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<document id="6B0744B19FE99BA3406712792B3C6273" ID-DOI="10.5852/ejt.2024.959.2677" ID-ISSN="2118-9773" ID-ZooBank="30872C13-516F-42FA-AFA7-30ADC6BF1BAF" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1729166637250" checkinUser="plazi" docAuthor="Rosa, Cecilia Lozano De La, Cupello, Mario &amp; Vaz-De-Mello, Fernando Z." docDate="2024" docId="1D50766630365208EF06FEDF745A02BD" docLanguage="en" docName="EJT.2024.959.1-272.pdf" docOrigin="European Journal of Taxonomy 959 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2677/12401" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Dichotomius triquetrus" docType="treatment" docVersion="1" lastPageNumber="109" masterDocId="E1690E1E305A5264ED17FFA7770C013F" masterDocTitle="The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution" masterLastPageNumber="272" masterPageNumber="1" pageNumber="109" updateTime="1729168044082" updateUser="GgImagineBatch" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="6ACF37E9094BF152193885970F19D1AD">The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution</mods:title>
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<mods:namePart id="119EC849A291A9F2CB6656943336E981">Rosa, Cecilia Lozano De La</mods:namePart>
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<mods:affiliation id="E04B65812689783AE5AD34921CEA28EC">Instituto Venezolano de Investigaciones Científicas, Centro de Estudios Botánicos y Agroforestales, Laboratorio de Ecología Espacial, Maracaibo, estado Zulia, Venezuela.</mods:affiliation>
<mods:nameIdentifier id="5D1393731FC7959B64BAF48440445478" type="email">lozanoceci@gmail.com</mods:nameIdentifier>
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<mods:namePart id="AF1A7FC1B69EFA2D1262770C2AF89212">Cupello, Mario</mods:namePart>
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<mods:affiliation id="0E02C57C971C12CDB80BAC45EE12AD03">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Curitiba, Paraná, Brazil. Currently at Department of Entomology, Texas A &amp; M University, College Station, TX, 77843, USA.</mods:affiliation>
<mods:nameIdentifier id="C69EC13CFA83CD60A4E8F7B693647D95" type="email">mcupello@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="AA87E67D2BBAD4880BC130ACF1B09CA0">Vaz-De-Mello, Fernando Z.</mods:namePart>
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<mods:affiliation id="17912494CBCA4222159BA6C1B9283D65">Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, Cuiabá, Mato Grosso, Brazil.</mods:affiliation>
<mods:nameIdentifier id="1EF4D73A66438BCF61928AD65CFB2F4F" type="email">vazdemello@gmail.com</mods:nameIdentifier>
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<mods:date id="85D5171B7DA7A8DCA400539DC93C17F1">2024</mods:date>
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<paragraph id="9546C77030365208EF06FEDF732E00AC" blockId="108.[529,1058,376,403]" box="[529,1058,376,403]" pageId="108" pageNumber="109">
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<taxonomicName id="52F9BCF330365208EF06FEDF732E00AC" ID-CoL="35ND7" authority="(Luederwaldt, 1923)" baseAuthorityName="Luederwaldt" baseAuthorityYear="1923" box="[529,1058,376,403]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="triquetrus">
<emphasis id="A78D1B6230365208EF06FEDF742600AD" bold="true" box="[529,810,376,403]" italics="true" pageId="108" pageNumber="109">Dichotomius triquetrus</emphasis>
(
<bibRefCitation id="F168BA8130365208EE2EFEDF731600AC" author="Luederwaldt H." box="[825,1050,376,403]" pageId="108" pageNumber="109" refId="ref128666" refString="Luederwaldt H. 1923. Neue Pinotus Arten. Revista do Museu Paulista 14: [2] - 13." type="journal volume" year="1923">Luederwaldt, 1923</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB30365208EDAAFE1C751A039D" pageId="108" pageNumber="109" type="reference_group">
<paragraph id="9546C77030365208EDAAFE1C741C0325" blockId="108.[189,1399,442,539]" pageId="108" pageNumber="109">
<treatmentCitationGroup id="B5E9E05E30365208EDAAFE1C741C0325" pageId="108" pageNumber="109">
<treatmentCitation id="1458E16130365208EDAAFE1C75A200EB" author="Luederwaldt H." box="[189,686,442,468]" page="11" pageId="108" pageNumber="109" year="1923">
<taxonomicName id="52F9BCF330365208EDAAFE1C75A200EB" ID-CoL="4HZWV" authority="Luederwaldt, 1923: 11" authorityName="Luederwaldt" authorityPageNumber="11" authorityYear="1923" box="[189,686,442,468]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="triquetrus">
