272 lines
35 KiB
XML
272 lines
35 KiB
XML
<document id="63BF86127862E43176D13D34C75011E6" ID-CLB-Dataset="55716" ID-DOI="10.5852/ejt.2022.817.1763" ID-GBIF-Dataset="daf1b8a9-0042-4b39-8f9c-df87cb1603f5" ID-ISSN="2118-9773" ID-Zenodo-Dep="6518289" ID-ZooBank="22C5B528-F7D7-40F2-8877-C30FF6057157" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1651674041341" checkinUser="felipe" docAuthor="Tshernyshev, Sergei E." docDate="2022" docId="1E7A87E150515C16FD1496973330688A" docLanguage="en" docName="ejt-817_58-77.pdf" docOrigin="European Journal of Taxonomy 817" docSource="http://dx.doi.org/10.5852/ejt.2022.817.1763" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Tonycolotes Tshernyshev 2022, gen. nov." docType="treatment" docUuid="4B5B95CC-00CD-47E8-9A6F-9B83E85850C9" docUuidSource="ZooBank" docVersion="9" lastPageNumber="67" masterDocId="E243FF9950595C1FFF8D9279370A6F24" masterDocTitle="Soft-winged flower beetles (Coleoptera: Malachiidae) of the United Arab Emirates" masterLastPageNumber="77" masterPageNumber="58" pageNumber="66" updateTime="1699329578614" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="1EA9993B2CADAD3F006F07586AE9472C">
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<mods:title id="04A66D63A231CF2DB3FD231D4A63C0B0">Soft-winged flower beetles (Coleoptera: Malachiidae) of the United Arab Emirates</mods:title>
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</mods:titleInfo>
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<mods:name id="5503687C9FF4DE1A4FC7EF813BF593D2" type="personal">
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<mods:namePart id="3A6B1CFF4A0D4EAAB984E890A2751152">Tshernyshev, Sergei E.</mods:namePart>
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<mods:affiliation id="B4BA5059C50ABD97A184AF5F9096CA0E">Institute of Systematics and Ecology of Animals, Siberian Branch of the Russian Academy of Sciences, Frunze Street 11, 630091 Novosibirsk, Russia. Tomsk State University, Lenina prospekt 36, Tomsk 634050, Russia. Email: sch-sch @ mail. ru</mods:affiliation>
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<mods:nameIdentifier id="4439A640CAA9BCB32FE5D972C910ABDB" type="email">sch-sch@mail.ru</mods:nameIdentifier>
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<mods:title id="C72BDB409EA68DE0A9AF066E783E5E94">European Journal of Taxonomy</mods:title>
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<mods:part id="DDF385751B18195F2DB914CE6AA8180B">
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<mods:date id="4104D811369A64916630384D0BDB766B">2022</mods:date>
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<mods:number id="8B9A267E754C4BD071ECEFE894BC9CFF">2022-04-29</mods:number>
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<mods:number id="495C429CD7CABAB5E9459634DDA7962A">817</mods:number>
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<mods:url id="B21406AB01E6A0F9B01F3E1E66B6D2CD">http://dx.doi.org/10.5852/ejt.2022.817.1763</mods:url>
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<treatment id="1E7A87E150515C16FD1496973330688A" ID-DOI="http://doi.org/10.5281/zenodo.6521829" ID-GBIF-Taxon="195191941" ID-Zenodo-Dep="6521829" LSID="urn:lsid:zoobank.org:act:4B5B95CC-00CD-47E8-9A6F-9B83E85850C9" httpUri="http://treatment.plazi.org/id/1E7A87E150515C16FD1496973330688A" lastPageId="9" lastPageNumber="67" pageId="8" pageNumber="66">
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<subSubSection id="DEC9657C50515C17FD14969734916A2D" box="[665,923,1262,1289]" pageId="8" pageNumber="66" type="nomenclature">
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<paragraph id="966C36F750515C17FD14969734916A2D" blockId="8.[365,1223,1262,1324]" box="[665,923,1262,1289]" pageId="8" pageNumber="66">
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<heading id="CD24819B50515C17FD14969734916A2D" box="[665,923,1262,1289]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="66" reason="2">
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<taxonomicName id="51D34D7450515C17FD14969734236A2D" authority="Tshernyshev, 2022" authorityName="Tshernyshev" authorityYear="2022" box="[665,809,1262,1289]" class="Insecta" family="Malachiidae" genus="Tonycolotes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="66" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="A4A7EAE550515C17FD14969734236A2D" bold="true" box="[665,809,1262,1289]" italics="true" pageId="8" pageNumber="66">Tonycolotes</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="BF94579E50515C17FCBD969634916A2D" box="[816,923,1263,1289]" pageId="8" pageNumber="66" rank="genus">gen. nov.</taxonomicNameLabel>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="DEC9657C50515C17FEE0976833CD6A08" box="[365,1223,1297,1324]" pageId="8" pageNumber="66" type="description">
