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<document id="D0B55B8ABD6C85A319470BE56C44AAA7" ID-DOI="10.5852/ejt.2024.959.2677" ID-ISSN="2118-9773" ID-Zenodo-Dep="13945464" ID-ZooBank="30872C13-516F-42FA-AFA7-30ADC6BF1BAF" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1729166637250" checkinUser="plazi" docAuthor="Rosa, Cecilia Lozano De La, Cupello, Mario &amp; Vaz-De-Mello, Fernando Z." docDate="2024" docId="1D507666304B5277ECEAF978752E036B" docLanguage="en" docName="EJT.2024.959.1-272.pdf" docOrigin="European Journal of Taxonomy 959 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2677/12401" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Ateuchus ambiguus Martinez &amp; Martinez 1990" docType="treatment" docVersion="2" lastPageNumber="20" masterDocId="E1690E1E305A5264ED17FFA7770C013F" masterDocTitle="The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution" masterLastPageNumber="272" masterPageNumber="1" pageNumber="18" updateTime="1729169574746" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="6D8C642E53C91B20F5C16A6CEA574549">
<mods:title id="E41697A26208E111751D0E40547C78D7">The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution</mods:title>
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<mods:roleTerm id="40E5C8536A021746FAE725999AA4A164">Author</mods:roleTerm>
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<mods:namePart id="02B5A30F5485BCFE1C3A0739CCF4483D">Rosa, Cecilia Lozano De La</mods:namePart>
<mods:nameIdentifier id="86B1CD6E28A53FD41D5BEF61257ED8F3" type="LSID">36CFEA67-71C5-4F3C-97F8-B57E43C811CC</mods:nameIdentifier>
<mods:affiliation id="DF640B11F00E6AA771D82D69CBC71098">Instituto Venezolano de Investigaciones Científicas, Centro de Estudios Botánicos y Agroforestales, Laboratorio de Ecología Espacial, Maracaibo, estado Zulia, Venezuela.</mods:affiliation>
<mods:nameIdentifier id="E44B8CE1B5D496D9803880E0D3EC2336" type="email">lozanoceci@gmail.com</mods:nameIdentifier>
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<mods:namePart id="F0D732F942ECC0CF6C312EC32BF360E0">Cupello, Mario</mods:namePart>
<mods:nameIdentifier id="A7D989DAE176A980F0A236D45FC6C145" type="LSID">BDB03C18-7095-4EAA-8BDD-03CB4F79676D</mods:nameIdentifier>
<mods:affiliation id="6ED4AAA687A74E7D89AE074662E8D42B">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Curitiba, Paraná, Brazil. Currently at Department of Entomology, Texas A &amp; M University, College Station, TX, 77843, USA.</mods:affiliation>
<mods:nameIdentifier id="A0504DAD0399DA5742152B106414ECE5" type="email">mcupello@hotmail.com</mods:nameIdentifier>
</mods:name>
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<mods:roleTerm id="9A4D67BCF39A6971801A8F7C8D4276B2">Author</mods:roleTerm>
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<mods:namePart id="5335BA3AD518CA2CF64A94AFA16A04BA">Vaz-De-Mello, Fernando Z.</mods:namePart>
<mods:nameIdentifier id="35D344AE50A96025CDB30FED31DDE337" type="LSID">2BC093C2-3E2B-466B-B31C-9F3D9FA844BC</mods:nameIdentifier>
<mods:affiliation id="3D37C095BFFE92C5A6DD5B9064E06457">Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, Cuiabá, Mato Grosso, Brazil.</mods:affiliation>
<mods:nameIdentifier id="F45155333A1807AA5E63EE978338EAEF" type="email">vazdemello@gmail.com</mods:nameIdentifier>
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<mods:date id="3B072A8F8DCAAE91CCE2674264B444CC">2024</mods:date>
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<treatment id="1D507666304B5277ECEAF978752E036B" ID-DOI="http://doi.org/10.5281/zenodo.13945579" ID-Zenodo-Dep="13945579" LSID="urn:lsid:plazi:treatment:1D507666304B5277ECEAF978752E036B" httpUri="http://treatment.plazi.org/id/1D507666304B5277ECEAF978752E036B" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18">
<subSubSection id="DDE394FB304B5275ECEAF978733B07C5" box="[509,1079,1759,1786]" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="9546C770304B5275ECEAF978733B07C5" blockId="17.[509,1079,1759,1818]" box="[509,1079,1759,1786]" pageId="17" pageNumber="18">
