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<document id="B8A50332992E44F9D1BD5153FA54577E" ID-DOI="10.11646/zootaxa.4097.1.11" ID-GBIF-Dataset="47edc57f-1dbf-4578-b4fd-d044c7aca3dc" ID-ISSN="1175-5326" ID-Zenodo-Dep="270986" ID-ZooBank="C03779A8-EFF5-4181-AB6D-3343FF7F46D5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459408888046" checkinUser="plazi" docAuthor="Mohrig, Werner, Kauschke, Ellen &amp; Broadley, Adam" docDate="2016" docId="1E5CB66CFF94BC11FF76FAE5B6BFFF4C" docLanguage="en" docName="zootaxa.4097.1.11.pdf.imf" docOrigin="Zootaxa 4097 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudolycoriella skusei Mohrig, Kauschke &amp; Broadley, 2016, sp. nov." docType="treatment" docVersion="10" lastPageNumber="141" masterDocId="E265CE14FF96BC13FFE1FA72B52CFA4B" masterDocTitle="Pseudolycoriella skusei sp. nov. (Diptera: Sciaridae), a new dark-winged fungus gnat from Norfolk Island and Australia" masterLastPageNumber="142" masterPageNumber="139" pageNumber="141" updateTime="1698186847501" updateUser="plazi">
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<mods:title id="95AFF6CC1A555BACB33FED9F171C0371">Pseudolycoriella skusei sp. nov. (Diptera: Sciaridae), a new dark-winged fungus gnat from Norfolk Island and Australia</mods:title>
</mods:titleInfo>
<mods:name id="36B8357DF5708C33B5B1BFDB206192D0" type="personal">
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<mods:namePart id="BCF20CF936F208430D2ADFC27E86A959">Mohrig, Werner</mods:namePart>
</mods:name>
<mods:name id="9914A7BB1E01E9AD9FD21BFA2526CDC0" type="personal">
<mods:role id="FC4E1C9D383BAB8360B1D7ABFCE061BE">
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<mods:namePart id="97E9FC2BCAD7E16CD284DF1E1F758193">Kauschke, Ellen</mods:namePart>
</mods:name>
<mods:name id="2D7C06529EE66D55CC0C0E5996651F50" type="personal">
<mods:role id="43FA4727A82191E0BBBD308DEC092822">
<mods:roleTerm id="77F40B0B36C3425FE9EC94421AD9F418">Author</mods:roleTerm>
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<mods:namePart id="D5A75E9AEAC30CFDCB41FDF37419E6EC">Broadley, Adam</mods:namePart>
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<mods:title id="52EDF11990345BB884961327DC18D0A2">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="11D5D7E41B52B37F826B2001EF99C77B">
<mods:date id="E90F545CE29348AE43CDAFD13583F2AD">2016</mods:date>
<mods:detail id="A67AE16704F201315A7C2C9C4CFD701E" type="volume">
<mods:number id="46FE87E482D1E76E3171EA377297F0A5">4097</mods:number>
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<mods:number id="63769EF76C066086660B5F6844E31286">1</mods:number>
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<mods:classification id="FAD28D83C76077B50A4BBAC92B3F02FA">journal article</mods:classification>
<mods:identifier id="D4C96B8AC88AA7ED10EC69D009EAFD99" type="DOI">10.11646/zootaxa.4097.1.11</mods:identifier>
<mods:identifier id="F5427C58586DCE0A93F2997E606A755E" type="GBIF-Dataset">47edc57f-1dbf-4578-b4fd-d044c7aca3dc</mods:identifier>
<mods:identifier id="3A97C088C2407FD9498FC6AFA7522DA8" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="0BBB7C71202E3B8F13B1FA3E293AD729" type="Zenodo-Dep">270986</mods:identifier>
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</mods:mods>
