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<document id="43D141BB27D656771DD4ABAB5DEF4236" ID-DOI="10.5852/ejt.2024.959.2677" ID-ISSN="2118-9773" ID-Zenodo-Dep="13945464" ID-ZooBank="30872C13-516F-42FA-AFA7-30ADC6BF1BAF" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1729166637250" checkinUser="plazi" docAuthor="Rosa, Cecilia Lozano De La, Cupello, Mario &amp; Vaz-De-Mello, Fernando Z." docDate="2024" docId="1D5076663032520DEF21F9D1731E070F" docLanguage="en" docName="EJT.2024.959.1-272.pdf" docOrigin="European Journal of Taxonomy 959 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2677/12401" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Dichotomius protectus" docType="treatment" docVersion="5" lastPageNumber="106" masterDocId="E1690E1E305A5264ED17FFA7770C013F" masterDocTitle="The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution" masterLastPageNumber="272" masterPageNumber="1" pageNumber="105" updateTime="1730324035732" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="8617B32742DDC00049E904C5D51B5476">The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution</mods:title>
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<mods:namePart id="46064E757A28DD4EA381585F6DFA4EA7">Rosa, Cecilia Lozano De La</mods:namePart>
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<mods:affiliation id="EB02B9F4C60E5570B117A72B3A540353">Instituto Venezolano de Investigaciones Científicas, Centro de Estudios Botánicos y Agroforestales, Laboratorio de Ecología Espacial, Maracaibo, estado Zulia, Venezuela.</mods:affiliation>
<mods:nameIdentifier id="4D767A591EC5EC9462B4B515B828F480" type="email">lozanoceci@gmail.com</mods:nameIdentifier>
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<mods:namePart id="5AFE6F72594A7A7798AC7A3D9B7728A5">Cupello, Mario</mods:namePart>
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<mods:affiliation id="AD134300EEB5B8B2E64A16FE94599549">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Curitiba, Paraná, Brazil. Currently at Department of Entomology, Texas A &amp; M University, College Station, TX, 77843, USA.</mods:affiliation>
<mods:nameIdentifier id="BF95BD12AB77B1B6D1F9BCE3B503E334" type="email">mcupello@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="FFDF9516A778A681ECD9A43DF7C3D441">Vaz-De-Mello, Fernando Z.</mods:namePart>
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<mods:affiliation id="F0DF11A9F0CA8A652D234588395CEB7F">Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, Cuiabá, Mato Grosso, Brazil.</mods:affiliation>
<mods:nameIdentifier id="379BBA7DB8A75E62D16CB4022139793F" type="email">vazdemello@gmail.com</mods:nameIdentifier>
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<subSubSection id="DDE394FB3032520CEF21F9D174F107AF" box="[566,1021,1654,1681]" pageId="104" pageNumber="105" type="nomenclature">
<paragraph id="9546C7703032520CEF21F9D174F107AF" blockId="104.[566,1021,1654,1716]" box="[566,1021,1654,1681]" pageId="104" pageNumber="105">
<heading id="CE0E701C3032520CEF21F9D174F107AF" box="[566,1021,1654,1681]" centered="true" fontSize="11" level="2" pageId="104" pageNumber="105" reason="2">
<taxonomicName id="52F9BCF33032520CEF21F9D174F107AF" ID-CoL="7ZN4Y" authority="(Harold, 1867)" baseAuthorityName="Harold" baseAuthorityYear="1867" box="[566,1021,1654,1681]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="protectus">
<emphasis id="A78D1B623032520CEF21F9D1744407AE" bold="true" box="[566,840,1654,1681]" italics="true" pageId="104" pageNumber="105">Dichotomius protectus</emphasis>
(Harold, 1867)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB3032520CEFF1F93E7443078B" box="[742,847,1689,1716]" pageId="104" pageNumber="105" type="description">
<paragraph id="9546C7703032520CEFF1F93E7443078B" blockId="104.[566,1021,1654,1716]" box="[742,847,1689,1716]" pageId="104" pageNumber="105">
<figureCitation id="0DC2DBF53032520CEFF1F93E7443078B" box="[742,847,1689,1716]" captionStart="Fig" captionStartId="106.[189,232,1963,1989]" captionTargetBox="[277,1301,278,1881]" captionTargetId="figure-17@106.[265,1322,265,1920]" captionTargetPageId="106" captionText="Fig. 42. Species distribution. A. Dichotomius protectus (Harold, 1867). B. Dichotomius robustus (Luederwaldt, 1935). Green diamond = CEMT collection data; black circle = literature data." figureDoi="http://doi.org/10.5281/zenodo.13945581" httpUri="https://zenodo.org/record/13945581/files/figure.png" pageId="104" pageNumber="105">Fig. 42A</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB3032520DEDAAF97B727B03BC" lastPageId="105" lastPageNumber="106" pageId="104" pageNumber="105" type="reference_group">
<paragraph id="9546C7703032520CEDAAF97B74020626" blockId="104.[189,1399,1755,1887]" pageId="104" pageNumber="105">
<treatmentCitationGroup id="B5E9E05E3032520CEDAAF97B74020626" pageId="104" pageNumber="105">
<treatmentCitation id="1458E1613032520CEDAAF97B757607CA" author="Harold E." box="[189,634,1755,1781]" page="98" pageId="104" pageNumber="105" year="1867">
<taxonomicName id="52F9BCF33032520CEDAAF97B757607CA" ID-CoL="7Y825" authority="Harold, 1867 d: 98" authorityName="Harold" authorityPageNumber="98" authorityYear="1867" box="[189,634,1755,1781]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="protectus">
<emphasis id="A78D1B623032520CEDAAF97B768307CA" box="[189,399,1756,1781]" italics="true" pageId="104" pageNumber="105">Pinotus protectus</emphasis>
<bibRefCitation id="F168BA813032520CEC8CF97C757607CA" author="Harold E." box="[411,634,1755,1781]" pageId="104" pageNumber="105" pagination="94 - 100" refId="ref123272" refString="Harold E. 1867 d. Diagnosen neuer Coprophagen. Coleopterologische Hefte 2: 94 - 100. Available from https: // www. biodiversitylibrary. org / page / 16069437 [accessed 16 Feb. 2024]." type="journal article" year="1867">Harold, 1867d: 98</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description). Type locality:
<collectingCountry id="EDEE87E03032520CE95FF97C73CC07CA" box="[1096,1216,1755,1781]" name="Colombia" pageId="104" pageNumber="105">Colombia</collectingCountry>
. Name-bearing type:
<typeStatus id="4A4279D23032520CEC3BF958769F0626" box="[300,403,1791,1817]" pageId="104" pageNumber="105" type="syntype">syntypes</typeStatus>
(MNHN), examined by FZVM.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C7703032520CEDAAF88574EB0661" blockId="104.[189,1399,1755,1887]" pageId="104" pageNumber="105">
<treatmentCitationGroup id="B5E9E05E3032520CEDAAF88574EB0661" pageId="104" pageNumber="105">
<treatmentCitation id="1458E1613032520CEDAAF885742A0604" author="Luederwaldt H." box="[189,806,1825,1851]" page="10" pageId="104" pageNumber="105" year="1923">
<taxonomicName id="52F9BCF33032520CEDAAF885742A0604" ID-CoL="7Z9H5" authority="Luederwaldt, 1923: 10" authorityName="Luederwaldt" authorityPageNumber="10" authorityYear="1923" box="[189,806,1825,1851]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="subSpecies" species="protectus" subSpecies="tridentatus">
<emphasis id="A78D1B623032520CEDAAF885751E0604" box="[189,530,1825,1851]" italics="true" pageId="104" pageNumber="105">Pinotus protectus tridentatus</emphasis>
<bibRefCitation id="F168BA813032520CEF0CF886742A0604" author="Luederwaldt H." box="[539,806,1825,1851]" pageId="104" pageNumber="105" refId="ref128666" refString="Luederwaldt H. 1923. Neue Pinotus Arten. Revista do Museu Paulista 14: [2] - 13." type="journal volume" year="1923">Luederwaldt, 1923: 10</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description). Type locality:
<collectingCountry id="EDEE87E03032520CE9F5F88672480604" box="[1250,1348,1825,1851]" name="Ecuador" pageId="104" pageNumber="105">Ecuador</collectingCountry>
and
<collectingCountry id="EDEE87E03032520CEDFCF8E376640661" box="[235,360,1860,1886]" name="Venezuela" pageId="104" pageNumber="105">Venezuela</collectingCountry>