<heading id="CE0E701C304B5275ECEAF978733B07C5" box="[509,1079,1759,1786]" centered="true" fontSize="11" level="2" pageId="17" pageNumber="18" reason="2">
<taxonomicName id="52F9BCF3304B5275ECEAF978733B07C5" ID-CoL="7ZLWV" authority="Martinez &amp; Martinez, 1990" authorityName="Martinez &amp; Martinez" authorityYear="1990" box="[509,1079,1759,1786]" class="Insecta" family="Scarabaeidae" genus="Ateuchus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="ambiguus">
<emphasis id="A78D1B62304B5275ECEAF97875E107C5" bold="true" box="[509,749,1759,1786]" italics="true" pageId="17" pageNumber="18">Ateuchus ambiguus</emphasis>
<bibRefCitation id="F168BA81304B5275EFE3F947733B07C5" author="Martinez A. &amp; Martinez A." box="[756,1079,1760,1786]" pageId="17" pageNumber="18" pagination="159 - 163" refId="ref130132" refString="Martinez A. &amp; Martinez A. 1990. Dos nuevas especies de Ateuchus Weber (Col. Scarab. Ateuchina). Boletin de Entomologia Venezolana (N. S.) 5 (20): 159 - 163." type="journal article" year="1990">Martínez &amp; Martínez, 1990</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB304B5275EFF9F8A774490625" box="[750,837,1792,1818]" pageId="17" pageNumber="18" type="description">
<paragraph id="9546C770304B5275EFF9F8A774490625" blockId="17.[509,1079,1759,1818]" box="[750,837,1792,1818]" pageId="17" pageNumber="18">
<figureCitation id="0DC2DBF5304B5275EFF9F8A774490625" box="[750,837,1792,1818]" captionStart="Fig" captionStartId="18.[189,232,1962,1988]" captionTargetBox="[269,1300,279,1899]" captionTargetId="figure-17@18.[257,1331,266,1914]" captionTargetPageId="18" captionText="Fig. 8. Species distribution. A. Ateuchus alvarezi (Martínez &amp; Halffter, 1986). B. Ateuchus ambiguus Martínez &amp; Martínez, 1990. Green diamond = CEMT collection data; black circle = literature data." figureDoi="http://doi.org/10.5281/zenodo.13945482" httpUri="https://zenodo.org/record/13945482/files/figure.png" pageId="17" pageNumber="18">Fig. 8B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB304B5275EDAAF89875C006E4" pageId="17" pageNumber="18" type="reference_group">
<paragraph id="9546C770304B5275EDAAF898748206A5" blockId="17.[189,1401,1855,1946]" pageId="17" pageNumber="18">
<treatmentCitationGroup id="B5E9E05E304B5275EDAAF898748206A5" pageId="17" pageNumber="18">
<treatmentCitation id="1458E161304B5275EDAAF89874380666" author="Martinez A. &amp; Martinez A." box="[189,820,1855,1881]" page="161" pageId="17" pageNumber="18" year="1990">
<taxonomicName id="52F9BCF3304B5275EDAAF89874380666" ID-CoL="7ZLWV" authority="Martinez &amp; Martinez, 1990: 161" authorityName="Martinez &amp; Martinez" authorityPageNumber="161" authorityYear="1990" box="[189,820,1855,1881]" class="Insecta" family="Scarabaeidae" genus="Ateuchus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="ambiguus">
<emphasis id="A78D1B62304B5275EDAAF89876A80666" box="[189,420,1855,1881]" italics="true" pageId="17" pageNumber="18">Ateuchus ambiguus</emphasis>
<bibRefCitation id="F168BA81304B5275ECB9F89874380666" author="Martinez A. &amp; Martinez A." box="[430,820,1855,1881]" pageId="17" pageNumber="18" pagination="159 - 163" refId="ref130132" refString="Martinez A. &amp; Martinez A. 1990. Dos nuevas especies de Ateuchus Weber (Col. Scarab. Ateuchina). Boletin de Entomologia Venezolana (N. S.) 5 (20): 159 - 163." type="journal article" year="1990">Martínez &amp; Martínez, 1990: 161</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description). Type locality:
<collectingCountry id="EDEE87E0304B5275E9EFF89872790666" box="[1272,1397,1855,1881]" name="Venezuela" pageId="17" pageNumber="18">Venezuela</collectingCountry>
:
<collectingRegion id="573D0992304B5275EDFBF8F876490646" box="[236,325,1887,1913]" country="Venezuela" name="Merida" pageId="17" pageNumber="18">Mérida</collectingRegion>
: San Rafael del Páramo de Mucuchíes, Laguna Grande,
<quantity id="52016A95304B5275E903F8C7737A0645" box="[1044,1142,1888,1914]" metricMagnitude="3" metricUnit="m" metricValue="3.8" pageId="17" pageNumber="18" unit="m" value="3800.0">3800 m</quantity>
. Name-bearing type:
<typeStatus id="4A4279D2304B5275EDFDF827765D06A5" box="[234,337,1920,1946]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
(MACN), confirmed to be there by MC in 2014.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C770304B5275EDAAF86675C006E4" blockId="17.[189,716,1985,2012]" box="[189,716,1985,2012]" pageId="17" pageNumber="18">
<treatmentCitationGroup id="B5E9E05E304B5275EDAAF86675C006E4" box="[189,716,1985,2012]" pageId="17" pageNumber="18">
<emphasis id="A78D1B62304B5275EDAAF86676B406E4" box="[189,440,1985,2011]" italics="true" pageId="17" pageNumber="18">
<taxonomicName id="52F9BCF3304B5275EDAAF86676AE06E4" ID-CoL="7ZLWV" authorityName="Martinez &amp; Martinez" authorityYear="1990" box="[189,418,1985,2011]" class="Insecta" family="Scarabaeidae" genus="Ateuchus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="ambiguus">Ateuchus ambiguus</taxonomicName>
–
</emphasis>
<treatmentCitation id="1458E161304B5275ECA8F866758A06E3" author="Krajcik M." box="[447,646,1985,2012]" page="49" pageId="17" pageNumber="18" year="2012">
<bibRefCitation id="F168BA81304B5275ECA8F866758A06E3" author="Krajcik M." box="[447,646,1985,2012]" pageId="17" pageNumber="18" pagination="1 - 278" refId="ref126632" refString="Krajcik M. 2012. Checklist of the World Scarabaeoidea. Animma. X Supplement 5: 1 - 278." type="journal article" year="2012">Krajcik 2012: 49</bibRefCitation>
</treatmentCitation>
(list).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="C18697F830485276EDAAF80D724406D8" ID-DOI="http://doi.org/10.5281/zenodo.13945482" ID-Zenodo-Dep="13945482" httpUri="https://zenodo.org/record/13945482/files/figure.png" pageId="18" pageNumber="19" startId="18.[189,232,1962,1988]" targetBox="[269,1300,279,1899]" targetPageId="18">
<paragraph id="9546C77030485276EDAAF80D724406D8" blockId="18.[189,1398,1962,2024]" pageId="18" pageNumber="19">
<emphasis id="A78D1B6230485276EDAAF80D760106FB" bold="true" box="[189,269,1962,1988]" pageId="18" pageNumber="19">Fig. 8.</emphasis>
Species distribution.
<emphasis id="A78D1B6230485276EF05F80D752A06FB" bold="true" box="[530,550,1962,1988]" pageId="18" pageNumber="19">A</emphasis>
.
<taxonomicName id="52F9BCF330485276EF22F80D736C06FB" authority="(Martinez &amp; Halffter, 1986)" baseAuthorityName="Martinez &amp; Halffter" baseAuthorityYear="1986" box="[565,1120,1962,1988]" class="Insecta" family="Scarabaeidae" genus="Ateuchus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="alvarezi">
<emphasis id="A78D1B6230485276EF22F80D740A06FB" box="[565,774,1962,1988]" italics="true" pageId="18" pageNumber="19">Ateuchus alvarezi</emphasis>
(
<bibRefCitation id="F168BA8130485276EE00F80D735A06FB" author="Martinez A. &amp; Halffter G." box="[791,1110,1962,1988]" pageId="18" pageNumber="19" pagination="19 - 40" refId="ref130097" refString="Martinez A. &amp; Halffter G. 1986. Situacion del genero Canthidium. Acta Zoologica Mexicana 18: 19 - 40. https: // doi. org / 10.21829 / azm. 1986.13182047" type="journal article" year="1986">Martínez &amp; Halffter, 1986</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="A78D1B6230485276E979F80D738C06FB" bold="true" box="[1134,1152,1962,1988]" pageId="18" pageNumber="19">B</emphasis>
.
<taxonomicName id="52F9BCF330485276E987F80D750F06D8" authority="Martinez &amp; Martinez, 1990" authorityName="Martinez &amp; Martinez" authorityYear="1990" class="Insecta" family="Scarabaeidae" genus="Ateuchus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="ambiguus">
<emphasis id="A78D1B6230485276E987F80D727A06FB" box="[1168,1398,1962,1988]" italics="true" pageId="18" pageNumber="19">Ateuchus ambiguus</emphasis>
<bibRefCitation id="F168BA8130485276EDAAF86A750F06D8" author="Martinez A. &amp; Martinez A." box="[189,515,1997,2023]" pageId="18" pageNumber="19" pagination="159 - 163" refId="ref130132" refString="Martinez A. &amp; Martinez A. 1990. Dos nuevas especies de Ateuchus Weber (Col. Scarab. Ateuchina). Boletin de Entomologia Venezolana (N. S.) 5 (20): 159 - 163." type="journal article" year="1990">Martínez &amp; Martínez, 1990</bibRefCitation>
</taxonomicName>
. Green diamond = CEMT collection data; black circle = literature data.