<treatment id="1E5CB66CFF94BC11FF76FAE5B6BFFF4C" ID-DOI="http://doi.org/10.5281/zenodo.6079482" ID-GBIF-Taxon="119329260" ID-Zenodo-Dep="6079482" LSID="urn:lsid:plazi:treatment:1E5CB66CFF94BC11FF76FAE5B6BFFF4C" httpUri="http://treatment.plazi.org/id/1E5CB66CFF94BC11FF76FAE5B6BFFF4C" lastPageNumber="141" pageId="2" pageNumber="141">
<subSubSection id="DEEF54F1FF94BC11FF76FAE5B75CFAF9" box="[151,624,151,178]" pageId="2" pageNumber="141" type="nomenclature">
<paragraph id="964A077AFF94BC11FF76FAE5B75CFAF9" blockId="2.[151,1437,151,1287]" box="[151,624,151,178]" pageId="2" pageNumber="141">
<heading id="CD02B016FF94BC11FF76FAE5B75CFAF9" bold="true" box="[151,624,151,178]" fontSize="11" level="1" pageId="2" pageNumber="141" reason="1">
<emphasis id="A481DB68FF94BC11FF76FAE5B75CFAF9" bold="true" box="[151,624,151,178]" pageId="2" pageNumber="141">
<taxonomicName id="51F57CF9FF94BC11FF76FAE5B49DFAFA" ID-CoL="8DJ6F" box="[151,433,151,177]" class="Insecta" family="Sciaridae" genus="Pseudolycoriella" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="species" species="skusei" status="sp. nov.">
<emphasis id="A481DB68FF94BC11FF76FAE5B49DFAFA" bold="true" box="[151,433,151,177]" italics="true" pageId="2" pageNumber="141">Pseudolycoriella skusei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BFB26613FF94BC11FE59FAEAB73AFAF9" box="[440,534,152,178]" pageId="2" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="0ECE1BFFFF94BC11FDC4FAEAB749FAF9" box="[549,613,152,178]" captionStart="FIGURE 1" captionStartId="1.[151,244,1943,1965]" captionTargetBox="[296,1282,206,1874]" captionTargetId="figure@1.[275,1306,194,1894]" captionTargetPageId="1" captionText="FIGURE 1. Male Pseudolycoriella skusei sp. nov. (holotype). (A) Gonostylus, (B) Hypopygium, ventral view. (C) antennal flagellomeres I IV, (D) antennal flagellomere IV, (E) apical part of fore tibia, (F) apical tarsal segment with toothed claws (photobased drawings)." httpUri="https://zenodo.org/record/270987/files/figure.png" pageId="2" pageNumber="141">Fig 1</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DEEF54F1FF94BC11FF76FAC8B776FB34" pageId="2" pageNumber="141" type="materials_examined">
<paragraph id="964A077AFF94BC11FF76FAC8B776FB34" blockId="2.[151,1437,151,1287]" pageId="2" pageNumber="141">
<emphasis id="A481DB68FF94BC11FF76FAC8B456FA9A" bold="true" box="[151,378,186,209]" pageId="2" pageNumber="141">Material examined.</emphasis>
<materialsCitation id="269D0D27FF94BC11FE65FAC8B1F3FB72" ID-GBIF-Occurrence="1262421572" collectingDate="2012-12-25" collectingMethod="yellow pan trap" collectionCode="NI" collectorName="L. Mound &amp; &amp; A. Wells" country="Norfolk Island" latitude="-29.0147222" location="National Park" longitude="167.9463889" pageId="2" pageNumber="141" specimenCount="1" specimenCount-male="1" stateProvince="NORFOLK ISLAND" typeStatus="holotype">
<emphasis id="A481DB68FF94BC11FE65FAC8B4DEFA9A" bold="true" box="[388,498,186,209]" pageId="2" pageNumber="141">
<typeStatus id="494EB9D8FF94BC11FE65FAC8B4C1FA9A" box="[388,493,186,209]" pageId="2" pageNumber="141" type="holotype">Holotype</typeStatus>
,
</emphasis>
<specimenCount id="80F3CCF3FF94BC11FE1FFAC8B719FA9A" box="[510,565,186,209]" pageId="2" pageNumber="141" type="male">male</specimenCount>
.
