageNumber="62">Tonyattalus</emphasis>
|
||
</taxonomicName>
|
||
’, is composed of two words, ‘
|
||
<emphasis id="A4A7EAE5505D5C1BFCF697C734B96AF3" box="[891,947,1470,1495]" italics="true" pageId="4" pageNumber="62">Tony</emphasis>
|
||
’
|
||
<emphasis id="A4A7EAE5505D5C1BFC4B97C734DF6AF3" box="[966,981,1470,1495]" italics="true" pageId="4" pageNumber="62">–</emphasis>
|
||
nickname of Antonius van Harten, enthusiastic organiser and participant of the
|
||
<collectingCountry id="EEC47667505D5C1BFD559799341E6ADE" box="[728,788,1504,1530]" name="United Arab Emirates" pageId="4" pageNumber="62">UAE</collectingCountry>
|
||
Insect Project, and ‘
|
||
<taxonomicName id="51D34D74505D5C1BFB9C9799336F6ADE" authorityName="Erichson" authorityYear="1840" box="[1041,1125,1504,1530]" class="Insecta" family="Melyridae" genus="Attalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="62" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A4A7EAE5505D5C1BFB9C9799336F6ADE" box="[1041,1125,1504,1530]" italics="true" pageId="4" pageNumber="62">Attalus</emphasis>
|
||
</taxonomicName>
|
||
’ – name of the genus, nominative to the tribe
|
||
<taxonomicName id="51D34D74505D5C1BFE40947A35216939" authorityName="Abeille de Perrin" authorityYear="1890" box="[461,555,1539,1565]" class="Insecta" family="Melyridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="62" phylum="Arthropoda" rank="tribe" tribe="Attalini">Attalini</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DEC9657C505D5C1BFF30943C35FF68A0" pageId="4" pageNumber="62" type="description">
|
||
<paragraph id="966C36F7505D5C1BFF30943C3653697B" blockId="4.[189,345,1605,1631]" box="[189,345,1605,1631]" pageId="4" pageNumber="62">
|
||
<heading id="CD24819B505D5C1BFF30943C3653697B" bold="true" box="[189,345,1605,1631]" fontSize="11" level="3" pageId="4" pageNumber="62" reason="3">
|
||
<emphasis id="A4A7EAE5505D5C1BFF30943C3653697B" bold="true" box="[189,345,1605,1631]" pageId="4" pageNumber="62">Comparison</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="966C36F7505D5C1BFF30940A35FF68A0" blockId="4.[188,1401,1651,1924]" pageId="4" pageNumber="62">
|
||
The species of the new genus should be compared with species of
|
||
<taxonomicName id="51D34D74505D5C1BFC2C940A33A469AA" authority="Erichson, 1840" authorityName="Erichson" authorityYear="1840" box="[929,1198,1651,1678]" class="Insecta" family="Melyridae" genus="Attalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="62" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A4A7EAE5505D5C1BFC2C940A34FF69A9" box="[929,1013,1651,1677]" italics="true" pageId="4" pageNumber="62">Attalus</emphasis>
|
||
Erichson, 1840
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="51D34D74505D5C1BFB35940A36886995" authority="Champion, 1921" authorityName="Champion" authorityYear="1921" class="Insecta" family="Melyridae" genus="Malachiomimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="62" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A4A7EAE5505D5C1BFB35940A327D69A9" box="[1208,1399,1651,1677]" italics="true" pageId="4" pageNumber="62">Malachiomimus</emphasis>
|
||
Champion, 1921
|
||
</taxonomicName>
|
||
, and
|
||
<emphasis id="A4A7EAE5505D5C1BFE3194EE35256995" box="[444,559,1687,1713]" italics="true" pageId="4" pageNumber="62">Nepachys</emphasis>
|
||
Thomson, 1859 from which they can be distinguished by a combination of the following characters: size small, ca
|
||
<quantity id="512B9B12505D5C1BFD2294C335FE69F0" box="[687,756,1722,1748]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="62" unit="mm" value="2.0">2 mm</quantity>
|
||
in length; the elytra abbreviated in both sexes and not covering abdomen completely; apices of elytra simple, lacking impressions; 2
|
||
<superScript id="61A69BBF505D5C1BFBEE94A5337E69CE" attach="left" box="[1123,1140,1756,1770]" fontSize="6" pageId="4" pageNumber="62">nd</superScript>
|
||