<emphasis id="A78D1B6230365208EDAAFE1C769900EB" box="[189,405,443,468]" italics="true" pageId="108" pageNumber="109">Pinotus triquetrus</emphasis>
<bibRefCitation id="F168BA8130365208EC88FE1D75A200EB" author="Luederwaldt H." box="[415,686,442,468]" pageId="108" pageNumber="109" refId="ref128666" refString="Luederwaldt H. 1923. Neue Pinotus Arten. Revista do Museu Paulista 14: [2] - 13." type="journal volume" year="1923">Luederwaldt, 1923: 11</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description). Type locality:
<collectingCountry id="EDEE87E030365208E963FE1D727F00EB" box="[1140,1395,442,468]" name="Trinidad and Tobago" pageId="108" pageNumber="109">Trinidad and Tobago</collectingCountry>
: Couva-Tabaquite-Talparo: Caparo. Name-bearing type:
<typeStatus id="4A4279D230365208EE6EFE7A74E900C8" box="[889,997,477,503]" pageId="108" pageNumber="109" type="lectotype">lectotype</typeStatus>
(MZSP), designated by
<bibRefCitation id="F168BA8130365208E814FE7975520325" author="Montoya-Molina S. &amp; Vaz-de-Mello F. Z." pageId="108" pageNumber="109" pagination="1 - 64" refId="ref131592" refString="Montoya-Molina S. &amp; Vaz-de-Mello F. Z. 2021. Taxonomic review of the Dichotomius (Luederwaldtinia) agenor species group (Coleoptera: Scarabaeidae: Scarabaeinae). European Journal of Taxonomy 734 (1): 1 - 64. https: // doi. org / 10.5852 / ejt. 2021.734.1233" type="journal article" year="2021">Montoya-Molina &amp; Vaz-de-Mello (2021)</bibRefCitation>
, not examined.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030365208EDAAFDE473240363" blockId="108.[189,1398,578,674]" box="[189,1064,578,604]" pageId="108" pageNumber="109">
<treatmentCitationGroup id="B5E9E05E30365208EDAAFDE473240363" box="[189,1064,578,604]" pageId="108" pageNumber="109">
<taxonomicName id="52F9BCF330365208EDAAFDE4769E0363" ID-CoL="4HZWV" authorityName="Luederwaldt" authorityYear="1923" box="[189,402,579,604]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="triquetrus">
<emphasis id="A78D1B6230365208EDAAFDE4769E0363" box="[189,402,579,604]" italics="true" pageId="108" pageNumber="109">Pinotus triquetrus</emphasis>
</taxonomicName>
<treatmentCitation id="1458E16130365208ECB8FDE575CD0363" author="Blackwelder R. E." box="[431,705,578,604]" page="208" pageId="108" pageNumber="109" year="1944">
<bibRefCitation id="F168BA8130365208ECB8FDE575CD0363" author="Blackwelder R. E." box="[431,705,578,604]" pageId="108" pageNumber="109" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 208</bibRefCitation>
</treatmentCitation>
(checklist, cited for
<collectingCountry id="EDEE87E030365208EEA5FDE573100363" box="[946,1052,578,604]" name="Trinidad and Tobago" pageId="108" pageNumber="109">Trinidad</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030365208EDAAFDC2751A039D" blockId="108.[189,1398,578,674]" pageId="108" pageNumber="109">
<treatmentCitationGroup id="B5E9E05E30365208EDAAFDC2751A039D" pageId="108" pageNumber="109">
<taxonomicName id="52F9BCF330365208EDAAFDC276DB0340" ID-CoL="35ND7" baseAuthorityName="Luederwaldt" baseAuthorityYear="1923" box="[189,471,613,639]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="triquetrus">
<emphasis id="A78D1B6230365208EDAAFDC276DB0340" box="[189,471,613,639]" italics="true" pageId="108" pageNumber="109">Dichotomius triquetrus</emphasis>
</taxonomicName>
<treatmentCitation id="1458E16130365208EF10FDC273100340" author="Montoya-Molina S. &amp; Vaz-de-Mello F. Z." box="[519,1052,613,639]" page="8" pageId="108" pageNumber="109" year="2021">
<bibRefCitation id="F168BA8130365208EF10FDC273100340" author="Montoya-Molina S. &amp; Vaz-de-Mello F. Z." box="[519,1052,613,639]" pageId="108" pageNumber="109" pagination="1 - 64" refId="ref131592" refString="Montoya-Molina S. &amp; Vaz-de-Mello F. Z. 2021. Taxonomic review of the Dichotomius (Luederwaldtinia) agenor species group (Coleoptera: Scarabaeidae: Scarabaeinae). European Journal of Taxonomy 734 (1): 1 - 64. https: // doi. org / 10.5852 / ejt. 2021.734.1233" type="journal article" year="2021">Montoya-Molina &amp; Vaz-de-Mello 2021: 8</bibRefCitation>
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, 4748 (key, redescription, designation of
<typeStatus id="4A4279D230365208EC8EFD2F7506039D" box="[409,522,648,674]" pageId="108" pageNumber="109" type="lectotype">lectotype</typeStatus>
).