. Name-bearing type:
<typeStatus id="4A4279D23032520CEF73F8E275C70660" box="[612,715,1861,1887]" pageId="104" pageNumber="105" type="syntype">syntypes</typeStatus>
(MZSP), not examined.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C7703032520DEDAAF82072290079" blockId="104.[189,1398,1926,1988]" lastBlockId="105.[189,1400,265,644]" lastPageId="105" lastPageNumber="106" pageId="104" pageNumber="105">
<treatmentCitationGroup id="B5E9E05E3032520DEDAAF82072290079" lastPageId="105" lastPageNumber="106" pageId="104" pageNumber="105">
<taxonomicName id="52F9BCF33032520CEDAAF8207681069F" ID-CoL="7Y825" authorityName="Harold" authorityYear="1867" box="[189,397,1927,1952]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="protectus">
<emphasis id="A78D1B623032520CEDAAF8207681069F" box="[189,397,1927,1952]" italics="true" pageId="104" pageNumber="105">Pinotus protectus</emphasis>
</taxonomicName>
<treatmentCitation id="1458E1613032520CECA6F8217599069F" author="Harold E." box="[433,661,1926,1952]" page="130" pageId="104" pageNumber="105" year="1869">
<bibRefCitation id="F168BA813032520CECA6F8217599069F" author="Harold E." box="[433,661,1926,1952]" pageId="104" pageNumber="105" pagination="123 - 144" refId="ref123559" refString="Harold E. 1869 b. Revision des especes qui rentrent dans le genre Pinotus Erichs. L'Abeille 6: 123 - 144." type="journal article" year="1869">Harold 1869b: 130</bibRefCitation>
</treatmentCitation>
(redescription, cited for
<collectingCountry id="EDEE87E03032520CEED3F8217331069F" box="[964,1085,1926,1952]" name="Colombia" pageId="104" pageNumber="105">Colombia</collectingCountry>
); 1869d: 1010 (catalogue, cited for
<collectingCountry id="EDEE87E03032520CEC42F80E76C206FC" box="[341,462,1961,1987]" name="Colombia" pageId="104" pageNumber="105">Colombia</collectingCountry>
). —
<treatmentCitation id="1458E1613032520CEF11F80E75E506FC" author="Candeze E." box="[518,745,1961,1987]" page="330" pageId="104" pageNumber="105" year="1891">
<bibRefCitation id="F168BA813032520CEF11F80E75E506FC" author="Candeze E." box="[518,745,1961,1987]" pageId="104" pageNumber="105" refId="ref113476" refString="Candeze E. 1891. Voyage de M. E. Simon au Venezuela (Decembre 1887 - Avril 1888), 15 e Memoire (1). Coleopteres - Lamellicornes determines par le Dr E. Candeze. Annales de la Societe entomologique de France 60: [329] - 332. Available from https: // www. biodiversitylibrary. org / page / 32547689 [accessed 14 Feb. 2024]." type="journal volume" year="1891">Candèze 1891: 330</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="1458E1613032520CEE4CF80D73C306FC" author="Heyne A. &amp; Taschenberg O." box="[859,1231,1961,1988]" page="62" pageId="104" pageNumber="105" year="1908">
<bibRefCitation id="F168BA813032520CEE4CF80D73C306FC" author="Heyne A. &amp; Taschenberg O." box="[859,1231,1961,1988]" pageId="104" pageNumber="105" refId="ref124102" refString="Heyne A. &amp; Taschenberg O. 1908. Die exotischen Kafer in Wort und Bild. J. F. Schreiber, Esslingen and Munich. https: // doi. org / 10.5962 / bhl. title. 9363" type="book" year="1908">Heyne &amp; Taschenberg 1908: 62</bibRefCitation>
</treatmentCitation>
(list, cited for
<collectingCountry id="EDEE87E03033520DEDFDFEAE766F001C" box="[234,355,265,291]" name="Colombia" pageId="105" pageNumber="106">Colombia</collectingCountry>
).
<emphasis id="A78D1B623033520DEC63FEAD7682001C" box="[372,398,266,291]" italics="true" pageId="105" pageNumber="106"></emphasis>
<treatmentCitation id="1458E1613033520DEC83FEAE7543001C" author="Gillet J. J. E." box="[404,591,265,291]" page="61" pageId="105" pageNumber="106" year="1911">
<bibRefCitation id="F168BA813033520DEC83FEAE7543001C" author="Gillet J. J. E." box="[404,591,265,291]" pageId="105" pageNumber="106" refId="ref121235" refString="Gillet J. J. E. 1911 b. Coleopterorum Catalogus. Pars 38: Scarabaeidae: Coprinae I. W. Junk, Berlin. Available from https: // www. biodiversitylibrary. org / page / 48405679 [accesseed 16 Feb. 2024]." type="url" year="1911">Gillet 1911b: 61</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="1458E1613033520DEE1FFEAE74DE001B" author="Campos F." box="[776,978,265,292]" page="56" pageId="105" pageNumber="106" year="1921">
<bibRefCitation id="F168BA813033520DEE1FFEAE74DE001B" author="Campos F." box="[776,978,265,292]" pageId="105" pageNumber="106" pagination="26 - 100" refId="ref113384" refString="Campos F. 1921. Estudios sobre la fauna entomologica del Ecuador. Revista del Colegio Nacional Vicente Rocafuerte 6: 26 - 100." type="journal article" year="1921">Campos 1921: 56</bibRefCitation>
</treatmentCitation>
(cited for
<collectingCountry id="EDEE87E03033520DE950FEAE73A2001C" box="[1095,1198,265,291]" name="Ecuador" pageId="105" pageNumber="106">Ecuador</collectingCountry>
). —
<treatmentCitation id="1458E1613033520DE9F5FEAE76430079" author="Luederwaldt H." page="34" pageId="105" pageNumber="106" year="1929">
<bibRefCitation id="F168BA813033520DE9F5FEAE76430079" author="Luederwaldt H." pageId="105" pageNumber="106" pagination="603 - 775" refId="ref128687" refString="Luederwaldt H. 1929. As especies brasileiras do genero Pinotus, com algumas consideracoes tambem sobre outras especies. Revista do Museu Paulista 16: 603 - 775." type="journal article" year="1929">Luederwaldt 1929: 34</bibRefCitation>
</treatmentCitation>
(key). —
<treatmentCitation id="1458E1613033520DECDFFE8B75D60079" author="Blackwelder R. E." box="[456,730,300,326]" page="207" pageId="105" pageNumber="106" year="1944">
<bibRefCitation id="F168BA813033520DECDFFE8B75D60079" author="Blackwelder R. E." box="[456,730,300,326]" pageId="105" pageNumber="106" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 207</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="1458E1613033520DEE5BFE8B73360079" author="Gacharna C." box="[844,1082,300,327]" page="222" pageId="105" pageNumber="106" year="1951">
<bibRefCitation id="F168BA813033520DEE5BFE8B73360079" author="Gacharna C." box="[844,1082,300,327]" pageId="105" pageNumber="106" pagination="220 - 229" refId="ref119803" refString="Gacharna C. 1951. Catalogo de los coleopteros colombianos. Revista de la Academia Colombiana de Ciencias Fisicas y Naturales 8: 220 - 229." type="journal article" year="1951">Gacharná 1951: 222</bibRefCitation>
</treatmentCitation>
(list for
<collectingCountry id="EDEE87E03033520DE9B7FE8B72150079" box="[1184,1305,300,326]" name="Colombia" pageId="105" pageNumber="106">Colombia</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C7703033520DEDAAFEE8727B03BC" blockId="105.[189,1400,265,644]" pageId="105" pageNumber="106">
<treatmentCitationGroup id="B5E9E05E3033520DEDAAFEE8727B03BC" pageId="105" pageNumber="106">
<taxonomicName id="52F9BCF33033520DEDAAFEE875790056" ID-CoL="7ZN4Y" baseAuthorityName="Harold" baseAuthorityYear="1867" box="[189,629,335,362]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="105" pageNumber="106" phylum="Arthropoda" rank="species" species="protectus" subGenus="Dichotomius">
<emphasis id="A78D1B623033520DEDAAFEE8765F0056" box="[189,339,335,361]" italics="true" pageId="105" pageNumber="106">Dichotomius</emphasis>
(
<emphasis id="A78D1B623033520DEC74FEE876F50056" box="[355,505,335,361]" italics="true" pageId="105" pageNumber="106">Dichotomius</emphasis>
)
<emphasis id="A78D1B623033520DEF1EFEF775790056" box="[521,629,336,361]" italics="true" pageId="105" pageNumber="106">protectus</emphasis>
</taxonomicName>
<treatmentCitation id="1458E1613033520DEF87FEF774350055" author="Roze J. A." box="[656,825,335,362]" page="44" pageId="105" pageNumber="106" year="1955">
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(cited, checklist for