tarsomere in anterior legs of male with simple small transverse comb above; metathorax weakly swollen, with specific straight, edged and slightly lamellate outer side; aedeagus thin and strongly curved, ultimate ventrite with round emargination and thin sharp processes, the structures necessary to hold curved aedeagus during copulation, is typical of the tribe
|
||
<taxonomicName id="51D34D74505D5C1BFD19951335F868A0" authorityName="Abeille de Perrin" authorityYear="1890" box="[660,754,1898,1924]" class="Insecta" family="Melyridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="62" phylum="Arthropoda" rank="tribe" tribe="Attalini">Attalini</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DEC9657C505C5C1AFF30937035BE6B28" pageId="5" pageNumber="63" type="discussion">
|
||
<paragraph id="966C36F7505C5C1AFF309370360E6E07" blockId="5.[189,260,265,291]" box="[189,260,265,291]" pageId="5" pageNumber="63">
|
||
<heading id="CD24819B505C5C1AFF309370360E6E07" bold="true" box="[189,260,265,291]" fontSize="11" level="3" pageId="5" pageNumber="63" reason="3">
|
||
<emphasis id="A4A7EAE5505C5C1AFF309370360E6E07" bold="true" box="[189,260,265,291]" pageId="5" pageNumber="63">Notes</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="966C36F7505C5C1AFF30934E36E46DF0" blockId="5.[189,1399,311,724]" pageId="5" pageNumber="63">
|
||
The genus
|
||
<taxonomicName id="51D34D74505C5C1AFEB3934E36CD6E75" authority="Tshernyshev, 2022" authorityName="Tshernyshev" authorityYear="2022" box="[318,455,311,337]" class="Insecta" family="Melyridae" genus="Tonyattalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="63" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="A4A7EAE5505C5C1AFEB3934E36CD6E75" box="[318,455,311,337]" italics="true" pageId="5" pageNumber="63">Tonyattalus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BF94579E505C5C1AFE43934135336E76" box="[462,569,312,338]" pageId="5" pageNumber="63" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
includes two species known only from the Arabian Peninsula. Besides the nominal species, the other closely related species is
|
||
<taxonomicName id="51D34D74505C5C1AFCB99322338F6E50" authority="(Wittmer, 1980)" baseAuthorityName="Wittmer" baseAuthorityYear="1980" box="[820,1157,346,372]" class="Insecta" family="Melyridae" genus="Malachiomimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="63" phylum="Arthropoda" rank="species" species="buettikeri">
|
||
<emphasis id="A4A7EAE5505C5C1AFCB9932234CB6E50" box="[820,961,346,372]" italics="true" pageId="5" pageNumber="63">T. buettikeri</emphasis>
|
||
(
|
||
<bibRefCitation id="F2424B06505C5C1AFC5D932333776E50" author="Wittmer W." box="[976,1149,346,372]" pageId="5" pageNumber="63" pagination="114 - 118" refId="ref11519" refString="Wittmer W. 1980. Fauna of Saudi Arabia. Coleoptera: Fam. Drilidae, Malachiidae, Prionoceridae (2. Beitrag). Fauna of Saudi Arabia 2: 114 - 118." type="journal article" year="1980">Wittmer, 1980</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, originally described as
|
||
<taxonomicName id="51D34D74505C5C1AFF53930435E56EBC" authority="Wittmer, 1980" authorityName="Wittmer" authorityYear="1980" box="[222,751,381,408]" class="Insecta" family="Melyridae" genus="Attalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="63" phylum="Arthropoda" rank="species" species="buettikeri" subGenus="Nepachys">
|
||
<emphasis id="A4A7EAE5505C5C1AFF53930436386EB3" box="[222,306,381,407]" italics="true" pageId="5" pageNumber="63">Attalus</emphasis>
|
||
(
|
||
<emphasis id="A4A7EAE5505C5C1AFEC8930436B26EB3" box="[325,440,381,407]" italics="true" pageId="5" pageNumber="63">Nepachys</emphasis>
|
||
)
|
||
<emphasis id="A4A7EAE5505C5C1AFE46930435376EB3" box="[459,573,381,407]" italics="true" pageId="5" pageNumber="63">buettikeri</emphasis>
|
||
