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<mods:title id="8B758F81160B66C7B38D5910CE72B95B">The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution</mods:title>
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<mods:namePart id="6B77F6D47DD3F31E0B5C79B5E3802ED3">Rosa, Cecilia Lozano De La</mods:namePart>
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<mods:affiliation id="1B81A967BFD5BB2C7515034D15D2DCC7">Instituto Venezolano de Investigaciones Científicas, Centro de Estudios Botánicos y Agroforestales, Laboratorio de Ecología Espacial, Maracaibo, estado Zulia, Venezuela.</mods:affiliation>
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<mods:namePart id="54BB9D5B8BA2A7D301492A19D09B8449">Cupello, Mario</mods:namePart>
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<mods:affiliation id="53375896B72A6A59316E59505C62FB5B">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Curitiba, Paraná, Brazil. Currently at Department of Entomology, Texas A &amp; M University, College Station, TX, 77843, USA.</mods:affiliation>
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<mods:namePart id="48452EF040890712E3A9ECA354746517">Vaz-De-Mello, Fernando Z.</mods:namePart>
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<mods:affiliation id="F95E0F383C9E1EA27957F5EF0F1BEA36">Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, Cuiabá, Mato Grosso, Brazil.</mods:affiliation>
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<treatment id="1D507666309052AEEF14FA4075F306FF" ID-DOI="http://doi.org/10.5281/zenodo.14154847" ID-GBIF-Taxon="246298083" ID-Zenodo-Dep="14154847" LSID="urn:lsid:plazi:treatment:1D507666309052AEEF14FA4075F306FF" httpUri="http://treatment.plazi.org/id/1D507666309052AEEF14FA4075F306FF" lastPageNumber="203" pageId="202" pageNumber="203">
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<paragraph id="9546C770309052AEEF14FA40733C073E" blockId="202.[515,1072,1510,1537]" box="[515,1072,1510,1537]" pageId="202" pageNumber="203">
<heading id="CE0E701C309052AEEF14FA40733C073E" box="[515,1072,1510,1537]" centered="true" fontSize="11" level="2" pageId="202" pageNumber="203" reason="2">
Genus
<taxonomicName id="52F9BCF3309052AEEF41FA41733C073E" ID-CoL="7QQS" authority="Halffter &amp; Martinez, 1977" authorityName="Halffter &amp; Martinez" authorityYear="1977" box="[598,1072,1510,1537]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="202" pageNumber="203" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B62309052AEEF41FA4175F8073E" bold="true" box="[598,756,1510,1537]" italics="true" pageId="202" pageNumber="203">Sylvicanthon</emphasis>
<bibRefCitation id="F168BA81309052AEEFECFA40733C073E" author="Halffter G. &amp; Martinez A." box="[763,1072,1511,1537]" pageId="202" pageNumber="203" pagination="29 - 107" refId="ref122649" refString="Halffter G. &amp; Martinez A. 1977. Revision monografica de los Canthonina Americanos IV. Clave para generos y subgeneros. Folia Entomologica Mexicana 38: 29 - 107." type="journal article" year="1977">Halffter &amp; Martínez, 1977</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB309052AEEDAAF98F75F306FF" pageId="202" pageNumber="203" type="reference_group">
<paragraph id="9546C770309052AEEDAAF98F75A6075A" blockId="202.[189,1399,1576,1637]" pageId="202" pageNumber="203">
<treatmentCitationGroup id="B5E9E05E309052AEEDAAF98F75A6075A" pageId="202" pageNumber="203">
<treatmentCitation id="1458E161309052AEEDAAF98F75D8077D" author="Halffter G. &amp; Martinez A." box="[189,724,1576,1602]" page="61" pageId="202" pageNumber="203" year="1977">
<taxonomicName id="52F9BCF3309052AEEDAAF98F75D8077D" ID-CoL="7QQS" authority="Halffter &amp; Martinez, 1977: 61" authorityName="Halffter &amp; Martinez" authorityPageNumber="61" authorityYear="1977" box="[189,724,1576,1602]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="202" pageNumber="203" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B62309052AEEDAAF98F7659077D" box="[189,341,1576,1602]" italics="true" pageId="202" pageNumber="203">Sylvicanthon</emphasis>
<bibRefCitation id="F168BA81309052AEEC77F98F75D8077D" author="Halffter G. &amp; Martinez A." box="[352,724,1576,1602]" pageId="202" pageNumber="203" pagination="29 - 107" refId="ref122649" refString="Halffter G. &amp; Martinez A. 1977. Revision monografica de los Canthonina Americanos IV. Clave para generos y subgeneros. Folia Entomologica Mexicana 38: 29 - 107." type="journal article" year="1977">Halffter &amp; Martínez, 1977: 61</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description).