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<paragraph id="966C36F750515C17FEE0976833CD6A08" blockId="8.[365,1223,1262,1324]" box="[365,1223,1297,1324]" pageId="8" pageNumber="66">
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<uri id="E2423AF550515C17FEE0976833CD6A08" box="[365,1223,1297,1324]" pageId="8" pageNumber="66">
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urn:lsid:zoobank.org:act:
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<uuid id="E2750C2250515C17FD02976833CD6A08" box="[655,1223,1297,1324]" pageId="8" pageNumber="66">4B5B95CC-00CD-47E8-9A6F-9B83E85850C9</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="DEC9657C50515C17FF30972A33206AB8" pageId="8" pageNumber="66" type="type_taxon">
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<paragraph id="966C36F750515C17FF30972A36536A49" blockId="8.[189,345,1363,1389]" box="[189,345,1363,1389]" pageId="8" pageNumber="66">
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<heading id="CD24819B50515C17FF30972A36536A49" bold="true" box="[189,345,1363,1389]" fontSize="11" level="3" pageId="8" pageNumber="66" reason="3">
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<emphasis id="A4A7EAE550515C17FF30972A36536A49" bold="true" box="[189,345,1363,1389]" pageId="8" pageNumber="66">
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<typeStatus id="4968885550515C17FF30972A37F16A49" box="[189,251,1363,1389]" pageId="8" pageNumber="66">Type</typeStatus>
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species
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="966C36F750515C17FF3097F833206AB8" blockId="8.[189,1066,1409,1438]" box="[189,1066,1409,1438]" pageId="8" pageNumber="66">
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<taxonomicName id="51D34D7450515C17FF3097F835036AB8" authority="Svihla, 1987" authorityName="Svihla" authorityYear="1987" box="[189,521,1409,1438]" class="Insecta" family="Melyridae" genus="Colotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="66" phylum="Arthropoda" rank="species" species="kovari">
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<emphasis id="A4A7EAE550515C17FF3097F836616ABF" box="[189,363,1409,1435]" italics="true" pageId="8" pageNumber="66">Colotes kovari</emphasis>
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Švihla, 1987
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</taxonomicName>
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, designated by monotypy in the present paper.
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</paragraph>
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</subSubSection>
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<subSubSection id="DEC9657C50515C16FF3097BA32606C75" lastPageId="9" lastPageNumber="67" pageId="8" pageNumber="66" type="diagnosis">
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<paragraph id="966C36F750515C17FF3097BA363D6AF9" blockId="8.[189,311,1475,1501]" box="[189,311,1475,1501]" pageId="8" pageNumber="66">
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<heading id="CD24819B50515C17FF3097BA363D6AF9" bold="true" box="[189,311,1475,1501]" fontSize="11" level="3" pageId="8" pageNumber="66" reason="3">
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<emphasis id="A4A7EAE550515C17FF3097BA363D6AF9" bold="true" box="[189,311,1475,1501]" pageId="8" pageNumber="66">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="966C36F750515C17FF30978833FF69FB" blockId="8.[189,1399,1521,1759]" pageId="8" pageNumber="66">
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Moderately small-sized (ca
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<quantity id="512B9B1250515C17FD8F9788359A6928" box="[514,656,1521,1548]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="8" pageNumber="66" unit="mm" value="1.6" valueMax="1.7" valueMin="1.5">1.5–1.7 mm</quantity>
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) malachiid beetles with the body subparallel and oval (
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<figureCitation id="0EE82A7250515C17FA9C978837D4690B" captionStart="Figs 11–24" captionStartId="11.[189,242,1857,1883]" captionTargetBox="[220,1368,274,1787]" captionTargetId="figure-92@11.[219,1368,265,1801]" captionTargetPageId="11" captionText="Figs 11–24. Tonycolotes kovari (Švihla, 1987) gen. et. comb. nov., ♂ (SCH_ISEA). 11. External appearance, dorsal view. 12. External appearance, lateral view. 13. Head and pronotum, subdorsal view. 14. Head and pronotum, dorsal view. 15. Right antenna. 16–18. Scapus in different positions. 19. Left palp. 20. Left anterior leg. 21. Pygidium. 22. Ultimate abdominal ventrite. 23. Aedeagus, lateral view. 24. Tegmen. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6518297" httpUri="https://zenodo.org/record/6518297/files/figure.png" pageId="8" pageNumber="66">Figs 11– 12</figureCitation>