<bibRefCitation id="F2424B06505C5C1AFDC8930435E56EBC" author="Wittmer W." box="[581,751,381,408]" pageId="5" pageNumber="63" pagination="114 - 118" refId="ref11519" refString="Wittmer W. 1980. Fauna of Saudi Arabia. Coleoptera: Fam. Drilidae, Malachiidae, Prionoceridae (2. Beitrag). Fauna of Saudi Arabia 2: 114 - 118." type="journal article" year="1980">Wittmer, 1980</bibRefCitation>
|
||
</taxonomicName>
|
||
due to external characters of male, flabellate antennae and anterior tarsi possessing a comb above the 2
|
||
<superScript id="61A69BBF505C5C1AFC8693D934166E8A" attach="left" box="[779,796,416,430]" fontSize="6" pageId="5" pageNumber="63">nd</superScript>
|
||
tarsomere. Simple elytra lacking impressions and appendages in apices questions the generic attribution. Later,
|
||
<bibRefCitation id="F2424B06505C5C1AFC2C93BD33316EFA" author="Evers A. M. J." box="[929,1083,452,478]" pageId="5" pageNumber="63" pagination="1 - 57" refId="ref10729" refString="Evers A. M. J. 1989. Phylogenese der Malachiidae (Col.) der Welt Kladistik und Biogeographie aus der Sicht der Pangaea-Theorie. Entomologische Blatter fur Biologie und Systematik der Kafer 85: 1 - 57." type="journal article" year="1989">Evers (1989)</bibRefCitation>
|
||
transferred this species to the genus
|
||
<taxonomicName id="51D34D74505C5C1AFEB1939E35C46D25" authority="Champion, 1921" authorityName="Champion" authorityYear="1921" box="[316,718,487,513]" class="Insecta" family="Melyridae" genus="Malachiomimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="63" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A4A7EAE5505C5C1AFEB1939E36F16D25" box="[316,507,487,513]" italics="true" pageId="5" pageNumber="63">Malachiomimus</emphasis>
|
||
Champion, 1921
|
||
</taxonomicName>
|
||
based on the flabellate antennae of the male. This new combination is doubtful and confuses the systematics of the Apalochrini because of strong differences of a
|
||
<typeStatus id="49688855505C5C1AFF76905736246D6C" box="[251,302,558,584]" pageId="5" pageNumber="63">type</typeStatus>
|
||
species of the genus
|
||
<taxonomicName id="51D34D74505C5C1AFDB3905435F76D63" authorityName="Champion" authorityYear="1921" box="[574,765,557,583]" class="Insecta" family="Melyridae" genus="Malachiomimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="63" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A4A7EAE5505C5C1AFDB3905435F76D63" box="[574,765,557,583]" italics="true" pageId="5" pageNumber="63">Malachiomimus</emphasis>
|
||
</taxonomicName>
|
||
, occurring in Himalaya Mountains when compared with the Arabian ‘
|
||
<taxonomicName id="51D34D74505C5C1AFE14902836E76D4F" authorityName="Erichson" authorityYear="1840" box="[409,493,593,619]" class="Insecta" family="Melyridae" genus="Attalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="63" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A4A7EAE5505C5C1AFE14902836E76D4F" box="[409,493,593,619]" italics="true" pageId="5" pageNumber="63">Attalus</emphasis>
|
||
</taxonomicName>
|
||
’. The genus
|
||
<taxonomicName id="51D34D74505C5C1AFD02902834446D4F" authorityName="Champion" authorityYear="1921" box="[655,846,593,619]" class="Insecta" family="Melyridae" genus="Malachiomimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="63" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A4A7EAE5505C5C1AFD02902834446D4F" box="[655,846,593,619]" italics="true" pageId="5" pageNumber="63">Malachiomimus</emphasis>
|
||
</taxonomicName>
|
||
is characterized by its considerably large size (ca
|
||
<quantity id="512B9B12505C5C1AFF64900D36416DAA" box="[233,331,628,655]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="5" pageNumber="63" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5-6 mm</quantity>
|
||
), wide, enlarged and transverse tarsal comb in anterior legs of male and simple pygidium; the body is elongated and subparallel, and the abdomen is completely covered by elytra, typical of Malachiinae-type species.
