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<document id="BF5F2FC2D2BF6910B606EF7C0A9829B3" ID-CLB-Dataset="46106" ID-DOI="10.11646/zootaxa.3784.3.7" ID-GBIF-Dataset="9359e557-e386-40de-b54c-8f3cd8e68ecd" ID-ISSN="1175-5326" ID-Zenodo-Dep="227368" ID-ZooBank="29C45BDD-A813-494C-A3A1-A5AF4AEADA95" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460403511369" checkinUser="plazi" docAuthor="Jałoszyński, Paweł &amp; Ślipiński, Adam" docDate="2014" docId="1E313229FFC8FF92FF3CFF73DB01FD97" docLanguage="en" docName="zt03784p280.pdf" docOrigin="Zootaxa 3784 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ruthmuelleria Jałoszyński &amp; Ślipiński, 2014, gen. nov." docType="treatment" docVersion="9" lastPageNumber="278" masterDocId="E2084A51FFCAFF91FFABFFEBDF42FFF7" masterDocTitle="Ruthmuelleria, a new genus of Carinodulini (Coleoptera: Coccinellidae: Microweiseinae) from South Africa" masterLastPageNumber="280" masterPageNumber="275" pageNumber="277" updateTime="1698299545394" updateUser="ExternalLinkService">
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<mods:title id="5D7D6D94C900FDCCBE51792EF03D9553">Ruthmuelleria, a new genus of Carinodulini (Coleoptera: Coccinellidae: Microweiseinae) from South Africa</mods:title>
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<mods:namePart id="715B39F147C829217BFE42D8DEB92D1C">Jałoszyński, Paweł</mods:namePart>
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<mods:namePart id="62AD3FE4E04B358A4F8C03920049D82A">Ślipiński, Adam</mods:namePart>
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<mods:date id="8511B7BED86CE588720E95D4816B0B2F">2014</mods:date>
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<treatment id="1E313229FFC8FF92FF3CFF73DB01FD97" ID-DOI="http://doi.org/10.5281/zenodo.6138372" ID-GBIF-Taxon="119420603" ID-Zenodo-Dep="6138372" LSID="urn:lsid:plazi:treatment:1E313229FFC8FF92FF3CFF73DB01FD97" httpUri="http://treatment.plazi.org/id/1E313229FFC8FF92FF3CFF73DB01FD97" lastPageId="3" lastPageNumber="278" pageId="2" pageNumber="277">
<subSubSection id="DE82D0B4FFC8FF93FF3CFF73DD51FF45" box="[151,531,151,178]" pageId="2" pageNumber="277" type="nomenclature">
<paragraph id="9627833FFFC8FF93FF3CFF73DD51FF45" blockId="2.[151,531,151,178]" box="[151,531,151,178]" pageId="2" pageNumber="277">
<heading id="CD6F3453FFC8FF93FF3CFF73DD51FF45" bold="true" box="[151,531,151,178]" fontSize="11" level="1" pageId="2" pageNumber="277" reason="1">
<emphasis id="A4EC5F2DFFC8FF93FF3CFF73DD51FF45" bold="true" box="[151,531,151,178]" pageId="2" pageNumber="277">
Genus
<taxonomicName id="5198F8BCFFC8FF93FF45FF7CDEDCFF46" box="[238,414,151,177]" class="Insecta" family="Coccinellidae" genus="Ruthmuelleria" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="277" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="A4EC5F2DFFC8FF93FF45FF7CDEDCFF46" bold="true" box="[238,414,151,177]" italics="true" pageId="2" pageNumber="277">Ruthmuelleria</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BFDFE256FFC8FF93FE0EFF73DD51FF45" box="[421,531,152,178]" pageId="2" pageNumber="277" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DE82D0B4FFC8FF93FF3CFF32DD12FF19" box="[151,592,216,238]" pageId="2" pageNumber="277" type="materials_examined">
<paragraph id="9627833FFFC8FF93FF3CFF32DD12FF19" blockId="2.[151,592,216,238]" box="[151,592,216,238]" pageId="2" pageNumber="277">
<typeStatus id="49233D9DFFC8FF93FF3CFF32DF8BFF19" box="[151,201,217,238]" pageId="2" pageNumber="277">Type</typeStatus>
species:
<taxonomicName id="5198F8BCFFC8FF93FE8DFF33DD08FF19" box="[294,586,216,238]" class="Insecta" family="Coccinellidae" genus="Ruthmuelleria" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="277" phylum="Arthropoda" rank="species" species="grootdrifensis">
<emphasis id="A4EC5F2DFFC8FF93FE8DFF33DD08FF19" box="[294,586,216,238]" italics="true" pageId="2" pageNumber="277">Ruthmuelleria grootdrifensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="DE82D0B4FFC8FF93FF3CFEF1DE01FE34" pageId="2" pageNumber="277" type="diagnosis">
<paragraph id="9627833FFFC8FF93FF3CFEF1DE01FE34" blockId="2.[151,1437,282,811]" pageId="2" pageNumber="277">
<emphasis id="A4EC5F2DFFC8FF93FF3CFEF1DE53FEC4" bold="true" box="[151,273,282,307]" pageId="2" pageNumber="277">Diagnosis.</emphasis>
Antennae eight-segmented; head without supraorbital sulcus; prothorax elongate, with sublateral carinae running from base to anterior pits; pronotum with two pairs of pits located latero-dorsally; prosternum with a pair of pits near anterior part of pronotosternal suture; mesoventrite with a pair of postmesocoxal pits with openings shifted to anterior portion of metaventrite; metaventral postcoxal lines not recurved; abdominal postcoxal lines V-shaped.
</paragraph>
</subSubSection>
<subSubSection id="DE82D0B4FFC8FF92FF6CFE25DA48FE02" lastPageId="3" lastPageNumber="278" pageId="2" pageNumber="277" type="description">
<paragraph id="9627833FFFC8FF93FF6CFE25DE76FDFD" blockId="2.[151,1437,282,811]" pageId="2" pageNumber="277">
<emphasis id="A4EC5F2DFFC8FF93FF6CFE25DE1BFE10" bold="true" box="[199,345,462,487]" pageId="2" pageNumber="277">Description.</emphasis>
Body (
<figureCitation id="0EA39FBAFFC8FF93FE05FE25DEB2FE10" box="[430,496,462,487]" captionStart="FIGURES 1 2" captionStartId="2.[151,264,1586,1609]" captionTargetBox="[154,1436,853,1563]" captionTargetId="figure@2.[151,1436,853,1566]" captionTargetPageId="2" captionText="FIGURES 1 2. Ruthmuelleria grootdrifensis sp. n. (1) Dorsal habitus of male; (2) Distribution map." httpUri="https://zenodo.org/record/227369/files/figure.png" pageId="2" pageNumber="277">Fig. 1</figureCitation>
) elongate and strongly flattened, covered with short setae, colour light brown; BL around
<quantity id="51602EDAFFC8FF93FF47FE18DE72FDFD" box="[236,304,499,523]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="277" unit="mm" value="1.0">1 mm</quantity>
.
</paragraph>
<paragraph id="9627833FFFC8FF93FF6CFDFDDDF8FCDD" blockId="2.[151,1437,282,811]" pageId="2" pageNumber="277">
Head (
<figureCitation id="0EA39FBAFFC8FF93FEB9FDFDDE3FFDD8" box="[274,381,534,559]" captionStart="FIGURES 3 6" captionStartId="3.[151,264,1894,1917]" captionTargetBox="[159,1425,675,1857]" captionTargetId="figure@3.[151,1436,661,1873]" captionTargetPageId="3" captionText="FIGURES 3 6. Ruthmuelleria grootdrifensis sp. n. (3) Head capsule, dorsal view (mouthparts omitted); (4) Head capsule, ventral view; (5) Prothorax, dorsal view; (6) Prothorax, ventral view." httpUri="https://zenodo.org/record/227370/files/figure.png" pageId="2" pageNumber="277">Figs. 34</figureCitation>
) elongate; vertex and frontoclypeus weakly convex; frontoclypeus slightly longer than length of antennal sockets, with anterior margin shallowly emarginate; supraorbital sulcus and subantennal grooves absent; eyes moderately large and convex, coarsely faceted. Gular sutures obliterated. Antenna (
<figureCitation id="0EA39FBAFFC8FF93FB54FDB5DA05FD80" box="[1279,1351,606,631]" captionStart="FIGURES 3 6" captionStartId="3.[151,264,1894,1917]" captionTargetBox="[159,1425,675,1857]" captionTargetId="figure@3.[151,1436,661,1873]" captionTargetPageId="3" captionText="FIGURES 3 6. Ruthmuelleria grootdrifensis sp. n. (3) Head capsule, dorsal view (mouthparts omitted); (4) Head capsule, ventral view; (5) Prothorax, dorsal view; (6) Prothorax, ventral view." httpUri="https://zenodo.org/record/227370/files/figure.png" pageId="2" pageNumber="277">Fig. 4</figureCitation>
) eightsegmented, with a distinct club composed of large terminal antennomere; antennomeres VVII transverse. Mandibular apex bidentate; maxillary palp large, with cultriform terminal palpomere; submentum very narrow and broadened in anterior portion; mentum much broader than submentum, strongly elongate, subtrapezoidal; ligula shorter than labial palp, with a fringe of setae along anterior margin; labial palpi broadly separated, palpomeres cylindrical, apex of terminal palpomere truncate.
</paragraph>
<caption id="C2E7D3B7FFC8FF93FF3CF9D9DBDAF9BF" box="[151,1176,1586,1609]" httpUri="https://zenodo.org/record/227369/files/figure.png" pageId="2" pageNumber="277" targetBox="[154,1436,853,1563]" targetPageId="2">
<paragraph id="9627833FFFC8FF93FF3CF9D9DBDAF9BF" blockId="2.[151,1176,1586,1609]" box="[151,1176,1586,1609]" pageId="2" pageNumber="277">
<emphasis id="A4EC5F2DFFC8FF93FF3CF9D9DE78F9B0" bold="true" box="[151,314,1586,1609]" pageId="2" pageNumber="277">FIGURES 12.</emphasis>
<taxonomicName id="5198F8BCFFC8FF93FEEBF9D9DD26F9BF" box="[320,612,1586,1608]" class="Insecta" family="Coccinellidae" genus="Ruthmuelleria" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="277" phylum="Arthropoda" rank="species" species="grootdrifensis" status="sp. nov.">
<emphasis id="A4EC5F2DFFC8FF93FEEBF9D9DD26F9BF" box="[320,612,1586,1608]" italics="true" pageId="2" pageNumber="277">Ruthmuelleria grootdrifensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BFDFE256FFC8FF93FDC0F9D8DDE6F9BF" box="[619,676,1587,1608]" pageId="2" pageNumber="277" rank="species">
<emphasis id="A4EC5F2DFFC8FF93FDC0F9D8DDE6F9BF" bold="true" box="[619,676,1587,1608]" pageId="2" pageNumber="277">sp. n.</emphasis>
</taxonomicNameLabel>
(1) Dorsal habitus of male; (2) Distribution map.