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). Colouration yellow with brown bases of scapus, two triangular spots on elytra below the middle and metathoracic segments. Antennae filiform (
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<figureCitation id="0EE82A7250515C17FD7D9441344F6977" box="[752,837,1592,1619]" captionStart="Figs 11–24" captionStartId="11.[189,242,1857,1883]" captionTargetBox="[220,1368,274,1787]" captionTargetId="figure-92@11.[219,1368,265,1801]" captionTargetPageId="11" captionText="Figs 11–24. Tonycolotes kovari (Švihla, 1987) gen. et. comb. nov., ♂ (SCH_ISEA). 11. External appearance, dorsal view. 12. External appearance, lateral view. 13. Head and pronotum, subdorsal view. 14. Head and pronotum, dorsal view. 15. Right antenna. 16–18. Scapus in different positions. 19. Left palp. 20. Left anterior leg. 21. Pygidium. 22. Ultimate abdominal ventrite. 23. Aedeagus, lateral view. 24. Tegmen. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6518297" httpUri="https://zenodo.org/record/6518297/files/figure.png" pageId="8" pageNumber="66">Fig. 15</figureCitation>
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), long, extending over the middle of the elytra; in males 1
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<superScript id="61A69BBF50515C17FEB69422364C694D" attach="left" box="[315,326,1627,1641]" fontSize="6" pageId="8" pageNumber="66">st</superScript>
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antennomere (
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<figureCitation id="0EE82A7250515C17FE77942235886952" box="[506,642,1627,1654]" captionStart="Figs 11–24" captionStartId="11.[189,242,1857,1883]" captionTargetBox="[220,1368,274,1787]" captionTargetId="figure-92@11.[219,1368,265,1801]" captionTargetPageId="11" captionText="Figs 11–24. Tonycolotes kovari (Švihla, 1987) gen. et. comb. nov., ♂ (SCH_ISEA). 11. External appearance, dorsal view. 12. External appearance, lateral view. 13. Head and pronotum, subdorsal view. 14. Head and pronotum, dorsal view. 15. Right antenna. 16–18. Scapus in different positions. 19. Left palp. 20. Left anterior leg. 21. Pygidium. 22. Ultimate abdominal ventrite. 23. Aedeagus, lateral view. 24. Tegmen. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6518297" httpUri="https://zenodo.org/record/6518297/files/figure.png" pageId="8" pageNumber="66">Figs 16–18</figureCitation>
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) enlarged, strongly compressed at base and depressed at apex, oval, 2
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<superScript id="61A69BBF50515C17FE829407362A69A8" attach="left" box="[271,288,1662,1676]" fontSize="6" pageId="8" pageNumber="66">nd</superScript>
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antennomere oval, 1.3 × as long as the 3
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<superScript id="61A69BBF50515C17FC959407342C69A8" attach="left" box="[792,806,1662,1676]" fontSize="6" pageId="8" pageNumber="66">rd</superScript>
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antennomere, the 3
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<superScript id="61A69BBF50515C17FB989407332969A8" attach="left" box="[1045,1059,1662,1676]" fontSize="6" pageId="8" pageNumber="66">rd</superScript>
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antennomere triangular, not wider than the 4
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<superScript id="61A69BBF50515C17FE0894D83698698B" attach="none" box="[389,402,1697,1711]" fontSize="6" pageId="8" pageNumber="66">th</superScript>
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, 4
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<superScript id="61A69BBF50515C17FE3994D836CB698B" attach="none" box="[436,449,1697,1711]" fontSize="6" pageId="8" pageNumber="66">th</superScript>
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–10
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<superScript id="61A69BBF50515C17FE6094D836F0698B" attach="left" box="[493,506,1697,1711]" fontSize="6" pageId="8" pageNumber="66">th</superScript>
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segments oval-cylindrical, elongate, simple and equal in length, terminal antennomere oval, evenly rounded and narrowed at apex, 1.2 × as long as 10
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<superScript id="61A69BBF50515C17FBCD94BD334769F6" attach="left" box="[1088,1101,1732,1746]" fontSize="6" pageId="8" pageNumber="66">th</superScript>
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antennomere.