|
||
</paragraph>
|
||
<paragraph id="966C36F7505C5C1AFF30908535BE6B28" blockId="5.[188,1400,764,1037]" pageId="5" pageNumber="63">
|
||
After the revision of Himalayan and SE Asian species of the genera
|
||
<taxonomicName id="51D34D74505C5C1AFC71908532496C32" authority="Wittmer, 1966" authorityName="Wittmer" authorityYear="1966" box="[1020,1347,764,790]" class="Insecta" family="Melyridae" genus="Sceloattalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="63" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A4A7EAE5505C5C1AFC71908533876C32" box="[1020,1165,764,790]" italics="true" pageId="5" pageNumber="63">Sceloattalus</emphasis>
|
||
Wittmer, 1966
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="51D34D74505C5C1AFF309166352C6C1D" authority="Pic, 1921" authorityName="Pic" authorityYear="1921" box="[189,550,799,825]" class="Insecta" family="Melyridae" genus="Dromanthomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="63" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A4A7EAE5505C5C1AFF30916636A66C1D" box="[189,428,799,825]" italics="true" pageId="5" pageNumber="63">Dromanthomorphus</emphasis>
|
||
Pic, 1921
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="F2424B06505C5C1AFDBB9166342A6C1E" author="Tshernyshev S. E." box="[566,800,799,826]" pageId="5" pageNumber="63" pagination="358 - 374" refId="ref11163" refString="Tshernyshev S. E. 2015 a. A review of species of the genus Apalochrus Erichson (Coleoptera, Malachiidae). Zootaxa 3941 (3): 358 - 374. https: // doi. org / 10.11646 / zootaxa. 3941.3.3" type="journal article" year="2015">Tshernyshev 2015a</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="F2424B06505C5C1AFCA0916634736C1E" author="Tshernyshev S. E." box="[813,889,799,826]" pageId="5" pageNumber="63" pagination="389 - 405" refId="ref11208" refString="Tshernyshev S. E. 2015 b. Soft-winged flower beetles (Coleoptera, Malachiidae) of the Himalaya region, with notes on Apalochrini. In: Hartmann M. & Weipert J. (eds) Biodiversitat und Naturausstattung im Himalaya, Volume 5: 389 - 405. Verein der Freunde & Forderer des Naturkundemuseums Erfurt e. V." type="journal article" year="2015">2015b</bibRefCitation>
|
||
, 2016),
|
||
<taxonomicName id="51D34D74505C5C1AFC51916633916C1D" authorityName="Champion" authorityYear="1921" box="[988,1179,799,825]" class="Insecta" family="Melyridae" genus="Malachiomimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="63" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A4A7EAE5505C5C1AFC51916633916C1D" box="[988,1179,799,825]" italics="true" pageId="5" pageNumber="63">Malachiomimus</emphasis>
|
||
</taxonomicName>
|
||
is considered as a Himalayan endemic, so the presence of some species in the Arabian Peninsula looks very strange. The specific appearance with abbreviated elytra, of the so-called ‘Carphurinae-type’, is similar to species of
|
||
<taxonomicName id="51D34D74505C5C1AFF5091F1368B6C86" authorityName="Champion" authorityYear="1923" box="[221,385,904,930]" class="Insecta" family="Melyridae" genus="Carphuroides" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="63" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A4A7EAE5505C5C1AFF5091F1368B6C86" box="[221,385,904,930]" italics="true" pageId="5" pageNumber="63">Carphuroides</emphasis>
|
||
</taxonomicName>
|
||
, but the tarsal comb above the 2
|
||
<superScript id="61A69BBF505C5C1AFC8F91F134196CB2" attach="left" box="[770,787,904,918]" fontSize="6" pageId="5" pageNumber="63">nd</superScript>
|
||
tarsomere allows this species to be attributed to the subfamily
|
||
<taxonomicName id="51D34D74505C5C1AFEB491D536C06CE2" authorityName="Fleming" authorityYear="1821" box="[313,458,940,966]" class="Insecta" family="Melyridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="63" phylum="Arthropoda" rank="subFamily" subFamily="Malachiinae">Malachiinae</taxonomicName>
|
||
and compared only with representatives of the tribe
|
||
<taxonomicName id="51D34D74505C5C1AFBA491D5338D6CE2" authorityName="Abeille de Perrin" authorityYear="1890" box="[1065,1159,940,966]" class="Insecta" family="Melyridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="63" phylum="Arthropoda" rank="tribe" tribe="Attalini">Attalini</taxonomicName>
|
||
. Formerly, there was no genus with characters compatible with the Arabian ‘
|
||
<taxonomicName id="51D34D74505C5C1AFCD791B634A46CCD" authorityName="Erichson" authorityYear="1840" box="[858,942,975,1001]" class="Insecta" family="Melyridae" genus="Attalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="63" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A4A7EAE5505C5C1AFCD791B634A46CCD" box="[858,942,975,1001]" italics="true" pageId="5" pageNumber="63">Attalus</emphasis>