<collectingRegion id="5431C998FF94BC11FDA2FAC8B61EFA9A" box="[579,818,186,209]" pageId="2" pageNumber="141">NORFOLK ISLAND</collectingRegion>
: Norfolk Island
<location id="932A51A1FF94BC11FC0DFAC8B1AAFA9A" LSID="urn:lsid:plazi:treatment:1E5CB66CFF94BC11FF76FAE5B6BFFF4C:932A51A1FF94BC11FC0DFAC8B1AAFA9A" box="[1004,1158,186,209]" country="Norfolk Island" name="National Park" pageId="2" pageNumber="141" stateProvince="NORFOLK ISLAND">National Park</location>
, near
<location id="932A51A1FF94BC11FB2FFAC8B0B7FA9A" LSID="urn:lsid:plazi:treatment:1E5CB66CFF94BC11FF76FAE5B6BFFF4C:932A51A1FF94BC11FB2FFAC8B0B7FA9A" box="[1230,1435,186,209]" country="Norfolk Island" name="Selwyn Pine Road" pageId="2" pageNumber="141" stateProvince="NORFOLK ISLAND">Selwyn Pine Road</location>
entrance (
<geoCoordinate id="F3C161BDFF94BC11FEE5FAAFB45EFABF" box="[260,370,221,244]" pageId="2" pageNumber="141">29°0ʹ53ʺS</geoCoordinate>
,
<geoCoordinate id="F3C161BDFF94BC11FE63FAAFB721FABF" box="[386,525,221,244]" pageId="2" pageNumber="141">167°56ʹ47ʺE</geoCoordinate>
), collected
<date id="E24B21BAFF94BC11FD6EFAAFB625FABF" box="[655,777,221,244]" pageId="2" pageNumber="141" value="2012-12-25">
<collectingDate id="F20FD852FF94BC11FD6EFAAFB625FABF" box="[655,777,221,244]" pageId="2" pageNumber="141" value="2012-12-25">25.xii.2012</collectingDate>
</date>
by
<collectorName id="3B0062ACFF94BC11FCD8FAAFB11FFABF" box="[825,1075,221,244]" pageId="2" pageNumber="141">L. Mound &amp; A. Wells</collectorName>
from
<collectingMethod id="4FB47F6DFF94BC11FB9CFAAFB000FABF" box="[1149,1324,221,244]" pageId="2" pageNumber="141">yellow pan trap</collectingMethod>
set under Norfolk Island pine tree (
<taxonomicName id="51F57CF9FF94BC11FE5FFA8DB792FB5C" box="[446,702,255,279]" class="Pinopsida" family="Araucariaceae" genus="Araucaria" kingdom="Plantae" order="Pinales" pageId="2" pageNumber="141" phylum="Tracheophyta" rank="species" species="heterophylla">
<emphasis id="A481DB68FF94BC11FE5FFA8DB792FB5C" box="[446,702,255,279]" italics="true" pageId="2" pageNumber="141">Araucaria heterophylla</emphasis>
</taxonomicName>
) as part of Norfolk Island Quarantine Survey (
<collectionCode id="F0E49FBFFF94BC11FB00FB72B02CFB5C" box="[1249,1280,256,279]" pageId="2" pageNumber="141">NI</collectionCode>
_053-11). The specimen will be lodged in the Australian National Insect Collection, CSIRO, Canberra, Australia.
</materialsCitation>
<materialsCitation id="269D0D27FF94BC11FB08FB50B77AFB34" ID-GBIF-Occurrence="1262421573" collectingDate="1997-08-08" collectingMethod="from Malaise trap" collectionCode="PWMP" collectorName="J. Seymour" country="Australia" latitude="-16.58333" location="Mt. Lewis, 37 km WSW of Port Douglas" longitude="145.26667" pageId="2" pageNumber="141" specimenCount="1" specimenCount-male="1" stateProvince="Queensland" typeStatus="paratype">
<emphasis id="A481DB68FF94BC11FB08FB50B07BFB72" bold="true" box="[1257,1367,290,313]" pageId="2" pageNumber="141">
<typeStatus id="494EB9D8FF94BC11FB08FB50B07EFB72" box="[1257,1362,290,313]" pageId="2" pageNumber="141" type="paratype">Paratype</typeStatus>
,
</emphasis>
<specimenCount id="80F3CCF3FF94BC11FA80FB50B0B4FB72" box="[1377,1432,290,313]" pageId="2" pageNumber="141" type="male">male</specimenCount>
.