<typeStatus id="4A4279D2309052AEEEE7F98E7326077C" box="[1008,1066,1577,1603]" pageId="202" pageNumber="203">Type</typeStatus>
species:
<taxonomicName id="52F9BCF3309052AEE988F98F768B075A" ID-CoL="QPBJ" authority="Harold, 1869" authorityName="Harold" authorityYear="1869" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="202" pageNumber="203" phylum="Arthropoda" rank="species" species="candezei">
<emphasis id="A78D1B62309052AEE988F98F727B077D" box="[1183,1399,1576,1602]" italics="true" pageId="202" pageNumber="203">Canthon candezei</emphasis>
Harold, 1869
</taxonomicName>
, by original designation.
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</paragraph>
<paragraph id="9546C770309052AEEDAAF92A74BE06A2" blockId="202.[189,1399,1677,1985]" pageId="202" pageNumber="203">
<treatmentCitationGroup id="B5E9E05E309052AEEDAAF92A74BE06A2" pageId="202" pageNumber="203">
<taxonomicName id="52F9BCF3309052AEEDAAF92A76590798" ID-CoL="7QQS" authorityName="Halffter &amp; Martinez" authorityYear="1977" box="[189,341,1677,1703]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="202" pageNumber="203" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B62309052AEEDAAF92A76590798" box="[189,341,1677,1703]" italics="true" pageId="202" pageNumber="203">Sylvicanthon</emphasis>
</taxonomicName>
<treatmentCitation id="1458E161309052AEEC65F92A75EC0798" author="Halffter G. &amp; Edmonds W. D." box="[370,736,1677,1703]" page="139" pageId="202" pageNumber="203" year="1982">
<bibRefCitation id="F168BA81309052AEEC65F92A75EC0798" author="Halffter G. &amp; Edmonds W. D." box="[370,736,1677,1703]" pageId="202" pageNumber="203" refId="ref122441" refString="Halffter G. &amp; Edmonds W. D. 1982. The Nesting Behavior of Dung Beetles (Scarabaeinae). An Ecological and Evolutive Approach. Instituto de Ecologia, Mexico, D. F." type="book" year="1982">Halffter &amp; Edmonds 1982: 139</bibRefCitation>
</treatmentCitation>
(catalog, distribution). —
<treatmentCitation id="1458E161309052AEE90EF92A761907F5" author="Medina C. A. &amp; Lopera-Toro A." page="311" pageId="202" pageNumber="203" year="2000">
<bibRefCitation id="F168BA81309052AEE90EF92A761907F5" author="Medina C. A. &amp; Lopera-Toro A." pageId="202" pageNumber="203" pagination="299 - 315" refId="ref130725" refString="Medina C. A. &amp; Lopera-Toro A. 2000. Clave ilustrada para la identificacion de generos de escarabajos coprofagos (Coleoptera: Scarabaeinae) de Colombia. Caldasia 22: 299 - 315." type="journal article" year="2000">Medina &amp; Lopera-Toro 2000: 311</bibRefCitation>
</treatmentCitation>
(characteristics). —
<treatmentCitation id="1458E161309052AEEF0FF917743307F4" author="Vaz-de-Mello F. Z." box="[536,831,1712,1739]" page="195" pageId="202" pageNumber="203" year="2000">
