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<mods:title id="3A17275A6B881157C9F77E7C49C2C133">The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution</mods:title>
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<mods:namePart id="CBAEC0229258A1255FF9E3DB62C2A7A6">Rosa, Cecilia Lozano De La</mods:namePart>
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<mods:affiliation id="C42C72E44DD85F36D4F31C35D54AD026">Instituto Venezolano de Investigaciones Científicas, Centro de Estudios Botánicos y Agroforestales, Laboratorio de Ecología Espacial, Maracaibo, estado Zulia, Venezuela.</mods:affiliation>
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<mods:namePart id="69E0F87F582394E8CB527AA27361C291">Cupello, Mario</mods:namePart>
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<mods:affiliation id="39F72EB24CFCA7367D7BF1434EED7511">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Curitiba, Paraná, Brazil. Currently at Department of Entomology, Texas A &amp; M University, College Station, TX, 77843, USA.</mods:affiliation>
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<mods:affiliation id="4E6FFEA9788CC7DB90FB5F4FFDD49823">Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, Cuiabá, Mato Grosso, Brazil.</mods:affiliation>
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<taxonomicName id="52F9BCF330D752E9EF13FC7A732302C7" ID-CoL="7ZQ3D" authority="(Mannerheim, 1829)" baseAuthorityName="Mannerheim" baseAuthorityYear="1829" box="[516,1071,989,1016]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="141" pageNumber="142" phylum="Arthropoda" rank="species" species="appendiculatus">
<emphasis id="A78D1B6230D752E9EF13FC7A743902C7" bold="true" box="[516,821,989,1016]" italics="true" pageId="141" pageNumber="142">Ontherus appendiculatus</emphasis>
(
<bibRefCitation id="F168BA8130D752E9EE52FC7A732B02C7" author="Mannerheim C. G. de" box="[837,1063,989,1016]" pageId="141" pageNumber="142" refId="ref129230" refString="Mannerheim C. G. de. 1829. Description de quarante nouvelles especes de scarabeides du Bresil; avec figures. Nouveaux Memoires de la Societe Imperiale des Naturalistes de Moscou 1: [29] - 80. Available from https: // www. biodiversitylibrary. org / page / 33157898 [accessed 16 Feb. 2024]." type="journal volume" year="1829">Mannerheim, 1829</bibRefCitation>
)
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<figureCitation id="0DC2DBF530D752E9EFF1FBA674410523" box="[742,845,1025,1052]" captionStart="Fig" captionStartId="142.[188,231,1959,1985]" captionTargetBox="[271,1306,278,1899]" captionTargetId="figure-18@142.[259,1327,265,1917]" captionTargetPageId="142" captionText="Fig. 55. Species distribution. A. Ontherus aphodioides Burmeister, 1874. B. Ontherus appendiculatus (Mannerheim, 1829). Green diamond = CEMT collection data; black circle = literature data." figureDoi="http://doi.org/10.5281/zenodo.13945617" httpUri="https://zenodo.org/record/13945617/files/figure.png" pageId="141" pageNumber="142">Fig. 55B</figureCitation>
</paragraph>
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<subSubSection id="DDE394FB30D752EBEDAAFBE476BE0493" lastPageId="143" lastPageNumber="144" pageId="141" pageNumber="142" type="reference_group">
<paragraph id="9546C77030D752E9EDAAFBE474CD0540" blockId="141.[189,1400,1090,1715]" pageId="141" pageNumber="142">
<treatmentCitationGroup id="B5E9E05E30D752E9EDAAFBE474CD0540" pageId="141" pageNumber="142">
<treatmentCitation id="1458E16130D752E9EDAAFBE475FF0563" author="Mannerheim C. G. de" box="[189,755,1090,1117]" page="43" pageId="141" pageNumber="142" year="1829">
<taxonomicName id="52F9BCF330D752E9EDAAFBE475FF0563" ID-CoL="7VDJK" authority="Mannerheim, 1829: 43" authorityName="Mannerheim" authorityPageNumber="43" authorityYear="1829" box="[189,755,1090,1117]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="141" pageNumber="142" phylum="Arthropoda" rank="species" species="appendiculatus">
<emphasis id="A78D1B6230D752E9EDAAFBE476C10563" box="[189,461,1090,1117]" italics="true" pageId="141" pageNumber="142">Copris appendiculatus</emphasis>
<bibRefCitation id="F168BA8130D752E9ECCCFBE575FF0563" author="Mannerheim C. G. de" box="[475,755,1090,1116]" pageId="141" pageNumber="142" refId="ref129230" refString="Mannerheim C. G. de. 1829. Description de quarante nouvelles especes de scarabeides du Bresil; avec figures. Nouveaux Memoires de la Societe Imperiale des Naturalistes de Moscou 1: [29] - 80. Available from https: // www. biodiversitylibrary. org / page / 33157898 [accessed 16 Feb. 2024]." type="journal volume" year="1829">Mannerheim, 1829: 43</bibRefCitation>
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(original description). Type locality: 
<collectingCountry id="EDEE87E030D752E9E9DDFBE572190563" box="[1226,1301,1090,1116]" name="Brazil" pageId="141" pageNumber="142">Brazil</collectingCountry>
. Name-bearing type: 
<typeStatus id="4A4279D230D752E9EC9BFBC276FF0540" box="[396,499,1125,1151]" pageId="141" pageNumber="142" type="holotype">holotype</typeStatus>
(ZMAS) (
<treatmentCitation id="1458E16130D752E9EF79FBC274090540" author="Genier F." box="[622,773,1125,1151]" pageId="141" pageNumber="142" year="1996">
<bibRefCitation id="F168BA8130D752E9EF79FBC274090540" author="Genier F." box="[622,773,1125,1151]" pageId="141" pageNumber="142" pagination="1 - 169" refId="ref120431" refString="Genier F. 1996. A revision of the Neotropical genus Ontherus Erichson (Coleoptera: Scarabaeidae, Scarabaeinae). Memoirs of the Entomological Society of Canada 170: 1 - 169. https: // doi. org / 10.4039 / entm 128170 fv" type="journal article" year="1996">Génier 1996</bibRefCitation>
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), not examined.
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</paragraph>
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<treatmentCitationGroup id="B5E9E05E30D752E9EDAAFB2E741804DF" pageId="141" pageNumber="142">
<treatmentCitation id="1458E16130D752E9EDAAFB2E75A9059C" author="Blanchard E." box="[189,677,1160,1187]" page="181" pageId="141" pageNumber="142" year="1846">
<taxonomicName id="52F9BCF330D752E9EDAAFB2E75A9059C" ID-CoL="7VDM8" authority="Blanchard, 1846: 181" authorityName="Blanchard" authorityPageNumber="181" authorityYear="1846" box="[189,677,1160,1187]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="141" pageNumber="142" phylum="Arthropoda" rank="species" species="rotundatus">
<emphasis id="A78D1B6230D752E9EDAAFB2E769A059D" box="[189,406,1160,1187]" italics="true" pageId="141" pageNumber="142">Copris rotundatus</emphasis>
<bibRefCitation id="F168BA8130D752E9ECB6FB2F75A9059C" author="Blanchard E." box="[417,677,1160,1187]" pageId="141" pageNumber="142" pagination="153 - 168" refId="ref111318" refString="Blanchard E. 1846. [livraison 79: 153 - 168, pl. 28; livraison 80: 169 - 184, pl. 31]. In: Blanchard E. &amp; Brulle G. A. 1835 - 1846. Voyage dans l'Amerique Meridionale (le Bresil, la Republique orientale de L'Uruguay, la Republique Argentine, la Patagonie, la Republique du Chili, la Republique de Bolivia, la Republique du Perou), execute pendant les annees 1826, 1827, 1828, 1829, 1830, 1831, 1832 et 1833, par Alcide d'Orbigny. Tome Sixieme. 2. e partie: Insectes. P. Bertrand, Paris, and Ve. Lerrault, Strasbourg. [Both livraisons date from 1846. For details on the publication date of livraison 79, see Cupello et al. (2023 a: 89 - 90); the earliest evidence of publication of livraison 80 dates from 18 Dec. 1846 (Neal Evenhuis, pers. com. to MC, 3 Sep. 2019).]" type="journal article" year="1846">Blanchard, 1846: 181</bibRefCitation>
</taxonomicName>
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(original description). Type locality: 
<collectingCountry id="EDEE87E030D752E9E97AFB2F73EB059D" box="[1133,1255,1160,1186]" name="Argentina" pageId="141" pageNumber="142">Argentina</collectingCountry>
: 
<collectingRegion id="573D099230D752E9E9E2FB2F727E059D" box="[1269,1394,1160,1186]" country="Argentina" name="Corrientes" pageId="141" pageNumber="142">Corrientes</collectingRegion>
, and 
<collectingCountry id="EDEE87E030D752E9EC08FB0B768605F9" box="[287,394,1196,1222]" name="Uruguay" pageId="141" pageNumber="142">Uruguay</collectingCountry>
: 
<collectingRegion id="573D099230D752E9EC80FB0B752505F9" box="[407,553,1196,1222]" country="Uruguay" name="Montevideo" pageId="141" pageNumber="142">Montevideo</collectingRegion>
. Name-bearing type: 
<typeStatus id="4A4279D230D752E9EE3AFB0B749805F9" box="[813,916,1196,1222]" pageId="141" pageNumber="142" type="syntype">syntypes</typeStatus>
(MNHN), not examined. 
<treatmentCitation id="1458E16130D752E9E9D9FB0B727A05F9" author="Genier F." box="[1230,1398,1196,1222]" pageId="141" pageNumber="142" year="1996">
<bibRefCitation id="F168BA8130D752E9E9D9FB0B727A05F9" author="Genier F." box="[1230,1398,1196,1222]" pageId="141" pageNumber="142" pagination="1 - 169" refId="ref120431" refString="Genier F. 1996. A revision of the Neotropical genus Ontherus Erichson (Coleoptera: Scarabaeidae, Scarabaeinae). Memoirs of the Entomological Society of Canada 170: 1 - 169. https: // doi. org / 10.4039 / entm 128170 fv" type="journal article" year="1996">Génier (1996)</bibRefCitation>
</treatmentCitation>
indicated an MNHN specimen from Corrientes as the 
<typeStatus id="4A4279D230D752E9EE93FB6874E205D6" box="[900,1006,1231,1257]" pageId="141" pageNumber="142" type="holotype">holotype</typeStatus>
, but this is incorrect. 