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<treatmentCitation id="1458E1613033520DE9CDFEE8760500B2" author="Blanco J." page="40" pageId="105" pageNumber="106" year="1988">
<bibRefCitation id="F168BA813033520DE9CDFEE8760500B2" author="Blanco J." pageId="105" pageNumber="106" pagination="39 - 48" refId="ref111584" refString="Blanco J. 1988. Catalogo de los Scarabaeinae (Coleoptera) coprofagos y necrofagos, del Estado Tachira. Venezuela. Revista Cientifica Unet 2 (1): 39 - 48." type="journal article" year="1988">Blanco 1988: 40</bibRefCitation>
</treatmentCitation>
, 41 (catalogue). —
<treatmentCitation id="1458E1613033520DECE5FED575D900B3" author="Havranek D." box="[498,725,370,397]" page="61" pageId="105" pageNumber="106" year="1989">
<bibRefCitation id="F168BA813033520DECE5FED575D900B3" author="Havranek D." box="[498,725,370,397]" pageId="105" pageNumber="106" pagination="55 - 64" refId="ref123886" refString="Havranek D. 1989. Coleoptera coprofagos y necrofagos (Scarabaeidae, Silphidae) atraidos a cebos en bosques venezolanos. UNET 3: 55 - 64." type="journal article" year="1989">Havranek 1989: 61</bibRefCitation>
</treatmentCitation>
, 62 (list, comments). —
<treatmentCitation id="1458E1613033520DEEEFFED573D800B2" author="Escobar F." box="[1016,1236,370,397]" page="208" pageId="105" pageNumber="106" year="2000">
<bibRefCitation id="F168BA813033520DEEEFFED573D800B2" author="Escobar F." box="[1016,1236,370,397]" pageId="105" pageNumber="106" pagination="197 - 210" refId="ref118039" refString="Escobar F. 2000. Diversidad y distribution de los escarabajos del estiercol (Coleoptera: Scarabaeidae: Scarabaeinae) de Colombia. In: Martin-Piera F., Morrone J. J. &amp; Melic A. (eds.) Hacia un proyecto CYTED para el Inventario y Estimacion de la Diversidad Entomologica en Iberoamerica: PRIBES- 2000: 197 - 210. m 3 m: Monografias Tercer Milenio, vol. 1. Sociedad Entomologica Aragonesa (SEA), Zaragoza." type="book chapter" year="2000">Escobar 2000: 208</bibRefCitation>
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(checklist for
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). —
<treatmentCitation id="1458E1613033520DEC8AFE3175B5008F" author="Medina C. A. &amp; Lopera-Toro A. &amp; Vitolo A. &amp; Gill B." box="[413,697,406,432]" page="138" pageId="105" pageNumber="106" year="2001">
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Medina
<emphasis id="A78D1B623033520DEF17FE307536008F" box="[512,570,406,432]" italics="true" pageId="105" pageNumber="106">et al.</emphasis>
2001: 138
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</treatmentCitation>
(checklist for
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). —
<treatmentCitation id="1458E1613033520DE90EFE31727A008E" author="Hamel-Leigue A. &amp; Mann D. J. &amp; Vaz-de-Mello F. Z. &amp; Herzog S. K." box="[1049,1398,406,433]" page="15" pageId="105" pageNumber="106" year="2006">
<bibRefCitation id="F168BA813033520DE90EFE31727A008E" author="Hamel-Leigue A. &amp; Mann D. J. &amp; Vaz-de-Mello F. Z. &amp; Herzog S. K." box="[1049,1398,406,433]" pageId="105" pageNumber="106" pagination="1 - 18" refId="ref122830" refString="Hamel-Leigue A., Mann D. J., Vaz-de-Mello F. Z. &amp; Herzog S. K. 2006. Hacia un inventario de los escarabajos peloteros (Coleoptera: Scarabaeinae) de Bolivia: primera compilacion de los generos y especies registrados para el pais. Revista Boliviana de Ecologia y Conservacion Ambiental 20: 1 - 18." type="journal article" year="2006">
Hamel-Leigue
<emphasis id="A78D1B623033520DE9DCFE30720A008F" box="[1227,1286,406,432]" italics="true" pageId="105" pageNumber="106">et al.</emphasis>
2006: 15
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(list, cited for
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). —
<treatmentCitation id="1458E1613033520DEF09FE1E748400EC" author="Pulido-Herrera L. A. &amp; Zunino M." box="[542,904,441,467]" page="307" pageId="105" pageNumber="106" year="2007">
<bibRefCitation id="F168BA813033520DEF09FE1E748400EC" author="Pulido-Herrera L. A. &amp; Zunino M." box="[542,904,441,467]" firstAuthor="Pulido-Herrera" pageId="105" pageNumber="106" refId="ref136342" refString="Pulido-Herrera L. A. &amp; Zunino M. 2007. Catalogo preliminar de los Onthophagini de America (Coleoptera: Scarabaeine [sic]). In: Zunino M. &amp; Melic A. (eds) Escarabajos, Diversidad y Conservacion Biologica. Ensayos en Homenaje a Gonzalo Halffter: 93 - 129. Monografias 3 ercer Milenio m 3 m Vol. 7, Sociedad Entomologica Aragonesa, Zaragoza." type="book" year="2007">
Pulido-Herrera
<emphasis id="A78D1B623033520DEFC5FE1D740100EC" box="[722,781,441,467]" italics="true" pageId="105" pageNumber="106">et al.</emphasis>
2007: 307
</bibRefCitation>
</treatmentCitation>
(cited for Andean region of
<collectingCountry id="EDEE87E03033520DE9D9FE1E724B00EC" box="[1230,1351,441,467]" name="Colombia" pageId="105" pageNumber="106">Colombia</collectingCountry>
). —
<treatmentCitation id="1458E1613033520DEDFEFE7B754F00C9" author="Orozco J. &amp; Perez M." box="[233,579,476,502]" page="38" pageId="105" pageNumber="106" year="2008">
<bibRefCitation id="F168BA813033520DEDFEFE7B754F00C9" author="Orozco J. &amp; Perez M." box="[233,579,476,502]" firstAuthor="Perez" journalOrPublisher="Revista Brasileira de Entomologia" pageId="105" pageNumber="106" pagination="36 - 40" part="52" publicationUrl="https://doi.org/10.1590/S0085-56262008000100007" refId="ref134201" refString="Orozco J. &amp; Perez M. 2008. Escarabajos coprofagos (Coleoptera, Scarabaeoidea) del Parque Nacional Los Estoraques (Norte de Santander, Colombia). Revista Brasileira de Entomologia 52: 36 - 40. https: // doi. org / 10.1590 / S 0085 - 56262008000100007" title="Escarabajos coprofagos (Coleoptera, Scarabaeoidea) del Parque Nacional Los Estoraques (Norte de Santander, Colombia)" type="journal article" year="2008">Orozco &amp; Pérez 2008: 3839</bibRefCitation>
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(list for
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). —
<treatmentCitation id="1458E1613033520DEE4AFE7B727D00C9" author="Sarmiento-Garces R. &amp; Amat-Garcia G." box="[861,1393,476,502]" page="290" pageId="105" pageNumber="106" year="2009">
<bibRefCitation id="F168BA813033520DEE4AFE7B727D00C9" author="Sarmiento-Garces R. &amp; Amat-Garcia G." box="[861,1393,476,502]" pageId="105" pageNumber="106" pagination="285 - 296" refId="ref138101" refString="Sarmiento-Garces R. &amp; Amat-Garcia G. 2009. Escarabajos del genero Dichotomius Hope, 1838 (Scarabaeidae: Scarabaeinae) en la amazonia colombiana. Revista de la Academia Colombiana de Ciencias Exactas, Fisicas y Naturales 33 (127): 285 - 296." type="journal article" year="2009">Sarmiento-Garcés &amp; Amat-García 2009: 290</bibRefCitation>
</treatmentCitation>
, 295 (diagnosis, key);
<treatmentCitation id="1458E1613033520DECF2FDA775790326" author="Sarmiento-Garces R. &amp; Amat-Garcia G." box="[485,629,511,538]" page="64" pageId="105" pageNumber="106" year="2014">
<bibRefCitation id="F168BA813033520DECF2FDA775790326" author="Sarmiento-Garces R. &amp; Amat-Garcia G." box="[485,629,511,538]" pageId="105" pageNumber="106" refId="ref138148" refString="Sarmiento-Garces R. &amp; Amat-Garcia G. 2014. Escarabajos del Genero Dichotomius Hope 1838 (Scarabaeidae: Scarabaeinae) en Colombia. Fauna de Colombia, Monografia n ° 4, Universidad Nacional de Colombia, Bogota. https: // doi. org / 10.18257 / raccefyn. 33 (127). 2009.2364" type="book" year="2014">2014: 51, 64</bibRefCitation>
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(key, diagnosis). —
<treatmentCitation id="1458E1613033520DEE73FE5873730325" author="Carvajal V. &amp; Villamarin S. &amp; Ortega A. M." box="[868,1151,511,538]" page="320" pageId="105" pageNumber="106" year="2011">