<bibRefCitation id="F168BA81309052AEEF0FF917743307F4" author="Vaz-de-Mello F. Z." box="[536,831,1712,1739]" pageId="202" pageNumber="203" pagination="183 - 195" refId="ref140384" refString="Vaz-de-Mello F. Z. 2000. Estado atual de conhecimentos dos Scarabaeidae s. str. (Coleoptera: Scarabaeoidea) do Brasil. In: Martin-Piera F., Morrone J. J. &amp; Melic A. (eds) Hacia un Proyecto CYTED para el Inventario y Estimacion de la Diversidad Entomologica en Iberoamerica: PRIBES- 2000: 183 - 195. m 3 m: Monografias Tercer Milenio vol. 1, Sociedad Entomologica Aragonesa (SEA), Zaragoza." type="book chapter" year="2000">Vaz-de-Mello 2000: 195</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E0309052AEEEE1F917734F07F5" box="[1014,1091,1712,1738]" name="Brazil" pageId="202" pageNumber="203">Brazil</collectingCountry>
). —
<treatmentCitation id="1458E161309052AEE993F917761A07D2" author="Medina C. A. &amp; Lopera-Toro A. &amp; Vitolo A. &amp; Gill B." page="137" pageId="202" pageNumber="203" year="2001">
<bibRefCitation id="F168BA81309052AEE993F917761A07D2" author="Medina C. A. &amp; Lopera-Toro A. &amp; Vitolo A. &amp; Gill B." pageId="202" pageNumber="203" pagination="131 - 144" refId="ref130799" refString="Medina C. A., Lopera-Toro A., Vitolo A. &amp; Gill B. 2001. Escarabajos coprofagos (Coleoptera: Scarabaeinae) de Colombia. Biota Colombiana 2: 131 - 144." type="journal article" year="2001">
Medina
<emphasis id="A78D1B62309052AEE9FEF916722407F5" box="[1257,1320,1712,1738]" italics="true" pageId="202" pageNumber="203">et al.</emphasis>
2001: 137
</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E0309052AEECDBF974754907D2" box="[460,581,1747,1773]" name="Colombia" pageId="202" pageNumber="203">Colombia</collectingCountry>
);
<treatmentCitation id="1458E161309052AEEF4BF97475CB07D1" author="Medina C. A. &amp; Scholtz C. H. &amp; Gill B. D." box="[604,711,1747,1774]" page="65" pageId="202" pageNumber="203" year="2003">
<bibRefCitation id="F168BA81309052AEEF4BF97475CB07D1" author="Medina C. A. &amp; Scholtz C. H. &amp; Gill B. D." box="[604,711,1747,1774]" pageId="202" pageNumber="203" pagination="23 - 68" refId="ref130892" refString="Medina C. A., Scholtz C. H. &amp; Gill B. D. 2003. Morphological variation and systematics of Canthon and related genera of new world Canthonini dung beetles. Deutsche Entomologische Zeitschrift 50 (1): 23 - 68. https: // doi. org / 10.1002 / mmnd. 20030500105" type="journal article" year="2003">2003: 65</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="1458E161309052AEEEBFF974721C07D1" author="Hamel-Leigue A. &amp; Mann D. J. &amp; Vaz-de-Mello F. Z. &amp; Herzog S. K." box="[936,1296,1747,1774]" page="15" pageId="202" pageNumber="203" year="2006">
<bibRefCitation id="F168BA81309052AEEEBFF974721C07D1" author="Hamel-Leigue A. &amp; Mann D. J. &amp; Vaz-de-Mello F. Z. &amp; Herzog S. K." box="[936,1296,1747,1774]" pageId="202" pageNumber="203" pagination="1 - 18" refId="ref122830" refString="Hamel-Leigue A., Mann D. J., Vaz-de-Mello F. Z. &amp; Herzog S. K. 2006. Hacia un inventario de los escarabajos peloteros (Coleoptera: Scarabaeinae) de Bolivia: primera compilacion de los generos y especies registrados para el pais. Revista Boliviana de Ecologia y Conservacion Ambiental 20: 1 - 18." type="journal article" year="2006">
Hamel-Leigue
<emphasis id="A78D1B62309052AEE94BF973739707D2" box="[1116,1179,1747,1773]" italics="true" pageId="202" pageNumber="203">et al.</emphasis>
2006: 15
</bibRefCitation>
</treatmentCitation>
(list for
<collectingCountry id="EDEE87E0309052AEEDFDF951764A062F" box="[234,326,1782,1808]" name="Bolivia" pageId="202" pageNumber="203">Bolivia</collectingCountry>
). —
<treatmentCitation id="1458E161309052AEEC6CF95175D8062F" author="Vaz-de-Mello F. Z. &amp; Edmonds W. D. &amp; Ocampo F. C. &amp; Schoolmeesters P." box="[379,724,1782,1809]" page="26" pageId="202" pageNumber="203" year="2011">
<bibRefCitation id="F168BA81309052AEEC6CF95175D8062F" author="Vaz-de-Mello F. Z. &amp; Edmonds W. D. &amp; Ocampo F. C. &amp; Schoolmeesters P." box="[379,724,1782,1809]" pageId="202" pageNumber="203" pagination="1 - 73" refId="ref140802" refString="Vaz-de-Mello F. Z., Edmonds W. D., Ocampo F. C., Schoolmeesters P. 2011 a. A multilingual key to the genera and subgenera of the subfamily Scarabaeinae of the New World (Coleoptera: Scarabaeidae). Zootaxa 2854 (1): 1 - 73. https: // doi. org / 10.11646 / zootaxa. 2854.1.1" type="journal article" year="2011">
Vaz-de-Mello
<emphasis id="A78D1B62309052AEEF37F9507550062F" box="[544,604,1782,1808]" italics="true" pageId="202" pageNumber="203">et al.</emphasis>
2011a: 26
</bibRefCitation>
</treatmentCitation>
(key). —
<treatmentCitation id="1458E161309052AEEE5EF9517364062E" author="Carvajal V. &amp; Villamarin S. &amp; Ortega A. M." box="[841,1128,1782,1809]" page="117" pageId="202" pageNumber="203" year="2011">
<bibRefCitation id="F168BA81309052AEEE5EF9517364062E" author="Carvajal V. &amp; Villamarin S. &amp; Ortega A. M." box="[841,1128,1782,1809]" pageId="202" pageNumber="203" refId="ref113831" refString="Carvajal V., Villamarin S. &amp; Ortega A. M. 2011. Escarabajos del Ecuador. Principales Generos. Instituto de Ciencias Biologicas, Escuela Politecnica Nacional, Quito." type="book" year="2011">
Carvajal
<emphasis id="A78D1B62309052AEEEA6F95074E0062F" box="[945,1004,1782,1808]" italics="true" pageId="202" pageNumber="203">et al.</emphasis>
2011: 117
</bibRefCitation>
</treatmentCitation>
, 316 (diagnosis, list for
<collectingCountry id="EDEE87E0309052AEEDFEF8BD765C060B" box="[233,336,1818,1844]" name="Ecuador" pageId="202" pageNumber="203">Ecuador</collectingCountry>
). —
<treatmentCitation id="1458E161309052AEEC90F8BD75B2060B" author="Solis A. &amp; Kohlmann B." box="[391,702,1818,1844]" page="3" pageId="202" pageNumber="203" year="2012">
<bibRefCitation id="F168BA81309052AEEC90F8BD75B2060B" author="Solis A. &amp; Kohlmann B." box="[391,702,1818,1844]" pageId="202" pageNumber="203" pagination="1 - 32" refId="ref139201" refString="Solis A. &amp; Kohlmann B. 2012. Checklist and distribution atlas of the Scarabaeinae of Costa Rica. Zootaxa 3482: 1 - 32. https: // doi. org / 10.11646 / zootaxa. 3482.1.1" type="journal article" year="2012">Solís &amp; Kohlmann 2012: 3</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E0309052AEEE73F8BD74E9060B" box="[868,997,1818,1844]" name="Costa Rica" pageId="202" pageNumber="203">Costa Rica</collectingCountry>
, cited as synonym of
<taxonomicName id="52F9BCF3309052AEE9CBF8BD724D060B" ID-CoL="62L63" authorityName="Hoffmannsegg" authorityYear="1817" box="[1244,1345,1818,1844]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="202" pageNumber="203" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B62309052AEE9CBF8BD724D060B" box="[1244,1345,1818,1844]" italics="true" pageId="202" pageNumber="203">Canthon</emphasis>
</taxonomicName>
). —
<treatmentCitation id="1458E161309052AEEDFEF89A75890668" author="Boilly O. &amp; Vaz-de-Mello F. Z." box="[233,645,1853,1879]" page="107" pageId="202" pageNumber="203" year="2013">
<bibRefCitation id="F168BA81309052AEEDFEF89A75890668" author="Boilly O. &amp; Vaz-de-Mello F. Z." box="[233,645,1853,1879]" pageId="202" pageNumber="203" pagination="103 - 112" refId="ref111991" refString="Boilly O. &amp; Vaz-de-Mello F. Z. 2013. Les Scarabaeinae de Guyane: cle illustree des genres. ACOREP- France 7: 103 - 112." type="journal article" year="2013">Boilly &amp; Vaz-de-Mello 2013: 107</bibRefCitation>
</treatmentCitation>
(key). —
<treatmentCitation id="1458E161309052AEEE10F89A73330668" author="Chamorro W. &amp; Marin-Armijos D. &amp; Granda V. &amp; Vaz-de-Mello F. Z." box="[775,1087,1853,1879]" page="76" pageId="202" pageNumber="203" year="2018">
<bibRefCitation id="F168BA81309052AEEE10F89A73330668" author="Chamorro W. &amp; Marin-Armijos D. &amp; Granda V. &amp; Vaz-de-Mello F. Z." box="[775,1087,1853,1879]" pageId="202" pageNumber="203" pagination="72 - 100" refId="ref114035" refString="Chamorro W., Marin-Armijos D., Granda V. &amp; Vaz-de-Mello F. Z. 2018. Listado de especies y clave de generos y subgeneros de escarabajos estercoleros (Coleoptera: Scarabaeidae: Scarabaeinae) presentes y presuntos para Ecuador. Revista Colombiana de Entomologia 44 (1): 72 - 100. https: // doi. org / 10.25100 / socolen. v 44 i 1.6545" type="journal article" year="2018">
Chamorro
<emphasis id="A78D1B62309052AEEE9EF89974CB0668" box="[905,967,1853,1879]" italics="true" pageId="202" pageNumber="203">et al.</emphasis>
2018: 76
</bibRefCitation>
</treatmentCitation>
, 98 (list for
<collectingCountry id="EDEE87E0309052AEE9CBF89A724F0668" box="[1244,1347,1853,1879]" name="Ecuador" pageId="202" pageNumber="203">Ecuador</collectingCountry>
). —
<treatmentCitation id="1458E161309052AEEDFDF8C7759D0645" author="Vaz-de-Mello F. Z. &amp; Cupello M." box="[234,657,1888,1914]" page="20" pageId="202" pageNumber="203" year="2018">