<treatmentCitation id="1458E16130D752E9E9E9FB6876340433" author="Blanchard E." pageId="141" pageNumber="142" year="1846">
<bibRefCitation id="F168BA8130D752E9E9E9FB6876340433" author="Blanchard E." pageId="141" pageNumber="142" pagination="153 - 168" refId="ref111318" refString="Blanchard E. 1846. [livraison 79: 153 - 168, pl. 28; livraison 80: 169 - 184, pl. 31]. In: Blanchard E. &amp; Brulle G. A. 1835 - 1846. Voyage dans l'Amerique Meridionale (le Bresil, la Republique orientale de L'Uruguay, la Republique Argentine, la Patagonie, la Republique du Chili, la Republique de Bolivia, la Republique du Perou), execute pendant les annees 1826, 1827, 1828, 1829, 1830, 1831, 1832 et 1833, par Alcide d'Orbigny. Tome Sixieme. 2. e partie: Insectes. P. Bertrand, Paris, and Ve. Lerrault, Strasbourg. [Both livraisons date from 1846. For details on the publication date of livraison 79, see Cupello et al. (2023 a: 89 - 90); the earliest evidence of publication of livraison 80 dates from 18 Dec. 1846 (Neal Evenhuis, pers. com. to MC, 3 Sep. 2019).]" type="journal article" year="1846">Blanchard (1846)</bibRefCitation>
</treatmentCitation>
clearly had more than 
<specimenCount id="83FF0CF930D752E9EF51FB5475EA0433" box="[582,742,1266,1293]" count="1" pageId="141" pageNumber="142" type="generic">one specimen</specimenCount>
in front of him for the description of 
<taxonomicName id="52F9BCF330D752E9E98AFB5472310433" ID-CoL="7VDM8" authorityName="Blanchard" authorityYear="1846" box="[1181,1341,1266,1293]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="141" pageNumber="142" phylum="Arthropoda" rank="species" species="rotundatus">
<emphasis id="A78D1B6230D752E9E98AFB5472310433" box="[1181,1341,1266,1293]" italics="true" pageId="141" pageNumber="142">C. rotundatus</emphasis>
</taxonomicName>
‒ he gave a length range instead of a precise figure (“Long. 9 à 10 millim.”) and cited the examination of material from both Corrientes and 
<collectingRegion id="573D099230D752E9EFB9FA9F7430046D" box="[686,828,1336,1362]" country="Uruguay" name="Montevideo" pageId="141" pageNumber="142">Montevideo</collectingRegion>
‒ and, hence, the name-bearing type cannot be a 
<typeStatus id="4A4279D230D752E9EC16FAFB76640449" box="[257,360,1372,1398]" pageId="141" pageNumber="142" type="holotype">holotype</typeStatus>
by monotypy. A 
<typeStatus id="4A4279D230D752E9EF2FFAFB75930449" box="[568,671,1372,1398]" pageId="141" pageNumber="142" type="holotype">holotype</typeStatus>
by original designation is not the case either, for Blanchard, in the spirit of the time, did not designate a sole specimen as the name-bearing type. Instead of a 
<typeStatus id="4A4279D230D752E9EC15FA0576600483" box="[258,364,1442,1468]" pageId="141" pageNumber="142" type="holotype">holotype</typeStatus>
, the name-bearing type is a series of at least 
<specimenCount id="83FF0CF930D752E9EE83FA0473380482" box="[916,1076,1443,1469]" count="2" pageId="141" pageNumber="142" type="generic" typeStatus="syntype">two syntypes</specimenCount>
, one from each of the two localities mentioned in the original description.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030D752E9EDAAFA4E73730719" blockId="141.[189,1400,1090,1715]" pageId="141" pageNumber="142">
<treatmentCitationGroup id="B5E9E05E30D752E9EDAAFA4E73730719" pageId="141" pageNumber="142">
<treatmentCitation id="1458E16130D752E9EDAAFA4E7577073C" author="Blanchard E." box="[189,635,1512,1539]" page="181" pageId="141" pageNumber="142" year="1846">
<taxonomicName id="52F9BCF330D752E9EDAAFA4E7577073C" ID-CoL="7VDLY" authority="Blanchard, 1846: 181" authorityName="Blanchard" authorityPageNumber="181" authorityYear="1846" box="[189,635,1512,1539]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="141" pageNumber="142" phylum="Arthropoda" rank="species" species="polynice">
<emphasis id="A78D1B6230D752E9EDAAFA4E767B073D" box="[189,375,1512,1539]" italics="true" pageId="141" pageNumber="142">Copris polynice</emphasis>
<bibRefCitation id="F168BA8130D752E9EC69FA4F7577073C" author="Blanchard E." box="[382,635,1512,1539]" pageId="141" pageNumber="142" pagination="153 - 168" refId="ref111318" refString="Blanchard E. 1846. [livraison 79: 153 - 168, pl. 28; livraison 80: 169 - 184, pl. 31]. In: Blanchard E. &amp; Brulle G. A. 1835 - 1846. Voyage dans l'Amerique Meridionale (le Bresil, la Republique orientale de L'Uruguay, la Republique Argentine, la Patagonie, la Republique du Chili, la Republique de Bolivia, la Republique du Perou), execute pendant les annees 1826, 1827, 1828, 1829, 1830, 1831, 1832 et 1833, par Alcide d'Orbigny. Tome Sixieme. 2. e partie: Insectes. P. Bertrand, Paris, and Ve. Lerrault, Strasbourg. [Both livraisons date from 1846. For details on the publication date of livraison 79, see Cupello et al. (2023 a: 89 - 90); the earliest evidence of publication of livraison 80 dates from 18 Dec. 1846 (Neal Evenhuis, pers. com. to MC, 3 Sep. 2019).]" type="journal article" year="1846">Blanchard, 1846: 181</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description). Type locality: 
<collectingCountry id="EDEE87E030D752E9E926FA4F7380073D" box="[1073,1164,1512,1538]" name="Bolivia" pageId="141" pageNumber="142">Bolivia</collectingCountry>
: 
<collectingRegion id="573D099230D752E9E980FA4E7210073C" box="[1175,1308,1513,1539]" country="Bolivia" name="Santa Cruz" pageId="141" pageNumber="142">Santa Cruz</collectingRegion>
. Name-bearing type: 
<typeStatus id="4A4279D230D752E9EC9BF9AB76F40719" box="[396,504,1548,1574]" pageId="141" pageNumber="142" type="lectotype">lectotype</typeStatus>
(MNHN), designated by 
<treatmentCitation id="1458E16130D752E9EE33F9AB74C00719" author="Genier F." box="[804,972,1548,1574]" pageId="141" pageNumber="142" year="1996">
<bibRefCitation id="F168BA8130D752E9EE33F9AB74C00719" author="Genier F." box="[804,972,1548,1574]" pageId="141" pageNumber="142" pagination="1 - 169" refId="ref120431" refString="Genier F. 1996. A revision of the Neotropical genus Ontherus Erichson (Coleoptera: Scarabaeidae, Scarabaeinae). Memoirs of the Entomological Society of Canada 170: 1 - 169. https: // doi. org / 10.4039 / entm 128170 fv" type="journal article" year="1996">Génier (1996)</bibRefCitation>
</treatmentCitation>
, not examined.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030D752E9EDAAF98873150753" blockId="141.[189,1400,1090,1715]" pageId="141" pageNumber="142">
<treatmentCitationGroup id="B5E9E05E30D752E9EDAAF98873150753" pageId="141" pageNumber="142">
<treatmentCitation id="1458E16130D752E9EDAAF988759E0775" author="Erichson W. F." box="[189,658,1583,1610]" page="564" pageId="141" pageNumber="142" year="1848">
<taxonomicName id="52F9BCF330D752E9EDAAF988759E0775" authority="Erichson, 1848: 564" authorityName="Erichson" authorityPageNumber="564" authorityYear="1848" box="[189,658,1583,1610]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="141" pageNumber="142" phylum="Arthropoda" rank="species" species="quadratus">
<emphasis id="A78D1B6230D752E9EDAAF988769C0776" box="[189,400,1583,1609]" italics="true" pageId="141" pageNumber="142">Copris quadratus</emphasis>
<bibRefCitation id="F168BA8130D752E9EC8BF988759E0775" author="Erichson W. F." box="[412,658,1583,1610]" pageId="141" pageNumber="142" pagination="553 - 1260" refId="ref117915" refString="Erichson W. F. 1848. Insecten. In: Schomburgk R. Reisen in Britisch-Guiana in den Jahren 1840 - 1844. Dritter Theil: 553 - 1260. J. J. Weber, Leipzig. https: // doi. org / 10.5962 / bhl. title. 109982" type="book chapter" year="1848">Erichson, 1848: 564</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description). Type locality: 
<collectingCountry id="EDEE87E030D752E9E977F98873B30776" box="[1120,1215,1583,1609]" name="Guyana" pageId="141" pageNumber="142">Guyana</collectingCountry>
. Name-bearing type: 
<typeStatus id="4A4279D230D752E9EC3BF9F576940753" box="[300,408,1618,1644]" pageId="141" pageNumber="142" type="lectotype">lectotype</typeStatus>
(MFNB), designated by 
<treatmentCitation id="1458E16130D752E9EFA9F9F5746A0753" author="Genier F." box="[702,870,1618,1644]" pageId="141" pageNumber="142" year="1996">
<bibRefCitation id="F168BA8130D752E9EFA9F9F5746A0753" author="Genier F." box="[702,870,1618,1644]" pageId="141" pageNumber="142" pagination="1 - 169" refId="ref120431" refString="Genier F. 1996. A revision of the Neotropical genus Ontherus Erichson (Coleoptera: Scarabaeidae, Scarabaeinae). Memoirs of the Entomological Society of Canada 170: 1 - 169. https: // doi. org / 10.4039 / entm 128170 fv" type="journal article" year="1996">Génier (1996)</bibRefCitation>
</treatmentCitation>
, not examined.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030D752E9EDAAF9D273F7078C" blockId="141.[189,1400,1090,1715]" pageId="141" pageNumber="142">
<treatmentCitationGroup id="B5E9E05E30D752E9EDAAF9D273F7078C" pageId="141" pageNumber="142">
<treatmentCitation id="1458E16130D752E9EDAAF9D275CE07B0" author="Burmeister H." box="[189,706,1653,1679]" page="127" pageId="141" pageNumber="142" year="1874">
<taxonomicName id="52F9BCF330D752E9EDAAF9D275CE07B0" ID-CoL="7XM89" authority="Burmeister, 1874: 127" authorityName="Burmeister" authorityPageNumber="127" authorityYear="1874" box="[189,706,1653,1679]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="141" pageNumber="142" phylum="Arthropoda" rank="species" species="contractus">
<emphasis id="A78D1B6230D752E9EDAAF9D276A307B0" box="[189,431,1653,1679]" italics="true" pageId="141" pageNumber="142">Ontherus contractus</emphasis>
<bibRefCitation id="F168BA8130D752E9ECAEF9D275CE07B0" author="Burmeister H." box="[441,706,1653,1679]" pageId="141" pageNumber="142" pagination="120 - 133" refId="ref113171" refString="Burmeister H. 1874. Lamellicornia Argentina [Fortsetzung]. Entomologische Zeitung Stettin 35: 120 - 133. Available from https: // www. biodiversitylibrary. org / page / 9236010 [accessed 14 Feb. 2024]." type="journal article" year="1874">Burmeister, 1874: 127</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description). Type locality: 
<collectingCountry id="EDEE87E030D752E9E969F9D273F407B0" box="[1150,1272,1653,1679]" name="Argentina" pageId="141" pageNumber="142">Argentina</collectingCountry>
: 
<collectingRegion id="573D099230D752E9E814F9D2727907B0" box="[1283,1397,1653,1679]" country="Argentina" name="Tucuman" pageId="141" pageNumber="142">Tucumán</collectingRegion>
. Name-bearing type: 
<typeStatus id="4A4279D230D752E9ECCAF93F7545078D" box="[477,585,1688,1714]" pageId="141" pageNumber="142" type="lectotype">lectotype</typeStatus>
(MACN), designated by 
<bibRefCitation id="F168BA8130D752E9EE64F93F7310078D" author="Genier F." box="[883,1052,1688,1714]" pageId="141" pageNumber="142" pagination="1 - 169" refId="ref120431" refString="Genier F. 1996. A revision of the Neotropical genus Ontherus Erichson (Coleoptera: Scarabaeidae, Scarabaeinae). Memoirs of the Entomological Society of Canada 170: 1 - 169. https: // doi. org / 10.4039 / entm 128170 fv" type="journal article" year="1996">
<treatmentCitation id="1458E16130D752E9EE64F93F731E078D" author="Genier F." box="[883,1042,1688,1714]" pageId="141" pageNumber="142" year="1996">Génier (1996</treatmentCitation>
)
</bibRefCitation>
, examined by MC.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030D752EBEDAAF97D764000B2" blockId="141.[189,1399,1754,1992]" lastBlockId="143.[189,1399,265,1452]" lastPageId="143" lastPageNumber="144" pageId="141" pageNumber="142">
<treatmentCitationGroup id="B5E9E05E30D752EBEDAAF97D764000B2" lastPageId="143" lastPageNumber="144" pageId="141" pageNumber="142">
<emphasis id="A78D1B6230D752E9EDAAF97D750907CB" box="[189,517,1754,1780]" italics="true" pageId="141" pageNumber="142">
<taxonomicName id="52F9BCF330D752E9EDAAF97D76E607CB" ID-CoL="7ZQ3D" baseAuthorityName="Mannerheim" baseAuthorityYear="1829" box="[189,490,1754,1780]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="141" pageNumber="142" phylum="Arthropoda" rank="species" species="appendiculatus">Ontherus appendiculatus</taxonomicName>
<emphasis id="A78D1B6230D752E9ECE1F97D750907CB" bold="true" box="[502,517,1754,1780]" italics="true" pageId="141" pageNumber="142">–</emphasis>
</emphasis>
<treatmentCitation id="1458E16130D752E9EF06F97D740407CB" author="Harold E." box="[529,776,1754,1781]" page="1008" pageId="141" pageNumber="142" year="1869">
<bibRefCitation id="F168BA8130D752E9EF06F97D740407CB" author="Harold E." box="[529,776,1754,1781]" pageId="141" pageNumber="142" refId="ref123629" refString="Harold E. 1869 d. Catalogus coleopterorum. Hucusque descriptorum, synonymicus et systematicus. Tom. IV. Scarabaeidae. E. H. Gummi, Munich. https: // doi. org / 10.5962 / bhl. title. 9089" type="book" year="1869">Harold 1869d: 1008</bibRefCitation>
</treatmentCitation>
(catalogue, cited for Tijuco). — 
<treatmentCitation id="1458E16130D752E9E9BFF97D727A07CB" author="Bruch C." box="[1192,1398,1754,1780]" page="186" pageId="141" pageNumber="142" year="1911">
<bibRefCitation id="F168BA8130D752E9E9BFF97D727A07CB" author="Bruch C." box="[1192,1398,1754,1780]" pageId="141" pageNumber="142" pagination="143 - 225" refId="ref112509" refString="Bruch C. 1911. Catalogo sistematico de los coleopteros de la Republica Argentina. Revista del Museo de La Plata 17: 143 - 225. https: // doi. org / 10.5962 / bhl. title. 8799" type="journal article" year="1911">Bruch 1911: 186</bibRefCitation>
</treatmentCitation>
(catalogue, cited for Chaco, Santa Fe, Tucumán); 
<treatmentCitation id="1458E16130D752E9EE4AF95A74D60627" author="Bruch C." box="[861,986,1789,1816]" page="541" pageId="141" pageNumber="142" year="1915">
<bibRefCitation id="F168BA8130D752E9EE4AF95A74D60627" author="Bruch C." box="[861,986,1789,1816]" pageId="141" pageNumber="142" pagination="538 - 573" refId="ref112550" refString="Bruch C. 1915. Suplemento al catalogo sistematico de los coleopteros de la Republica Argentina. I (addenda, corrigenda y resumen). Revista del Museo de La Plata 19 (2): 538 - 573. Available from http: // sedici. unlp. edu. ar / handle / 10915 / 155145 [accessed 14 Feb. 2024]." type="journal article" year="1915">1915: 541</bibRefCitation>
</treatmentCitation>