<collectingCountry id="EEE247EAFF94BC11FF76FB37B41EFB17" box="[151,306,325,348]" name="Australia" pageId="2" pageNumber="141" value="Australia">AUSTRALIA</collectingCountry>
: collected
<date id="E24B21BAFF94BC11FE4CFB37B70DFB17" box="[429,545,325,348]" pageId="2" pageNumber="141" value="1997-08-08">
<collectingDate id="F20FD852FF94BC11FE4CFB37B70DFB17" box="[429,545,325,348]" pageId="2" pageNumber="141" value="1997-08-08">8.viii.1997</collectingDate>
</date>
by
<collectorName id="3B0062ACFF94BC11FDB5FB37B7FDFB17" box="[596,721,325,348]" pageId="2" pageNumber="141">J. Seymour</collectorName>
<collectingMethod id="4FB47F6DFF94BC11FD3CFB37B686FB17" box="[733,938,325,348]" pageId="2" pageNumber="141">from Malaise trap</collectingMethod>
,
<location id="932A51A1FF94BC11FC5BFB37B0BBFB17" LSID="urn:lsid:plazi:treatment:1E5CB66CFF94BC11FF76FAE5B6BFFF4C:932A51A1FF94BC11FC5BFB37B0BBFB17" box="[954,1431,325,348]" country="Australia" name="Mt. Lewis, 37 km WSW of Port Douglas" pageId="2" pageNumber="141" stateProvince="Queensland">
Mt. Lewis,
<quantity id="510DAA9FFF94BC11FBDEFB37B1A4FB17" box="[1087,1160,325,348]" metricMagnitude="4" metricUnit="m" metricValue="3.7" pageId="2" pageNumber="141" unit="km" value="37.0">37 km</quantity>
WSW of Port Douglas
</location>
,
<collectingRegion id="5431C998FF94BC11FF76FB1AB439FB34" box="[151,277,360,383]" pageId="2" pageNumber="141">Queensland</collectingRegion>
(
<geoCoordinate id="F3C161BDFF94BC11FEC2FB1AB456FB34" box="[291,378,360,383]" pageId="2" pageNumber="141">16°35ʹS</geoCoordinate>
,
<geoCoordinate id="F3C161BDFF94BC11FE67FB1AB4CBFB34" box="[390,487,360,383]" pageId="2" pageNumber="141">145°16ʹE</geoCoordinate>
)(
<collectionCode id="F0E49FBFFF94BC11FE16FB1AB762FB34" box="[503,590,360,383]" collectionName="PWMP" pageId="2" pageNumber="141">PWMP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DEEF54F1FF94BC11FF26FBF8B7B7F984" pageId="2" pageNumber="141" type="description">
<paragraph id="964A077AFF94BC11FF26FBF8B73FF867" blockId="2.[151,1437,151,1287]" pageId="2" pageNumber="141">
<emphasis id="A481DB68FF94BC11FF26FBF8B467FBEA" bold="true" box="[199,331,394,417]" pageId="2" pageNumber="141">Description</emphasis>
.
<emphasis id="A481DB68FF94BC11FEBAFBF8B4B8FBEA" bold="true" box="[347,404,394,417]" pageId="2" pageNumber="141">Male</emphasis>
.
<emphasis id="A481DB68FF94BC11FE44FBF8B4C7FBEA" bold="true" box="[421,491,394,417]" pageId="2" pageNumber="141">Head:</emphasis>
Dark brown, ovoid, mouth parts short, clypeus bulbous; eye bridge 3
<emphasis id="A481DB68FF94BC11FB15FBF8B02DFBEA" box="[1268,1281,394,417]" italics="true" pageId="2" pageNumber="141"></emphasis>
4 facets wide; antenna rather long, with yellowish-brown scape and pedicel; flagellomeres blackish-brown; necks pale, with darker ends (bicoloured); 4th flagellomere with a l/w-index of 2.2, setae bristle-like and as long as half of the diameter of basal part. Palp long, three-segmented; basal segment with 3
<emphasis id="A481DB68FF94BC11FCE1FB80B621F842" box="[768,781,498,521]" italics="true" pageId="2" pageNumber="141"></emphasis>
4 bristles and without a deepened patch of sensillae; third segment longer than basal segment.
</paragraph>
<paragraph id="964A077AFF94BC11FF26F84AB4DFF96A" blockId="2.[151,1437,151,1287]" pageId="2" pageNumber="141">
<emphasis id="A481DB68FF94BC11FF26F84AB408F804" bold="true" box="[199,292,568,591]" pageId="2" pageNumber="141">Thorax:</emphasis>
Brown; scutum with short and fine pubescence and three darker stripes; scutellum without longer marginal setae; postpronotum bare; katepisternum with darker corner. Wing brownish, with distinct veins; R1 = 3/4 R; R5 with ventral macrotrichia in the distal third; C longer than 1/2 w; y longer than x, without macrotrichia; Cu-stem shorter than x; posterior wing veins without macrotrichia. Haltere brown. Fore coxa yellowish-brown, mid and hind coxa brown; femora yellowish-brown, tibiae and tarsal segments darker brown; inner apex of fore tibia with an irregular row-like patch of bristles, not distinctly bordered; spurs of middle and hind tibia of same size, brownish and as long as the diameter of apex; claws toothed.