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</paragraph>
</subSubSection>
<subSubSection id="DDE394FB30365208EDAAFD6E745A02BD" pageId="108" pageNumber="109" type="distribution">
<paragraph id="9546C77030365208EDAAFD6E765A03DC" blockId="108.[189,342,713,739]" box="[189,342,713,739]" pageId="108" pageNumber="109">
<heading id="CE0E701C30365208EDAAFD6E765A03DC" bold="true" box="[189,342,713,739]" fontSize="11" level="3" pageId="108" pageNumber="109" reason="3">
<emphasis id="A78D1B6230365208EDAAFD6E765A03DC" bold="true" box="[189,342,713,739]" pageId="108" pageNumber="109">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="9546C77030365208EDAAFD5F7566022D" blockId="108.[189,618,760,786]" box="[189,618,760,786]" pageId="108" pageNumber="109">
<collectingCountry id="EDEE87E030365208EDAAFD5F763A022D" box="[189,310,760,786]" name="Venezuela" pageId="108" pageNumber="109">Venezuela</collectingCountry>
and
<collectingCountry id="EDEE87E030365208EC79FD5F7569022D" box="[366,613,760,786]" name="Trinidad and Tobago" pageId="108" pageNumber="109">Trinidad and Tobago</collectingCountry>
.
</paragraph>
<paragraph id="9546C77030365208EDAAFC9E7695026B" blockId="108.[189,409,825,852]" box="[189,409,825,852]" pageId="108" pageNumber="109">
<heading id="CE0E701C30365208EDAAFC9E7695026B" bold="true" box="[189,409,825,852]" fontSize="11" level="3" pageId="108" pageNumber="109" reason="3">
<emphasis id="A78D1B6230365208EDAAFC9E7695026B" bold="true" box="[189,409,825,852]" pageId="108" pageNumber="109">Literature record</emphasis>
</heading>
</paragraph>
<paragraph id="9546C77030365208EDAAFCCF745A02BD" blockId="108.[189,854,872,898]" box="[189,854,872,898]" pageId="108" pageNumber="109">
<bibRefCitation id="F168BA8130365208EDAAFCCF75B602BD" author="Montoya-Molina S. &amp; Vaz-de-Mello F. Z." box="[189,698,872,898]" pageId="108" pageNumber="109" pagination="1 - 64" refId="ref131592" refString="Montoya-Molina S. &amp; Vaz-de-Mello F. Z. 2021. Taxonomic review of the Dichotomius (Luederwaldtinia) agenor species group (Coleoptera: Scarabaeidae: Scarabaeinae). European Journal of Taxonomy 734 (1): 1 - 64. https: // doi. org / 10.5852 / ejt. 2021.734.1233" type="journal article" year="2021">Montoya-Molina &amp; Vaz-de-Mello 2021: 47</bibRefCitation>
(
<collectingCountry id="EDEE87E030365208EFDDFCCF744602BD" box="[714,842,872,898]" name="Venezuela" pageId="108" pageNumber="109">Venezuela</collectingCountry>
).
</paragraph>
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</treatment>
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