(catalogue). — 
<treatmentCitation id="1458E16130D752E9E9BCF95A727A0627" author="Gillet J. J. E." box="[1195,1398,1789,1816]" page="57" pageId="141" pageNumber="142" year="1911">
<bibRefCitation id="F168BA8130D752E9E9BCF95A727A0627" author="Gillet J. J. E." box="[1195,1398,1789,1816]" pageId="141" pageNumber="142" refId="ref121235" refString="Gillet J. J. E. 1911 b. Coleopterorum Catalogus. Pars 38: Scarabaeidae: Coprinae I. W. Junk, Berlin. Available from https: // www. biodiversitylibrary. org / page / 48405679 [accesseed 16 Feb. 2024]." type="url" year="1911">Gillet 1911b: 57</bibRefCitation>
</treatmentCitation>
(catalogue, cited for 
<collectingCountry id="EDEE87E030D752E9ECF0F88775380605" box="[487,564,1824,1850]" name="Brazil" pageId="141" pageNumber="142">Brazil</collectingCountry>
). — 
<treatmentCitation id="1458E16130D752E9EF64F88674310604" author="Lucas R." box="[627,829,1824,1851]" page="459" pageId="141" pageNumber="142" year="1920">
<bibRefCitation id="F168BA8130D752E9EF64F88674310604" author="Lucas R." box="[627,829,1824,1851]" pageId="141" pageNumber="142" refId="ref128561" refString="Lucas R. 1920. Catalogus alphabeticus generum et subgenerum Coleopterorum orbis terrarum totius (famil., trib., subtr., sect. incl.). Pars I. Nicolaische, Berlin." type="book" year="1920">Lucas 1920: 459</bibRefCitation>
</treatmentCitation>
(catalogue). — 
<treatmentCitation id="1458E16130D752E9E911F88672090605" author="Boucomont A." box="[1030,1285,1824,1851]" page="6" pageId="141" pageNumber="142" year="1928">
<bibRefCitation id="F168BA8130D752E9E911F88672090605" author="Boucomont A." box="[1030,1285,1824,1851]" pageId="141" pageNumber="142" pagination="1 - 8" refId="ref112356" refString="Boucomont A. 1928 b. Wissenschaftliche Ergebnisse der schwedischen entomologischen Reise des Herrn. Dr. A. Roman in Amazonas 1914 - 1915 (und 1923 - 1924), 12. Scarabaeidae. Arkiv for Zoologi 19 A (22): 1 - 8. https: // doi. org / 10.5962 / bhl. part. 1064" type="journal article" year="1928">Boucomont 1928b: 6</bibRefCitation>
</treatmentCitation>
(list). — 
<treatmentCitation id="1458E16130D752E9EDFDF8E376F60661" author="Luederwaldt H." box="[234,506,1860,1886]" page="107" pageId="141" pageNumber="142" year="1930">
<bibRefCitation id="F168BA8130D752E9EDFDF8E376F60661" author="Luederwaldt H." box="[234,506,1860,1886]" pageId="141" pageNumber="142" pagination="105 - 108" refId="ref128717" refString="Luederwaldt H. 1930. Especies novas de Ontherus. Archivos do Instituto Biologico 3: 105 - 108." type="journal article" year="1930">Luederwaldt 1930: 107</bibRefCitation>
</treatmentCitation>
(comments); 
<treatmentCitation id="1458E16130D752E9EF8CF8E374110661" author="Luederwaldt H." box="[667,797,1860,1886]" page="371" pageId="141" pageNumber="142" year="1931">
<bibRefCitation id="F168BA8130D752E9EF8CF8E374110661" author="Luederwaldt H." box="[667,797,1860,1886]" firstAuthor="a" pageId="141" pageNumber="142" pagination="363 - 422" refId="ref128737" refString="Luederwaldt H. 1931 a. O genero Ontherus (Coleop.) (Lamellic-Coprid. - Pinot.) com uma chave, para a determinacao dos pinotides americanos. Revista do Museu Paulista 17: 363 - 422 + 1 pl." type="journal article" year="1931">1931a: 371</bibRefCitation>
</treatmentCitation>
–372, 381–391 (key, distribution, redescription). — 
<treatmentCitation id="1458E16130D752E9EDFEF8C0752806BD" author="Pessoa S. B. &amp; Lane F." box="[233,548,1895,1922]" page="457" pageId="141" pageNumber="142" year="1941">
<bibRefCitation id="F168BA8130D752E9EDFEF8C0752806BD" author="Pessoa S. B. &amp; Lane F." box="[233,548,1895,1922]" pageId="141" pageNumber="142" pagination="389 - 504" refId="ref135859" refString="Pessoa S. B. &amp; Lane F. 1941. Coleopteros necrofagos de interesse medico-legal: ensaio monografico sobre a familia Scarabaeidae de S. Paulo e regioes vizinhas. Arquivos de Zoologia do Estado de Sao Paulo 2: 389 - 504. https: // doi. org / 10.11606 / issn. 2176 - 7793.19412389 - 504" type="journal article" year="1941">Pessôa &amp; Lane 1941: 457</bibRefCitation>
</treatmentCitation>
, 458 (key, redescription, distribution). — 
<treatmentCitation id="1458E16130D752E9E937F8C0723B06BE" author="Blackwelder R. E." box="[1056,1335,1895,1921]" page="206" pageId="141" pageNumber="142" year="1944">
<bibRefCitation id="F168BA8130D752E9E937F8C0723B06BE" author="Blackwelder R. E." box="[1056,1335,1895,1921]" pageId="141" pageNumber="142" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 206</bibRefCitation>
</treatmentCitation>
(list, cited for 
<collectingCountry id="EDEE87E030D752E9EC40F82D76A8069B" box="[343,420,1930,1956]" name="Brazil" pageId="141" pageNumber="142">Brazil</collectingCountry>
). — 
<treatmentCitation id="1458E16130D752E9ECC9F82D75AF069B" author="Blanco J." box="[478,675,1930,1956]" page="44" pageId="141" pageNumber="142" year="1987">
<bibRefCitation id="F168BA8130D752E9ECC9F82D75AF069B" author="Blanco J." box="[478,675,1930,1956]" pageId="141" pageNumber="142" pagination="39 - 46" refId="ref111553" refString="Blanco J. 1987. Catalogo de los Scarabaeidae (Coleoptera) coprofagos y necrofagos, del estado Tachira. Venezuela. Revista Cientifica Unet 1: 39 - 46." type="journal article" year="1987">Blanco 1987: 44</bibRefCitation>
</treatmentCitation>
(catalogue). — 
<treatmentCitation id="1458E16130D752E9EE71F82D732B069B" author="Genier F." box="[870,1063,1930,1956]" page="78" pageId="141" pageNumber="142" year="1996">
<bibRefCitation id="F168BA8130D752E9EE71F82D732B069B" author="Genier F." box="[870,1063,1930,1956]" pageId="141" pageNumber="142" pagination="1 - 169" refId="ref120431" refString="Genier F. 1996. A revision of the Neotropical genus Ontherus Erichson (Coleoptera: Scarabaeidae, Scarabaeinae). Memoirs of the Entomological Society of Canada 170: 1 - 169. https: // doi. org / 10.4039 / entm 128170 fv" type="journal article" year="1996">Génier 1996: 78</bibRefCitation>
</treatmentCitation>
(revision, cited as synonym of 
<taxonomicName id="52F9BCF330D752E9EC19F80976E506F8" ID-CoL="7ZQ3D" baseAuthorityName="Mannerheim" baseAuthorityYear="1829" box="[270,489,1965,1992]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="141" pageNumber="142" phylum="Arthropoda" rank="species" species="appendiculatus">
<emphasis id="A78D1B6230D752E9EC19F80976E506F8" box="[270,489,1965,1992]" italics="true" pageId="141" pageNumber="142">O. appendiculatus</emphasis>
</taxonomicName>
). — 
<treatmentCitation id="1458E16130D752E9EF38F80A741806F8" author="Escobar F." box="[559,788,1965,1991]" page="208" pageId="141" pageNumber="142" year="2000">
<bibRefCitation id="F168BA8130D752E9EF38F80A741806F8" author="Escobar F." box="[559,788,1965,1991]" pageId="141" pageNumber="142" pagination="197 - 210" refId="ref118039" refString="Escobar F. 2000. Diversidad y distribution de los escarabajos del estiercol (Coleoptera: Scarabaeidae: Scarabaeinae) de Colombia. In: Martin-Piera F., Morrone J. J. &amp; Melic A. (eds.) Hacia un proyecto CYTED para el Inventario y Estimacion de la Diversidad Entomologica en Iberoamerica: PRIBES- 2000: 197 - 210. m 3 m: Monografias Tercer Milenio, vol. 1. Sociedad Entomologica Aragonesa (SEA), Zaragoza." type="book chapter" year="2000">Escobar 2000: 208</bibRefCitation>
</treatmentCitation>
(checklist for 
<collectingCountry id="EDEE87E030D752E9EEDCF80A734806F8" box="[971,1092,1965,1991]" name="Colombia" pageId="141" pageNumber="142">Colombia</collectingCountry>
). — 
<treatmentCitation id="1458E16130D752EBE993F80A761A001C" author="Medina C. A. &amp; Lopera-Toro A. &amp; Vitolo A. &amp; Gill B." lastPageId="143" lastPageNumber="144" page="138" pageId="141" pageNumber="142" year="2001">
<bibRefCitation id="F168BA8130D752EBE993F80A761A001C" author="Medina C. A. &amp; Lopera-Toro A. &amp; Vitolo A. &amp; Gill B." lastPageId="143" lastPageNumber="144" pageId="141" pageNumber="142" pagination="131 - 144" refId="ref130799" refString="Medina C. A., Lopera-Toro A., Vitolo A. &amp; Gill B. 2001. Escarabajos coprofagos (Coleoptera: Scarabaeinae) de Colombia. Biota Colombiana 2: 131 - 144." type="journal article" year="2001">
Medina 
<emphasis id="A78D1B6230D752E9E9FEF809722406F8" box="[1257,1320,1965,1991]" italics="true" pageId="141" pageNumber="142">et al.</emphasis>
2001: 138
</bibRefCitation>
</treatmentCitation>
(checklist for 
<collectingCountry id="EDEE87E030D552EBECC6FEAE7546001C" box="[465,586,265,291]" name="Colombia" pageId="143" pageNumber="144">Colombia</collectingCountry>
). — 
<treatmentCitation id="1458E16130D552EBEF99FEAE7318001C" author="Pulido-Herrera L. A. &amp; Zunino M." box="[654,1044,265,291]" page="307" pageId="143" pageNumber="144" year="2007">
<bibRefCitation id="F168BA8130D552EBEF99FEAE7318001C" author="Pulido-Herrera L. A. &amp; Zunino M." box="[654,1044,265,291]" firstAuthor="Pulido-Herrera" pageId="143" pageNumber="144" refId="ref136342" refString="Pulido-Herrera L. A. &amp; Zunino M. 2007. Catalogo preliminar de los Onthophagini de America (Coleoptera: Scarabaeine [sic]). In: Zunino M. &amp; Melic A. (eds) Escarabajos, Diversidad y Conservacion Biologica. Ensayos en Homenaje a Gonzalo Halffter: 93 - 129. Monografias 3 ercer Milenio m 3 m Vol. 7, Sociedad Entomologica Aragonesa, Zaragoza." type="book" year="2007">
Pulido-Herrera 
<emphasis id="A78D1B6230D552EBEE5CFEAD7487001C" box="[843,907,265,291]" italics="true" pageId="143" pageNumber="144">et al.</emphasis>
2007: 307
</bibRefCitation>
</treatmentCitation>
(cited for Andean region of 
<collectingCountry id="EDEE87E030D552EBEDFDFE8B766F0079" box="[234,355,300,326]" name="Colombia" pageId="143" pageNumber="144">Colombia</collectingCountry>
). — 
<treatmentCitation id="1458E16130D552EBECB2FE8B74590079" author="Medina C. A. &amp; Pulido-Herrera L. A." box="[421,853,300,326]" page="60" pageId="143" pageNumber="144" year="2009">
<bibRefCitation id="F168BA8130D552EBECB2FE8B74590079" author="Medina C. A. &amp; Pulido-Herrera L. A." box="[421,853,300,326]" pageId="143" pageNumber="144" pagination="55 - 62" refId="ref130761" refString="Medina C. A. &amp; Pulido-Herrera L. A. 2009. Escarabajos coprofagos (Coleoptera: Scarabaeinae) de la Orinoquia Colombiana. Biota Colombiana 10 (1 - 2): 55 - 62." type="journal article" year="2009">Medina &amp; Pulido-Herrera 2009: 60</bibRefCitation>
</treatmentCitation>
(diversity). — 
<treatmentCitation id="1458E16130D552EBE90EFE8B727A0079" author="Ferrer-Paris J. &amp; Sanchez-Mercado A. &amp; Rodriguez J. P." box="[1049,1398,300,326]" page="109" pageId="143" pageNumber="144" year="2013">
<bibRefCitation id="F168BA8130D552EBE90EFE8B727A0079" author="Ferrer-Paris J. &amp; Sanchez-Mercado A. &amp; Rodriguez J. P." box="[1049,1398,300,326]" pageId="143" pageNumber="144" pagination="89 - 110" refId="ref119203" refString="Ferrer-Paris J. R, Sanchez-Mercado A. &amp; Rodriguez J. P. 2013. Optimizacion del muestreo de invertebrados tropicales: un ejemplo con escarabajos coprofagos (Coleoptera: Scarabaeinae) en Venezuela. Revista de Biologia Tropical 61 (1): 89 - 110. https: // doi. org / 10.15517 / rbt. v 61 i 1.10941" type="journal article" year="2013">
Ferrer-Paris 
<emphasis id="A78D1B6230D552EBE9A7FE8A73FC0079" box="[1200,1264,300,326]" italics="true" pageId="143" pageNumber="144">et al.</emphasis>
2013: 109
</bibRefCitation>
</treatmentCitation>
(list). — 
<treatmentCitation id="1458E16130D552EBEC42FEF775440055" author="Franca F. M. &amp; Korasaki V. &amp; Louzada J. &amp; Vaz-de-Mello F. Z." box="[341,584,335,362]" page="3" pageId="143" pageNumber="144" year="2016">
<bibRefCitation id="F168BA8130D552EBEC42FEF775440055" author="Franca F. M. &amp; Korasaki V. &amp; Louzada J. &amp; Vaz-de-Mello F. Z." box="[341,584,335,362]" pageId="143" pageNumber="144" pagination="1 - 9" refId="ref119681" refString="Franca F. M., Korasaki V., Louzada J. &amp; Vaz-de-Mello F. Z. 2016. First report on dung beetles in intra-Amazonian savannahs in Roraima, Brazil. Biota Neotropica 16 (1): 1 - 9. https: // doi. org / 10.1590 / 1676 - 0611 - BN- 2015 - 0034" type="journal article" year="2016">
<collectingCountry id="EDEE87E030D552EBEC42FEF776A90055" box="[341,421,336,362]" name="France" pageId="143" pageNumber="144">França</collectingCountry>
<emphasis id="A78D1B6230D552EBECBBFEF776EB0056" box="[428,487,335,361]" italics="true" pageId="143" pageNumber="144">et al.</emphasis>
2016: 3
</bibRefCitation>
</treatmentCitation>
(report). — 
<treatmentCitation id="1458E16130D552EBEFCBFEE874EA0056" author="Giraldo C. &amp; Montoya S. &amp; Escobar F." box="[732,998,335,362]" page="62" pageId="143" pageNumber="144" year="2018">
<bibRefCitation id="F168BA8130D552EBEFCBFEE874EA0056" author="Giraldo C. &amp; Montoya S. &amp; Escobar F." box="[732,998,335,362]" pageId="143" pageNumber="144" refId="ref121556" refString="Giraldo C., Montoya S. &amp; Escobar F. 2018. Escarabajos del Estiercol en Paisajes Ganaderos de Colombia. Fundacion CIPAV, Cali, Colombia." type="book" year="2018">
Giraldo 
<emphasis id="A78D1B6230D552EBEE2AFEF7747B0056" box="[829,887,335,361]" italics="true" pageId="143" pageNumber="144">et al.</emphasis>
2018: 62
</bibRefCitation>
</treatmentCitation>
(guide). — 
<treatmentCitation id="1458E16130D552EBE961FEE8727A0056" author="Nieto M. J. A. &amp; Giraldo E. C. &amp; Quevedo V. C. J. &amp; Chara J. &amp; Medina A. C." box="[1142,1398,335,362]" page="136" pageId="143" pageNumber="144" year="2020">
<bibRefCitation id="F168BA8130D552EBE961FEE8727A0056" author="Nieto M. J. A. &amp; Giraldo E. C. &amp; Quevedo V. C. J. &amp; Chara J. &amp; Medina A. C." box="[1142,1398,335,362]" pageId="143" pageNumber="144" pagination="134 - 141" refId="ref132145" refString="Nieto M. J. A., Giraldo E. C., Quevedo V. C. J., Chara J. &amp; Medina A. C. 2020. Escarabajos estercoleros asociados a sistemas de ganaderia sostenible en diferentes regiones de Colombia. Biota Colombiana 21: 134 - 141. https: // doi. org / 10.21068 / c 2020. v 21 n 02 a 09" type="journal article" year="2020">
Nieto 
<emphasis id="A78D1B6230D552EBE9A9FEF773F50056" box="[1214,1273,335,361]" italics="true" pageId="143" pageNumber="144">et al.</emphasis>
2020: 136
</bibRefCitation>
</treatmentCitation>
(report).