<bibRefCitation id="F168BA813033520DEE73FE5873730325" author="Carvajal V. &amp; Villamarin S. &amp; Ortega A. M." box="[868,1151,511,538]" pageId="105" pageNumber="106" refId="ref113831" refString="Carvajal V., Villamarin S. &amp; Ortega A. M. 2011. Escarabajos del Ecuador. Principales Generos. Instituto de Ciencias Biologicas, Escuela Politecnica Nacional, Quito." type="book" year="2011">
Carvajal
<emphasis id="A78D1B623033520DEEDCFDA773080326" box="[971,1028,511,537]" italics="true" pageId="105" pageNumber="106">et al.</emphasis>
2011: 320
</bibRefCitation>
</treatmentCitation>
(list for
<collectingCountry id="EDEE87E03033520DE9F7FE58724B0326" box="[1248,1351,511,537]" name="Ecuador" pageId="105" pageNumber="106">Ecuador</collectingCountry>
). —
<treatmentCitation id="1458E1613033520DEDFEFD8576A20302" author="Krajcik M." box="[233,430,546,573]" page="92" pageId="105" pageNumber="106" year="2012">
<bibRefCitation id="F168BA813033520DEDFEFD8576A20302" author="Krajcik M." box="[233,430,546,573]" pageId="105" pageNumber="106" pagination="1 - 278" refId="ref126632" refString="Krajcik M. 2012. Checklist of the World Scarabaeoidea. Animma. X Supplement 5: 1 - 278." type="journal article" year="2012">Krajcik 2012: 92</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="1458E1613033520DEF36FD8574660302" author="Ferrer-Paris J. &amp; Sanchez-Mercado A. &amp; Rodriguez J. P." box="[545,874,546,573]" page="109" pageId="105" pageNumber="106" year="2013">
<bibRefCitation id="F168BA813033520DEF36FD8574660302" author="Ferrer-Paris J. &amp; Sanchez-Mercado A. &amp; Rodriguez J. P." box="[545,874,546,573]" pageId="105" pageNumber="106" pagination="89 - 110" refId="ref119203" refString="Ferrer-Paris J. R, Sanchez-Mercado A. &amp; Rodriguez J. P. 2013. Optimizacion del muestreo de invertebrados tropicales: un ejemplo con escarabajos coprofagos (Coleoptera: Scarabaeinae) en Venezuela. Revista de Biologia Tropical 61 (1): 89 - 110. https: // doi. org / 10.15517 / rbt. v 61 i 1.10941" type="journal article" year="2013">
Ferrer-Paris
<emphasis id="A78D1B623033520DEFA5FD8475E10303" box="[690,749,546,572]" italics="true" pageId="105" pageNumber="106">et al.</emphasis>
2013: 109
</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="1458E1613033520DEECDFD8573F20302" author="Ratcliffe B. C. &amp; Jameson M. L. &amp; Figueroa L. &amp; Cave R. D. &amp; Paulsen M. J. &amp; Cano E. B. &amp; Beza-Beza C. &amp; Ferbans L. &amp; Reyes-Castillo P." box="[986,1278,546,573]" page="197" pageId="105" pageNumber="106" year="2015">
<bibRefCitation id="F168BA813033520DEECDFD8573F20302" author="Ratcliffe B. C. &amp; Jameson M. L. &amp; Figueroa L. &amp; Cave R. D. &amp; Paulsen M. J. &amp; Cano E. B. &amp; Beza-Beza C. &amp; Ferbans L. &amp; Reyes-Castillo P." box="[986,1278,546,573]" pageId="105" pageNumber="106" pagination="186 - 207" refId="ref137018" refString="Ratcliffe B. C., Jameson M. L., Figueroa L., Cave R. D., Paulsen M. J., Cano E. B., Beza-Beza C., Jimenez- Ferbans L., Reyes-Castillo P. 2015. Beetles (Coleoptera) of Peru: a survey of the families. Journal of the Kansas Entomological Society 88 (2): 186 - 207." type="journal article" year="2015">
Ratcliffe
<emphasis id="A78D1B623033520DE951FD84738D0303" box="[1094,1153,546,572]" italics="true" pageId="105" pageNumber="106">et al.</emphasis>
2015: 197
</bibRefCitation>
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(checklist for
<collectingCountry id="EDEE87E03033520DEC02FDE1765C035F" box="[277,336,582,608]" name="Peru" pageId="105" pageNumber="106">Peru</collectingCountry>
). —
<treatmentCitation id="1458E1613033520DEC9DFDE175B5035F" author="Chamorro W. &amp; Marin-Armijos D. &amp; Granda V. &amp; Vaz-de-Mello F. Z." box="[394,697,582,608]" page="94" pageId="105" pageNumber="106" year="2018">
<bibRefCitation id="F168BA813033520DEC9DFDE175B5035F" author="Chamorro W. &amp; Marin-Armijos D. &amp; Granda V. &amp; Vaz-de-Mello F. Z." box="[394,697,582,608]" pageId="105" pageNumber="106" pagination="72 - 100" refId="ref114035" refString="Chamorro W., Marin-Armijos D., Granda V. &amp; Vaz-de-Mello F. Z. 2018. Listado de especies y clave de generos y subgeneros de escarabajos estercoleros (Coleoptera: Scarabaeidae: Scarabaeinae) presentes y presuntos para Ecuador. Revista Colombiana de Entomologia 44 (1): 72 - 100. https: // doi. org / 10.25100 / socolen. v 44 i 1.6545" type="journal article" year="2018">
Chamorro
<emphasis id="A78D1B623033520DEF1BFDE07544035F" box="[524,584,582,608]" italics="true" pageId="105" pageNumber="106">et al.</emphasis>
2018: 94
</bibRefCitation>
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(list for
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);
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<bibRefCitation id="F168BA813033520DEEB7FDE1731B035F" author="Chamorro W. &amp; Marin-Armijos D. &amp; Asenjo A. &amp; Vaz-De-Mello F. Z." box="[928,1047,582,608]" pageId="105" pageNumber="106" pagination="1 - 343" refId="ref114107" refString="Chamorro W., Marin-Armijos D., Asenjo A. &amp; Vaz-De-Mello F. Z. 2019. Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records. ZooKeys 826: 1 - 343. https: // doi. org / 10.3897 / zookeys. 826.26488" type="journal article" year="2019">2019: 119</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="1458E1613033520DE9CFFDE1765303BC" author="Giraldo C. &amp; Montoya S. &amp; Escobar F." page="100" pageId="105" pageNumber="106" year="2018">
<bibRefCitation id="F168BA813033520DE9CFFDE1765303BC" author="Giraldo C. &amp; Montoya S. &amp; Escobar F." pageId="105" pageNumber="106" refId="ref121556" refString="Giraldo C., Montoya S. &amp; Escobar F. 2018. Escarabajos del Estiercol en Paisajes Ganaderos de Colombia. Fundacion CIPAV, Cali, Colombia." type="book" year="2018">
Giraldo
<emphasis id="A78D1B623033520DE82BFDE07274035F" box="[1340,1400,582,608]" italics="true" pageId="105" pageNumber="106">et al.</emphasis>
2018: 100
</bibRefCitation>
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(guide). —
<treatmentCitation id="1458E1613033520DECFDFDCE740203BB" author="Pardo-Diaz C. &amp; Lopera T. A. &amp; Pena S. A. &amp; Sarmiento-Garces R. &amp; Sanchez-Herrera M. &amp; Salazar C." box="[490,782,617,644]" page="5" pageId="105" pageNumber="106" year="2019">
<bibRefCitation id="F168BA813033520DECFDFDCE740203BB" author="Pardo-Diaz C. &amp; Lopera T. A. &amp; Pena S. A. &amp; Sarmiento-Garces R. &amp; Sanchez-Herrera M. &amp; Salazar C." box="[490,782,617,644]" pageId="105" pageNumber="106" refId="ref134659" refString="Pardo-Diaz C., Lopera T. A., Pena S. A., Sarmiento-Garces R., Sanchez-Herrera M. &amp; Salazar C. 2019. Taxonomic reassessment of the genus Dichotomius (Coleoptera: Scarabaeinae) through integrative taxonomy. PeerJ 7: e 7332. https: // doi. org / 10.7717 / peerj. 7332" type="journal volume" year="2019">
Pardo-Díaz
<emphasis id="A78D1B623033520DEF63FDCD75A303BC" box="[628,687,617,643]" italics="true" pageId="105" pageNumber="106">et al.</emphasis>
2019: 5
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, 7, 11 (comments). —
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Nieto
<emphasis id="A78D1B623033520DE94DFDCD739903BC" box="[1114,1173,617,643]" italics="true" pageId="105" pageNumber="106">et al.</emphasis>
2020: 136
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(report).
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<emphasis id="A78D1B623033520DEDAAFD0D76A603FB" bold="true" box="[189,426,682,708]" pageId="105" pageNumber="106">Material examined</emphasis>
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<collectingRegion id="573D09923033520DEC90FD7E76EA03CC" box="[391,486,729,755]" country="Venezuela" name="Aragua" pageId="105" pageNumber="106">Aragua</collectingRegion>
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a.s.l.
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;
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;
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<collectionCode id="F3E85FB53033520DE9BAFD7E73F203CC" box="[1197,1278,729,755]" pageId="105" pageNumber="106">CEMT</collectionCode>
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<materialsCitation id="2591CD2D3033520DE80FFD7E75D90206" collectedFrom="forest" collectingDate="1971-02-21" collectingDateMax="1971-02-25" collectingDateMin="1971-02-21" collectingMethod="human dung trap" collectionCode="CEMT" collectorName="S. Peck" country="Venezuela" elevation="1500" location="Estacion Biologica Rancho Grande" municipality="Parque Nacional Henri Pittier" pageId="105" pageNumber="106" specimenCount="2" stateProvince="Aragua">
<specimenCount id="83FF0CF93033520DE80FFD7E727E03CC" box="[1304,1394,729,755]" pageId="105" pageNumber="106">2 specs</specimenCount>
;
<collectingMunicipality id="75225D0A3033520DEDAAFD5A752A0229" box="[189,550,764,791]" pageId="105" pageNumber="106">Parque Nacional Henri Pittier</collectingMunicipality>
,
<location id="902691AB3033520DEF22FD5B74EC0229" LSID="urn:lsid:plazi:treatment:1D5076663032520DEF21F9D1731E070F:902691AB3033520DEF22FD5B74EC0229" box="[565,992,764,790]" country="Venezuela" municipality="Parque Nacional Henri Pittier" name="Estacion Biologica Rancho Grande" pageId="105" pageNumber="106" stateProvince="Aragua">Estación Biológica Rancho Grande</location>
;
<elevation id="1ED420433033520DEEE7FD5B739C0229" box="[1008,1168,764,791]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="105" pageNumber="106" unit="m" value="1500.0">
<quantity id="52016A953033520DEEE7FD5B73400228" box="[1008,1100,764,791]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="105" pageNumber="106" unit="m" value="1500.0">1500 m</quantity>
a.s.l.
</elevation>
;
<date id="E147E1B03033520DE9B6FD5B727E0229" box="[1185,1394,764,791]" pageId="105" pageNumber="106" value="1971-02-21" valueMax="1971-02-25" valueMin="1971-02-21">
<collectingDate id="F10318583033520DE9B6FD5B727E0229" box="[1185,1394,764,791]" pageId="105" pageNumber="106" value="1971-02-21" valueMax="1971-02-25" valueMin="1971-02-21">2125 Feb. 1971</collectingDate>
</date>
;
<collectorName id="380CA2A63033520DEDAAFCB8761A0206" box="[189,278,799,825]" pageId="105" pageNumber="106">S. Peck</collectorName>
leg.;
<collectingMethod id="4CB8BF673033520DEC4EFCB875280205" box="[345,548,799,826]" pageId="105" pageNumber="106">human dung trap</collectingMethod>
;
<collectedFrom id="26E5848D3033520DEF26FCB875740206" box="[561,632,799,825]" pageId="105" pageNumber="106">forest</collectedFrom>
;
<collectionCode id="F3E85FB53033520DEF93FCB875D90206" box="[644,725,799,825]" pageId="105" pageNumber="106">CEMT</collectionCode>
</materialsCitation>
<materialsCitation id="2591CD2D3033520DEFF8FCB875A20262" collectedFrom="forest" collectingDate="1971-02-22" collectingDateMax="1971-02-25" collectingDateMin="1971-02-22" collectingMethod="carrion" collectionCode="2 specs, CEMT" collectorName="S. Peck" country="Venezuela" elevation="2000" location="road Tiara - Caracas" pageId="105" pageNumber="106" specimenCount="1" stateProvince="Aragua">
<collectionCode id="F3E85FB53033520DEFF8FCB874450205" box="[751,841,799,826]" pageId="105" pageNumber="106">2 specs</collectionCode>
;
<location id="902691AB3033520DEE41FCB873310206" LSID="urn:lsid:plazi:treatment:1D5076663032520DEF21F9D1731E070F:902691AB3033520DEE41FCB873310206" box="[854,1085,799,825]" country="Venezuela" name="road Tiara - Caracas" pageId="105" pageNumber="106" stateProvince="Aragua">road TiaraCaracas</location>
;
<elevation id="1ED420433033520DE95DFCB873E80206" box="[1098,1252,799,826]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="105" pageNumber="106" unit="m" value="2000.0">
<quantity id="52016A953033520DE95DFCB873AF0205" box="[1098,1187,799,826]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="105" pageNumber="106" unit="m" value="2000.0">2000 m</quantity>
a.s.l.
</elevation>
;
<date id="E147E1B03033520DE9E5FCB877F70262" pageId="105" pageNumber="106" value="1971-02-22" valueMax="1971-02-25" valueMin="1971-02-22">
<collectingDate id="F10318583033520DE9E5FCB877F70262" pageId="105" pageNumber="106" value="1971-02-22" valueMax="1971-02-25" valueMin="1971-02-22">2225 Feb. 1971</collectingDate>
</date>
;
<collectorName id="380CA2A63033520DEC1FFCE4766D0263" box="[264,353,834,861]" pageId="105" pageNumber="106">S. Peck</collectorName>
leg.;
<collectingMethod id="4CB8BF673033520DECB3FCE576F70263" box="[420,507,834,860]" pageId="105" pageNumber="106">carrion</collectingMethod>
;
<collectedFrom id="26E5848D3033520DEF1FFCE575430263" box="[520,591,834,860]" pageId="105" pageNumber="106">forest</collectedFrom>
;
<collectionCode id="F3E85FB53033520DEF4CFCE475A20262" box="[603,686,835,861]" pageId="105" pageNumber="106">CEMT</collectionCode>
</materialsCitation>
.
<materialsCitation id="2591CD2D3033520DEFC7FCE572500262" box="[720,1372,834,861]" collectingDate="1949-12" collectionCode="CEMT" country="Venezuela" location="Caracas" pageId="105" pageNumber="106" specimenCount="1" stateProvince="Distrito Capital">
<emphasis id="A78D1B623033520DEFC7FCE574990263" bold="true" box="[720,917,834,860]" pageId="105" pageNumber="106">
<collectingRegion id="573D09923033520DEFC7FCE574990263" box="[720,917,834,860]" country="Venezuela" name="Distrito Capital" pageId="105" pageNumber="106">Distrito Capital</collectingRegion>
</emphasis>
<specimenCount id="83FF0CF93033520DEEA7FCE473090262" box="[944,1029,835,861]" count="1" pageId="105" pageNumber="106" type="generic">1 spec.</specimenCount>
;
<location id="902691AB3033520DE905FCE4737F0262" LSID="urn:lsid:plazi:treatment:1D5076663032520DEF21F9D1731E070F:902691AB3033520DE905FCE4737F0262" box="[1042,1139,835,861]" country="Venezuela" name="Caracas" pageId="105" pageNumber="106" stateProvince="Distrito Capital">Caracas</location>
;
<date id="E147E1B03033520DE968FCE473F00262" box="[1151,1276,835,861]" pageId="105" pageNumber="106" value="1949-12">
<collectingDate id="F10318583033520DE968FCE473F00262" box="[1151,1276,835,861]" pageId="105" pageNumber="106" value="1949-12">Dec. 1949</collectingDate>
</date>
;
<collectionCode id="F3E85FB53033520DE81EFCE472500262" box="[1289,1372,835,861]" pageId="105" pageNumber="106">CEMT</collectionCode>
</materialsCitation>
.
<materialsCitation id="2591CD2D3033520DEDAAFCC2727B02BF" box="[189,1399,869,897]" collectedFrom="dung" collectingDate="1995-07" collectionCode="CEMT" collectorName="A. de Ascencao" country="Venezuela" elevation="2200" location="Merida" municipality="Monte Zerpa" pageId="105" pageNumber="106" specimenCount="2" stateProvince="Merida">
<emphasis id="A78D1B623033520DEDAAFCC2761602BF" bold="true" box="[189,282,869,896]" pageId="105" pageNumber="106">
<collectingRegion id="573D09923033520DEDAAFCC2761602BF" box="[189,282,869,896]" country="Venezuela" name="Merida" pageId="105" pageNumber="106">Mérida</collectingRegion>
</emphasis>
<specimenCount id="83FF0CF93033520DEC21FCC1769D02BF" box="[310,401,870,896]" pageId="105" pageNumber="106">2 specs</specimenCount>
;
<collectingMunicipality id="75225D0A3033520DEC88FCC1753002BF" box="[415,572,870,896]" pageId="105" pageNumber="106">Monte Zerpa</collectingMunicipality>
,
<location id="902691AB3033520DEF5DFCC175AF02BF" LSID="urn:lsid:plazi:treatment:1D5076663032520DEF21F9D1731E070F:902691AB3033520DEF5DFCC175AF02BF" box="[586,675,870,896]" country="Venezuela" municipality="Monte Zerpa" name="Merida" pageId="105" pageNumber="106" stateProvince="Merida">Merida</location>
;
<elevation id="1ED420433033520DEFA7FCC1744002BF" box="[688,844,870,896]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="105" pageNumber="106" unit="m" value="2200.0">
<quantity id="52016A953033520DEFA7FCC1740702BF" box="[688,779,870,896]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="105" pageNumber="106" unit="m" value="2200.0">2200 m</quantity>
a.s.l.