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</paragraph>
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<paragraph id="966C36F750515C16FF30957F33546E63" blockId="8.[187,1399,1798,2000]" lastBlockId="9.[189,1398,265,327]" lastPageId="9" lastPageNumber="67" pageId="8" pageNumber="66">
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Head with same width or slightly narrower as pronotum, slightly elongate and prognathous, eyes small, slightly protruding over the contour of head; frons flat; inter-ocular area flat and slightly impressed, glabrous, lacking depression or protuberances, genae short and straight, lacking excavations or depressions; clypeus narrow and short, transverse, straight; labrum short and transverse; palpi in male with stout and complicatedly sculptured terminal and slightly enlarged cupulate penultimate palpomeres, in female the terminal palpomere subquadrate, slightly flattened, small, the penultimate palpomere small and subtriangular. Pronotum (
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<figureCitation id="0EE82A7250505C16FD95937035976E07" box="[536,669,265,291]" captionStart="Figs 11–24" captionStartId="11.[189,242,1857,1883]" captionTargetBox="[220,1368,274,1787]" captionTargetId="figure-92@11.[219,1368,265,1801]" captionTargetPageId="11" captionText="Figs 11–24. Tonycolotes kovari (Švihla, 1987) gen. et. comb. nov., ♂ (SCH_ISEA). 11. External appearance, dorsal view. 12. External appearance, lateral view. 13. Head and pronotum, subdorsal view. 14. Head and pronotum, dorsal view. 15. Right antenna. 16–18. Scapus in different positions. 19. Left palp. 20. Left anterior leg. 21. Pygidium. 22. Ultimate abdominal ventrite. 23. Aedeagus, lateral view. 24. Tegmen. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6518297" httpUri="https://zenodo.org/record/6518297/files/figure.png" pageId="9" pageNumber="67">Figs 11–13</figureCitation>
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) transverse, narrow, with rounded lateral sides, not sinuate but depressed at base; scutellum small, triangular, with curved and adpressed apex.
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</paragraph>
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<paragraph id="966C36F750505C16FF30931732606C75" blockId="9.[189,1399,366,850]" pageId="9" pageNumber="67">
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Elytra subparallel, evenly expanded just behind the base and evenly rounded posteriorly, apices simple; humeri indistinct and small; surface densely punctured, covered with light dense adpressed short hairs. Anterior tarsi simple, four-segmented, slightly compressed, lacking comb above the 2
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<superScript id="61A69BBF50505C16FB6393CA33F56EE5" attach="left" box="[1262,1279,435,449]" fontSize="6" pageId="9" pageNumber="67">nd</superScript>
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segment, 3
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<superScript id="61A69BBF50505C16FF4193AF37D06EC0" attach="left" box="[204,218,470,484]" fontSize="6" pageId="9" pageNumber="67">rd</superScript>
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tarsomere is the smallest, 1
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<superScript id="61A69BBF50505C16FDA193AE353D6EC1" attach="left" box="[556,567,471,485]" fontSize="6" pageId="9" pageNumber="67">st</superScript>
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is the longest, twice as long as 2
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<superScript id="61A69BBF50505C16FC5993AF34EF6EC0" attach="none" box="[980,997,470,484]" fontSize="6" pageId="9" pageNumber="67">nd</superScript>
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, 2
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<superScript id="61A69BBF50505C16FB8B93AF331D6EC0" attach="left" box="[1030,1047,470,484]" fontSize="6" pageId="9" pageNumber="67">nd</superScript>
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and 3
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<superScript id="61A69BBF50505C16FBE893AF33796EC0" attach="left" box="[1125,1139,470,484]" fontSize="6" pageId="9" pageNumber="67">rd</superScript>
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of similar shape, the terminal tarsomere strongly narrower; in anterior and posterior legs the 1
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<superScript id="61A69BBF50505C16FB979383332F6D2C" attach="left" box="[1050,1061,506,520]" fontSize="6" pageId="9" pageNumber="67">st</superScript>