</treatmentCitationGroup>
</paragraph>
<caption id="C18697F830D452EAEDABF80073F406DB" ID-DOI="http://doi.org/10.5281/zenodo.13945617" ID-Zenodo-Dep="13945617" httpUri="https://zenodo.org/record/13945617/files/figure.png" pageId="142" pageNumber="143" startId="142.[188,231,1959,1985]" targetBox="[271,1306,278,1899]" targetPageId="142">
<paragraph id="9546C77030D452EAEDABF80073F406DB" blockId="142.[188,1397,1959,2021]" pageId="142" pageNumber="143">
<emphasis id="A78D1B6230D452EAEDABF800761706FE" bold="true" box="[188,283,1959,1985]" pageId="142" pageNumber="143">Fig. 55.</emphasis>
Species distribution. 
<emphasis id="A78D1B6230D452EAEF0AF800753D06FE" bold="true" box="[541,561,1959,1985]" pageId="142" pageNumber="143">A</emphasis>
. 
<taxonomicName id="52F9BCF330D452EAEF56F800731106FE" authority="Burmeister, 1874" authorityName="Burmeister" authorityYear="1874" box="[577,1053,1959,1985]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="142" pageNumber="143" phylum="Arthropoda" rank="species" species="aphodioides">
<emphasis id="A78D1B6230D452EAEF56F800744906FE" box="[577,837,1959,1985]" italics="true" pageId="142" pageNumber="143">Ontherus aphodioides</emphasis>
<bibRefCitation id="F168BA8130D452EAEE59F800731106FE" author="Burmeister H." box="[846,1053,1959,1985]" pageId="142" pageNumber="143" pagination="120 - 133" refId="ref113171" refString="Burmeister H. 1874. Lamellicornia Argentina [Fortsetzung]. Entomologische Zeitung Stettin 35: 120 - 133. Available from https: // www. biodiversitylibrary. org / page / 9236010 [accessed 14 Feb. 2024]." type="journal article" year="1874">Burmeister, 1874</bibRefCitation>
</taxonomicName>
. 
<emphasis id="A78D1B6230D452EAE93EF800733106FE" bold="true" box="[1065,1085,1959,1985]" pageId="142" pageNumber="143">B</emphasis>
. 
<taxonomicName id="52F9BCF330D452EAE95AF80076B806DA" authority="(Mannerheim, 1829)" baseAuthorityName="Mannerheim" baseAuthorityYear="1829" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="142" pageNumber="143" phylum="Arthropoda" rank="species" species="appendiculatus">
<emphasis id="A78D1B6230D452EAE95AF800727906FE" box="[1101,1397,1959,1985]" italics="true" pageId="142" pageNumber="143">Ontherus appendiculatus</emphasis>
(
<bibRefCitation id="F168BA8130D452EAEDD2F86D76A606DA" author="Mannerheim C. G. de" box="[197,426,1994,2021]" pageId="142" pageNumber="143" refId="ref129230" refString="Mannerheim C. G. de. 1829. Description de quarante nouvelles especes de scarabeides du Bresil; avec figures. Nouveaux Memoires de la Societe Imperiale des Naturalistes de Moscou 1: [29] - 80. Available from https: // www. biodiversitylibrary. org / page / 33157898 [accessed 16 Feb. 2024]." type="journal volume" year="1829">Mannerheim, 1829</bibRefCitation>
)
</taxonomicName>
. Green diamond = CEMT collection data; black circle = literature data.
</paragraph>
</caption>
<paragraph id="9546C77030D552EBEDAAFE31730E0302" blockId="143.[189,1399,265,1452]" pageId="143" pageNumber="144">
<treatmentCitationGroup id="B5E9E05E30D552EBEDAAFE31730E0302" pageId="143" pageNumber="144">
<taxonomicName id="52F9BCF330D552EBEDAAFE31769F008F" ID-CoL="7VDLY" box="[189,403,406,432]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="polynice">
<emphasis id="A78D1B6230D552EBEDAAFE31769F008F" box="[189,403,406,432]" italics="true" pageId="143" pageNumber="144">Ontherus polynice</emphasis>
</taxonomicName>
– 
<treatmentCitation id="1458E16130D552EBECB9FE317574008E" author="Harold E." box="[430,632,406,433]" page="59" pageId="143" pageNumber="144" year="1869">
<bibRefCitation id="F168BA8130D552EBECB9FE317574008E" author="Harold E." box="[430,632,406,433]" pageId="143" pageNumber="144" pagination="46 - 70" refId="ref123583" refString="Harold E. 1869 c. Ueber Coprophage Lamellicornien mit besonderer Berucksichtigung der Pariser Sammlungen. Coleopterologische Hefte 5: 46 - 70. Available from https: // www. biodiversitylibrary. org / page / 16069721 [accessed 16 Feb. 2024]." type="journal article" year="1869">Harold 1869c: 59</bibRefCitation>
</treatmentCitation>
(comments); 
<treatmentCitation id="1458E16130D552EBEE0EFE3174A0008F" author="Harold E." box="[793,940,406,432]" page="1008" pageId="143" pageNumber="144" year="1869">
<bibRefCitation id="F168BA8130D552EBEE0EFE3174A0008F" author="Harold E." box="[793,940,406,432]" firstAuthor="d" pageId="143" pageNumber="144" refId="ref123629" refString="Harold E. 1869 d. Catalogus coleopterorum. Hucusque descriptorum, synonymicus et systematicus. Tom. IV. Scarabaeidae. E. H. Gummi, Munich. https: // doi. org / 10.5962 / bhl. title. 9089" type="book" year="1869">1869d: 1008</bibRefCitation>
</treatmentCitation>
(catalogue, cited for 
<collectingCountry id="EDEE87E030D552EBE9B6FE3173F1008F" box="[1185,1277,406,432]" name="Bolivia" pageId="143" pageNumber="144">Bolivia</collectingCountry>
). — 
<treatmentCitation id="1458E16130D552EBE823FE31765700EB" author="Gillet J. J. E." page="57" pageId="143" pageNumber="144" year="1911">
<bibRefCitation id="F168BA8130D552EBE823FE31765700EB" author="Gillet J. J. E." pageId="143" pageNumber="144" pagination="315 - 319" refId="ref121188" refString="Gillet J. J. E. 1911 a. Lamellicornes coprophages nouveaux ou peu connus d'Amerique du Sud. Annales de la Societe entomologique de Belgique 55: 315 - 319. https: // doi. org / 10.5962 / bhl. part. 2978" type="journal article" year="1911">Gillet 1911a: 57</bibRefCitation>
</treatmentCitation>
(catalogue, cited as synonym of 
<taxonomicName id="52F9BCF330D552EBEFC4FE1E74A600EC" ID-CoL="7ZQ3D" baseAuthorityName="Mannerheim" baseAuthorityYear="1829" box="[723,938,441,467]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="appendiculatus">
<emphasis id="A78D1B6230D552EBEFC4FE1E74A600EC" box="[723,938,441,467]" italics="true" pageId="143" pageNumber="144">O. appendiculatus</emphasis>
</taxonomicName>
). — 
<treatmentCitation id="1458E16130D552EBEEF3FE1E720D00EC" author="Luederwaldt H." box="[996,1281,441,467]" page="389" pageId="143" pageNumber="144" year="1931">
<bibRefCitation id="F168BA8130D552EBEEF3FE1E720D00EC" author="Luederwaldt H." box="[996,1281,441,467]" pageId="143" pageNumber="144" pagination="297 - 304" refId="ref128781" refString="Luederwaldt H. 1931 b. Novos subsidios para o conhecimento do genero Pinotus. Revista de Entomologia 1 (3): 297 - 304." type="journal article" year="1931">Luederwaldt 1931b: 389</bibRefCitation>
</treatmentCitation>
(revision, cited as synonym of 
<taxonomicName id="52F9BCF330D552EBECC2FE7A75A000C9" ID-CoL="7ZQ3D" baseAuthorityName="Mannerheim" baseAuthorityYear="1829" box="[469,684,476,503]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="appendiculatus">
<emphasis id="A78D1B6230D552EBECC2FE7A75A000C9" box="[469,684,476,503]" italics="true" pageId="143" pageNumber="144">O. appendiculatus</emphasis>
</taxonomicName>
). — 
<treatmentCitation id="1458E16130D552EBEFF2FE7B730000C8" author="Pessoa S. B. &amp; Lane F." box="[741,1036,476,503]" page="458" pageId="143" pageNumber="144" year="1941">
<bibRefCitation id="F168BA8130D552EBEFF2FE7B730000C8" author="Pessoa S. B. &amp; Lane F." box="[741,1036,476,503]" pageId="143" pageNumber="144" pagination="389 - 504" refId="ref135859" refString="Pessoa S. B. &amp; Lane F. 1941. Coleopteros necrofagos de interesse medico-legal: ensaio monografico sobre a familia Scarabaeidae de S. Paulo e regioes vizinhas. Arquivos de Zoologia do Estado de Sao Paulo 2: 389 - 504. https: // doi. org / 10.11606 / issn. 2176 - 7793.19412389 - 504" type="journal article" year="1941">Pessôa &amp; Lane 1941: 458</bibRefCitation>
</treatmentCitation>
(monograph, cited as synonym of 
<taxonomicName id="52F9BCF330D552EBEC1DFDA776ED0326" ID-CoL="7ZQ3D" baseAuthorityName="Mannerheim" baseAuthorityYear="1829" box="[266,481,511,538]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="appendiculatus">
<emphasis id="A78D1B6230D552EBEC1DFDA776ED0326" box="[266,481,511,538]" italics="true" pageId="143" pageNumber="144">O. appendiculatus</emphasis>
</taxonomicName>
). — 
<treatmentCitation id="1458E16130D552EBEF08FE58743D0326" author="Blackwelder R. E." box="[543,817,511,538]" page="206" pageId="143" pageNumber="144" year="1944">
<bibRefCitation id="F168BA8130D552EBEF08FE58743D0326" author="Blackwelder R. E." box="[543,817,511,538]" pageId="143" pageNumber="144" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 206</bibRefCitation>
</treatmentCitation>
(list, cited as synonym of 
<taxonomicName id="52F9BCF330D552EBE97EFDA7724C0326" ID-CoL="7ZQ3D" baseAuthorityName="Mannerheim" baseAuthorityYear="1829" box="[1129,1344,511,538]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="appendiculatus">
<emphasis id="A78D1B6230D552EBE97EFDA7724C0326" box="[1129,1344,511,538]" italics="true" pageId="143" pageNumber="144">O. appendiculatus</emphasis>
</taxonomicName>
). — 
<treatmentCitation id="1458E16130D552EBEDFCFD8576A50302" author="Genier F." box="[235,425,546,573]" page="78" pageId="143" pageNumber="144" year="1996">
<bibRefCitation id="F168BA8130D552EBEDFCFD8576A50302" author="Genier F." box="[235,425,546,573]" pageId="143" pageNumber="144" pagination="1 - 169" refId="ref120431" refString="Genier F. 1996. A revision of the Neotropical genus Ontherus Erichson (Coleoptera: Scarabaeidae, Scarabaeinae). Memoirs of the Entomological Society of Canada 170: 1 - 169. https: // doi. org / 10.4039 / entm 128170 fv" type="journal article" year="1996">Génier 1996: 78</bibRefCitation>
</treatmentCitation>
(revision, cited as synonym of 
<taxonomicName id="52F9BCF330D552EBEE0DFD8474FD0303" ID-CoL="7ZQ3D" baseAuthorityName="Mannerheim" baseAuthorityYear="1829" box="[794,1009,546,573]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="appendiculatus">
<emphasis id="A78D1B6230D552EBEE0DFD8474FD0303" box="[794,1009,546,573]" italics="true" pageId="143" pageNumber="144">O. appendiculatus</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030D552EBEDAAFDE1748A035F" blockId="143.[189,1399,265,1452]" box="[189,902,582,609]" pageId="143" pageNumber="144">
<treatmentCitationGroup id="B5E9E05E30D552EBEDAAFDE1748A035F" box="[189,902,582,609]" pageId="143" pageNumber="144">
<taxonomicName id="52F9BCF330D552EBEDAAFDE176F3035F" ID-CoL="7VDLY" box="[189,511,582,608]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="polynice" subGenus="Copris">
<emphasis id="A78D1B6230D552EBEDAAFDE17626035F" box="[189,298,582,608]" italics="true" pageId="143" pageNumber="144">Ontherus</emphasis>
(
<emphasis id="A78D1B6230D552EBEC2CFDE17687035F" box="[315,395,582,608]" italics="true" pageId="143" pageNumber="144">Copris</emphasis>
) 
<emphasis id="A78D1B6230D552EBEC8BFDE176F3035F" box="[412,511,582,608]" italics="true" pageId="143" pageNumber="144">polynice</emphasis>
</taxonomicName>
– 
<treatmentCitation id="1458E16130D552EBEF0AFDE175E6035E" author="Harold E." box="[541,746,582,609]" page="59" pageId="143" pageNumber="144" year="1869">
<bibRefCitation id="F168BA8130D552EBEF0AFDE175E6035E" author="Harold E." box="[541,746,582,609]" pageId="143" pageNumber="144" pagination="46 - 70" refId="ref123583" refString="Harold E. 1869 c. Ueber Coprophage Lamellicornien mit besonderer Berucksichtigung der Pariser Sammlungen. Coleopterologische Hefte 5: 46 - 70. Available from https: // www. biodiversitylibrary. org / page / 16069721 [accessed 16 Feb. 2024]." type="journal article" year="1869">Harold 1869c: 59</bibRefCitation>
</treatmentCitation>
(comments).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030D552EBEDAAFDCE721A020C" blockId="143.[189,1399,265,1452]" pageId="143" pageNumber="144">
<treatmentCitationGroup id="B5E9E05E30D552EBEDAAFDCE721A020C" pageId="143" pageNumber="144">
<taxonomicName id="52F9BCF330D552EBEDAAFDCE76A503BC" authorityName="Erichson" authorityYear="1848" box="[189,425,617,643]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="quadratus">
<emphasis id="A78D1B6230D552EBEDAAFDCE76A503BC" box="[189,425,617,643]" italics="true" pageId="143" pageNumber="144">Ontherus quadratus</emphasis>
</taxonomicName>
– 
<treatmentCitation id="1458E16130D552EBECD1FDCE759F03BB" author="Harold E." box="[454,659,617,644]" page="59" pageId="143" pageNumber="144" year="1869">
<bibRefCitation id="F168BA8130D552EBECD1FDCE759F03BB" author="Harold E." box="[454,659,617,644]" pageId="143" pageNumber="144" pagination="46 - 70" refId="ref123583" refString="Harold E. 1869 c. Ueber Coprophage Lamellicornien mit besonderer Berucksichtigung der Pariser Sammlungen. Coleopterologische Hefte 5: 46 - 70. Available from https: // www. biodiversitylibrary. org / page / 16069721 [accessed 16 Feb. 2024]." type="journal article" year="1869">Harold 1869c: 59</bibRefCitation>
</treatmentCitation>
(comments); 
<treatmentCitation id="1458E16130D552EBEE21FDCE74C603BC" author="Harold E." box="[822,970,617,644]" page="1008" pageId="143" pageNumber="144" year="1869">
<bibRefCitation id="F168BA8130D552EBEE21FDCE74C603BC" author="Harold E." box="[822,970,617,644]" firstAuthor="d" pageId="143" pageNumber="144" refId="ref123629" refString="Harold E. 1869 d. Catalogus coleopterorum. Hucusque descriptorum, synonymicus et systematicus. Tom. IV. Scarabaeidae. E. H. Gummi, Munich. https: // doi. org / 10.5962 / bhl. title. 9089" type="book" year="1869">1869d: 1008</bibRefCitation>
</treatmentCitation>
(catalogue, cited for 
<collectingCountry id="EDEE87E030D552EBE9D3FDCE721D03BC" box="[1220,1297,617,643]" name="Brazil" pageId="143" pageNumber="144">Brazil</collectingCountry>
); 
<treatmentCitation id="1458E16130D552EBE832FDCE761A0399" author="Harold E." page="181" pageId="143" pageNumber="144" year="1875">
<bibRefCitation id="F168BA8130D552EBE832FDCE761A0399" author="Harold E." firstAuthor="b" pageId="143" pageNumber="144" pagination="181 - 183" refId="ref123769" refString="Harold E. 1875 b. Miscellen. Coleopterologische Hefte 13: 181 - 183. Available from https: // www. biodiversitylibrary. org / page / 25118793 [accessed 16 Feb. 2024]." type="journal article" year="1875">1875b: 181</bibRefCitation>
</treatmentCitation>
(comments). — 
<treatmentCitation id="1458E16130D552EBECF0FD2B75BD0399" author="Bruch C." box="[487,689,652,678]" page="187" pageId="143" pageNumber="144" year="1911">
<bibRefCitation id="F168BA8130D552EBECF0FD2B75BD0399" author="Bruch C." box="[487,689,652,678]" pageId="143" pageNumber="144" pagination="143 - 225" refId="ref112509" refString="Bruch C. 1911. Catalogo sistematico de los coleopteros de la Republica Argentina. Revista del Museo de La Plata 17: 143 - 225. https: // doi. org / 10.5962 / bhl. title. 8799" type="journal article" year="1911">Bruch 1911: 187</bibRefCitation>
</treatmentCitation>
(catalogue, cited as synonym of 
<taxonomicName id="52F9BCF330D552EBE95FFD2B72310399" ID-CoL="7VDM8" box="[1096,1341,652,678]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="rotundatus">
<emphasis id="A78D1B6230D552EBE95FFD2B72310399" box="[1096,1341,652,678]" italics="true" pageId="143" pageNumber="144">Ontherus rotundatus</emphasis>
</taxonomicName>
). — 
<treatmentCitation id="1458E16130D552EBEDFDFD0876AB03F5" author="Gillet J. J. E." box="[234,423,687,714]" page="57" pageId="143" pageNumber="144" year="1911">
<bibRefCitation id="F168BA8130D552EBEDFDFD0876AB03F5" author="Gillet J. J. E." box="[234,423,687,714]" pageId="143" pageNumber="144" pagination="315 - 319" refId="ref121188" refString="Gillet J. J. E. 1911 a. Lamellicornes coprophages nouveaux ou peu connus d'Amerique du Sud. Annales de la Societe entomologique de Belgique 55: 315 - 319. https: // doi. org / 10.5962 / bhl. part. 2978" type="journal article" year="1911">Gillet 1911a: 57</bibRefCitation>
</treatmentCitation>
(catalogue, cited as synonym of 
<taxonomicName id="52F9BCF330D552EBEE30FD08734303F6" ID-CoL="7ZQ3D" baseAuthorityName="Mannerheim" baseAuthorityYear="1829" box="[807,1103,687,713]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="appendiculatus">
<emphasis id="A78D1B6230D552EBEE30FD08734303F6" box="[807,1103,687,713]" italics="true" pageId="143" pageNumber="144">Ontherus appendiculatus</emphasis>
</taxonomicName>
). — 
<treatmentCitation id="1458E16130D552EBE99CFD08761A03D2" author="Luederwaldt H." page="389" pageId="143" pageNumber="144" year="1931">
<bibRefCitation id="F168BA8130D552EBE99CFD08761A03D2" author="Luederwaldt H." pageId="143" pageNumber="144" pagination="363 - 422" refId="ref128737" refString="Luederwaldt H. 1931 a. O genero Ontherus (Coleop.) (Lamellic-Coprid. - Pinot.) com uma chave, para a determinacao dos pinotides americanos. Revista do Museu Paulista 17: 363 - 422 + 1 pl." type="journal article" year="1931">Luederwaldt 1931a: 389</bibRefCitation>
</treatmentCitation>
(revision, cited as synonym of 
<taxonomicName id="52F9BCF330D552EBEF98FD74746A03D3" ID-CoL="7ZQ3D" baseAuthorityName="Mannerheim" baseAuthorityYear="1829" box="[655,870,722,749]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="appendiculatus">
<emphasis id="A78D1B6230D552EBEF98FD74746A03D3" box="[655,870,722,749]" italics="true" pageId="143" pageNumber="144">O. appendiculatus</emphasis>
</taxonomicName>
). — 
<treatmentCitation id="1458E16130D552EBEEB2FD7473DB03D2" author="Pessoa S. B. &amp; Lane F." box="[933,1239,722,749]" page="458" pageId="143" pageNumber="144" year="1941">
<bibRefCitation id="F168BA8130D552EBEEB2FD7473DB03D2" author="Pessoa S. B. &amp; Lane F." box="[933,1239,722,749]" pageId="143" pageNumber="144" pagination="389 - 504" refId="ref135859" refString="Pessoa S. B. &amp; Lane F. 1941. Coleopteros necrofagos de interesse medico-legal: ensaio monografico sobre a familia Scarabaeidae de S. Paulo e regioes vizinhas. Arquivos de Zoologia do Estado de Sao Paulo 2: 389 - 504. https: // doi. org / 10.11606 / issn. 2176 - 7793.19412389 - 504" type="journal article" year="1941">Pessôa &amp; Lane 1941: 458</bibRefCitation>
</treatmentCitation>
(monograph, cited as synonym of 
<taxonomicName id="52F9BCF330D552EBECE6FD5175C0022F" ID-CoL="7ZQ3D" baseAuthorityName="Mannerheim" baseAuthorityYear="1829" box="[497,716,758,784]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="appendiculatus">
<emphasis id="A78D1B6230D552EBECE6FD5175C0022F" box="[497,716,758,784]" italics="true" pageId="143" pageNumber="144">O. appendiculatus</emphasis>
</taxonomicName>
). — 
<treatmentCitation id="1458E16130D552EBEE04FD517323022F" author="Blackwelder R. E." box="[787,1071,758,784]" page="206" pageId="143" pageNumber="144" year="1944">
<bibRefCitation id="F168BA8130D552EBEE04FD517323022F" author="Blackwelder R. E." box="[787,1071,758,784]" pageId="143" pageNumber="144" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 206</bibRefCitation>
</treatmentCitation>
(list, cited as synonym of 
<taxonomicName id="52F9BCF330D552EBEDFDFCBE76CD020C" ID-CoL="7ZQ3D" baseAuthorityName="Mannerheim" baseAuthorityYear="1829" box="[234,449,793,819]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="appendiculatus">
<emphasis id="A78D1B6230D552EBEDFDFCBE76CD020C" box="[234,449,793,819]" italics="true" pageId="143" pageNumber="144">O. appendiculatus</emphasis>
</taxonomicName>
). — 
<treatmentCitation id="1458E16130D552EBECE8FCBE75B1020C" author="Genier F." box="[511,701,793,819]" page="78" pageId="143" pageNumber="144" year="1996">
<bibRefCitation id="F168BA8130D552EBECE8FCBE75B1020C" author="Genier F." box="[511,701,793,819]" pageId="143" pageNumber="144" pagination="1 - 169" refId="ref120431" refString="Genier F. 1996. A revision of the Neotropical genus Ontherus Erichson (Coleoptera: Scarabaeidae, Scarabaeinae). Memoirs of the Entomological Society of Canada 170: 1 - 169. https: // doi. org / 10.4039 / entm 128170 fv" type="journal article" year="1996">Génier 1996: 78</bibRefCitation>
</treatmentCitation>
(revision, cited as synonym of 
<taxonomicName id="52F9BCF330D552EBE939FCBE7209020C" ID-CoL="7ZQ3D" baseAuthorityName="Mannerheim" baseAuthorityYear="1829" box="[1070,1285,793,820]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="appendiculatus">
<emphasis id="A78D1B6230D552EBE939FCBE734F020C" box="[1070,1091,793,819]" italics="true" pageId="143" pageNumber="144">O</emphasis>
. 
<emphasis id="A78D1B6230D552EBE945FCBE7209020C" box="[1106,1285,793,819]" italics="true" pageId="143" pageNumber="144">appendiculatus</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030D552EBEDAAFC9B730E0539" blockId="143.[189,1399,265,1452]" pageId="143" pageNumber="144">
<treatmentCitationGroup id="B5E9E05E30D552EBEDAAFC9B730E0539" pageId="143" pageNumber="144">
<taxonomicName id="52F9BCF330D552EBEDAAFC9B76BE0269" ID-CoL="7VDM8" box="[189,434,828,854]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="rotundatus">
<emphasis id="A78D1B6230D552EBEDAAFC9B76BE0269" box="[189,434,828,854]" italics="true" pageId="143" pageNumber="144">Ontherus rotundatus</emphasis>
</taxonomicName>
– 
<treatmentCitation id="1458E16130D552EBECC2FC9B75AB0268" author="Harold E." box="[469,679,828,855]" page="59" pageId="143" pageNumber="144" year="1869">
<bibRefCitation id="F168BA8130D552EBECC2FC9B75AB0268" author="Harold E." box="[469,679,828,855]" pageId="143" pageNumber="144" pagination="46 - 70" refId="ref123583" refString="Harold E. 1869 c. Ueber Coprophage Lamellicornien mit besonderer Berucksichtigung der Pariser Sammlungen. Coleopterologische Hefte 5: 46 - 70. Available from https: // www. biodiversitylibrary. org / page / 16069721 [accessed 16 Feb. 2024]." type="journal article" year="1869">Harold 1869c: 59</bibRefCitation>
</treatmentCitation>
(comments); 
<treatmentCitation id="1458E16130D552EBEE47FC9B74EA0269" author="Harold E." box="[848,998,828,855]" page="1008" pageId="143" pageNumber="144" year="1869">
<bibRefCitation id="F168BA8130D552EBEE47FC9B74EA0269" author="Harold E." box="[848,998,828,855]" firstAuthor="d" pageId="143" pageNumber="144" refId="ref123629" refString="Harold E. 1869 d. Catalogus coleopterorum. Hucusque descriptorum, synonymicus et systematicus. Tom. IV. Scarabaeidae. E. H. Gummi, Munich. https: // doi. org / 10.5962 / bhl. title. 9089" type="book" year="1869">1869d: 1008</bibRefCitation>
</treatmentCitation>
(catalogue, cited for 
<collectingRegion id="573D099230D552EBE9FDFC9B72660269" box="[1258,1386,828,854]" country="Argentina" name="Corrientes" pageId="143" pageNumber="144">Corrientes</collectingRegion>
); 
<treatmentCitation id="1458E16130D552EBEDFDFCF8767C0245" author="Harold E." box="[234,368,863,890]" page="181" pageId="143" pageNumber="144" year="1875">
<bibRefCitation id="F168BA8130D552EBEDFDFCF8767C0245" author="Harold E." box="[234,368,863,890]" firstAuthor="b" pageId="143" pageNumber="144" pagination="181 - 183" refId="ref123769" refString="Harold E. 1875 b. Miscellen. Coleopterologische Hefte 13: 181 - 183. Available from https: // www. biodiversitylibrary. org / page / 25118793 [accessed 16 Feb. 2024]." type="journal article" year="1875">1875b: 181</bibRefCitation>
</treatmentCitation>
(comments). — 
<treatmentCitation id="1458E16130D552EBEF2DFCF8740C0245" author="Bruch C." box="[570,768,863,890]" page="187" pageId="143" pageNumber="144" year="1911">
<bibRefCitation id="F168BA8130D552EBEF2DFCF8740C0245" author="Bruch C." box="[570,768,863,890]" pageId="143" pageNumber="144" pagination="143 - 225" refId="ref112509" refString="Bruch C. 1911. Catalogo sistematico de los coleopteros de la Republica Argentina. Revista del Museo de La Plata 17: 143 - 225. https: // doi. org / 10.5962 / bhl. title. 8799" type="journal article" year="1911">Bruch 1911: 187</bibRefCitation>
</treatmentCitation>
(catalogue, cited for 
<collectingRegion id="573D099230D552EBEEE9FCF873720246" box="[1022,1150,863,889]" country="Argentina" name="Corrientes" pageId="143" pageNumber="144">Corrientes</collectingRegion>
). — 
<treatmentCitation id="1458E16130D552EBE9A0FCF8727A0245" author="Gillet J. J. E." box="[1207,1398,863,890]" page="57" pageId="143" pageNumber="144" year="1911">
<bibRefCitation id="F168BA8130D552EBE9A0FCF8727A0245" author="Gillet J. J. E." box="[1207,1398,863,890]" pageId="143" pageNumber="144" pagination="315 - 319" refId="ref121188" refString="Gillet J. J. E. 1911 a. Lamellicornes coprophages nouveaux ou peu connus d'Amerique du Sud. Annales de la Societe entomologique de Belgique 55: 315 - 319. https: // doi. org / 10.5962 / bhl. part. 2978" type="journal article" year="1911">Gillet 1911a: 57</bibRefCitation>
</treatmentCitation>
(catalogue, cited as synonym of 
<taxonomicName id="52F9BCF330D552EBEF65FC25749502A3" ID-CoL="7ZQ3D" baseAuthorityName="Mannerheim" baseAuthorityYear="1829" box="[626,921,898,924]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="appendiculatus">
<emphasis id="A78D1B6230D552EBEF65FC25749502A3" box="[626,921,898,924]" italics="true" pageId="143" pageNumber="144">Ontherus appendiculatus</emphasis>
</taxonomicName>
). — 
<treatmentCitation id="1458E16130D552EBEECCFC2573F102A2" author="Luederwaldt H." box="[987,1277,898,925]" page="389" pageId="143" pageNumber="144" year="1931">
<bibRefCitation id="F168BA8130D552EBEECCFC2573F102A2" author="Luederwaldt H." box="[987,1277,898,925]" pageId="143" pageNumber="144" pagination="363 - 422" refId="ref128737" refString="Luederwaldt H. 1931 a. O genero Ontherus (Coleop.) (Lamellic-Coprid. - Pinot.) com uma chave, para a determinacao dos pinotides americanos. Revista do Museu Paulista 17: 363 - 422 + 1 pl." type="journal article" year="1931">Luederwaldt 1931a: 389</bibRefCitation>
</treatmentCitation>
(revision, cited as synonym of 
<taxonomicName id="52F9BCF330D552EBECC2FC0175A002FF" ID-CoL="7ZQ3D" baseAuthorityName="Mannerheim" baseAuthorityYear="1829" box="[469,684,934,960]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="appendiculatus">
<emphasis id="A78D1B6230D552EBECC2FC0175A002FF" box="[469,684,934,960]" italics="true" pageId="143" pageNumber="144">O. appendiculatus</emphasis>
</taxonomicName>
). — 
<treatmentCitation id="1458E16130D552EBEFF2FC01730002FE" author="Pessoa S. B. &amp; Lane F." box="[741,1036,934,961]" page="458" pageId="143" pageNumber="144" year="1941">
<bibRefCitation id="F168BA8130D552EBEFF2FC01730002FE" author="Pessoa S. B. &amp; Lane F." box="[741,1036,934,961]" pageId="143" pageNumber="144" pagination="389 - 504" refId="ref135859" refString="Pessoa S. B. &amp; Lane F. 1941. Coleopteros necrofagos de interesse medico-legal: ensaio monografico sobre a familia Scarabaeidae de S. Paulo e regioes vizinhas. Arquivos de Zoologia do Estado de Sao Paulo 2: 389 - 504. https: // doi. org / 10.11606 / issn. 2176 - 7793.19412389 - 504" type="journal article" year="1941">Pessôa &amp; Lane 1941: 458</bibRefCitation>
</treatmentCitation>
(monograph, cited as synonym of 
<taxonomicName id="52F9BCF330D552EBEC1DFC6E76ED02DC" ID-CoL="7ZQ3D" baseAuthorityName="Mannerheim" baseAuthorityYear="1829" box="[266,481,969,995]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="appendiculatus">
<emphasis id="A78D1B6230D552EBEC1DFC6E76ED02DC" box="[266,481,969,995]" italics="true" pageId="143" pageNumber="144">O. appendiculatus</emphasis>
</taxonomicName>
). — 
<treatmentCitation id="1458E16130D552EBEF08FC6E743D02DC" author="Blackwelder R. E." box="[543,817,969,995]" page="206" pageId="143" pageNumber="144" year="1944">
<bibRefCitation id="F168BA8130D552EBEF08FC6E743D02DC" author="Blackwelder R. E." box="[543,817,969,995]" pageId="143" pageNumber="144" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 206</bibRefCitation>
</treatmentCitation>
(list, cited as synonym of 
<taxonomicName id="52F9BCF330D552EBE97EFC6E724C02DC" ID-CoL="7ZQ3D" baseAuthorityName="Mannerheim" baseAuthorityYear="1829" box="[1129,1344,969,995]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="appendiculatus">
<emphasis id="A78D1B6230D552EBE97EFC6E724C02DC" box="[1129,1344,969,995]" italics="true" pageId="143" pageNumber="144">O. appendiculatus</emphasis>
</taxonomicName>
). — 
<treatmentCitation id="1458E16130D552EBEDFCFC4B76A50539" author="Genier F." box="[235,425,1004,1030]" page="78" pageId="143" pageNumber="144" year="1996">
<bibRefCitation id="F168BA8130D552EBEDFCFC4B76A50539" author="Genier F." box="[235,425,1004,1030]" pageId="143" pageNumber="144" pagination="1 - 169" refId="ref120431" refString="Genier F. 1996. A revision of the Neotropical genus Ontherus Erichson (Coleoptera: Scarabaeidae, Scarabaeinae). Memoirs of the Entomological Society of Canada 170: 1 - 169. https: // doi. org / 10.4039 / entm 128170 fv" type="journal article" year="1996">Génier 1996: 78</bibRefCitation>
</treatmentCitation>
(revision, cited as synonym of 
<taxonomicName id="52F9BCF330D552EBEE0DFC4A74FD0539" ID-CoL="7ZQ3D" baseAuthorityName="Mannerheim" baseAuthorityYear="1829" box="[794,1009,1004,1031]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="appendiculatus">
<emphasis id="A78D1B6230D552EBEE0DFC4A74FD0539" box="[794,1009,1004,1031]" italics="true" pageId="143" pageNumber="144">O. appendiculatus</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030D552EBEDAAFBB77569054F" blockId="143.[189,1399,265,1452]" pageId="143" pageNumber="144">
<treatmentCitationGroup id="B5E9E05E30D552EBEDAAFBB77569054F" pageId="143" pageNumber="144">
<taxonomicName id="52F9BCF330D552EBEDAAFBB7769A0516" ID-CoL="7VDM8" authorityName="Blanchard" authorityYear="1846" box="[189,406,1039,1066]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="rotundatus">
<emphasis id="A78D1B6230D552EBEDAAFBB7769A0516" box="[189,406,1039,1066]" italics="true" pageId="143" pageNumber="144">Copris rotundatus</emphasis>
</taxonomicName>
– 
<treatmentCitation id="1458E16130D552EBECABFBA875C90515" author="Burmeister H." box="[444,709,1039,1066]" page="127" pageId="143" pageNumber="144" year="1874">
<bibRefCitation id="F168BA8130D552EBECABFBA875C90515" author="Burmeister H." box="[444,709,1039,1066]" pageId="143" pageNumber="144" pagination="120 - 133" refId="ref113171" refString="Burmeister H. 1874. Lamellicornia Argentina [Fortsetzung]. Entomologische Zeitung Stettin 35: 120 - 133. Available from https: // www. biodiversitylibrary. org / page / 9236010 [accessed 14 Feb. 2024]." type="journal article" year="1874">Burmeister 1874: 127</bibRefCitation>
</treatmentCitation>
(comments, cited as 
<taxonomicName id="52F9BCF330D552EBEED9FBB773900516" box="[974,1180,1039,1066]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="rotundata">
<emphasis id="A78D1B6230D552EBEED9FBB773900516" box="[974,1180,1039,1066]" italics="true" pageId="143" pageNumber="144">Copris rotundata</emphasis>
</taxonomicName>
). — 
<treatmentCitation id="1458E16130D552EBE9F6FBA8761A0572" author="Bruch C." page="187" pageId="143" pageNumber="144" year="1911">
<bibRefCitation id="F168BA8130D552EBE9F6FBA8761A0572" author="Bruch C." pageId="143" pageNumber="144" pagination="143 - 225" refId="ref112509" refString="Bruch C. 1911. Catalogo sistematico de los coleopteros de la Republica Argentina. Revista del Museo de La Plata 17: 143 - 225. https: // doi. org / 10.5962 / bhl. title. 8799" type="journal article" year="1911">Bruch 1911: 187</bibRefCitation>
</treatmentCitation>
(catalogue, cited as synonym of 
<taxonomicName id="52F9BCF330D552EBEFB2FB95749B0573" ID-CoL="7VDM8" box="[677,919,1074,1100]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="rotundatus">
<emphasis id="A78D1B6230D552EBEFB2FB95749B0573" box="[677,919,1074,1100]" italics="true" pageId="143" pageNumber="144">Ontherus rotundatus</emphasis>
</taxonomicName>
). — 
<treatmentCitation id="1458E16130D552EBEECFFB9573960572" author="Genier F." box="[984,1178,1074,1101]" page="78" pageId="143" pageNumber="144" year="1996">
<bibRefCitation id="F168BA8130D552EBEECFFB9573960572" author="Genier F." box="[984,1178,1074,1101]" pageId="143" pageNumber="144" pagination="1 - 169" refId="ref120431" refString="Genier F. 1996. A revision of the Neotropical genus Ontherus Erichson (Coleoptera: Scarabaeidae, Scarabaeinae). Memoirs of the Entomological Society of Canada 170: 1 - 169. https: // doi. org / 10.4039 / entm 128170 fv" type="journal article" year="1996">Génier 1996: 78</bibRefCitation>
</treatmentCitation>
(revision, cited as synonym of 
<taxonomicName id="52F9BCF330D552EBEC6AFBF17558054F" ID-CoL="7ZQ3D" baseAuthorityName="Mannerheim" baseAuthorityYear="1829" box="[381,596,1110,1136]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="appendiculatus">
<emphasis id="A78D1B6230D552EBEC6AFBF17558054F" box="[381,596,1110,1136]" italics="true" pageId="143" pageNumber="144">O. appendiculatus</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030D552EBEDAAFBDE74A70589" blockId="143.[189,1399,265,1452]" pageId="143" pageNumber="144">
<treatmentCitationGroup id="B5E9E05E30D552EBEDAAFBDE74A70589" pageId="143" pageNumber="144">
<taxonomicName id="52F9BCF330D552EBEDAAFBDE767505AC" ID-CoL="7VDLY" authorityName="Blanchard" authorityYear="1846" box="[189,377,1145,1171]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="polynice">
<emphasis id="A78D1B6230D552EBEDAAFBDE767505AC" box="[189,377,1145,1171]" italics="true" pageId="143" pageNumber="144">Copris polynice</emphasis>
</taxonomicName>
– 
<treatmentCitation id="1458E16130D552EBEC8AFBDE756905AC" author="Bruch C." box="[413,613,1145,1171]" page="187" pageId="143" pageNumber="144" year="1911">
<bibRefCitation id="F168BA8130D552EBEC8AFBDE756905AC" author="Bruch C." box="[413,613,1145,1171]" pageId="143" pageNumber="144" pagination="143 - 225" refId="ref112509" refString="Bruch C. 1911. Catalogo sistematico de los coleopteros de la Republica Argentina. Revista del Museo de La Plata 17: 143 - 225. https: // doi. org / 10.5962 / bhl. title. 8799" type="journal article" year="1911">Bruch 1911: 187</bibRefCitation>
</treatmentCitation>
(catalogue, cited as synonym of 
<taxonomicName id="52F9BCF330D552EBEEE4FBDE73EA05AC" ID-CoL="7VDM8" box="[1011,1254,1145,1171]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="rotundatus">
<emphasis id="A78D1B6230D552EBEEE4FBDE73EA05AC" box="[1011,1254,1145,1171]" italics="true" pageId="143" pageNumber="144">Ontherus rotundatus</emphasis>
</taxonomicName>
). — 
<treatmentCitation id="1458E16130D552EBE830FBDE765F0589" author="Genier F." page="78" pageId="143" pageNumber="144" year="1996">
<bibRefCitation id="F168BA8130D552EBE830FBDE765F0589" author="Genier F." pageId="143" pageNumber="144" pagination="1 - 169" refId="ref120431" refString="Genier F. 1996. A revision of the Neotropical genus Ontherus Erichson (Coleoptera: Scarabaeidae, Scarabaeinae). Memoirs of the Entomological Society of Canada 170: 1 - 169. https: // doi. org / 10.4039 / entm 128170 fv" type="journal article" year="1996">Génier 1996: 78</bibRefCitation>
</treatmentCitation>
(revision, cited as synonym of 
<taxonomicName id="52F9BCF330D552EBEFD4FB3A74960589" ID-CoL="7ZQ3D" baseAuthorityName="Mannerheim" baseAuthorityYear="1829" box="[707,922,1180,1207]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="appendiculatus">
<emphasis id="A78D1B6230D552EBEFD4FB3A74960589" box="[707,922,1180,1207]" italics="true" pageId="143" pageNumber="144">O. appendiculatus</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030D552EBEDAAFB1873B8041F" blockId="143.[189,1399,265,1452]" pageId="143" pageNumber="144">
<treatmentCitationGroup id="B5E9E05E30D552EBEDAAFB1873B8041F" pageId="143" pageNumber="144">
<taxonomicName id="52F9BCF330D552EBEDAAFB1876BC05E6" ID-CoL="7XM89" authorityName="Burmeister" authorityYear="1874" box="[189,432,1215,1241]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="contractus">
<emphasis id="A78D1B6230D552EBEDAAFB1876BC05E6" box="[189,432,1215,1241]" italics="true" pageId="143" pageNumber="144">Ontherus contractus</emphasis>
</taxonomicName>
– 
<treatmentCitation id="1458E16130D552EBECC4FB18759105E5" author="Bruch C." box="[467,669,1215,1242]" page="187" pageId="143" pageNumber="144" year="1911">
<bibRefCitation id="F168BA8130D552EBECC4FB18759105E5" author="Bruch C." box="[467,669,1215,1242]" pageId="143" pageNumber="144" pagination="143 - 225" refId="ref112509" refString="Bruch C. 1911. Catalogo sistematico de los coleopteros de la Republica Argentina. Revista del Museo de La Plata 17: 143 - 225. https: // doi. org / 10.5962 / bhl. title. 8799" type="journal article" year="1911">Bruch 1911: 187</bibRefCitation>
</treatmentCitation>
(catalogue, cited for 
<collectingRegion id="573D099230D552EBEEB5FB67731905E5" box="[930,1045,1216,1242]" country="Argentina" name="Tucuman" pageId="143" pageNumber="144">Tucumán</collectingRegion>
). — 
<treatmentCitation id="1458E16130D552EBE945FB18727A05E5" author="Luederwaldt H." box="[1106,1398,1215,1242]" page="389" pageId="143" pageNumber="144" year="1931">
<bibRefCitation id="F168BA8130D552EBE945FB18727A05E5" author="Luederwaldt H." box="[1106,1398,1215,1242]" pageId="143" pageNumber="144" pagination="363 - 422" refId="ref128737" refString="Luederwaldt H. 1931 a. O genero Ontherus (Coleop.) (Lamellic-Coprid. - Pinot.) com uma chave, para a determinacao dos pinotides americanos. Revista do Museu Paulista 17: 363 - 422 + 1 pl." type="journal article" year="1931">Luederwaldt 1931a: 389</bibRefCitation>
</treatmentCitation>
(revision, cited as synonym of 
<taxonomicName id="52F9BCF330D552EBEF62FB44745F05C3" ID-CoL="7ZQ3D" baseAuthorityName="Mannerheim" baseAuthorityYear="1829" box="[629,851,1250,1277]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="appendiculatus">
<emphasis id="A78D1B6230D552EBEF62FB44745F05C3" box="[629,851,1250,1277]" italics="true" pageId="143" pageNumber="144">O. appendiculatus</emphasis>
</taxonomicName>
). — 
<treatmentCitation id="1458E16130D552EBEE8AFB4573B005C3" author="Blackwelder R. E." box="[925,1212,1250,1277]" page="206" pageId="143" pageNumber="144" year="1944">
<bibRefCitation id="F168BA8130D552EBEE8AFB4573B005C3" author="Blackwelder R. E." box="[925,1212,1250,1277]" pageId="143" pageNumber="144" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 206</bibRefCitation>
</treatmentCitation>
(list, cited for 
<collectingCountry id="EDEE87E030D552EBEDFDFAA17668041F" box="[234,356,1286,1312]" name="Argentina" pageId="143" pageNumber="144">Argentina</collectingCountry>
). — 
<treatmentCitation id="1458E16130D552EBEC8BFAA17557041F" author="Genier F." box="[412,603,1286,1312]" page="78" pageId="143" pageNumber="144" year="1996">
<bibRefCitation id="F168BA8130D552EBEC8BFAA17557041F" author="Genier F." box="[412,603,1286,1312]" pageId="143" pageNumber="144" pagination="1 - 169" refId="ref120431" refString="Genier F. 1996. A revision of the Neotropical genus Ontherus Erichson (Coleoptera: Scarabaeidae, Scarabaeinae). Memoirs of the Entomological Society of Canada 170: 1 - 169. https: // doi. org / 10.4039 / entm 128170 fv" type="journal article" year="1996">Génier 1996: 78</bibRefCitation>
</treatmentCitation>
(revision, cited as synonym of 
<taxonomicName id="52F9BCF330D552EBEEDBFAA173AF041F" ID-CoL="7ZQ3D" baseAuthorityName="Mannerheim" baseAuthorityYear="1829" box="[972,1187,1286,1312]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="appendiculatus">
<emphasis id="A78D1B6230D552EBEEDBFAA174ED041F" box="[972,993,1286,1312]" italics="true" pageId="143" pageNumber="144">O</emphasis>
. 
<emphasis id="A78D1B6230D552EBEEE7FAA173AF041F" box="[1008,1187,1286,1312]" italics="true" pageId="143" pageNumber="144">appendiculatus</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030D552EBEDAAFA8E7428047C" blockId="143.[189,1399,265,1452]" box="[189,804,1321,1348]" pageId="143" pageNumber="144">
<treatmentCitationGroup id="B5E9E05E30D552EBEDAAFA8E7428047C" box="[189,804,1321,1348]" pageId="143" pageNumber="144">
<taxonomicName id="52F9BCF330D552EBEDAAFA8E76CB047C" ID-CoL="7VDJK" authorityName="Mannerheim" authorityYear="1829" box="[189,455,1321,1347]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="appendiculatus">
<emphasis id="A78D1B6230D552EBEDAAFA8E76CB047C" box="[189,455,1321,1347]" italics="true" pageId="143" pageNumber="144">Copris appendiculatus</emphasis>
</taxonomicName>
– 
<treatmentCitation id="1458E16130D552EBECF2FA8E75AF047C" author="Genier F." box="[485,675,1321,1347]" page="78" pageId="143" pageNumber="144" year="1996">
<bibRefCitation id="F168BA8130D552EBECF2FA8E75AF047C" author="Genier F." box="[485,675,1321,1347]" pageId="143" pageNumber="144" pagination="1 - 169" refId="ref120431" refString="Genier F. 1996. A revision of the Neotropical genus Ontherus Erichson (Coleoptera: Scarabaeidae, Scarabaeinae). Memoirs of the Entomological Society of Canada 170: 1 - 169. https: // doi. org / 10.4039 / entm 128170 fv" type="journal article" year="1996">Génier 1996: 78</bibRefCitation>
</treatmentCitation>
(revision).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030D552EBEDAAFAEB76BE0493" blockId="143.[189,1399,265,1452]" pageId="143" pageNumber="144">
<treatmentCitationGroup id="B5E9E05E30D552EBEDAAFAEB76BE0493" pageId="143" pageNumber="144">
<taxonomicName id="52F9BCF330D552EBEDAAFAEB75650459" ID-CoL="7ZQ3D" baseAuthorityName="Mannerheim" baseAuthorityYear="1829" box="[189,617,1356,1382]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="appendiculatus" subGenus="Ontherus">
<emphasis id="A78D1B6230D552EBEDAAFAEB76260459" box="[189,298,1356,1382]" italics="true" pageId="143" pageNumber="144">Ontherus</emphasis>
(
<emphasis id="A78D1B6230D552EBEC2EFAEB76AA0459" box="[313,422,1356,1382]" italics="true" pageId="143" pageNumber="144">Ontherus</emphasis>
) 
<emphasis id="A78D1B6230D552EBECA1FAEB75650459" box="[438,617,1356,1382]" italics="true" pageId="143" pageNumber="144">appendiculatus</emphasis>
</taxonomicName>
– 
<treatmentCitation id="1458E16130D552EBEF96FAEB74370459" author="Genier F." box="[641,827,1356,1382]" page="78" pageId="143" pageNumber="144" year="1996">
<bibRefCitation id="F168BA8130D552EBEF96FAEB74370459" author="Genier F." box="[641,827,1356,1382]" pageId="143" pageNumber="144" pagination="1 - 169" refId="ref120431" refString="Genier F. 1996. A revision of the Neotropical genus Ontherus Erichson (Coleoptera: Scarabaeidae, Scarabaeinae). Memoirs of the Entomological Society of Canada 170: 1 - 169. https: // doi. org / 10.4039 / entm 128170 fv" type="journal article" year="1996">Génier 1996: 78</bibRefCitation>
</treatmentCitation>
(revision). — 
<treatmentCitation id="1458E16130D552EBEEF3FAEB73F30459" author="Vaz-de-Mello F. Z." box="[996,1279,1356,1382]" page="194" pageId="143" pageNumber="144" year="2000">
<bibRefCitation id="F168BA8130D552EBEEF3FAEB73F30459" author="Vaz-de-Mello F. Z." box="[996,1279,1356,1382]" pageId="143" pageNumber="144" pagination="183 - 195" refId="ref140384" refString="Vaz-de-Mello F. Z. 2000. Estado atual de conhecimentos dos Scarabaeidae s. str. (Coleoptera: Scarabaeoidea) do Brasil. In: Martin-Piera F., Morrone J. J. &amp; Melic A. (eds) Hacia un Proyecto CYTED para el Inventario y Estimacion de la Diversidad Entomologica en Iberoamerica: PRIBES- 2000: 183 - 195. m 3 m: Monografias Tercer Milenio vol. 1, Sociedad Entomologica Aragonesa (SEA), Zaragoza." type="book chapter" year="2000">Vaz-de-Mello 2000: 194</bibRefCitation>
</treatmentCitation>
(checklist for 
<collectingCountry id="EDEE87E030D552EBEC04FAC8766C04B6" box="[275,352,1391,1417]" name="Brazil" pageId="143" pageNumber="144">Brazil</collectingCountry>
). — 
<treatmentCitation id="1458E16130D552EBEC8BFAC8740C04B5" author="Vaz-de-Mello F. Z. &amp; Silva R. L. R. &amp; Nunes L. G. O. A. &amp; Correa P. R. O. A." box="[412,768,1391,1418]" page="88" pageId="143" pageNumber="144" year="2011">
<bibRefCitation id="F168BA8130D552EBEC8BFAC8740C04B5" author="Vaz-de-Mello F. Z. &amp; Silva R. L. R. &amp; Nunes L. G. O. A. &amp; Correa P. R. O. A." box="[412,768,1391,1418]" pageId="143" pageNumber="144" pagination="77 - 102" refId="ref140870" refString="Vaz-de-Mello F. Z., Silva R. L. R., Nunes L. G. O. A. &amp; Correa P. R. O. A. 2011 b. Os besouros rola-bosta (Insecta: Coleoptera: Scarabaeidae: Scarabaeinae) da Fazenda Sao Nicolau. In: Rodrigues D. J., Izzo T. J. &amp; Battirola L. D. (eds.) Descobrindo a Amazonica Meridional: Biodiversidade da Fazenda Sao Nicolau: 77 - 102. Pau e Prosa Comunicacao, Cuiaba." type="book chapter" year="2011">
Vaz-de-Mello 
<emphasis id="A78D1B6230D552EBEF50FAD7758E04B6" box="[583,642,1391,1417]" italics="true" pageId="143" pageNumber="144">et al.</emphasis>
2011b: 88
</bibRefCitation>
</treatmentCitation>
(list). — 
<treatmentCitation id="1458E16130D552EBEE61FAC873FF04B5" ID-DOI="http://doi.org/10.5281/zenodo.3684689" ID-Zenodo-Dep="3684689" author="Hielkema A. J. &amp; Hielkema M. A." box="[886,1267,1391,1418]" httpUri="http://treatment.plazi.org/id/03D7FF1CFFF2FFD7FF48FDC2328DFE81" page="56" pageId="143" pageNumber="144" year="2019">
<bibRefCitation id="F168BA8130D552EBEE61FAC873FF04B5" author="Hielkema A. J. &amp; Hielkema M. A." box="[886,1267,1391,1418]" pageId="143" pageNumber="144" pagination="1 - 306" refId="ref124225" refString="Hielkema A. J. &amp; Hielkema M. A. 2019. An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas. Insecta Mundi 732 (732): 1 - 306." type="journal article" year="2019">Hielkema &amp; Hielkema 2019: 56</bibRefCitation>
</treatmentCitation>
(catalogue for the Guianas).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB30D552EBEDAAFA7375C3077F" pageId="143" pageNumber="144" type="materials_examined">
<paragraph id="9546C77030D552EBEDAAFA7376A604D1" blockId="143.[189,426,1492,1518]" box="[189,426,1492,1518]" pageId="143" pageNumber="144">
<heading id="CE0E701C30D552EBEDAAFA7376A604D1" bold="true" box="[189,426,1492,1518]" fontSize="11" level="3" pageId="143" pageNumber="144" reason="3">
<emphasis id="A78D1B6230D552EBEDAAFA7376A604D1" bold="true" box="[189,426,1492,1518]" pageId="143" pageNumber="144">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="9546C77030D552EBEDAAF9A475C3077F" blockId="143.[189,1399,1538,1601]" pageId="143" pageNumber="144">
<materialsCitation id="2591CD2D30D552EBEDAAF9A475C7077F" ID-GBIF-Occurrence="4979307400" collectingDate="2006-08" collectionCode="CEMT" collectorName="T. Good" country="Venezuela" elevation="550" latitude="7.7741" location="San Joaquin de Navay" longLatPrecision="7" longitude="-71.6675" municipality="Libertador" pageId="143" pageNumber="144" specimenCount="1" stateProvince="Tachira">
<collectingCountry id="EDEE87E030D552EBEDAAF9A476670722" box="[189,363,1539,1565]" name="Venezuela" pageId="143" pageNumber="144">VENEZUELA</collectingCountry>
‒ 
<emphasis id="A78D1B6230D552EBEC83F9A576FB0722" bold="true" box="[404,503,1538,1565]" pageId="143" pageNumber="144">
<collectingRegion id="573D099230D552EBEC83F9A576FB0722" box="[404,503,1538,1565]" country="Venezuela" name="Tachira" pageId="143" pageNumber="144">Táchira</collectingRegion>
</emphasis>
• 
<specimenCount id="83FF0CF930D552EBEF0BF9A475740722" box="[540,632,1539,1565]" count="1" pageId="143" pageNumber="144" type="generic">1 spec.</specimenCount>
; 
<collectingMunicipality id="75225D0A30D552EBEF9DF9A574040723" box="[650,776,1538,1564]" pageId="143" pageNumber="144">Libertador</collectingMunicipality>
, 
<location id="902691AB30D552EBEE0EF9A473390722" LSID="urn:lsid:plazi:treatment:1D50766630D752F4EF13FC7A76C10084:902691AB30D552EBEE0EF9A473390722" box="[793,1077,1538,1565]" country="Venezuela" latitude="7.7741" longLatPrecision="7" longitude="-71.6675" municipality="Libertador" name="San Joaquin de Navay" pageId="143" pageNumber="144" stateProvince="Tachira">San Joaquín de Navay</location>
; 
<geoCoordinate id="F0CDA1B730D552EBE950F9A473CA0722" box="[1095,1222,1539,1565]" degrees="7.7741" direction="north" orientation="latitude" pageId="143" pageNumber="144" precision="5" value="7.7741">7.7741° N</geoCoordinate>
, 
<geoCoordinate id="F0CDA1B730D552EBE9CBF9A5727E0722" box="[1244,1394,1538,1565]" degrees="71.6675" direction="west" orientation="longitude" pageId="143" pageNumber="144" precision="5" value="-71.6675">71.6675° W</geoCoordinate>
; 
<elevation id="1ED4204330D552EBEDAAF9817649077F" box="[189,325,1574,1601]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="143" pageNumber="144" unit="m" value="550.0">
<quantity id="52016A9530D552EBEDAAF981760B077F" box="[189,263,1574,1601]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="143" pageNumber="144" unit="m" value="550.0">550 m</quantity>
a.s.l.