|
||
</taxonomicName>
|
||
’; therefore, a new genus,
|
||
<taxonomicName id="51D34D74505C5C1AFB6091B6327C6CCD" authority="Tshernyshev, 2022" authorityName="Tshernyshev" authorityYear="2022" box="[1261,1398,975,1001]" class="Insecta" family="Melyridae" genus="Tonyattalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="63" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="A4A7EAE5505C5C1AFB6091B6327C6CCD" box="[1261,1398,975,1001]" italics="true" pageId="5" pageNumber="63">Tonyattalus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BF94579E505C5C1AFF30918A36226B29" box="[189,296,1011,1037]" pageId="5" pageNumber="63" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
is proposed and described above.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DEC9657C505C5C1AFF30964A35F06BBB" pageId="5" pageNumber="63" type="reference_group">
|
||
<paragraph id="966C36F7505C5C1AFF30964A34136B6A" blockId="5.[189,793,1075,1102]" box="[189,793,1075,1102]" pageId="5" pageNumber="63">
|
||
<heading id="CD24819B505C5C1AFF30964A34136B6A" bold="true" box="[189,793,1075,1102]" fontSize="11" level="3" pageId="5" pageNumber="63" reason="3">
|
||
<emphasis id="A4A7EAE5505C5C1AFF30964A34136B6A" bold="true" box="[189,793,1075,1102]" pageId="5" pageNumber="63">
|
||
List of species in the genus
|
||
<taxonomicName id="51D34D74505C5C1AFD99964A35A86B6A" authority="Tshernyshev, 2022" authorityName="Tshernyshev" authorityYear="2022" box="[532,674,1075,1102]" class="Insecta" family="Melyridae" genus="Tonyattalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="63" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="A4A7EAE5505C5C1AFD99964A35A86B6A" bold="true" box="[532,674,1075,1102]" italics="true" pageId="5" pageNumber="63">Tonyattalus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BF94579E505C5C1AFD27964D34136B6A" box="[682,793,1076,1102]" pageId="5" pageNumber="63" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="966C36F7505C5C1AFF30961B33216B59" blockId="5.[189,1067,1122,1184]" box="[189,1067,1122,1149]" pageId="5" pageNumber="63">
|
||
1.
|
||
<taxonomicName id="51D34D74505C5C1AFF57961B35AA6B58" authority="(Wittmer, 1980)" authorityName="Tshernyshev" authorityYear="2022" baseAuthorityName="Wittmer" baseAuthorityYear="1980" box="[218,672,1122,1148]" class="Insecta" family="Melyridae" genus="Tonyattalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="63" phylum="Arthropoda" rank="species" species="buettikeri" status="gen. et comb nov.">
|
||
<emphasis id="A4A7EAE5505C5C1AFF57961B36D76B58" box="[218,477,1122,1148]" italics="true" pageId="5" pageNumber="63">Tonyattalus buettikeri</emphasis>
|
||
(
|
||
<bibRefCitation id="F2424B06505C5C1AFE61961B35926B58" author="Wittmer W." box="[492,664,1122,1148]" pageId="5" pageNumber="63" pagination="114 - 118" refId="ref11519" refString="Wittmer W. 1980. Fauna of Saudi Arabia. Coleoptera: Fam. Drilidae, Malachiidae, Prionoceridae (2. Beitrag). Fauna of Saudi Arabia 2: 114 - 118." type="journal article" year="1980">Wittmer, 1980</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BF94579E505C5C1AFD25961A34726B59" box="[680,888,1122,1149]" pageId="5" pageNumber="63" rank="species">gen. et comb nov.</taxonomicNameLabel>
|
||
(
|
||
<collectingCountry id="EEC47667505C5C1AFC0A961B33296B59" box="[903,1059,1122,1149]" name="Saudi Arabia" pageId="5" pageNumber="63">Saudi Arabia</collectingCountry>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="966C36F7505C5C1AFF3096FC35F06BBB" blockId="5.[189,1067,1122,1184]" box="[189,762,1157,1184]" pageId="5" pageNumber="63">
|
||
2.
|
||
<taxonomicName id="51D34D74505C5C1AFF5796FC36E16BBB" authority="Tshernyshev, 2022" authorityName="Tshernyshev" authorityYear="2022" box="[218,491,1157,1183]" class="Insecta" family="Melyridae" genus="Tonyattalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="63" phylum="Arthropoda" rank="species" species="vanharteni" status="gen. et sp. nov.">
|
||
<emphasis id="A4A7EAE5505C5C1AFF5796FC36E16BBB" box="[218,491,1157,1183]" italics="true" pageId="5" pageNumber="63">Tonyattalus vanharteni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BF94579E505C5C1AFE7E96FF35A96B84" box="[499,675,1158,1184]" pageId="5" pageNumber="63" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
(
|
||
<collectingCountry id="EEC47667505C5C1AFD3F96FC35F86BBB" box="[690,754,1157,1183]" name="United Arab Emirates" pageId="5" pageNumber="63">UAE</collectingCountry>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |