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<document id="A4AE97D3D25DDE5A0AA2F22C2F8D06C0" ID-DOI="10.5852/ejt.2024.959.2677" ID-ISSN="2118-9773" ID-Zenodo-Dep="13945464" ID-ZooBank="30872C13-516F-42FA-AFA7-30ADC6BF1BAF" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1729166637250" checkinUser="plazi" docAuthor="Rosa, Cecilia Lozano De La, Cupello, Mario &amp; Vaz-De-Mello, Fernando Z." docDate="2024" docId="1D50766630FB52C6EF32FBA7752E00E1" docLanguage="en" docName="EJT.2024.959.1-272.pdf" docOrigin="European Journal of Taxonomy 959 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2677/12401" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Onthophagus rhinophyllus Harold 1868" docType="treatment" docVersion="2" lastPageNumber="163" masterDocId="E1690E1E305A5264ED17FFA7770C013F" masterDocTitle="The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution" masterLastPageNumber="272" masterPageNumber="1" pageNumber="162" updateTime="1729169574746" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="5D79EB1D094A3748529372E8940B75F8">The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution</mods:title>
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<mods:namePart id="3837902E203BEB5036912357363189DA">Cupello, Mario</mods:namePart>
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<mods:affiliation id="D71C7F5990002ABA315E25B71BAD39FB">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Curitiba, Paraná, Brazil. Currently at Department of Entomology, Texas A &amp; M University, College Station, TX, 77843, USA.</mods:affiliation>
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Harold, 1868
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<emphasis id="A78D1B6230FB52C5EDAAFBC276F50540" box="[189,505,1125,1151]" italics="true" pageId="161" pageNumber="162">Onthophagus rhinophyllus</emphasis>
<bibRefCitation id="F168BA8130FB52C5EF15FBC275D70540" author="Harold E." box="[514,731,1125,1151]" pageId="161" pageNumber="162" pagination="79 - 86" refId="ref123468" refString="Harold E. 1868 e. Diagnosen neuer Coprophagen. Coleopterologische Hefte 4: 79 - 86. Available from https: // www. biodiversitylibrary. org / page / 16069646 [accessed 16 Feb. 2024]." type="journal article" year="1868">Harold, 1868e: 84</bibRefCitation>
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(original description). Type locality:
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. Name-bearing type: a single known
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<emphasis id="A78D1B6230FB52C5EDAAFB6D76F805DB" box="[189,500,1226,1252]" italics="true" pageId="161" pageNumber="162">Onthophagus rhinophyllus</emphasis>
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<bibRefCitation id="F168BA8130FB52C5EF1AFB6D75A705DB" author="Harold E." box="[525,683,1226,1252]" pageId="161" pageNumber="162" pagination="46 - 70" refId="ref123583" refString="Harold E. 1869 c. Ueber Coprophage Lamellicornien mit besonderer Berucksichtigung der Pariser Sammlungen. Coleopterologische Hefte 5: 46 - 70. Available from https: // www. biodiversitylibrary. org / page / 16069721 [accessed 16 Feb. 2024]." type="journal article" year="1869">Harold 1869c</bibRefCitation>
(cited); 1880: 31 (comments). —
<treatmentCitation id="1458E16130FB52C5E926FB6D721C05DB" author="Candeze E." box="[1073,1296,1226,1252]" page="330" pageId="161" pageNumber="162" year="1891">
<bibRefCitation id="F168BA8130FB52C5E926FB6D721C05DB" author="Candeze E." box="[1073,1296,1226,1252]" pageId="161" pageNumber="162" refId="ref113476" refString="Candeze E. 1891. Voyage de M. E. Simon au Venezuela (Decembre 1887 - Avril 1888), 15 e Memoire (1). Coleopteres - Lamellicornes determines par le Dr E. Candeze. Annales de la Societe entomologique de France 60: [329] - 332. Available from https: // www. biodiversitylibrary. org / page / 32547689 [accessed 14 Feb. 2024]." type="journal volume" year="1891">Candèze 1891: 330</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="1458E16130FB52C5EDFDFB4A75410438" author="Gillet J. J. E. &amp; Boucomont A." box="[234,589,1261,1288]" page="208" pageId="161" pageNumber="162" year="1927">
<bibRefCitation id="F168BA8130FB52C5EDFDFB4A75410438" author="Gillet J. J. E. &amp; Boucomont A." box="[234,589,1261,1288]" pageId="161" pageNumber="162" refId="ref121284" refString="Gillet J. J. E. &amp; Boucomont A. 1927. Scarabaeidae: Coprinae, Termitotroginae (Partes 38 et 90). W. Junk, Berlin. [We are not sure about the publication date of this book. The title page gives the range &quot; 1911 - 1927 &quot;, which could be interpreted as indicating that the book was published in parts between those years. What we suspect, however, is that 1911 refers to the first part of the Coprinae catalogue, published by Gillet (1911 b) - i. e., our previous reference -, whereas the continuation of the Coprinae catalogue, now added by the Termitotroginae, was published at once in 1927 with the joint-authorship of Gillet and Boucomont.]" type="book" year="1927">Gillet &amp; Boucomont 1927: 208</bibRefCitation>
</treatmentCitation>
(catalogue).—
<treatmentCitation id="1458E16130FB52C5EE15FB49730E0438" author="Boucomont A." box="[770,1026,1261,1288]" page="299" pageId="161" pageNumber="162" year="1932">
<bibRefCitation id="F168BA8130FB52C5EE15FB49730E0438" author="Boucomont A." box="[770,1026,1261,1288]" pageId="161" pageNumber="162" pagination="293 - 332" refId="ref112419" refString="Boucomont A. 1932. Synopsis des Onthophagus d'Amerique du Sud (Col. Scarab.). Annales de la Societe entomologique de France 101: 293 - 332. https: // doi. org / 10.1080 / 21686351.1932.12280217" type="journal article" year="1932">Boucomont 1932: 299</bibRefCitation>
</treatmentCitation>
(synopsis, key). —
<treatmentCitation id="1458E16130FB52C5E9F5FB4A76520414" author="Blackwelder R. E." page="212" pageId="161" pageNumber="162" year="1944">
<bibRefCitation id="F168BA8130FB52C5E9F5FB4A76520414" author="Blackwelder R. E." pageId="161" pageNumber="162" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 212</bibRefCitation>
</treatmentCitation>
(list, cited for
<collectingCountry id="EDEE87E030FB52C5EF07FAB775850415" box="[528,649,1296,1322]" name="Colombia" pageId="161" pageNumber="162">Colombia</collectingCountry>
). —
<treatmentCitation id="1458E16130FB52C5EFD1FAB6747F0415" author="Roze J. A." box="[710,883,1296,1323]" page="46" pageId="161" pageNumber="162" year="1955">
<bibRefCitation id="F168BA8130FB52C5EFD1FAB6747F0415" author="Roze J. A." box="[710,883,1296,1323]" pageId="161" pageNumber="162" pagination="39 - 63" refId="ref137914" refString="Roze J. A. 1955. Lista preliminar de la familia Scarabaeidae sensu lato de Venezuela. Boletin del Museo de Ciencias Naturales 1 (1): 39 - 63." type="journal article" year="1955">Roze 1955: 46</bibRefCitation>
</treatmentCitation>
(cited, checklist for
<collectingCountry id="EDEE87E030FB52C5E97DFAB773EF0415" box="[1130,1251,1296,1322]" name="Colombia" pageId="161" pageNumber="162">Colombia</collectingCountry>
). —
<treatmentCitation id="1458E16130FB52C5E80AFAB776AC0471" author="Medina C. A. &amp; Lopera-Toro A. &amp; Vitolo A. &amp; Gill B." page="140" pageId="161" pageNumber="162" year="2001">
<bibRefCitation id="F168BA8130FB52C5E80AFAB776AC0471" author="Medina C. A. &amp; Lopera-Toro A. &amp; Vitolo A. &amp; Gill B." pageId="161" pageNumber="162" pagination="131 - 144" refId="ref130799" refString="Medina C. A., Lopera-Toro A., Vitolo A. &amp; Gill B. 2001. Escarabajos coprofagos (Coleoptera: Scarabaeinae) de Colombia. Biota Colombiana 2: 131 - 144." type="journal article" year="2001">
Medina
<emphasis id="A78D1B6230FB52C5EDFCFA9376290471" box="[235,293,1332,1358]" italics="true" pageId="161" pageNumber="162">et al.</emphasis>
2001: 140
</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E030FB52C5EF50FA9375CC0471" box="[583,704,1332,1358]" name="Colombia" pageId="161" pageNumber="162">Colombia</collectingCountry>
). —
<treatmentCitation id="1458E16130FB52C5EFE1FA9374F40472" author="Delgado L. &amp; Peraza L. N. &amp; Deloya C." box="[758,1016,1331,1358]" page="6" pageId="161" pageNumber="162" year="2006">
<bibRefCitation id="F168BA8130FB52C5EFE1FA9374F40472" author="Delgado L. &amp; Peraza L. N. &amp; Deloya C." box="[758,1016,1331,1358]" pageId="161" pageNumber="162" pagination="6 - 9" refId="ref116516" refString="Delgado L., Peraza L. N. &amp; Deloya C. 2006. Onthophagus yucatanus, a new species of the Clypeatus group from Mexico and Guatemala (Coleoptera: Scarabaeidae). Florida Entomologist 89 (1): 6 - 9. https: // doi. org / 10.1653 / 0015 - 4040 (2006) 89 [6: OYANSO] 2.0. CO; 2" type="journal article" year="2006">
Delgado
<emphasis id="A78D1B6230FB52C5EE48FA9374940471" box="[863,920,1332,1358]" italics="true" pageId="161" pageNumber="162">et al.</emphasis>
2006: 6
</bibRefCitation>
</treatmentCitation>
(comments); 2012: 327 (list). —
<treatmentCitation id="1458E16130FB52C5EDFEFAF0759D044E" author="Pulido-Herrera L. A. &amp; Zunino M." box="[233,657,1367,1393]" page="114" pageId="161" pageNumber="162" year="2007">
<bibRefCitation id="F168BA8130FB52C5EDFEFAF0759D044E" author="Pulido-Herrera L. A. &amp; Zunino M." box="[233,657,1367,1393]" pageId="161" pageNumber="162" refId="ref136342" refString="Pulido-Herrera L. A. &amp; Zunino M. 2007. Catalogo preliminar de los Onthophagini de America (Coleoptera: Scarabaeine [sic]). In: Zunino M. &amp; Melic A. (eds) Escarabajos, Diversidad y Conservacion Biologica. Ensayos en Homenaje a Gonzalo Halffter: 93 - 129. Monografias 3 ercer Milenio m 3 m Vol. 7, Sociedad Entomologica Aragonesa, Zaragoza." type="book" year="2007">Pulido-Herrera &amp; Zunino 2007: 114</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="1458E16130FB52C5EE46FAF07391044E" author="Ferrer-Paris J. &amp; Sanchez-Mercado A. &amp; Rodriguez J. P." box="[849,1181,1367,1393]" page="108" pageId="161" pageNumber="162" year="2013">
<bibRefCitation id="F168BA8130FB52C5EE46FAF07391044E" author="Ferrer-Paris J. &amp; Sanchez-Mercado A. &amp; Rodriguez J. P." box="[849,1181,1367,1393]" pageId="161" pageNumber="162" pagination="89 - 110" refId="ref119203" refString="Ferrer-Paris J. R, Sanchez-Mercado A. &amp; Rodriguez J. P. 2013. Optimizacion del muestreo de invertebrados tropicales: un ejemplo con escarabajos coprofagos (Coleoptera: Scarabaeinae) en Venezuela. Revista de Biologia Tropical 61 (1): 89 - 110. https: // doi. org / 10.15517 / rbt. v 61 i 1.10941" type="journal article" year="2013">
Ferrer-Paris
<emphasis id="A78D1B6230FB52C5EEF2FAFF732C044E" box="[997,1056,1367,1393]" italics="true" pageId="161" pageNumber="162">et al.</emphasis>
2013: 108
</bibRefCitation>
</treatmentCitation>
(list). —
<bibRefCitation id="F168BA8130FB52C5E807FAF0766704AA" author="Ratcliffe B. C. &amp; Jameson M. L. &amp; Figueroa L. &amp; Cave R. D. &amp; Paulsen M. J. &amp; Cano E. B. &amp; Beza-Beza C. &amp; Ferbans L. &amp; Reyes-Castillo P." pageId="161" pageNumber="162" pagination="186 - 207" refId="ref137018" refString="Ratcliffe B. C., Jameson M. L., Figueroa L., Cave R. D., Paulsen M. J., Cano E. B., Beza-Beza C., Jimenez- Ferbans L., Reyes-Castillo P. 2015. Beetles (Coleoptera) of Peru: a survey of the families. Journal of the Kansas Entomological Society 88 (2): 186 - 207." type="journal article" year="2015">
Ratcliffe
<emphasis id="A78D1B6230FB52C5EDFCFADC762904AB" box="[235,293,1402,1428]" italics="true" pageId="161" pageNumber="162">et al.</emphasis>
2015
</bibRefCitation>
: (list for
<collectingCountry id="EDEE87E030FB52C5ECC2FADD751C04AB" box="[469,528,1402,1428]" name="Peru" pageId="161" pageNumber="162">Peru</collectingCountry>
). —
<treatmentCitation id="1458E16130FB52C5EF5FFADD74CC04AB" ID-DOI="http://doi.org/10.5281/zenodo.3683616" ID-Zenodo-Dep="3683616" author="Hielkema A. J. &amp; Hielkema M. A." box="[584,960,1402,1428]" httpUri="http://treatment.plazi.org/id/03D7FF1CFF96FFB0FF48FF0D34DDFAD8" page="92" pageId="161" pageNumber="162" year="2019">
<bibRefCitation id="F168BA8130FB52C5EF5FFADD74CC04AB" author="Hielkema A. J. &amp; Hielkema M. A." box="[584,960,1402,1428]" pageId="161" pageNumber="162" pagination="1 - 306" refId="ref124225" refString="Hielkema A. J. &amp; Hielkema M. A. 2019. An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas. Insecta Mundi 732 (732): 1 - 306." type="journal article" year="2019">Hielkema &amp; Hielkema 2019: 92</bibRefCitation>
</treatmentCitation>
(catalogue for the Guianas).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB30FB52C5EDAAFA1C766007AE" pageId="161" pageNumber="162" type="materials_examined">
<paragraph id="9546C77030FB52C5EDAAFA1C76A604EA" blockId="161.[189,426,1467,1493]" box="[189,426,1467,1493]" pageId="161" pageNumber="162">
<heading id="CE0E701C30FB52C5EDAAFA1C76A604EA" bold="true" box="[189,426,1467,1493]" fontSize="11" level="3" pageId="161" pageNumber="162" reason="3">
<emphasis id="A78D1B6230FB52C5EDAAFA1C76A604EA" bold="true" box="[189,426,1467,1493]" pageId="161" pageNumber="162">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="9546C77030FB52C5EDAAFA4D766007AE" blockId="161.[188,1399,1514,1681]" pageId="161" pageNumber="162">
<materialsCitation id="2591CD2D30FB52C5EDAAFA4D73E3073B" box="[189,1263,1514,1541]" collectingDate="2004-07-06" collectorName="D Garcia" country="Venezuela" elevation="50" location="Coyagua" pageId="161" pageNumber="162" specimenCount="1" stateProvince="Aragua">
<collectingCountry id="EDEE87E030FB52C5EDAAFA4D7667073B" box="[189,363,1514,1540]" name="Venezuela" pageId="161" pageNumber="162">VENEZUELA</collectingCountry>
<emphasis id="A78D1B6230FB52C5EC90FA4D76EA073B" bold="true" box="[391,486,1514,1540]" pageId="161" pageNumber="162">
<collectingRegion id="573D099230FB52C5EC90FA4D76EA073B" box="[391,486,1514,1540]" country="Venezuela" name="Aragua" pageId="161" pageNumber="162">Aragua</collectingRegion>
</emphasis>
<specimenCount id="83FF0CF930FB52C5ECE9FA4D755F073A" box="[510,595,1514,1541]" count="1" pageId="161" pageNumber="162" type="generic">1 spec.</specimenCount>
;
<location id="902691AB30FB52C5EF48FA4D75C7073B" LSID="urn:lsid:plazi:treatment:1D50766630FB52C6EF32FBA7752E00E1:902691AB30FB52C5EF48FA4D75C7073B" box="[607,715,1514,1540]" country="Venezuela" name="Coyagua" pageId="161" pageNumber="162" stateProvince="Aragua">Coyagua</location>
;
<elevation id="1ED4204330FB52C5EFC1FA4D745D073B" box="[726,849,1514,1541]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="161" pageNumber="162" unit="m" value="50.0">
<quantity id="52016A9530FB52C5EFC1FA4D741E073A" box="[726,786,1514,1541]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="161" pageNumber="162" unit="m" value="50.0">50 m</quantity>
a.s.l.
</elevation>
;
<date id="E147E1B030FB52C5EE48FA4D74EF073B" box="[863,995,1514,1540]" pageId="161" pageNumber="162" value="2004-07-06">
<collectingDate id="F103185830FB52C5EE48FA4D74EF073B" box="[863,995,1514,1540]" pageId="161" pageNumber="162" value="2004-07-06">6 Jul. 2004</collectingDate>
</date>
;
<collectorName id="380CA2A630FB52C5EEF8FA4C7356073B" box="[1007,1114,1514,1541]" pageId="161" pageNumber="162">D García</collectorName>
leg.; CEMT
</materialsCitation>
.
<materialsCitation id="2591CD2D30FB52C5E819FA4D746C0775" collectingDate="2009-07-19" collectorName="M. Asmussen &amp; P. Colmenares &amp; H. Martinez" country="Venezuela" elevation="1380" latitude="10.386639" location="Aroa" longLatPrecision="1" longitude="-68.84706" pageId="161" pageNumber="162" specimenCount="1" stateProvince="Yaracuy">
<emphasis id="A78D1B6230FB52C5E819FA4D727A073B" bold="true" box="[1294,1398,1514,1540]" pageId="161" pageNumber="162">
<collectingRegion id="573D099230FB52C5E819FA4D727A073B" box="[1294,1398,1514,1540]" country="Venezuela" name="Yaracuy" pageId="161" pageNumber="162">Yaracuy</collectingRegion>
</emphasis>
• 4 specs;
<collectingRegion id="573D099230FB52C5EC2CF9AA76990718" box="[315,405,1549,1575]" country="Venezuela" name="Bolivar" pageId="161" pageNumber="162">Bolívar</collectingRegion>
,
<location id="902691AB30FB52C5ECB6F9AA76D30718" LSID="urn:lsid:plazi:treatment:1D50766630FB52C6EF32FBA7752E00E1:902691AB30FB52C5ECB6F9AA76D30718" box="[417,479,1549,1575]" country="Venezuela" latitude="10.386639" longLatPrecision="1" longitude="-68.84706" name="Aroa" pageId="161" pageNumber="162" stateProvince="Yaracuy">Aroa</location>
;
<geoCoordinate id="F0CDA1B730FB52C5ECFAF9AB75960717" box="[493,666,1548,1576]" degrees="10" direction="north" minutes="23" orientation="latitude" pageId="161" pageNumber="162" precision="1" seconds="11.9" value="10.386639">10°2311.9″ N</geoCoordinate>
,
<geoCoordinate id="F0CDA1B730FB52C5EFBFF9AB74670717" box="[680,875,1548,1576]" degrees="68" direction="west" minutes="50" orientation="longitude" pageId="161" pageNumber="162" precision="1" seconds="49.41" value="-68.84706">68°5049.41″ W</geoCoordinate>
;
<elevation id="1ED4204330FB52C5EE6EF9AA731B0718" box="[889,1047,1549,1576]" metricMagnitude="3" metricUnit="m" metricValue="1.38" pageId="161" pageNumber="162" unit="m" value="1380.0">
<quantity id="52016A9530FB52C5EE6EF9AA74D80717" box="[889,980,1549,1576]" metricMagnitude="3" metricUnit="m" metricValue="1.38" pageId="161" pageNumber="162" unit="m" value="1380.0">1380 m</quantity>
a.s.l.
</elevation>
;
<date id="E147E1B030FB52C5E931F9AA73B30718" box="[1062,1215,1549,1575]" pageId="161" pageNumber="162" value="2009-07-19">
<collectingDate id="F103185830FB52C5E931F9AA73B30718" box="[1062,1215,1549,1575]" pageId="161" pageNumber="162" value="2009-07-19">19 Jul. 2009</collectingDate>
</date>
;
<collectorName id="380CA2A630FB52C5E9DAF9A9727F0718" box="[1229,1395,1549,1576]" pageId="161" pageNumber="162">M. Asmüssen</collectorName>
,
<collectorName id="380CA2A630FB52C5EDAAF996766F0775" box="[189,355,1584,1611]" pageId="161" pageNumber="162">P. Colmenares</collectorName>
and
<collectorName id="380CA2A630FB52C5EC8FF99675280775" box="[408,548,1584,1611]" pageId="161" pageNumber="162">H. Martínez</collectorName>
leg.; human faeces; CEMT
</materialsCitation>
<materialsCitation id="2591CD2D30FB52C5EE62F997766407AE" collectingDate="2009-07-19" collectorName="M. Asmussen &amp; P. Colmenares &amp; H. Martinez" country="Venezuela" elevation="710" latitude="10.0" location="Aroa" longLatPrecision="21" longitude="-68.0" pageId="161" pageNumber="162" specimenCount="1" stateProvince="Bolivar">
<specimenCount id="83FF0CF930FB52C5EE62F99774C40774" box="[885,968,1584,1611]" count="1" pageId="161" pageNumber="162" type="generic">1 spec.</specimenCount>
;
<collectingRegion id="573D099230FB52C5EEC5F99773200775" box="[978,1068,1584,1610]" country="Venezuela" name="Bolivar" pageId="161" pageNumber="162">Bolívar</collectingRegion>
,
<location id="902691AB30FB52C5E923F996737E0774" LSID="urn:lsid:plazi:treatment:1D50766630FB52C6EF32FBA7752E00E1:902691AB30FB52C5E923F996737E0774" box="[1076,1138,1585,1611]" country="Venezuela" latitude="10.0" longLatPrecision="21" longitude="-68.0" name="Aroa" pageId="161" pageNumber="162" stateProvince="Bolivar">Aroa</location>
;
<geoCoordinate id="F0CDA1B730FB52C5E96AF98873FF0774" box="[1149,1267,1583,1611]" degrees="10" direction="north" minutes="0" orientation="latitude" pageId="161" pageNumber="162" precision="15" seconds="0" value="10.0">10°00″ N</geoCoordinate>
,
<geoCoordinate id="F0CDA1B730FB52C5E9EAF988727B0774" box="[1277,1399,1583,1611]" degrees="68" direction="west" minutes="0" orientation="longitude" pageId="161" pageNumber="162" precision="15" seconds="0" value="-68.0">68°00″ W</geoCoordinate>
[incorrect coordinates];
<quantity id="52016A9530FB52C5ECC1F9F3752E0751" box="[470,546,1620,1646]" metricMagnitude="2" metricUnit="m" metricValue="7.1" pageId="161" pageNumber="162" unit="m" value="710.0">
<elevation id="1ED4204330FB52C5ECC1F9F3752E0751" box="[470,546,1620,1646]" metricMagnitude="2" metricUnit="m" metricValue="7.1" pageId="161" pageNumber="162" unit="m" value="710.0">710 m</elevation>
</quantity>
;
<date id="E147E1B030FB52C5EF39F9F375CD0751" box="[558,705,1619,1646]" pageId="161" pageNumber="162" value="2009-07-19">
<collectingDate id="F103185830FB52C5EF39F9F375CD0751" box="[558,705,1619,1646]" pageId="161" pageNumber="162" value="2009-07-19">19 Jul. 2009</collectingDate>
</date>
;
<collectorName id="380CA2A630FB52C5EFDBF9F374630751" box="[716,879,1620,1646]" pageId="161" pageNumber="162">M. Asmüssen</collectorName>
,
<collectorName id="380CA2A630FB52C5EE6EF9F3732D0752" box="[889,1057,1619,1646]" pageId="161" pageNumber="162">P. Colmenares</collectorName>
and
<collectorName id="380CA2A630FB52C5E94EF9F373EA0751" box="[1113,1254,1620,1646]" pageId="161" pageNumber="162">H. Martínez</collectorName>
leg.; human faeces; CEMT
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB30FB52C6EDAAF91F752E00E1" lastPageId="162" lastPageNumber="163" pageId="161" pageNumber="162" type="distribution">
<paragraph id="9546C77030FB52C5EDAAF91F765A07ED" blockId="161.[189,342,1720,1746]" box="[189,342,1720,1746]" pageId="161" pageNumber="162">
<heading id="CE0E701C30FB52C5EDAAF91F765A07ED" bold="true" box="[189,342,1720,1746]" fontSize="11" level="3" pageId="161" pageNumber="162" reason="3">
<emphasis id="A78D1B6230FB52C5EDAAF91F765A07ED" bold="true" box="[189,342,1720,1746]" pageId="161" pageNumber="162">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="9546C77030FB52C5EDAAF9407539063E" blockId="161.[189,565,1767,1793]" box="[189,565,1767,1793]" pageId="161" pageNumber="162">
<collectingCountry id="EDEE87E030FB52C5EDAAF9407638063E" box="[189,308,1767,1793]" name="Colombia" pageId="161" pageNumber="162">Colombia</collectingCountry>
,
<collectingCountry id="EDEE87E030FB52C5EC28F94076B7063E" box="[319,443,1767,1793]" name="Venezuela" pageId="161" pageNumber="162">Venezuela</collectingCountry>
, and
<collectingCountry id="EDEE87E030FB52C5ECEFF940753E063E" box="[504,562,1767,1793]" name="Peru" pageId="161" pageNumber="162">Peru</collectingCountry>
.
</paragraph>
<paragraph id="9546C77030FB52C5EDAAF88F76E1067D" blockId="161.[189,493,1832,1858]" box="[189,493,1832,1858]" pageId="161" pageNumber="162">
<heading id="CE0E701C30FB52C5EDAAF88F76E1067D" bold="true" box="[189,493,1832,1858]" fontSize="11" level="3" pageId="161" pageNumber="162" reason="3">
<emphasis id="A78D1B6230FB52C5EDAAF88F76E1067D" bold="true" box="[189,493,1832,1858]" pageId="161" pageNumber="162">
Subregions of
<collectingCountry id="EDEE87E030FB52C5EC78F88F76E1067D" box="[367,493,1832,1858]" name="Venezuela" pageId="161" pageNumber="162">Venezuela</collectingCountry>
</emphasis>
</heading>
</paragraph>
<paragraph id="9546C77030FB52C5EDAAF8F0722B064E" blockId="161.[189,1319,1879,1905]" box="[189,1319,1879,1905]" pageId="161" pageNumber="162">
System of hills and low sierras Lara-Falcón, Central
<collectingRegion id="573D099230FB52C5EE3EF8F07460064E" box="[809,876,1879,1905]" country="New Zealand" name="West Coast" pageId="161" pageNumber="162">Coast</collectingRegion>
Mountain Range, and Guiana Shield.
</paragraph>
<paragraph id="9546C77030F852C6EDAAFEAF76A8001C" blockId="162.[189,420,264,291]" box="[189,420,264,291]" pageId="162" pageNumber="163">
<heading id="CE0E701C30F852C6EDAAFEAF76A8001C" bold="true" box="[189,420,264,291]" fontSize="11" level="3" pageId="162" pageNumber="163" reason="3">
<emphasis id="A78D1B6230F852C6EDAAFEAF76A8001C" bold="true" box="[189,420,264,291]" pageId="162" pageNumber="163">Literature records</emphasis>
</heading>
</paragraph>
<paragraph id="9546C77030F852C6EDAAFE90752E00E1" blockId="162.[189,1399,311,478]" pageId="162" pageNumber="163">
<bibRefCitation id="F168BA8130F852C6EDAAFE90769E006E" author="Harold E." box="[189,402,311,337]" pageId="162" pageNumber="163" pagination="79 - 86" refId="ref123468" refString="Harold E. 1868 e. Diagnosen neuer Coprophagen. Coleopterologische Hefte 4: 79 - 86. Available from https: // www. biodiversitylibrary. org / page / 16069646 [accessed 16 Feb. 2024]." type="journal article" year="1868">Harold 1868e: 84</bibRefCitation>
(
<collectingCountry id="EDEE87E030F852C6ECB1FE90752A006E" box="[422,550,311,337]" name="Venezuela" pageId="162" pageNumber="163">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA8130F852C6EF71FE90741E006E" author="Candeze E." box="[614,786,311,337]" pageId="162" pageNumber="163" refId="ref113476" refString="Candeze E. 1891. Voyage de M. E. Simon au Venezuela (Decembre 1887 - Avril 1888), 15 e Memoire (1). Coleopteres - Lamellicornes determines par le Dr E. Candeze. Annales de la Societe entomologique de France 60: [329] - 332. Available from https: // www. biodiversitylibrary. org / page / 32547689 [accessed 14 Feb. 2024]." type="journal volume" year="1891">Candèze 1891</bibRefCitation>
(
<collectingCountry id="EDEE87E030F852C6EE32FE9074A8006E" box="[805,932,311,337]" name="Venezuela" pageId="162" pageNumber="163">Venezuela</collectingCountry>
:
<collectingRegion id="573D099230F852C6EEA4FE907326006E" box="[947,1066,311,337]" country="Venezuela" name="Carabobo" pageId="162" pageNumber="163">Carabobo</collectingRegion>
: San-Esteban). —
<bibRefCitation id="F168BA8130F852C6E806FE9076C3004B" author="Gillet J. J. E. &amp; Boucomont A." pageId="162" pageNumber="163" refId="ref121284" refString="Gillet J. J. E. &amp; Boucomont A. 1927. Scarabaeidae: Coprinae, Termitotroginae (Partes 38 et 90). W. Junk, Berlin. [We are not sure about the publication date of this book. The title page gives the range &quot; 1911 - 1927 &quot;, which could be interpreted as indicating that the book was published in parts between those years. What we suspect, however, is that 1911 refers to the first part of the Coprinae catalogue, published by Gillet (1911 b) - i. e., our previous reference -, whereas the continuation of the Coprinae catalogue, now added by the Termitotroginae, was published at once in 1927 with the joint-authorship of Gillet and Boucomont.]" type="book" year="1927">Gillet &amp; Boucomont 1927: 208</bibRefCitation>
(
<collectingCountry id="EDEE87E030F852C6ECF3FEFD7568004B" box="[484,612,346,372]" name="Venezuela" pageId="162" pageNumber="163">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA8130F852C6EFBFFEFC74B6004B" author="Boucomont A." box="[680,954,346,373]" pageId="162" pageNumber="163" pagination="293 - 332" refId="ref112419" refString="Boucomont A. 1932. Synopsis des Onthophagus d'Amerique du Sud (Col. Scarab.). Annales de la Societe entomologique de France 101: 293 - 332. https: // doi. org / 10.1080 / 21686351.1932.12280217" type="journal article" year="1932">Boucomont 1932: 329</bibRefCitation>
(
<collectingCountry id="EDEE87E030F852C6EED8FEFD7343004B" box="[975,1103,346,372]" name="Venezuela" pageId="162" pageNumber="163">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA8130F852C6E985FEFD77E500A7" author="Blackwelder R. E." pageId="162" pageNumber="163" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 212</bibRefCitation>
(
<collectingCountry id="EDEE87E030F852C6EDE9FEDA767200A8" box="[254,382,381,407]" name="Venezuela" pageId="162" pageNumber="163">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA8130F852C6ECD5FEDA75EC00A8" author="Delgado L. &amp; Peraza L. N. &amp; Deloya C." box="[450,736,381,407]" pageId="162" pageNumber="163" pagination="6 - 9" refId="ref116516" refString="Delgado L., Peraza L. N. &amp; Deloya C. 2006. Onthophagus yucatanus, a new species of the Clypeatus group from Mexico and Guatemala (Coleoptera: Scarabaeidae). Florida Entomologist 89 (1): 6 - 9. https: // doi. org / 10.1653 / 0015 - 4040 (2006) 89 [6: OYANSO] 2.0. CO; 2" type="journal article" year="2006">
Delgado
<emphasis id="A78D1B6230F852C6EF25FED9757F00A8" box="[562,627,381,407]" italics="true" pageId="162" pageNumber="163">et al.</emphasis>
2006: 6
</bibRefCitation>
(
<collectingCountry id="EDEE87E030F852C6EFE3FEDA747800A8" box="[756,884,381,407]" name="Venezuela" pageId="162" pageNumber="163">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA8130F852C6EEAFFEDA727B00A7" author="Pulido-Herrera L. A. &amp; Zunino M." box="[952,1399,381,408]" pageId="162" pageNumber="163" refId="ref136342" refString="Pulido-Herrera L. A. &amp; Zunino M. 2007. Catalogo preliminar de los Onthophagini de America (Coleoptera: Scarabaeine [sic]). In: Zunino M. &amp; Melic A. (eds) Escarabajos, Diversidad y Conservacion Biologica. Ensayos en Homenaje a Gonzalo Halffter: 93 - 129. Monografias 3 ercer Milenio m 3 m Vol. 7, Sociedad Entomologica Aragonesa, Zaragoza." type="book" year="2007">Pulido-Herrera &amp; Zunino 2007: 114</bibRefCitation>
(
<collectingCountry id="EDEE87E030F852C6EDD2FE0676490084" box="[197,325,417,443]" name="Venezuela" pageId="162" pageNumber="163">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA8130F852C6EC95FE0675D40084" author="Ferrer-Paris J. &amp; Sanchez-Mercado A. &amp; Rodriguez J. P." box="[386,728,417,443]" pageId="162" pageNumber="163" pagination="89 - 110" refId="ref119203" refString="Ferrer-Paris J. R, Sanchez-Mercado A. &amp; Rodriguez J. P. 2013. Optimizacion del muestreo de invertebrados tropicales: un ejemplo con escarabajos coprofagos (Coleoptera: Scarabaeinae) en Venezuela. Revista de Biologia Tropical 61 (1): 89 - 110. https: // doi. org / 10.15517 / rbt. v 61 i 1.10941" type="journal article" year="2013">
Ferrer-Paris
<emphasis id="A78D1B6230F852C6EF0FFE0675590084" box="[536,597,417,443]" italics="true" pageId="162" pageNumber="163">et al.</emphasis>
2013: 109
</bibRefCitation>
(
<collectingRegion id="573D099230F852C6EFFCFE0674450084" box="[747,841,417,443]" country="Venezuela" name="Bolivar" pageId="162" pageNumber="163">Bolívar</collectingRegion>
:
<collectingCountry id="EDEE87E030F852C6EE40FE06748D0084" box="[855,897,417,443]" name="Iceland" pageId="162" pageNumber="163">Isla</collectingCountry>
de Anacoco; Miranda: Altos de Pipe and Yaracuy: Hacienda Guáquira).
</paragraph>
</subSubSection>
</treatment>
</document>