<bibRefCitation id="F168BA81309052AEEDFDF8C7759D0645" author="Vaz-de-Mello F. Z. &amp; Cupello M." box="[234,657,1888,1914]" pageId="202" pageNumber="203" pagination="33 - 76" refId="ref140606" refString="Vaz-de-Mello F. Z. &amp; Cupello M. 2018. The type specimens of South American dung beetles. Part I: on the species described in the genus Canthon Hoffmannsegg, 1817 by the German entomologist Adolf Schmidt (1856 - 1923) (Coleoptera, Scarabaeidae, Scarabaeinae). Spixiana 41 (1): 33 - 76." type="journal article" year="2018">Cupello &amp; Vaz-de-Mello 2018: 20</bibRefCitation>
</treatmentCitation>
, 56 (revision, redescription, key);
<treatmentCitation id="1458E161309052AEE92CF8C773BA0645" author="Cupello M. &amp; Vaz-de-Mello F. Z." box="[1083,1206,1888,1914]" page="239" pageId="202" pageNumber="203" year="2019">
<bibRefCitation id="F168BA81309052AEE92CF8C773BA0645" box="[1083,1206,1888,1914]" pageId="202" pageNumber="203" year="2019">2019: 239</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="1458E161309052AEEDFDF824756E06A2" ID-DOI="http://doi.org/10.5281/zenodo.3683692" ID-Zenodo-Dep="3683692" author="Hielkema A. J. &amp; Hielkema M. A." box="[234,610,1923,1949]" httpUri="http://treatment.plazi.org/id/03D7FF1CFF9FFFB9FF48FEFD33BBFA08" page="83" pageId="202" pageNumber="203" year="2019">
<bibRefCitation id="F168BA81309052AEEDFDF824756E06A2" author="Hielkema A. J. &amp; Hielkema M. A." box="[234,610,1923,1949]" pageId="202" pageNumber="203" pagination="1 - 306" refId="ref124225" refString="Hielkema A. J. &amp; Hielkema M. A. 2019. An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas. Insecta Mundi 732 (732): 1 - 306." type="journal article" year="2019">Hielkema &amp; Hielkema 2019: 83</bibRefCitation>
</treatmentCitation>
(catalogue for the Guianas).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C770309052AEEDAAF80175F306FF" blockId="202.[189,1399,1677,1985]" box="[189,767,1958,1985]" pageId="202" pageNumber="203">
<treatmentCitationGroup id="B5E9E05E309052AEEDAAF80175F306FF" box="[189,767,1958,1985]" pageId="202" pageNumber="203">
<taxonomicName id="52F9BCF3309052AEEDAAF80176D906FE" box="[189,469,1958,1985]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="202" pageNumber="203" phylum="Arthropoda" rank="subGenus" subGenus="Sylvicanthon">
<emphasis id="A78D1B62309052AEEDAAF801762E06FF" box="[189,290,1958,1984]" italics="true" pageId="202" pageNumber="203">Canthon</emphasis>
(
<emphasis id="A78D1B62309052AEEC24F80176C706FF" box="[307,459,1958,1984]" italics="true" pageId="202" pageNumber="203">Sylvicanthon</emphasis>
)
</taxonomicName>
<treatmentCitation id="1458E161309052AEECE5F80175B406FF" author="Krajcik M." box="[498,696,1958,1985]" page="63" pageId="202" pageNumber="203" year="2012">
<bibRefCitation id="F168BA81309052AEECE5F80175B406FF" author="Krajcik M." box="[498,696,1958,1985]" pageId="202" pageNumber="203" pagination="1 - 278" refId="ref126632" refString="Krajcik M. 2012. Checklist of the World Scarabaeoidea. Animma. X Supplement 5: 1 - 278." type="journal article" year="2012">Krajcik 2012: 63</bibRefCitation>
</treatmentCitation>
(list).
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</paragraph>
</subSubSection>
</treatment>
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