</elevation>
;
<date id="E147E1B03033520DEE4BFCC174C002BE" box="[860,972,870,897]" pageId="105" pageNumber="106" value="1995-07">
<collectingDate id="F10318583033520DEE4BFCC174C002BE" box="[860,972,870,897]" pageId="105" pageNumber="106" value="1995-07">Jul. 1995</collectingDate>
</date>
;
<collectorName id="380CA2A63033520DEECEFCC1739802BF" box="[985,1172,870,896]" pageId="105" pageNumber="106">A. de Ascençao</collectorName>
leg.;
<collectedFrom id="26E5848D3033520DE9CDFCC1721402BF" box="[1242,1304,870,896]" pageId="105" pageNumber="106">dung</collectedFrom>
;
<collectionCode id="F3E85FB53033520DE831FCC1727B02BF" box="[1318,1399,870,896]" pageId="105" pageNumber="106">CEMT</collectionCode>
</materialsCitation>
<materialsCitation id="2591CD2D3033520DEDC5FC2E7432029C" box="[210,830,905,931]" collectingDate="2003-11" collectionCode="CEMT" collectorName="G. Arriagada" country="Venezuela" location="Zea" pageId="105" pageNumber="106" specimenCount="1" stateProvince="Merida">
<specimenCount id="83FF0CF93033520DEDC5FC2E7626029C" box="[210,298,905,931]" count="1" pageId="105" pageNumber="106" type="generic">1 spec.</specimenCount>
;
<location id="902691AB3033520DEC2EFC2E7665029C" LSID="urn:lsid:plazi:treatment:1D5076663032520DEF21F9D1731E070F:902691AB3033520DEC2EFC2E7665029C" box="[313,361,905,931]" country="Venezuela" name="Zea" pageId="105" pageNumber="106" stateProvince="Merida">Zea</location>
;
<date id="E147E1B03033520DEC6FFC2E76F5029C" box="[376,505,905,931]" pageId="105" pageNumber="106" value="2003-11">
<collectingDate id="F10318583033520DEC6FFC2E76F5029C" box="[376,505,905,931]" pageId="105" pageNumber="106" value="2003-11">Nov. 2003</collectingDate>
</date>
;
<collectorName id="380CA2A63033520DEF1FFC2E75AF029C" box="[520,675,905,931]" pageId="105" pageNumber="106">G. Arriágada</collectorName>
leg.;
<collectionCode id="F3E85FB53033520DEFFCFC2E7432029C" box="[747,830,905,931]" pageId="105" pageNumber="106">CEMT</collectionCode>
</materialsCitation>
.
<materialsCitation id="2591CD2D3033520DEE71FC2E759002D6" collectingDate="2009-07" collectionCode="CEMT" collectorName="F. Z. Vaz-de-Mello" country="Venezuela" county="Altos de Pipe" elevation="1500" latitude="10.3975" location="Instituto Venezolano de Investigaciones Cientificas" longLatPrecision="21" longitude="-66.97083" municipality="Instituto Venezolano de Investigaciones Cientificas" pageId="105" pageNumber="106" specimenCount="2" stateProvince="Miranda">
<emphasis id="A78D1B623033520DEE71FC2E74D9029C" bold="true" box="[870,981,905,931]" pageId="105" pageNumber="106">
<collectingRegion id="573D09923033520DEE71FC2E74D9029C" box="[870,981,905,931]" country="Venezuela" name="Miranda" pageId="105" pageNumber="106">Miranda</collectingRegion>
</emphasis>
<specimenCount id="83FF0CF93033520DEEE3FC2E735C029C" box="[1012,1104,905,931]" pageId="105" pageNumber="106">2 specs</specimenCount>
;
<collectingCounty id="7C27BFFC3033520DE949FC2E720A029C" box="[1118,1286,905,931]" pageId="105" pageNumber="106">Altos de Pipe</collectingCounty>
,
<location id="902691AB3033520DE802FC2E75A602F9" LSID="urn:lsid:plazi:treatment:1D5076663032520DEF21F9D1731E070F:902691AB3033520DE802FC2E75A602F9" country="Venezuela" county="Altos de Pipe" latitude="10.3975" longLatPrecision="21" longitude="-66.97083" municipality="Instituto Venezolano de Investigaciones Cientificas" name="Instituto Venezolano de Investigaciones Cientificas" pageId="105" pageNumber="106" stateProvince="Miranda">Instituto Venezolano de Investigaciones Científicas</location>
(“
<location id="902691AB3033520DEFD3FC0B747E02F9" LSID="urn:lsid:plazi:treatment:1D5076663032520DEF21F9D1731E070F:902691AB3033520DEFD3FC0B747E02F9" box="[708,882,940,966]" country="Venezuela" county="Altos de Pipe" latitude="10.3975" longLatPrecision="21" longitude="-66.97083" municipality="Instituto Venezolano de Investigaciones Cientificas" name="Campus IVIC" pageId="105" pageNumber="106" stateProvince="Miranda">Campus IVIC</location>
”);
<geoCoordinate id="F0CDA1B73033520DEE86FC0C732A02F8" box="[913,1062,939,967]" degrees="10" direction="north" minutes="23" orientation="latitude" pageId="105" pageNumber="106" precision="15" seconds="51" value="10.3975">10°2351″ N</geoCoordinate>
,
<geoCoordinate id="F0CDA1B73033520DE925FC0C73C102F8" box="[1074,1229,939,967]" degrees="66" direction="west" minutes="58" orientation="longitude" pageId="105" pageNumber="106" precision="15" seconds="15" value="-66.97083">66°5815″ W</geoCoordinate>
;
<elevation id="1ED420433033520DE9CFFC0B727C02F9" box="[1240,1392,940,967]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="105" pageNumber="106" unit="m" value="1500.0">
<quantity id="52016A953033520DE9CFFC0B723D02F8" box="[1240,1329,940,967]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="105" pageNumber="106" unit="m" value="1500.0">1500 m</quantity>
a.s.l.
</elevation>
;
<date id="E147E1B03033520DEDAAFC68762602D6" box="[189,298,975,1001]" pageId="105" pageNumber="106" value="2009-07">
<collectingDate id="F10318583033520DEDAAFC68762602D6" box="[189,298,975,1001]" pageId="105" pageNumber="106" value="2009-07">Jul. 2009</collectingDate>
</date>
;
<collectorName id="380CA2A63033520DEC22FC77750602D6" box="[309,522,975,1002]" pageId="105" pageNumber="106">F.Z. Vaz-de-Mello</collectorName>
leg.;
<collectionCode id="F3E85FB53033520DEF5EFC68759002D6" box="[585,668,975,1001]" pageId="105" pageNumber="106">CEMT</collectionCode>
</materialsCitation>
.
<materialsCitation id="2591CD2D3033520DEFADFC6974F50532" collectingDate="1989-01-17" collectingMethod="human faeces trap" collectionCode="CEMT" collectorName="D. Havranek" country="Venezuela" elevation="600" location="Futura Presa Las Cuevas" municipality="near Pregonero" pageId="105" pageNumber="106" specimenCount="1" stateProvince="Tachira">
<emphasis id="A78D1B623033520DEFADFC69741102D6" bold="true" box="[698,797,974,1001]" pageId="105" pageNumber="106">
<collectingRegion id="573D09923033520DEFADFC69741102D6" box="[698,797,974,1001]" country="Venezuela" name="Tachira" pageId="105" pageNumber="106">Táchira</collectingRegion>
</emphasis>
<specimenCount id="83FF0CF93033520DEE23FC68748B02D5" box="[820,903,975,1002]" count="1" pageId="105" pageNumber="106" type="generic">1 spec.</specimenCount>
;
<collectingMunicipality id="75225D0A3033520DEE85FC77734A02D5" box="[914,1094,976,1002]" pageId="105" pageNumber="106">near Pregonero</collectingMunicipality>
,
<location id="902691AB3033520DE947FC77727E02D6" LSID="urn:lsid:plazi:treatment:1D5076663032520DEF21F9D1731E070F:902691AB3033520DE947FC77727E02D6" box="[1104,1394,975,1002]" country="Venezuela" municipality="near Pregonero" name="Futura Presa Las Cuevas" pageId="105" pageNumber="106" stateProvince="Tachira">Futura Presa Las Cuevas</location>
;
<elevation id="1ED420433033520DEDAAFC5576490533" box="[189,325,1010,1037]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="105" pageNumber="106" unit="m" value="600.0">
<quantity id="52016A953033520DEDAAFC55760A0532" box="[189,262,1010,1037]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="105" pageNumber="106" unit="m" value="600.0">600 m</quantity>
a.s.l.
</elevation>
;
<date id="E147E1B03033520DEC46FC5476EB0532" box="[337,487,1011,1037]" pageId="105" pageNumber="106" value="1989-01-17">
<collectingDate id="F10318583033520DEC46FC5476EB0532" box="[337,487,1011,1037]" pageId="105" pageNumber="106" value="1989-01-17">17 Jan. 1989</collectingDate>
</date>
;
<collectorName id="380CA2A63033520DECE5FC54758B0533" box="[498,647,1010,1037]" pageId="105" pageNumber="106">D. Havranek</collectorName>
leg.;
<collectingMethod id="4CB8BF673033520DEFD1FC5574910532" box="[710,925,1010,1037]" pageId="105" pageNumber="106">human faeces trap</collectingMethod>
;
<collectionCode id="F3E85FB53033520DEEBFFC5474F50532" box="[936,1017,1011,1037]" pageId="105" pageNumber="106">CEMT</collectionCode>
</materialsCitation>
<materialsCitation id="2591CD2D3033520DE918FC54726F050F" collectingDate="1989-08-16" collectingMethod="mini-cup trap" collectionCode="CEMT" collectorName="La Honda &amp; D. Havranek" country="Venezuela" elevation="1100" location="La Idea" municipality="near Pregonero" pageId="105" pageNumber="106" specimenCode="CEMT" specimenCount="2" stateProvince="Tachira">
<specimenCount id="83FF0CF93033520DE918FC54736B0532" box="[1039,1127,1011,1037]" pageId="105" pageNumber="106">2 specs</specimenCount>
;
<collectingMunicipality id="75225D0A3033520DE965FC54722A0532" box="[1138,1318,1011,1037]" pageId="105" pageNumber="106">near Pregonero</collectingMunicipality>
, presa/
<collectorName id="380CA2A63033520DEDAAFBB1763F050F" box="[189,307,1046,1072]" pageId="105" pageNumber="106">La Honda</collectorName>
(
<location id="902691AB3033520DEC53FBB176A9050F" LSID="urn:lsid:plazi:treatment:1D5076663032520DEF21F9D1731E070F:902691AB3033520DEC53FBB176A9050F" box="[324,421,1046,1072]" country="Venezuela" municipality="near Pregonero" name="La Idea" pageId="105" pageNumber="106" stateProvince="Tachira">La Idea</location>
);
<elevation id="1ED420433033520DECABFBB1755B050F" box="[444,599,1046,1072]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="105" pageNumber="106" unit="m" value="1100.0">
<quantity id="52016A953033520DECABFBB17519050F" box="[444,533,1046,1072]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="105" pageNumber="106" unit="m" value="1100.0">1100 m</quantity>
a.s.l.
</elevation>
;
<date id="E147E1B03033520DEF71FBB17400050F" box="[614,780,1046,1072]" pageId="105" pageNumber="106" value="1989-08-16">
<collectingDate id="F10318583033520DEF71FBB17400050F" box="[614,780,1046,1072]" pageId="105" pageNumber="106" value="1989-08-16">16 Aug. 1989</collectingDate>
</date>
;
<collectorName id="380CA2A63033520DEE0DFBB174BD050F" box="[794,945,1046,1072]" pageId="105" pageNumber="106">D. Havranek</collectorName>
leg.;
<collectingMethod id="4CB8BF673033520DEEE0FBB17394050F" box="[1015,1176,1046,1072]" pageId="105" pageNumber="106">mini-cup trap</collectingMethod>
C(T7-6);
<specimenCode id="C55F6F0B3033520DE805FBB1726F050F" box="[1298,1379,1046,1072]" pageId="105" pageNumber="106">CEMT</specimenCode>
</materialsCitation>
<materialsCitation id="2591CD2D3033520DEDAAFB9E74B4056C" box="[189,952,1081,1107]" collectingDate="1987-08" collectionCode="CEMT" collectorName="J. Blanco" country="Venezuela" location="Via Chorro del Indio" pageId="105" pageNumber="106" specimenCount="1" stateProvince="Tachira">
<specimenCount id="83FF0CF93033520DEDAAFB9E761F056C" box="[189,275,1081,1107]" count="1" pageId="105" pageNumber="106" type="generic">1 spec.</specimenCount>
;
<location id="902691AB3033520DEC37FB9E7510056C" LSID="urn:lsid:plazi:treatment:1D5076663032520DEF21F9D1731E070F:902691AB3033520DEC37FB9E7510056C" box="[288,540,1081,1107]" country="Venezuela" name="Via Chorro del Indio" pageId="105" pageNumber="106" stateProvince="Tachira">Vía Chorro del Indio</location>
;
<date id="E147E1B03033520DEF30FB9E75AB056C" box="[551,679,1081,1107]" pageId="105" pageNumber="106" value="1987-08">
<collectingDate id="F10318583033520DEF30FB9E75AB056C" box="[551,679,1081,1107]" pageId="105" pageNumber="106" value="1987-08">Aug. 1987</collectingDate>
</date>
;
<collectorName id="380CA2A63033520DEFA3FB9E742E056C" box="[692,802,1081,1107]" pageId="105" pageNumber="106">J. Blanco</collectorName>
leg.;
<collectionCode id="F3E85FB53033520DEE70FB9E74B4056C" box="[871,952,1081,1107]" pageId="105" pageNumber="106">CEMT</collectionCode>
</materialsCitation>
<materialsCitation id="2591CD2D3033520DEEC4FB9E75060549" collectingDate="1987-08" collectionCode="CEMT" collectorName="J. Blanco" country="Venezuela" location="Via Chorro del Indio" pageId="105" pageNumber="106" specimenCount="1" stateProvince="Tachira">
<specimenCount id="83FF0CF93033520DEEC4FB9E7325056C" box="[979,1065,1081,1107]" count="1" pageId="105" pageNumber="106" type="generic">1 spec.</specimenCount>
;
<location id="902691AB3033520DE922FB9E723E056C" LSID="urn:lsid:plazi:treatment:1D5076663032520DEF21F9D1731E070F:902691AB3033520DE922FB9E723E056C" box="[1077,1330,1081,1107]" country="Venezuela" name="Via Chorro del Indio" pageId="105" pageNumber="106" stateProvince="Tachira">Vía Chorro del Indio</location>
;
<date id="E147E1B03033520DE82AFB9E77F00549" pageId="105" pageNumber="106" value="1987-08">
<collectingDate id="F10318583033520DE82AFB9E77F00549" pageId="105" pageNumber="106" value="1987-08">Aug. 1987</collectingDate>
</date>
;
<collectorName id="380CA2A63033520DEC1FFBFA76790549" box="[264,373,1116,1143]" pageId="105" pageNumber="106">J. Blanco</collectorName>
leg.;
<collectionCode id="F3E85FB53033520DECA0FBFB75060549" box="[439,522,1116,1142]" pageId="105" pageNumber="106">CEMT</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB3033520DEDAAFB3A743B0468" pageId="105" pageNumber="106" type="distribution">
<paragraph id="9546C7703033520DEDAAFB3A765A0588" blockId="105.[189,342,1181,1207]" box="[189,342,1181,1207]" pageId="105" pageNumber="106">
<heading id="CE0E701C3033520DEDAAFB3A765A0588" bold="true" box="[189,342,1181,1207]" fontSize="11" level="3" pageId="105" pageNumber="106" reason="3">
<emphasis id="A78D1B623033520DEDAAFB3A765A0588" bold="true" box="[189,342,1181,1207]" pageId="105" pageNumber="106">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="9546C7703033520DEDAAFB6B740605D9" blockId="105.[189,778,1228,1255]" box="[189,778,1228,1255]" pageId="105" pageNumber="106">
<collectingCountry id="EDEE87E03033520DEDAAFB6B763805D9" box="[189,308,1228,1254]" name="Colombia" pageId="105" pageNumber="106">Colombia</collectingCountry>
,
<collectingCountry id="EDEE87E03033520DEC28FB6B76B705D9" box="[319,443,1228,1254]" name="Venezuela" pageId="105" pageNumber="106">Venezuela</collectingCountry>
,
<collectingCountry id="EDEE87E03033520DECD1FB6B752605D9" box="[454,554,1228,1254]" name="Ecuador" pageId="105" pageNumber="106">Ecuador</collectingCountry>
,
<collectingCountry id="EDEE87E03033520DEF22FB6A756205D8" box="[565,622,1229,1255]" name="Peru" pageId="105" pageNumber="106">Peru</collectingCountry>
, and
<collectingCountry id="EDEE87E03033520DEFBCFB6B740905D9" box="[683,773,1228,1254]" name="Bolivia" pageId="105" pageNumber="106">Bolivia</collectingCountry>
.
</paragraph>
<paragraph id="9546C7703033520DEDAAFAAA76E10417" blockId="105.[189,493,1293,1320]" box="[189,493,1293,1320]" pageId="105" pageNumber="106">
<heading id="CE0E701C3033520DEDAAFAAA76E10417" bold="true" box="[189,493,1293,1320]" fontSize="11" level="3" pageId="105" pageNumber="106" reason="3">
<emphasis id="A78D1B623033520DEDAAFAAA76E10417" bold="true" box="[189,493,1293,1320]" pageId="105" pageNumber="106">
Subregions of
<collectingCountry id="EDEE87E03033520DEC78FAA976E10417" box="[367,493,1294,1320]" name="Venezuela" pageId="105" pageNumber="106">Venezuela</collectingCountry>
</emphasis>
</heading>
</paragraph>
<paragraph id="9546C7703033520DEDAAFA9B743B0468" blockId="105.[189,823,1340,1367]" box="[189,823,1340,1367]" pageId="105" pageNumber="106">
Andes mountains and Central
<collectingRegion id="573D09923033520DEF37FA9B756F0469" box="[544,611,1340,1366]" country="New Zealand" name="West Coast" pageId="105" pageNumber="106">Coast</collectingRegion>
Mountain Range.
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB3033520DEDAAFADA731E070F" pageId="105" pageNumber="106" type="reference_group">
<paragraph id="9546C7703033520DEDAAFADA76A804A7" blockId="105.[189,420,1405,1432]" box="[189,420,1405,1432]" pageId="105" pageNumber="106">
<heading id="CE0E701C3033520DEDAAFADA76A804A7" bold="true" box="[189,420,1405,1432]" fontSize="11" level="3" pageId="105" pageNumber="106" reason="3">
<emphasis id="A78D1B623033520DEDAAFADA76A804A7" bold="true" box="[189,420,1405,1432]" pageId="105" pageNumber="106">Literature records</emphasis>
</heading>
</paragraph>
<paragraph id="9546C7703033520DEDAAFA0B731E070F" blockId="105.[188,1398,1452,1584]" pageId="105" pageNumber="106">
<bibRefCitation id="F168BA813033520DEDAAFA0B768704F9" author="Gillet J. J. E." box="[189,395,1452,1478]" pageId="105" pageNumber="106" refId="ref121235" refString="Gillet J. J. E. 1911 b. Coleopterorum Catalogus. Pars 38: Scarabaeidae: Coprinae I. W. Junk, Berlin. Available from https: // www. biodiversitylibrary. org / page / 48405679 [accesseed 16 Feb. 2024]." type="url" year="1911">Gillet 1911b: 61</bibRefCitation>
(
<collectingCountry id="EDEE87E03033520DECB6FA0B752D04F9" box="[417,545,1452,1478]" name="Venezuela" pageId="105" pageNumber="106">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA813033520DEF7FFA0B747504F9" author="Luederwaldt H." box="[616,889,1452,1478]" pageId="105" pageNumber="106" pagination="603 - 775" refId="ref128687" refString="Luederwaldt H. 1929. As especies brasileiras do genero Pinotus, com algumas consideracoes tambem sobre outras especies. Revista do Museu Paulista 16: 603 - 775." type="journal article" year="1929">Luederwaldt 1929: 34</bibRefCitation>
(
<collectingCountry id="EDEE87E03033520DEE87FA0B731C04F9" box="[912,1040,1452,1478]" name="Venezuela" pageId="105" pageNumber="106">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA813033520DE941FA0B727A04F9" author="Blackwelder R. E." box="[1110,1398,1452,1478]" pageId="105" pageNumber="106" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 207</bibRefCitation>
(
<collectingCountry id="EDEE87E03033520DEDD2FA68764904D6" box="[197,325,1487,1513]" name="Venezuela" pageId="105" pageNumber="106">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA813033520DEC6CFA77752904D6" author="Roze J. A." box="[379,549,1487,1514]" pageId="105" pageNumber="106" pagination="39 - 63" refId="ref137914" refString="Roze J. A. 1955. Lista preliminar de la familia Scarabaeidae sensu lato de Venezuela. Boletin del Museo de Ciencias Naturales 1 (1): 39 - 63." type="journal article" year="1955">Roze 1955: 44</bibRefCitation>
(
<collectingCountry id="EDEE87E03033520DEF23FA6875B804D6" box="[564,692,1487,1513]" name="Venezuela" pageId="105" pageNumber="106">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA813033520DEFFCFA6874A204D6" author="Blanco J." box="[747,942,1487,1513]" pageId="105" pageNumber="106" pagination="39 - 48" refId="ref111584" refString="Blanco J. 1988. Catalogo de los Scarabaeinae (Coleoptera) coprofagos y necrofagos, del Estado Tachira. Venezuela. Revista Cientifica Unet 2 (1): 39 - 48." type="journal article" year="1988">Blanco 1988: 40</bibRefCitation>
, 41 (
<collectingCountry id="EDEE87E03033520DEEF1FA68736904D6" box="[998,1125,1487,1513]" name="Venezuela" pageId="105" pageNumber="106">Venezuela</collectingCountry>
:
<collectingRegion id="573D09923033520DE978FA6873C204D6" box="[1135,1230,1487,1513]" country="Venezuela" name="Tachira" pageId="105" pageNumber="106">Táchira</collectingRegion>
). —
<bibRefCitation id="F168BA813033520DE813FA6876290733" author="Havranek D." pageId="105" pageNumber="106" pagination="55 - 64" refId="ref123886" refString="Havranek D. 1989. Coleoptera coprofagos y necrofagos (Scarabaeidae, Silphidae) atraidos a cebos en bosques venezolanos. UNET 3: 55 - 64." type="journal article" year="1989">Havranek 1989: 61</bibRefCitation>
(
<collectingCountry id="EDEE87E03033520DEC20FA5576BA0733" box="[311,438,1522,1548]" name="Venezuela" pageId="105" pageNumber="106">Venezuela</collectingCountry>
:
<collectingRegion id="573D09923033520DECD5FA5575530732" box="[450,607,1522,1549]" country="Venezuela" name="Tachira" pageId="105" pageNumber="106">Táchira state</collectingRegion>
). —
<bibRefCitation id="F168BA813033520DEF8CFA5574E10732" author="Ferrer-Paris J. &amp; Sanchez-Mercado A. &amp; Rodriguez J. P." box="[667,1005,1522,1549]" pageId="105" pageNumber="106" pagination="89 - 110" refId="ref119203" refString="Ferrer-Paris J. R, Sanchez-Mercado A. &amp; Rodriguez J. P. 2013. Optimizacion del muestreo de invertebrados tropicales: un ejemplo con escarabajos coprofagos (Coleoptera: Scarabaeinae) en Venezuela. Revista de Biologia Tropical 61 (1): 89 - 110. https: // doi. org / 10.15517 / rbt. v 61 i 1.10941" type="journal article" year="2013">
Ferrer-Paris
<emphasis id="A78D1B623033520DEE25FA5474690733" box="[818,869,1522,1548]" italics="true" pageId="105" pageNumber="106">et al</emphasis>
. 2013: 109
</bibRefCitation>
(
<collectingCountry id="EDEE87E03033520DEEE9FA5573710733" box="[1022,1149,1522,1548]" name="Venezuela" pageId="105" pageNumber="106">Venezuela</collectingCountry>
:
<collectingRegion id="573D09923033520DE99DFA5573EF0733" box="[1162,1251,1522,1548]" country="Venezuela" name="Merida" pageId="105" pageNumber="106">Mérida</collectingRegion>
: La Azulita-Jají;
<collectingRegion id="573D09923033520DEDE2F9B17651070F" box="[245,349,1558,1584]" country="Venezuela" name="Miranda" pageId="105" pageNumber="106">Miranda</collectingRegion>
: Altos de Pipe). —
<bibRefCitation id="F168BA813033520DEF55F9B1747B070F" author="Chamorro W. &amp; Marin-Armijos D. &amp; Asenjo A. &amp; Vaz-De-Mello F. Z." box="[578,887,1558,1584]" pageId="105" pageNumber="106" pagination="1 - 343" refId="ref114107" refString="Chamorro W., Marin-Armijos D., Asenjo A. &amp; Vaz-De-Mello F. Z. 2019. Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records. ZooKeys 826: 1 - 343. https: // doi. org / 10.3897 / zookeys. 826.26488" type="journal article" year="2019">
Chamorro
<emphasis id="A78D1B623033520DEFD7F9B175FF070F" box="[704,755,1558,1584]" italics="true" pageId="105" pageNumber="106">et al</emphasis>
. 2019: 119
</bibRefCitation>
(
<collectingCountry id="EDEE87E03033520DEE91F9B1730A070F" box="[902,1030,1558,1584]" name="Venezuela" pageId="105" pageNumber="106">Venezuela</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>