</paragraph>
<paragraph id="964A077AFF94BC11FF26F95FB7B7F984" blockId="2.[151,1437,151,1287]" pageId="2" pageNumber="141">
<emphasis id="A481DB68FF94BC11FF26F95FB413F90F" bold="true" box="[199,319,813,836]" pageId="2" pageNumber="141">Abdomen:</emphasis>
Brownish; tergal and sternal setae moderately short and sparse; Hypopygium dark brown, intergonocoxal space bare, inner ventral margin of gonocoxite sparsely haired, with longer setae in the apical half; gonostylus elongate, narrow, apically curved; with three distinct sub-apical dark spines of the same size and three longer whiplash hairs; the inner dorsal side with rather long setae. Tegmen pyramid-like, much higher than broad, membranous at the end; aedeagus long. Body length:
<quantity id="510DAA9FFF94BC11FDA3F9CAB7BBF984" box="[578,663,952,975]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="2" pageNumber="141" unit="mm" value="4.5">4.5 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="DEEF54F1FF94BC11FF26F9A8B1DCFE8A" pageId="2" pageNumber="141" type="discussion">
<paragraph id="964A077AFF94BC11FF26F9A8B1DCFE8A" blockId="2.[151,1437,151,1287]" pageId="2" pageNumber="141">
<emphasis id="A481DB68FF94BC11FF26F9A8B403F9BA" bold="true" box="[199,303,986,1009]" pageId="2" pageNumber="141">Remarks</emphasis>
. This large species is characterized by a long and narrow gonostylus with three subequal, long apical spines and two longer whiplash hairs; claws strongly toothed; antennal flagellomeres robust and blackish-brown, with rather long bicoloured necks; three dark stripes on mesonotum and a long pyramid-like tegmen with a membranous apex. Two species from New Caledonia (
<taxonomicName id="51F57CF9FF94BC11FDF6FE30B6E1FE12" authority="Vilkamaa, Hippa &amp; Mohrig" authorityName="Vilkamaa, Hippa &amp; Mohrig" box="[535,973,1090,1113]" class="Insecta" family="Sciaridae" genus="Pseudolycoriella" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="species" species="trispicata">
<emphasis id="A481DB68FF94BC11FDF6FE30B7B5FE12" box="[535,665,1090,1113]" italics="true" pageId="2" pageNumber="141">P. trispicata</emphasis>
Vilkamaa, Hippa &amp; Mohrig
</taxonomicName>
and
<emphasis id="A481DB68FF94BC11FBE5FE30B170FE12" box="[1028,1116,1090,1113]" italics="true" pageId="2" pageNumber="141">P. t e n ui s</emphasis>
Vilkamaa, Hippa &amp; Mohrig) also have three subapical spines, but differ significantly in having a shorter tegmen, shorter flagellomeres, and fewer whiplash hairs. The
<typeStatus id="494EB9D8FF94BC11FE8EFEFAB4E7FED4" box="[367,459,1160,1183]" pageId="2" pageNumber="141" type="paratype">paratype</typeStatus>
from Queensland differs in having a lesser contrast-rich body colour: the scape and pedicel are brownish as are the flagellomeres, all the coxae are dark brown and the legs are uniformly brownish.
</paragraph>
</subSubSection>
<subSubSection id="DEEF54F1FF94BC11FF26FEBFB6BFFF4C" pageId="2" pageNumber="141" type="etymology">
<paragraph id="964A077AFF94BC11FF26FEBFB6BFFF4C" blockId="2.[151,1437,151,1287]" pageId="2" pageNumber="141">
<emphasis id="A481DB68FF94BC11FF26FEBFB46CFEAF" bold="true" box="[199,320,1229,1252]" pageId="2" pageNumber="141">Etymology</emphasis>
. We dedicate this new species to Frederick Arthur Askew Skuse (
<date id="E24B21BAFF94BC11FBFCFEBFB1B7FEAF" box="[1053,1179,1229,1252]" pageId="2" pageNumber="141" value="1866" valueMax="1896">
1866
<emphasis id="A481DB68FF94BC11FBB5FEBFB14DFEAF" box="[1108,1121,1229,1252]" italics="true" pageId="2" pageNumber="141"></emphasis>
1896
</date>
), an English-Australian entomologist who was a pioneering worker on the Australian
<taxonomicName id="51F57CF9FF94BC11FCC8FE82B6A3FF4C" box="[809,911,1264,1287]" class="Insecta" family="Sciaridae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="family">Sciaridae</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>