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tarsomere is the longest and the penultimate is the shortest; claws thin, weakly curved and sharp, slightly widened at base; anterior tibiae impressed and flattened underneath in apical half, intermediate straight, posterior slightly curved; femora simple, not swollen or strongly curved, lacking indentation, flattened. Metathorax weakly swollen, transverse, lacking appendages or tuft of hairs, slightly bituberculate. Pygidium (
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<figureCitation id="0EE82A7250505C16FA9E90FE32676D85" box="[1299,1389,647,673]" captionStart="Figs 11–24" captionStartId="11.[189,242,1857,1883]" captionTargetBox="[220,1368,274,1787]" captionTargetId="figure-92@11.[219,1368,265,1801]" captionTargetPageId="11" captionText="Figs 11–24. Tonycolotes kovari (Švihla, 1987) gen. et. comb. nov., ♂ (SCH_ISEA). 11. External appearance, dorsal view. 12. External appearance, lateral view. 13. Head and pronotum, subdorsal view. 14. Head and pronotum, dorsal view. 15. Right antenna. 16–18. Scapus in different positions. 19. Left palp. 20. Left anterior leg. 21. Pygidium. 22. Ultimate abdominal ventrite. 23. Aedeagus, lateral view. 24. Tegmen. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6518297" httpUri="https://zenodo.org/record/6518297/files/figure.png" pageId="9" pageNumber="67">Fig. 21</figureCitation>
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) undivided, elongate, evenly narrowed and rounded, weakly emarginate in middle; ultimate ventrite (
|
||
<figureCitation id="0EE82A7250505C16FF4B90B736116DCC" box="[198,283,718,744]" captionStart="Figs 11–24" captionStartId="11.[189,242,1857,1883]" captionTargetBox="[220,1368,274,1787]" captionTargetId="figure-92@11.[219,1368,265,1801]" captionTargetPageId="11" captionText="Figs 11–24. Tonycolotes kovari (Švihla, 1987) gen. et. comb. nov., ♂ (SCH_ISEA). 11. External appearance, dorsal view. 12. External appearance, lateral view. 13. Head and pronotum, subdorsal view. 14. Head and pronotum, dorsal view. 15. Right antenna. 16–18. Scapus in different positions. 19. Left palp. 20. Left anterior leg. 21. Pygidium. 22. Ultimate abdominal ventrite. 23. Aedeagus, lateral view. 24. Tegmen. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6518297" httpUri="https://zenodo.org/record/6518297/files/figure.png" pageId="9" pageNumber="67">Fig. 22</figureCitation>
|
||
) bilaciniate, elongate and wide, evenly narrowed posteriorly, with narrow oval emargination in middle and with curved short apices, aedeagus specific, with curved under right-angle base and straight central lobe, apical lamella depressed, flat, with two sharp processes from both sides; inner sac lacking strong bristles (
|
||
<figureCitation id="0EE82A7250505C16FEFF914E36C06C75" box="[370,458,823,849]" captionStart="Figs 11–24" captionStartId="11.[189,242,1857,1883]" captionTargetBox="[220,1368,274,1787]" captionTargetId="figure-92@11.[219,1368,265,1801]" captionTargetPageId="11" captionText="Figs 11–24. Tonycolotes kovari (Švihla, 1987) gen. et. comb. nov., ♂ (SCH_ISEA). 11. External appearance, dorsal view. 12. External appearance, lateral view. 13. Head and pronotum, subdorsal view. 14. Head and pronotum, dorsal view. 15. Right antenna. 16–18. Scapus in different positions. 19. Left palp. 20. Left anterior leg. 21. Pygidium. 22. Ultimate abdominal ventrite. 23. Aedeagus, lateral view. 24. Tegmen. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6518297" httpUri="https://zenodo.org/record/6518297/files/figure.png" pageId="9" pageNumber="67">Fig. 23</figureCitation>
|
||
); tegmen wide and elongate with strongly shortened thin appendages (
|
||
<figureCitation id="0EE82A7250505C16FA88914E32566C75" box="[1285,1372,823,849]" captionStart="Figs 11–24" captionStartId="11.[189,242,1857,1883]" captionTargetBox="[220,1368,274,1787]" captionTargetId="figure-92@11.[219,1368,265,1801]" captionTargetPageId="11" captionText="Figs 11–24. Tonycolotes kovari (Švihla, 1987) gen. et. comb. nov., ♂ (SCH_ISEA). 11. External appearance, dorsal view. 12. External appearance, lateral view. 13. Head and pronotum, subdorsal view. 14. Head and pronotum, dorsal view. 15. Right antenna. 16–18. Scapus in different positions. 19. Left palp. 20. Left anterior leg. 21. Pygidium. 22. Ultimate abdominal ventrite. 23. Aedeagus, lateral view. 24. Tegmen. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6518297" httpUri="https://zenodo.org/record/6518297/files/figure.png" pageId="9" pageNumber="67">Fig. 24</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DEC9657C50505C16FF309100353D6B23" pageId="9" pageNumber="67" type="etymology">