</paragraph>
</caption>
<paragraph id="9627833FFFC8FF93FF6CF984DC12F8CA" blockId="2.[151,1436,1647,2032]" pageId="2" pageNumber="277">
Prothorax (
<figureCitation id="0EA39FBAFFC8FF93FEECF984DEF6F97F" box="[327,436,1647,1672]" captionStart="FIGURES 3 6" captionStartId="3.[151,264,1894,1917]" captionTargetBox="[159,1425,675,1857]" captionTargetId="figure@3.[151,1436,661,1873]" captionTargetPageId="3" captionText="FIGURES 3 6. Ruthmuelleria grootdrifensis sp. n. (3) Head capsule, dorsal view (mouthparts omitted); (4) Head capsule, ventral view; (5) Prothorax, dorsal view; (6) Prothorax, ventral view." httpUri="https://zenodo.org/record/227370/files/figure.png" pageId="2" pageNumber="277">Figs. 56</figureCitation>
) elongate, with three pairs of pits: two on laterodorsal portion of pronotum and one in anterior portion of prosternum. Pronotum (
<figureCitation id="0EA39FBAFFC8FF93FD38F97FDD99F95A" box="[659,731,1684,1709]" captionStart="FIGURES 3 6" captionStartId="3.[151,264,1894,1917]" captionTargetBox="[159,1425,675,1857]" captionTargetId="figure@3.[151,1436,661,1873]" captionTargetPageId="3" captionText="FIGURES 3 6. Ruthmuelleria grootdrifensis sp. n. (3) Head capsule, dorsal view (mouthparts omitted); (4) Head capsule, ventral view; (5) Prothorax, dorsal view; (6) Prothorax, ventral view." httpUri="https://zenodo.org/record/227370/files/figure.png" pageId="2" pageNumber="277">Fig. 5</figureCitation>
) weakly convex, lateral margin indistinctly microcrenulate; sublateral carinae extending from pronotal base to anterior pits; posterior margin arcuate and strongly expanding caudad. Prosternum (
<figureCitation id="0EA39FBAFFC8FF93FE25F937DE94F902" box="[398,470,1756,1781]" captionStart="FIGURES 3 6" captionStartId="3.[151,264,1894,1917]" captionTargetBox="[159,1425,675,1857]" captionTargetId="figure@3.[151,1436,661,1873]" captionTargetPageId="3" captionText="FIGURES 3 6. Ruthmuelleria grootdrifensis sp. n. (3) Head capsule, dorsal view (mouthparts omitted); (4) Head capsule, ventral view; (5) Prothorax, dorsal view; (6) Prothorax, ventral view." httpUri="https://zenodo.org/record/227370/files/figure.png" pageId="2" pageNumber="277">Fig. 6</figureCitation>
) in front of coxae about three times as long as coxal cavity, flattened; prosternal intercoxal process with nearly parallel lateral margins in posterior half, truncated at apex and extending beyond procoxal cavities, as narrow as about one third of coxal width.
</paragraph>
<paragraph id="9627833FFFC8FF93FF6CF8ACDC6AF872" blockId="2.[151,1436,1647,2032]" pageId="2" pageNumber="277">
Mesoventrite (
<figureCitation id="0EA39FBAFFC8FF93FED8F8ACDE82F897" box="[371,448,1863,1888]" captionStart="FIGURES 7 9" captionStartId="4.[151,264,1366,1389]" captionTargetBox="[160,1436,184,1332]" captionTargetId="figure@4.[151,1436,167,1345]" captionTargetPageId="4" captionText="FIGURES 7 9. Ruthmuelleria grootdrifensis sp. n. (7) Pterothorax and abdomen, ventral view; (8 9) Male genitalia." httpUri="https://zenodo.org/record/227371/files/figure.png" pageId="2" pageNumber="277">Fig. 7</figureCitation>
) flattened, procoxal rest not demarcated; mesoventral intercoxal process slightly broadened in posterior part, with posterior margin straight.
</paragraph>
<paragraph id="9627833FFFC8FF93FF6CF864DBFFF850" blockId="2.[151,1436,1647,2032]" box="[199,1213,1935,1960]" pageId="2" pageNumber="277">Mesoscutellum (not shown) well visible between bases of elytra, subtriangular, transverse.</paragraph>
<paragraph id="9627833FFFC8FF93FF6CF85FDD2BF818" blockId="2.[151,1436,1647,2032]" pageId="2" pageNumber="277">
Elytra (
<figureCitation id="0EA39FBAFFC8FF93FEB0F85FDE1FF83B" box="[283,349,1972,1997]" captionStart="FIGURES 1 2" captionStartId="2.[151,264,1586,1609]" captionTargetBox="[154,1436,853,1563]" captionTargetId="figure@2.[151,1436,853,1566]" captionTargetPageId="2" captionText="FIGURES 1 2. Ruthmuelleria grootdrifensis sp. n. (1) Dorsal habitus of male; (2) Distribution map." httpUri="https://zenodo.org/record/227369/files/figure.png" pageId="2" pageNumber="277">Fig. 1</figureCitation>
) oval; broadest slightly anterior to middle; epipleura incomplete, fused with elytral margin near posterior margin of abdominal ventrite II.
</paragraph>
<paragraph id="9627833FFFC9FF92FF6CFF7CDB60FF0F" blockId="3.[151,1436,151,608]" pageId="3" pageNumber="278">
Metaventrite (
<figureCitation id="0EA39FBAFFC9FF92FEC0FF7CDEF6FF47" box="[363,436,151,176]" captionStart="FIGURES 7 9" captionStartId="4.[151,264,1366,1389]" captionTargetBox="[160,1436,184,1332]" captionTargetId="figure@4.[151,1436,167,1345]" captionTargetPageId="4" captionText="FIGURES 7 9. Ruthmuelleria grootdrifensis sp. n. (7) Pterothorax and abdomen, ventral view; (8 9) Male genitalia." httpUri="https://zenodo.org/record/227371/files/figure.png" pageId="3" pageNumber="278">Fig. 7</figureCitation>
) strongly transverse, slightly convex and slightly shorter than abdominal ventrite I; postcoxal lines oblique, broadly separated medially, extending from middle width of mesocoxa to lateral margin of metaventrite; posterior margin of metaventrite between metacoxa nearly straight.
</paragraph>
<paragraph id="9627833FFFC9FF92FF6CFEEFDEB9FEEA" blockId="3.[151,1436,151,608]" box="[199,507,260,285]" pageId="3" pageNumber="278">Metathoracic wings absent.</paragraph>
<paragraph id="9627833FFFC9FF92FF6CFECCDDF3FE7F" blockId="3.[151,1436,151,608]" pageId="3" pageNumber="278">
Abdomen (
<figureCitation id="0EA39FBAFFC9FF92FEE3FECCDECFFEB7" box="[328,397,295,320]" captionStart="FIGURES 7 9" captionStartId="4.[151,264,1366,1389]" captionTargetBox="[160,1436,184,1332]" captionTargetId="figure@4.[151,1436,167,1345]" captionTargetPageId="4" captionText="FIGURES 7 9. Ruthmuelleria grootdrifensis sp. n. (7) Pterothorax and abdomen, ventral view; (8 9) Male genitalia." httpUri="https://zenodo.org/record/227371/files/figure.png" pageId="3" pageNumber="278">Fig. 7</figureCitation>
) with six ventrites, ventrite I slightly longer than II and III combined; postcoxal lines Vshaped and reaching posterior margin of ventrite. Pygidium and propygidium (
<figureCitation id="0EA39FBAFFC9FF92FB80FEA7DB2DFE93" box="[1067,1135,332,357]" captionStart="FIGURES 1 2" captionStartId="2.[151,264,1586,1609]" captionTargetBox="[154,1436,853,1563]" captionTargetId="figure@2.[151,1436,853,1566]" captionTargetPageId="2" captionText="FIGURES 1 2. Ruthmuelleria grootdrifensis sp. n. (1) Dorsal habitus of male; (2) Distribution map." httpUri="https://zenodo.org/record/227369/files/figure.png" pageId="3" pageNumber="278">Fig. 1</figureCitation>
) in all studied specimens strongly sclerotized and exposed in dorsal view.
</paragraph>
<paragraph id="9627833FFFC9FF92FF6CFE7EDC28FE5A" blockId="3.[151,1436,151,608]" box="[199,874,404,429]" pageId="3" pageNumber="278">
Legs (
<figureCitation id="0EA39FBAFFC9FF92FEA7FE7FDE35FE5A" box="[268,375,404,429]" captionStart-0="FIGURES 3 6" captionStart-1="FIGURES 7 9" captionStartId-0="3.[151,264,1894,1917]" captionStartId-1="4.[151,264,1366,1389]" captionTargetBox-0="[159,1425,675,1857]" captionTargetBox-1="[160,1436,184,1332]" captionTargetId-0="figure@3.[151,1436,661,1873]" captionTargetId-1="figure@4.[151,1436,167,1345]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 3 6. Ruthmuelleria grootdrifensis sp. n. (3) Head capsule, dorsal view (mouthparts omitted); (4) Head capsule, ventral view; (5) Prothorax, dorsal view; (6) Prothorax, ventral view." captionText-1="FIGURES 7 9. Ruthmuelleria grootdrifensis sp. n. (7) Pterothorax and abdomen, ventral view; (8 9) Male genitalia." httpUri-0="https://zenodo.org/record/227370/files/figure.png" httpUri-1="https://zenodo.org/record/227371/files/figure.png" pageId="3" pageNumber="278">Figs. 67</figureCitation>
) short but slender; all tarsi three-segmented.
</paragraph>
<paragraph id="9627833FFFC9FF92FF6CFE5CDA48FE02" blockId="3.[151,1436,151,608]" pageId="3" pageNumber="278">
Male genitalia (
<figureCitation id="0EA39FBAFFC9FF92FE20FE5CDD40FE27" box="[395,514,439,464]" captionStart="FIGURES 7 9" captionStartId="4.[151,264,1366,1389]" captionTargetBox="[160,1436,184,1332]" captionTargetId="figure@4.[151,1436,167,1345]" captionTargetPageId="4" captionText="FIGURES 7 9. Ruthmuelleria grootdrifensis sp. n. (7) Pterothorax and abdomen, ventral view; (8 9) Male genitalia." httpUri="https://zenodo.org/record/227371/files/figure.png" pageId="3" pageNumber="278">Figs. 89</figureCitation>
). Tegmen asymmetrical, penis guide broad and relatively short, parameres articulated, anterior strut very short. Penis about 0.7 times as long as abdomen, without basal capsule.
</paragraph>
</subSubSection>
<subSubSection id="DE82D0B4FFC9FF92FF6CFE14DFB0FDCB" pageId="3" pageNumber="278" type="distribution">
<paragraph id="9627833FFFC9FF92FF6CFE14DFB0FDCB" blockId="3.[151,1436,151,608]" pageId="3" pageNumber="278">
<emphasis id="A4EC5F2DFFC9FF92FF6CFE14DD6EFDEF" bold="true" box="[199,556,511,536]" pageId="3" pageNumber="278">Distribution and composition.</emphasis>
A single species is known to occur in the West Cape province of
<collectingCountry id="EE8FC3AFFFC9FF92FAA7FE14DADEFDEF" box="[1292,1436,511,536]" name="South Africa" pageId="3" pageNumber="278">South Africa</collectingCountry>
(
<figureCitation id="0EA39FBAFFC9FF92FF34FDCFDFA7FDCB" box="[159,229,548,573]" captionStart="FIGURES 1 2" captionStartId="2.[151,264,1586,1609]" captionTargetBox="[154,1436,853,1563]" captionTargetId="figure@2.[151,1436,853,1566]" captionTargetPageId="2" captionText="FIGURES 1 2. Ruthmuelleria grootdrifensis sp. n. (1) Dorsal habitus of male; (2) Distribution map." httpUri="https://zenodo.org/record/227369/files/figure.png" pageId="3" pageNumber="278">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="DE82D0B4FFC9FF92FF6CFDACDB01FD97" box="[199,1091,583,608]" pageId="3" pageNumber="278" type="etymology">
<paragraph id="9627833FFFC9FF92FF6CFDACDB01FD97" blockId="3.[151,1436,151,608]" box="[199,1091,583,608]" pageId="3" pageNumber="278">
<emphasis id="A4EC5F2DFFC9FF92FF6CFDACDE0CFD97" bold="true" box="[199,334,583,608]" pageId="3" pageNumber="278">Etymology.</emphasis>
This genus is named in honour of Ruth Müller, a curator at TMSA.
</paragraph>
</subSubSection>
</treatment>
</document>