</paragraph>
</caption>
<subSubSection id="DDE394FB30495277EDAAFEAF752E036B" pageId="19" pageNumber="20" type="distribution">
<paragraph id="9546C77030495277EDAAFEAF765A001D" blockId="19.[189,342,264,290]" box="[189,342,264,290]" pageId="19" pageNumber="20">
<heading id="CE0E701C30495277EDAAFEAF765A001D" bold="true" box="[189,342,264,290]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="A78D1B6230495277EDAAFEAF765A001D" bold="true" box="[189,342,264,290]" pageId="19" pageNumber="20">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="9546C77030495277EDAAFE90741D006E" blockId="19.[189,785,311,337]" box="[189,785,311,337]" pageId="19" pageNumber="20">
<collectingCountry id="EDEE87E030495277EDAAFE90763A006E" box="[189,310,311,337]" name="Venezuela" pageId="19" pageNumber="20">Venezuela</collectingCountry>
(endemic) (
<bibRefCitation id="F168BA8130495277ECD4FE907408006E" author="Martinez A. &amp; Martinez A." box="[451,772,311,337]" pageId="19" pageNumber="20" pagination="159 - 163" refId="ref130132" refString="Martinez A. &amp; Martinez A. 1990. Dos nuevas especies de Ateuchus Weber (Col. Scarab. Ateuchina). Boletin de Entomologia Venezolana (N. S.) 5 (20): 159 - 163." type="journal article" year="1990">Martínez &amp; Martínez 1990</bibRefCitation>
).
</paragraph>
<paragraph id="9546C77030495277EDAAFEDF76EE00AC" blockId="19.[189,482,376,403]" box="[189,482,376,403]" pageId="19" pageNumber="20">
<heading id="CE0E701C30495277EDAAFEDF76EE00AC" bold="true" box="[189,482,376,403]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="A78D1B6230495277EDAAFEDF76EE00AC" bold="true" box="[189,482,376,403]" pageId="19" pageNumber="20">
Subregion of
<collectingCountry id="EDEE87E030495277EC73FEDE76EE00AC" box="[356,482,377,403]" name="Venezuela" pageId="19" pageNumber="20">Venezuela</collectingCountry>
</emphasis>
</heading>
</paragraph>
<paragraph id="9546C77030495277EDAAFE00769D00FE" blockId="19.[189,401,423,449]" box="[189,401,423,449]" pageId="19" pageNumber="20">Andes mountains.</paragraph>
<paragraph id="9546C77030495277EDAAFE4F76A8033C" blockId="19.[189,420,488,515]" box="[189,420,488,515]" pageId="19" pageNumber="20">
<heading id="CE0E701C30495277EDAAFE4F76A8033C" bold="true" box="[189,420,488,515]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="A78D1B6230495277EDAAFE4F76A8033C" bold="true" box="[189,420,488,515]" pageId="19" pageNumber="20">Literature records</emphasis>
</heading>
</paragraph>
<paragraph id="9546C77030495277EDAAFDB0752E036B" blockId="19.[187,1398,535,596]" pageId="19" pageNumber="20">
<bibRefCitation id="F168BA8130495277EDAAFDB07520030E" author="Martinez A. &amp; Martinez A." box="[189,556,535,561]" pageId="19" pageNumber="20" pagination="159 - 163" refId="ref130132" refString="Martinez A. &amp; Martinez A. 1990. Dos nuevas especies de Ateuchus Weber (Col. Scarab. Ateuchina). Boletin de Entomologia Venezolana (N. S.) 5 (20): 159 - 163." type="journal article" year="1990">Martínez &amp; Martínez 1990: 161</bibRefCitation>
(
<collectingCountry id="EDEE87E030495277EF2DFDB075B5030E" box="[570,697,535,561]" name="Venezuela" pageId="19" pageNumber="20">Venezuela</collectingCountry>
:
<collectingRegion id="573D099230495277EFD6FDB07416030E" box="[705,794,535,561]" country="Venezuela" name="Merida" pageId="19" pageNumber="20">Mérida</collectingRegion>
: Páramo de Mucuchíes, Laguna Grande). —
<bibRefCitation id="F168BA8130495277E809FDB0762E036B" author="Krajcik M." pageId="19" pageNumber="20" pagination="1 - 278" refId="ref126632" refString="Krajcik M. 2012. Checklist of the World Scarabaeoidea. Animma. X Supplement 5: 1 - 278." type="journal article" year="2012">Krajcik 2012: 49</bibRefCitation>
(
<collectingCountry id="EDEE87E030495277EC25FD9D76BD036B" box="[306,433,570,596]" name="Venezuela" pageId="19" pageNumber="20">Venezuela</collectingCountry>
:
<collectingRegion id="573D099230495277ECABFD9D751A036B" box="[444,534,570,596]" country="Venezuela" name="Merida" pageId="19" pageNumber="20">Mérida</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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