|
||
<paragraph id="966C36F750505C16FF309100364F6CB7" blockId="9.[189,325,889,915]" box="[189,325,889,915]" pageId="9" pageNumber="67">
|
||
<heading id="CD24819B50505C16FF309100364F6CB7" bold="true" box="[189,325,889,915]" fontSize="11" level="3" pageId="9" pageNumber="67" reason="3">
|
||
<emphasis id="A4A7EAE550505C16FF309100364F6CB7" bold="true" box="[189,325,889,915]" pageId="9" pageNumber="67">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="966C36F750505C16FF3091DE353D6B23" blockId="9.[189,1399,935,1032]" pageId="9" pageNumber="67">
|
||
The new genus ‘
|
||
<taxonomicName id="51D34D7450505C16FE0E91DE351A6CE5" authority="Tshernyshev, 2022" authorityName="Tshernyshev" authorityYear="2022" box="[387,528,935,961]" class="Insecta" family="Melyridae" genus="Tonycolotes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="75" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="A4A7EAE550505C16FE0E91DE351A6CE5" box="[387,528,935,961]" italics="true" pageId="9" pageNumber="67">Tonycolotes</emphasis>
|
||
</taxonomicName>
|
||
’, is composed of two words, ‘
|
||
<emphasis id="A4A7EAE550505C16FCF191D134BE6CE5" box="[892,948,936,961]" italics="true" pageId="9" pageNumber="67">Tony</emphasis>
|
||
’
|
||
<emphasis id="A4A7EAE550505C16FC4A91D134DC6CE5" box="[967,982,936,961]" italics="true" pageId="9" pageNumber="67">–</emphasis>
|
||
nickname of Antonius van Harten, enthusiastic organiser and participant of the
|
||
<collectingCountry id="EEC4766750505C16FD4491B2340F6CC1" box="[713,773,971,997]" name="United Arab Emirates" pageId="9" pageNumber="67">UAE</collectingCountry>
|
||
Insect Project, and ‘
|
||
<taxonomicName id="51D34D7450505C16FC7591B333596CC0" baseAuthorityName="Tshernyshev" baseAuthorityYear="2018" box="[1016,1107,970,996]" class="Insecta" family="Melyridae" genus="Colotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A4A7EAE550505C16FC7591B333596CC0" box="[1016,1107,970,996]" italics="true" pageId="9" pageNumber="67">Colotes</emphasis>
|
||
</taxonomicName>
|
||
’, the name of the genus, nominative to the tribe
|
||
<taxonomicName id="51D34D7450505C16FE42919435396B23" authorityName="Abeille de Perrin" authorityYear="1890" box="[463,563,1005,1031]" class="Insecta" family="Melyridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="67" phylum="Arthropoda" rank="tribe" tribe="Colotini">Colotini</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DEC9657C50505C16FF30965634B46A6F" pageId="9" pageNumber="67" type="description">
|
||
<paragraph id="966C36F750505C16FF30965636536B6D" blockId="9.[189,345,1071,1097]" box="[189,345,1071,1097]" pageId="9" pageNumber="67">
|
||
<heading id="CD24819B50505C16FF30965636536B6D" bold="true" box="[189,345,1071,1097]" fontSize="11" level="3" pageId="9" pageNumber="67" reason="3">
|
||
<emphasis id="A4A7EAE550505C16FF30965636536B6D" bold="true" box="[189,345,1071,1097]" pageId="9" pageNumber="67">Comparison</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="966C36F750505C16FF30962734B46A6F" blockId="9.[189,1399,1117,1355]" pageId="9" pageNumber="67">
|
||
Due to the fact that the
|
||
<typeStatus id="4968885550505C16FE54962735066B5C" box="[473,524,1118,1144]" pageId="9" pageNumber="67">type</typeStatus>
|
||
species of the new genus was previously attributed to the genus
|
||
<taxonomicName id="51D34D7450505C16FA9B962432656B53" baseAuthorityName="Tshernyshev" baseAuthorityYear="2018" box="[1302,1391,1117,1143]" class="Insecta" family="Melyridae" genus="Colotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A4A7EAE550505C16FA9B962432656B53" box="[1302,1391,1117,1143]" italics="true" pageId="9" pageNumber="67">Colotes</emphasis>
|
||
</taxonomicName>
|
||
, it should be compared with
|
||
<taxonomicName id="51D34D7450505C16FD8496F8356E6BBF" baseAuthorityName="Tshernyshev" baseAuthorityYear="2018" box="[521,612,1153,1179]" class="Insecta" family="Melyridae" genus="Colotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A4A7EAE550505C16FD8496F8356E6BBF" box="[521,612,1153,1179]" italics="true" pageId="9" pageNumber="67">Colotes</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="51D34D7450505C16FD1396F834116BBF" authorityName="Wittmer" authorityYear="1979" box="[670,795,1153,1179]" class="Insecta" family="Melyridae" genus="Colotrema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A4A7EAE550505C16FD1396F834116BBF" box="[670,795,1153,1179]" italics="true" pageId="9" pageNumber="67">Colotrema</emphasis>