</elevation>
; 
<date id="E147E1B030D552EBEC44F98176C7077F" box="[339,459,1574,1600]" pageId="143" pageNumber="144" value="2006-08">
<collectingDate id="F103185830D552EBEC44F98176C7077F" box="[339,459,1574,1600]" pageId="143" pageNumber="144" value="2006-08">Aug 2006</collectingDate>
</date>
; 
<collectorName id="380CA2A630D552EBECC1F981753A077F" box="[470,566,1574,1600]" pageId="143" pageNumber="144">T. Good</collectorName>
leg.; 
<collectionCode id="F3E85FB530D552EBEF6FF98175C7077F" box="[632,715,1574,1600]" pageId="143" pageNumber="144">CEMT</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB30D552EBEDAAF9C074FE0658" pageId="143" pageNumber="144" type="distribution">
<paragraph id="9546C77030D552EBEDAAF9C0765A07BE" blockId="143.[189,342,1639,1665]" box="[189,342,1639,1665]" pageId="143" pageNumber="144">
<heading id="CE0E701C30D552EBEDAAF9C0765A07BE" bold="true" box="[189,342,1639,1665]" fontSize="11" level="3" pageId="143" pageNumber="144" reason="3">
<emphasis id="A78D1B6230D552EBEDAAF9C0765A07BE" bold="true" box="[189,342,1639,1665]" pageId="143" pageNumber="144">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="9546C77030D552EBEDAAF931765507EC" blockId="143.[187,1398,1686,1747]" pageId="143" pageNumber="144">
<collectingCountry id="EDEE87E030D552EBEDAAF9317638078F" box="[189,308,1686,1712]" name="Colombia" pageId="143" pageNumber="144">Colombia</collectingCountry>
, 
<collectingCountry id="EDEE87E030D552EBEC2AF93176B5078F" box="[317,441,1686,1712]" name="Venezuela" pageId="143" pageNumber="144">Venezuela</collectingCountry>
, 
<collectingCountry id="EDEE87E030D552EBECD5F931752C078F" box="[450,544,1686,1712]" name="Guyana" pageId="143" pageNumber="144">Guyana</collectingCountry>
, 
<collectingCountry id="EDEE87E030D552EBEF3DF9317591078F" box="[554,669,1686,1712]" name="Suriname" pageId="143" pageNumber="144">Suriname</collectingCountry>
, French Guiana, 
<collectingCountry id="EDEE87E030D552EBEE48F93174A6078F" box="[863,938,1686,1712]" name="Brazil" pageId="143" pageNumber="144">Brazil</collectingCountry>
, 
<collectingCountry id="EDEE87E030D552EBEEA3F9317314078F" box="[948,1048,1686,1712]" name="Ecuador" pageId="143" pageNumber="144">Ecuador</collectingCountry>
, 
<collectingCountry id="EDEE87E030D552EBE936F9317376078F" box="[1057,1146,1686,1712]" name="Bolivia" pageId="143" pageNumber="144">Bolivia</collectingCountry>
, 
<collectingCountry id="EDEE87E030D552EBE993F93173FE078F" box="[1156,1266,1686,1712]" name="Paraguay" pageId="143" pageNumber="144">Paraguay</collectingCountry>
, 
<collectingCountry id="EDEE87E030D552EBE9EDF931727E078F" box="[1274,1394,1686,1712]" name="Argentina" pageId="143" pageNumber="144">Argentina</collectingCountry>
, and 
<collectingCountry id="EDEE87E030D552EBEDFAF91E765A07EC" box="[237,342,1721,1747]" name="Uruguay" pageId="143" pageNumber="144">Uruguay</collectingCountry>
.
</paragraph>
<paragraph id="9546C77030D552EBEDD3F95D76F9062B" blockId="143.[196,501,1786,1812]" box="[196,501,1786,1812]" pageId="143" pageNumber="144">
<emphasis id="A78D1B6230D552EBEDD3F95D76F9062B" bold="true" box="[196,501,1786,1812]" pageId="143" pageNumber="144">
Subregions of 
<collectingCountry id="EDEE87E030D552EBEC60F95D76F9062B" box="[375,501,1786,1812]" name="Venezuela" pageId="143" pageNumber="144">Venezuela</collectingCountry>
</emphasis>
</paragraph>
<paragraph id="9546C77030D552EBEDAAF88E74FE0658" blockId="143.[189,1399,1833,1895]" pageId="143" pageNumber="144">
Plains, System of hills and low sierras Lara-Falcón, System of hills and low piedmont mountains of the Guiana Shield, Andes mountains, and Central 
<collectingRegion id="573D099230D552EBEFCCF8EB74120659" box="[731,798,1868,1894]" country="New Zealand" name="West Coast" pageId="143" pageNumber="144">Coast</collectingRegion>
Mountain Range.
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB30CA52F4EDAAFEAF76C10084" pageId="144" pageNumber="145" type="reference_group">
<paragraph id="9546C77030CA52F4EDAAFEAF76A8001C" blockId="144.[189,420,264,291]" box="[189,420,264,291]" pageId="144" pageNumber="145">
<heading id="CE0E701C30CA52F4EDAAFEAF76A8001C" bold="true" box="[189,420,264,291]" fontSize="11" level="3" pageId="144" pageNumber="145" reason="3">
<emphasis id="A78D1B6230CA52F4EDAAFEAF76A8001C" bold="true" box="[189,420,264,291]" pageId="144" pageNumber="145">Literature records</emphasis>
</heading>
</paragraph>
<paragraph id="9546C77030CA52F4EDAAFE9076C10084" blockId="144.[189,1399,311,443]" pageId="144" pageNumber="145">
<bibRefCitation id="F168BA8130CA52F4EDAAFE90768A006E" author="Blanco J." box="[189,390,311,337]" pageId="144" pageNumber="145" pagination="39 - 46" refId="ref111553" refString="Blanco J. 1987. Catalogo de los Scarabaeidae (Coleoptera) coprofagos y necrofagos, del estado Tachira. Venezuela. Revista Cientifica Unet 1: 39 - 46." type="journal article" year="1987">Blanco 1987: 44</bibRefCitation>
(
<collectingCountry id="EDEE87E030CA52F4EC8EFE907514006E" box="[409,536,311,337]" name="Venezuela" pageId="144" pageNumber="145">Venezuela</collectingCountry>
: 
<collectingRegion id="573D099230CA52F4EF32FE907588006E" box="[549,644,311,337]" country="Venezuela" name="Tachira" pageId="144" pageNumber="145">Táchira</collectingRegion>
). — 
<bibRefCitation id="F168BA8130CA52F4EFD4FE907484006E" author="Genier F." box="[707,904,311,337]" pageId="144" pageNumber="145" pagination="1 - 169" refId="ref120431" refString="Genier F. 1996. A revision of the Neotropical genus Ontherus Erichson (Coleoptera: Scarabaeidae, Scarabaeinae). Memoirs of the Entomological Society of Canada 170: 1 - 169. https: // doi. org / 10.4039 / entm 128170 fv" type="journal article" year="1996">Génier 1996: 78</bibRefCitation>
(
<collectingCountry id="EDEE87E030CA52F4EE8CFE907316006E" box="[923,1050,311,337]" name="Venezuela" pageId="144" pageNumber="145">Venezuela</collectingCountry>
: 
<collectingRegion id="573D099230CA52F4E930FE9073BD006E" box="[1063,1201,311,337]" country="Venezuela" name="Anzoategui" pageId="144" pageNumber="145">Anzoategui</collectingRegion>
, 
<collectingRegion id="573D099230CA52F4E9A8FE907207006E" box="[1215,1291,311,337]" country="Venezuela" name="Apure" pageId="144" pageNumber="145">Apure</collectingRegion>
, 
<collectingRegion id="573D099230CA52F4E80EFE90727E006E" box="[1305,1394,311,337]" country="Venezuela" name="Aragua" pageId="144" pageNumber="145">Aragua</collectingRegion>
, 
<collectingRegion id="573D099230CA52F4EDAAFEFD761B004B" box="[189,279,346,372]" country="Venezuela" name="Bolivar" pageId="144" pageNumber="145">Bolívar</collectingRegion>
, 
<collectingRegion id="573D099230CA52F4EC35FEFD769B004B" box="[290,407,346,372]" country="Venezuela" name="Carabobo" pageId="144" pageNumber="145">Carabobo</collectingRegion>
, 
<collectingRegion id="573D099230CA52F4ECB4FEFD7550004B" box="[419,604,346,372]" country="Venezuela" name="Distrito Capital" pageId="144" pageNumber="145">Distrito Capital</collectingRegion>
, 
<collectingRegion id="573D099230CA52F4EF70FEFD75C4004B" box="[615,712,346,372]" country="Venezuela" name="Guarico" pageId="144" pageNumber="145">Guárico</collectingRegion>
, 
<collectingRegion id="573D099230CA52F4EFC3FEFD7436004B" box="[724,826,346,372]" country="Venezuela" name="Miranda" pageId="144" pageNumber="145">Miranda</collectingRegion>
, 
<collectingRegion id="573D099230CA52F4EE51FEFC74B8004A" box="[838,948,347,373]" country="Venezuela" name="Monagas" pageId="144" pageNumber="145">Monagas</collectingRegion>
, 
<collectingRegion id="573D099230CA52F4EED6FEFC734A004A" box="[961,1094,347,373]" country="Venezuela" name="Portuguesa" pageId="144" pageNumber="145">Portuguesa</collectingRegion>
, 
<collectingRegion id="573D099230CA52F4E946FEFD73A5004B" box="[1105,1193,346,372]" country="Venezuela" name="Tachira" pageId="144" pageNumber="145">Tachira</collectingRegion>
and 
<collectingRegion id="573D099230CA52F4E9F7FEFD724A004B" box="[1248,1350,346,372]" country="Venezuela" name="Yaracuy" pageId="144" pageNumber="145">Yaracuy</collectingRegion>
). — 
<bibRefCitation id="F168BA8130CA52F4EDAAFED976D400A7" author="Franca F. M. &amp; Korasaki V. &amp; Louzada J. &amp; Vaz-de-Mello F. Z." box="[189,472,381,408]" pageId="144" pageNumber="145" pagination="1 - 9" refId="ref119681" refString="Franca F. M., Korasaki V., Louzada J. &amp; Vaz-de-Mello F. Z. 2016. First report on dung beetles in intra-Amazonian savannahs in Roraima, Brazil. Biota Neotropica 16 (1): 1 - 9. https: // doi. org / 10.1590 / 1676 - 0611 - BN- 2015 - 0034" type="journal article" year="2016">
<collectingCountry id="EDEE87E030CA52F4EDAAFED9760100A7" box="[189,269,382,408]" name="France" pageId="144" pageNumber="145">França</collectingCountry>
<emphasis id="A78D1B6230CA52F4EC00FED9765900A8" box="[279,341,381,407]" italics="true" pageId="144" pageNumber="145">et al.</emphasis>
2016: 4–5
</bibRefCitation>
(
<collectingCountry id="EDEE87E030CA52F4ECFDFEDA756600A8" box="[490,618,381,407]" name="Venezuela" pageId="144" pageNumber="145">Venezuela</collectingCountry>
). — 
<bibRefCitation id="F168BA8130CA52F4EFBFFEDA74F200A7" author="Ferrer-Paris J. &amp; Sanchez-Mercado A. &amp; Rodriguez J. P." box="[680,1022,381,408]" pageId="144" pageNumber="145" pagination="89 - 110" refId="ref119203" refString="Ferrer-Paris J. R, Sanchez-Mercado A. &amp; Rodriguez J. P. 2013. Optimizacion del muestreo de invertebrados tropicales: un ejemplo con escarabajos coprofagos (Coleoptera: Scarabaeinae) en Venezuela. Revista de Biologia Tropical 61 (1): 89 - 110. https: // doi. org / 10.15517 / rbt. v 61 i 1.10941" type="journal article" year="2013">
Ferrer-Paris 
<emphasis id="A78D1B6230CA52F4EE2AFED9747700A8" box="[829,891,381,407]" italics="true" pageId="144" pageNumber="145">et al.</emphasis>
2013: 109
</bibRefCitation>
(
<collectingCountry id="EDEE87E030CA52F4E907FEDA738300A8" box="[1040,1167,381,407]" name="Venezuela" pageId="144" pageNumber="145">Venezuela</collectingCountry>
: 
<collectingRegion id="573D099230CA52F4E98BFED973FE00A7" box="[1180,1266,382,408]" country="Venezuela" name="Aragua" pageId="144" pageNumber="145">Aragua</collectingRegion>
[
<collectingRegion id="573D099230CA52F4E814FEDA726000A8" box="[1283,1388,381,407]" country="Venezuela" name="Guarico" pageId="144" pageNumber="145">Guárico</collectingRegion>
]: Altagracia de Orituco).
</paragraph>
</subSubSection>
</treatment>
</document>