|
||
</taxonomicName>
|
||
, also occurring in the region, from which it can be distinguished by the following characters: the elytral apices simple, not impressed; the 1
|
||
<superScript id="61A69BBF50505C16FB4196DD33DD6B96" attach="left" box="[1228,1239,1188,1202]" fontSize="6" pageId="9" pageNumber="67">st</superScript>
|
||
antennomere strongly enlarge and modified, compressed at base and depressed at distal part; anterior tibiae impressed and flattened underneath in apical half, head flat, lacking depressions near genae; hind wings normally developed in both sexes; aedeagus with curved under right-angle base and straight central lobe, apical lamella depressed, flat, with two sharp processes from both sides.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DEC9657C50505C16FF30970B36B8681A" pageId="9" pageNumber="67" type="discussion">
|
||
<paragraph id="966C36F750505C16FF30970B360E6AA8" blockId="9.[189,260,1394,1420]" box="[189,260,1394,1420]" pageId="9" pageNumber="67">
|
||
<heading id="CD24819B50505C16FF30970B360E6AA8" bold="true" box="[189,260,1394,1420]" fontSize="11" level="3" pageId="9" pageNumber="67" reason="3">
|
||
<emphasis id="A4A7EAE550505C16FF30970B360E6AA8" bold="true" box="[189,260,1394,1420]" pageId="9" pageNumber="67">Notes</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="966C36F750505C16FF3097D836B8681A" blockId="9.[187,1399,1441,1854]" pageId="9" pageNumber="67">
|
||
Modified and enlarged palpi in male are typical of representatives of the tribe
|
||
<taxonomicName id="51D34D7450505C16FBD097D833CB6A9F" authorityName="Abeille de Perrin" authorityYear="1890" box="[1117,1217,1441,1467]" class="Insecta" family="Melyridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="67" phylum="Arthropoda" rank="tribe" tribe="Colotini">Colotini</taxonomicName>
|
||
. In the Arabian Peninsula, the tribe includes two genera,
|
||
<taxonomicName id="51D34D7450505C16FD2B97BD340B6AFA" baseAuthorityName="Tshernyshev" baseAuthorityYear="2018" box="[678,769,1476,1502]" class="Insecta" family="Melyridae" genus="Colotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A4A7EAE550505C16FD2B97BD340B6AFA" box="[678,769,1476,1502]" italics="true" pageId="9" pageNumber="67">Colotes</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="51D34D7450505C16FCB397BD34B16AFA" authorityName="Wittmer" authorityYear="1979" box="[830,955,1476,1502]" class="Insecta" family="Melyridae" genus="Colotrema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A4A7EAE550505C16FCB397BD34B16AFA" box="[830,955,1476,1502]" italics="true" pageId="9" pageNumber="67">Colotrema</emphasis>
|
||
</taxonomicName>
|
||
. The taxonomic structure of the tribe needs to be revised mainly because of taxonomic uncertainty in the nominal genus
|
||
<taxonomicName id="51D34D7450505C16FBFA979E36096900" authority="(Tshernyshev 2018)" baseAuthorityName="Tshernyshev" baseAuthorityYear="2018" class="Insecta" family="Melyridae" genus="Colotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A4A7EAE550505C16FBFA979E33D86925" box="[1143,1234,1511,1537]" italics="true" pageId="9" pageNumber="67">Colotes</emphasis>
|
||
(Tshernyshev 2018)
|
||
</taxonomicName>
|
||
. Definition of the subgeneric taxa on the basis of the only character, the shape of female palpi, results in a confusing conception of the genus which was initially identified by male special characters. Recently, the
|
||
<typeStatus id="4968885550505C16FED194283685694F" box="[348,399,1617,1643]" pageId="9" pageNumber="67">type</typeStatus>
|
||
species of the genus,
|
||
<taxonomicName id="51D34D7450505C16FD0194283310694F" authority="Erichson, 1840" authorityName="Erichson" authorityYear="1840" box="[652,1050,1617,1643]" class="Insecta" family="Melyridae" genus="Colotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="67" phylum="Arthropoda" rank="species" species="trinotatus">
|
||
<emphasis id="A4A7EAE550505C16FD019428346B694F" box="[652,865,1617,1643]" italics="true" pageId="9" pageNumber="67">Colotes trinotatus</emphasis>
|
||
Erichson, 1840
|
||
</taxonomicName>
|
||
(a junior synonym of
|
||
<taxonomicName id="51D34D7450505C16FA969428359769AB" authority="(Laporte de Castelnau, 1836))" authorityName="" baseAuthorityName="Laporte de Castelnau" baseAuthorityYear="1836" class="Insecta" family="Melyridae" genus="Colotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="67" phylum="Arthropoda" rank="species" species="maculatus">
|
||
<emphasis id="A4A7EAE550505C16FA969428363D69AA" italics="true" pageId="9" pageNumber="67">Colotes maculatus</emphasis>
|
||
(Laporte de Castelnau, 1836))
|
||
</taxonomicName>
|
||
has been re-described and illustrated in detail, and the generic characters were determined (Tshernyshev 2018); as a result, a new genus,
|
||
<emphasis id="A4A7EAE550505C16FBA494EE33DC6995" box="[1065,1238,1687,1713]" italics="true" pageId="9" pageNumber="67">Himalacolotes</emphasis>
|
||
Tshernyshev, 2018, was described for a group of Himalayan endemic species on the basis of similarity of external morphology, coloration of body and shape of male genitalia. The proposed new genus,
|
||
<taxonomicName id="51D34D7450505C16FB6794A4327D69D3" authority="Tshernyshev, 2022" authorityName="Tshernyshev" authorityYear="2022" box="[1258,1399,1757,1783]" class="Insecta" family="Melyridae" genus="Tonycolotes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="67" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="A4A7EAE550505C16FB6794A4327D69D3" box="[1258,1399,1757,1783]" italics="true" pageId="9" pageNumber="67">Tonycolotes</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BF94579E50505C16FF3095783620683F" box="[189,298,1793,1819]" pageId="9" pageNumber="67" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
, differs from the nominal species in a series of strong characters, allowing one to consider it a good separate genus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DEC9657C50505C16FF30951C3330688A" pageId="9" pageNumber="67" type="reference_group">
|
||
<paragraph id="966C36F750505C16FF30951C3411685B" blockId="9.[189,795,1893,1920]" box="[189,795,1893,1920]" pageId="9" pageNumber="67">
|
||
<heading id="CD24819B50505C16FF30951C3411685B" bold="true" box="[189,795,1893,1920]" fontSize="11" level="3" pageId="9" pageNumber="67" reason="3">
|
||
<emphasis id="A4A7EAE550505C16FF30951C3411685B" bold="true" box="[189,795,1893,1920]" pageId="9" pageNumber="67">
|
||
List of species in the genus
|
||
<taxonomicName id="51D34D7450505C16FD99951C35AE68A4" authority="Tshernyshev, 2022" authorityName="Tshernyshev" authorityYear="2022" box="[532,676,1893,1920]" class="Insecta" family="Malachiidae" genus="Tonycolotes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="66" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="A4A7EAE550505C16FD99951C35AE68A4" bold="true" box="[532,676,1893,1920]" italics="true" pageId="9" pageNumber="67">Tonycolotes</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BF94579E50505C16FD21951C3411685B" box="[684,795,1893,1919]" pageId="9" pageNumber="67" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="966C36F750505C16FF3095ED3330688A" blockId="9.[189,1082,1940,1968]" box="[189,1082,1940,1968]" pageId="9" pageNumber="67">
|
||
1.
|
||
<taxonomicName id="51D34D7450505C16FF5795ED366D688A" authority="Tshernyshev, 2022" authorityName="Tshernyshev" authorityYear="2022" box="[218,359,1940,1966]" class="Insecta" family="Malachiidae" genus="Tonycolotes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="75" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="A4A7EAE550505C16FF5795ED366D688A" box="[218,359,1940,1966]" italics="true" pageId="9" pageNumber="67">Tonycolotes</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicName id="51D34D7450505C16FEE395ED3563688A" authority="(Svihla, 1987)" baseAuthorityName="Svihla" baseAuthorityYear="1987" box="[366,617,1940,1968]" class="Insecta" family="Melyridae" genus="Colotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="67" phylum="Arthropoda" rank="species" species="kovari">
|
||
<emphasis id="A4A7EAE550505C16FEE395ED36B0688A" box="[366,442,1940,1966]" italics="true" pageId="9" pageNumber="67">kovari</emphasis>
|
||
(Švihla, 1987)
|
||
</taxonomicName>
|
||
gen. et. comb. nov. (
|
||
<collectingCountry id="EEC4766750505C16FCD295ED3498688A" box="[863,914,1940,1966]" name="Iran" pageId="9" pageNumber="67">Iran</collectingCountry>
|
||
,
|
||
<collectingCountry id="EEC4766750505C16FC1395ED34E3688A" box="[926,1001,1940,1966]" name="Oman" pageId="9" pageNumber="67">Oman</collectingCountry>
|
||
,
|
||
<collectingCountry id="EEC4766750505C16FC7995ED3338688A" box="[1012,1074,1940,1966]" name="United Arab Emirates" pageId="9" pageNumber="67">UAE</collectingCountry>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |