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<document id="8BB253987F0FA2D2C4C951C368F4AA35" ID-DOI="10.5852/ejt.2024.959.2677" ID-ISSN="2118-9773" ID-Zenodo-Dep="13945464" ID-ZooBank="30872C13-516F-42FA-AFA7-30ADC6BF1BAF" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1729166637250" checkinUser="plazi" docAuthor="Rosa, Cecilia Lozano De La, Cupello, Mario &amp; Vaz-De-Mello, Fernando Z." docDate="2024" docId="1D5076663013522EECE9F9D3765D027D" docLanguage="en" docName="EJT.2024.959.1-272.pdf" docOrigin="European Journal of Taxonomy 959 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2677/12401" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Deltochilum bordoni Halffter &amp; Martinez 1976" docType="treatment" docVersion="2" lastPageNumber="75" masterDocId="E1690E1E305A5264ED17FFA7770C013F" masterDocTitle="The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution" masterLastPageNumber="272" masterPageNumber="1" pageNumber="74" updateTime="1729169574746" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="C36D46F85E4442B31290B50DFBFDE4F1" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F442D7CAE42EAE91B9696904A67A711C">
<mods:title id="33D9DB398EB7E6B0DE71D957F849D82A">The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution</mods:title>
</mods:titleInfo>
<mods:name id="BB732C2518D27F13346FCE8B96D9AC8C" type="personal">
<mods:role id="EBC08CC47553B960D5717CFCBDE1FB0F">
<mods:roleTerm id="AF2B8C740D6D31E01C4D417C7B6DE570">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0744521FD883FBD42B5350079E96936F">Rosa, Cecilia Lozano De La</mods:namePart>
<mods:nameIdentifier id="BD3515D620579D966CAAE753218E4001" type="LSID">36CFEA67-71C5-4F3C-97F8-B57E43C811CC</mods:nameIdentifier>
<mods:affiliation id="8064067A69B93928A031724390C28868">Instituto Venezolano de Investigaciones Científicas, Centro de Estudios Botánicos y Agroforestales, Laboratorio de Ecología Espacial, Maracaibo, estado Zulia, Venezuela.</mods:affiliation>
<mods:nameIdentifier id="39B052A2074F5238F9EE5E7A92424B3F" type="email">lozanoceci@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="FCFE0E1C51130EB46B42558B39C155E1" type="personal">
<mods:role id="CDE6B7A45891837A7087F49346BCD6A3">
<mods:roleTerm id="5FC934BD371D9AB9A3D1143AC3BD0AC4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="149D33C5C0796614E4DA3159BEB1E721">Cupello, Mario</mods:namePart>
<mods:nameIdentifier id="C13606B479A13A9A1D718CCAFBCB8AFB" type="LSID">BDB03C18-7095-4EAA-8BDD-03CB4F79676D</mods:nameIdentifier>
<mods:affiliation id="28D8331230C3C91E88E9272B957E2466">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Curitiba, Paraná, Brazil. Currently at Department of Entomology, Texas A &amp; M University, College Station, TX, 77843, USA.</mods:affiliation>
<mods:nameIdentifier id="B556261B788E8193EC5AD38875608E33" type="email">mcupello@hotmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="AC13B629965BF4655CB63A3C732B575E" type="personal">
<mods:role id="03A45CC727EE90FA0705CB7CB0161463">
<mods:roleTerm id="08E85F0CE2BB14D2EE4C4F627637D2F9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="35D06C2E53F93AFCB9EFF7657CC44916">Vaz-De-Mello, Fernando Z.</mods:namePart>
<mods:nameIdentifier id="F735945475ECBFC77E89938FF33EE03E" type="LSID">2BC093C2-3E2B-466B-B31C-9F3D9FA844BC</mods:nameIdentifier>
<mods:affiliation id="A274DC2E443498E7E29ED3E91EF50E7D">Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, Cuiabá, Mato Grosso, Brazil.</mods:affiliation>
<mods:nameIdentifier id="86A91BCC360856891F934CE9A7718695" type="email">vazdemello@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="D8C449B9AAA6FDF126D04BDBE7A2A93A">text</mods:typeOfResource>
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<mods:title id="D3903FB66837C48B65CEA7E02FB4B3D1">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
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<mods:date id="450D8E8C5AF28CA263CBC8B39AEBCC07">2024</mods:date>
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<mods:number id="521A9D43BA0D19240509D2D1445EC671">2024-10-11</mods:number>
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<mods:detail id="75BCEFF8FD8333B58955E36837808ADF" type="volume">
<mods:number id="200A753EB47FB0659CC2825D0A3D926E">959</mods:number>
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<mods:identifier id="631B748BAC7F94DA0E49DF6286D06F33" type="DOI">10.5852/ejt.2024.959.2677</mods:identifier>
<mods:identifier id="D78444D9C721AD9142554FA95D271B5C" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="F42A87601251D3C5BB34A8C0EDF98292" type="Zenodo-Dep">13945464</mods:identifier>
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<treatment id="1D5076663013522EECE9F9D3765D027D" ID-DOI="http://doi.org/10.5281/zenodo.13945712" ID-Zenodo-Dep="13945712" LSID="urn:lsid:plazi:treatment:1D5076663013522EECE9F9D3765D027D" httpUri="http://treatment.plazi.org/id/1D5076663013522EECE9F9D3765D027D" lastPageId="74" lastPageNumber="75" pageId="73" pageNumber="74">
<subSubSection id="DDE394FB3013522DECE9F9D3733A07B1" box="[510,1078,1652,1679]" pageId="73" pageNumber="74" type="nomenclature">
<paragraph id="9546C7703013522DECE9F9D3733A07B1" blockId="73.[510,1078,1652,1712]" box="[510,1078,1652,1679]" pageId="73" pageNumber="74">
<heading id="CE0E701C3013522DECE9F9D3733A07B1" box="[510,1078,1652,1679]" centered="true" fontSize="11" level="2" pageId="73" pageNumber="74" reason="2">
<taxonomicName id="52F9BCF33013522DECE9F9D3733A07B1" ID-CoL="34KBW" authority="Halffter &amp; Martinez, 1976" authorityName="Halffter &amp; Martinez" authorityYear="1976" box="[510,1078,1652,1679]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="bordoni">
<emphasis id="A78D1B623013522DECE9F9D375F607B0" bold="true" box="[510,762,1652,1679]" italics="true" pageId="73" pageNumber="74">Deltochilum bordoni</emphasis>
Halffter &amp; Martínez, 1976
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB3013522DEFF1F9327443078F" box="[742,847,1685,1712]" pageId="73" pageNumber="74" type="description">
<paragraph id="9546C7703013522DEFF1F9327443078F" blockId="73.[510,1078,1652,1712]" box="[742,847,1685,1712]" pageId="73" pageNumber="74">
<figureCitation id="0DC2DBF53013522DEFF1F9327443078F" box="[742,847,1685,1712]" captionStart="Fig" captionStartId="75.[189,232,1963,1989]" captionTargetBox="[271,1304,278,1893]" captionTargetId="figure-12@75.[260,1327,265,1909]" captionTargetPageId="75" captionText="Fig. 29. Species distribution. A. Deltochilum bordoni Halffter &amp; Martínez, 1976. B. Deltochilum cristinae Martínez, 1991. Green diamond = CEMT collection data; black circle = literature data." figureDoi="http://doi.org/10.5281/zenodo.13945543" httpUri="https://zenodo.org/record/13945543/files/figure.png" pageId="73" pageNumber="74">Fig. 29A</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB3013522DEDAAF97276D1064A" pageId="73" pageNumber="74" type="reference_group">
<paragraph id="9546C7703013522DEDAAF9727229062F" blockId="73.[189,1399,1749,1808]" pageId="73" pageNumber="74">
<taxonomicName id="52F9BCF33013522DEDAAF972744F07CF" ID-CoL="34KBW" authority="Halffter &amp; Martinez, 1976: 39 - 45" authorityName="Halffter &amp; Martinez" authorityPageNumber="39 - 45" authorityYear="1976" box="[189,835,1749,1776]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="bordoni">
<emphasis id="A78D1B623013522DEDAAF97276BD07D0" box="[189,433,1749,1775]" italics="true" pageId="73" pageNumber="74">Deltochilum bordoni</emphasis>
Halffter &amp; Martínez, 1976: 39–45
</taxonomicName>
(original description). Type locality:
<collectingCountry id="EDEE87E03013522DE9E1F972727F07D0" box="[1270,1395,1749,1775]" name="Venezuela" pageId="73" pageNumber="74">Venezuela</collectingCountry>
:
<collectingRegion id="573D09923013522DEDFDF951764B062F" box="[234,327,1782,1808]" country="Venezuela" name="Bolivar" pageId="73" pageNumber="74">Bolívar</collectingRegion>
: Meseta de Sarisariñama. Name-bearing type:
<typeStatus id="4A4279D23013522DEE7DF95174DD062F" box="[874,977,1782,1808]" pageId="73" pageNumber="74" type="holotype">holotype</typeStatus>
(MACN), examined by MC.
</paragraph>
<paragraph id="9546C7703013522DEDAAF89076D1064A" blockId="73.[189,1398,1847,1909]" pageId="73" pageNumber="74">
<treatmentCitationGroup id="B5E9E05E3013522DEDAAF89076D1064A" pageId="73" pageNumber="74">
<taxonomicName id="52F9BCF33013522DEDAAF89076BA066E" ID-CoL="34KBW" authorityName="Halffter &amp; Martinez" authorityYear="1976" box="[189,438,1847,1873]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="bordoni">
<emphasis id="A78D1B623013522DEDAAF89076BA066E" box="[189,438,1847,1873]" italics="true" pageId="73" pageNumber="74">Deltochilum bordoni</emphasis>
</taxonomicName>
—
<treatmentCitation id="1458E1613013522DECFCF89075B6066D" author="Krajcik M." box="[491,698,1847,1874]" page="88" pageId="73" pageNumber="74" year="2012">
<bibRefCitation id="F168BA813013522DECFCF89075B6066D" author="Krajcik M." box="[491,698,1847,1874]" pageId="73" pageNumber="74" pagination="1 - 278" refId="ref126632" refString="Krajcik M. 2012. Checklist of the World Scarabaeoidea. Animma. X Supplement 5: 1 - 278." type="journal article" year="2012">Krajcik 2012: 88</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="1458E1613013522DEE2DF89073C8066E" ID-DOI="http://doi.org/10.5281/zenodo.3683487" ID-Zenodo-Dep="3683487" author="Hielkema A. J. &amp; Hielkema M. A." box="[826,1220,1847,1874]" httpUri="http://treatment.plazi.org/id/03D7FF1CFF86FFA0FF48FF4A305AFB5A" page="76" pageId="73" pageNumber="74" year="2019">
<bibRefCitation id="F168BA813013522DEE2DF89073C8066E" author="Hielkema A. J. &amp; Hielkema M. A." box="[826,1220,1847,1874]" pageId="73" pageNumber="74" pagination="1 - 306" refId="ref124225" refString="Hielkema A. J. &amp; Hielkema M. A. 2019. An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas. Insecta Mundi 732 (732): 1 - 306." type="journal article" year="2019">Hielkema &amp; Hielkema 2019: 76</bibRefCitation>
</treatmentCitation>
(bibliographic review for Guianas).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB3013522EEDAAF83B732800F0" lastPageId="74" lastPageNumber="75" pageId="73" pageNumber="74" type="materials_examined">
<paragraph id="9546C7703013522DEDAAF83B76A60689" blockId="73.[189,426,1948,1974]" box="[189,426,1948,1974]" pageId="73" pageNumber="74">
<heading id="CE0E701C3013522DEDAAF83B76A60689" bold="true" box="[189,426,1948,1974]" fontSize="11" level="3" pageId="73" pageNumber="74" reason="3">
<emphasis id="A78D1B623013522DEDAAF83B76A60689" bold="true" box="[189,426,1948,1974]" pageId="73" pageNumber="74">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="9546C7703013522DEDFDF86B765006D9" blockId="73.[234,348,1996,2022]" box="[234,348,1996,2022]" pageId="73" pageNumber="74">
<emphasis id="A78D1B623013522DEDFDF86B765006D9" bold="true" box="[234,348,1996,2022]" pageId="73" pageNumber="74">
<typeStatus id="4A4279D23013522DEDFDF86B765006D9" box="[234,348,1996,2022]" pageId="73" pageNumber="74" type="paratype">Paratype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="9546C7703010522EEDAAFEAE735D0078" blockId="74.[189,1398,265,327]" pageId="74" pageNumber="75">
<collectingCountry id="EDEE87E03010522EEDAAFEAE7667001C" box="[189,363,265,291]" name="Venezuela" pageId="74" pageNumber="75">VENEZUELA</collectingCountry>
‒
<specimenCount id="83FF0CF93010522EEC81FEAE76C6001C" box="[406,458,265,291]" count="1" pageId="74" pageNumber="75" type="female">1 ♀</specimenCount>
; “
<typeStatus id="4A4279D23010522EECFDFEAE7571001C" box="[490,637,265,291]" pageId="74" pageNumber="75" type="paratype">PARATIPO</typeStatus>
”; “
<taxonomicName id="52F9BCF33010522EEFB0FEAE7434001C" authorityName="Eschscholtz" authorityYear="1822" box="[679,824,265,291]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B623010522EEFB0FEAE7434001C" box="[679,824,265,291]" italics="true" pageId="74" pageNumber="75">Deltochilum</emphasis>
</taxonomicName>
(Delto- / hyboma)
<specimenCount id="83FF0CF93010522EE92FFEAE7343001C" box="[1080,1103,265,291]" count="1" pageId="74" pageNumber="75" type="female">♀</specimenCount>
. /
<taxonomicName id="52F9BCF33010522EE96CFEAE73DB001C" authorityName="Halffter &amp; Martinez" authorityYear="1976" box="[1147,1239,265,291]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="bordoni">
<emphasis id="A78D1B623010522EE96CFEAE73DB001C" box="[1147,1239,265,291]" italics="true" pageId="74" pageNumber="75">bordoni</emphasis>
</taxonomicName>
. / Halffter / A. Martínez- DET. 19 / 76”; “Kaieteur, BG. /
<date id="E147E1B03010522EEFC2FE8B74530078" box="[725,863,300,327]" pageId="74" pageNumber="75" value="1911-08-05">VIII.5 1911</date>
”; CEMT-00109591.
</paragraph>
<paragraph id="9546C7703010522EEDFDFEC976ED00B7" blockId="74.[189,1398,366,464]" box="[234,481,366,392]" pageId="74" pageNumber="75">
<emphasis id="A78D1B623010522EEDFDFEC976ED00B7" bold="true" box="[234,481,366,392]" pageId="74" pageNumber="75">Additional material</emphasis>
</paragraph>
<paragraph id="9546C7703010522EEDAAFE35732800F0" blockId="74.[189,1398,366,464]" pageId="74" pageNumber="75">
<materialsCitation id="2591CD2D3010522EEDAAFE3572680093" box="[189,1380,401,428]" collectingDate="1987-12-20" collectingDateMax="1987-12-27" collectingDateMin="1987-12-20" collectorName="B. D. Gill" country="Venezuela" location="El Dorado" pageId="74" pageNumber="75" specimenCount="1" stateProvince="Bolivar">
<collectingCountry id="EDEE87E03010522EEDAAFE3576670093" box="[189,363,402,428]" name="Venezuela" pageId="74" pageNumber="75">VENEZUELA</collectingCountry>
‒
<emphasis id="A78D1B623010522EEC9EFE3676E90093" bold="true" box="[393,485,401,428]" pageId="74" pageNumber="75">
<collectingRegion id="573D09923010522EEC9EFE3676E90093" box="[393,485,401,428]" country="Venezuela" name="Bolivar" pageId="74" pageNumber="75">Bolívar</collectingRegion>
</emphasis>
•
<specimenCount id="83FF0CF93010522EECE8FE3575590093" box="[511,597,402,428]" count="1" pageId="74" pageNumber="75" type="generic">1 spec.</specimenCount>
;
<quantity id="52016A953010522EEF75FE3575B70093" box="[610,699,402,428]" metricMagnitude="5" metricUnit="m" metricValue="1.2" pageId="74" pageNumber="75" unit="km" value="120.0">120 km</quantity>
S of
<location id="902691AB3010522EEFEAFE3574770093" LSID="urn:lsid:plazi:treatment:1D5076663013522EECE9F9D3765D027D:902691AB3010522EEFEAFE3574770093" box="[765,891,402,428]" country="Venezuela" name="El Dorado" pageId="74" pageNumber="75" stateProvince="Bolivar">El Dorado</location>
;
<date id="E147E1B03010522EEE90FE3573590093" box="[903,1109,402,428]" pageId="74" pageNumber="75" value="1987-12-20" valueMax="1987-12-27" valueMin="1987-12-20">
<collectingDate id="F10318583010522EEE90FE3573590093" box="[903,1109,402,428]" pageId="74" pageNumber="75" value="1987-12-20" valueMax="1987-12-27" valueMin="1987-12-20">20‒27 Dec. 1987</collectingDate>
</date>
;
<collectorName id="380CA2A63010522EE975FE3573DC0093" box="[1122,1232,402,428]" pageId="74" pageNumber="75">B.D. Gill</collectorName>
leg.; CEMT
</materialsCitation>
•
<materialsCitation id="2591CD2D3010522EEDAAFE12732C00F0" box="[189,1056,437,464]" collectingDate="1986-07-20" collectingDateMax="1986-08-07" collectingDateMin="1986-07-20" collectorName="B. Gill" country="Venezuela" location="El Dorado" pageId="74" pageNumber="75" specimenCount="1" stateProvince="Bolivar">
<specimenCount id="83FF0CF93010522EEDAAFE12761E00EF" box="[189,274,437,464]" count="1" pageId="74" pageNumber="75" type="generic">1 spec.</specimenCount>
;
<quantity id="52016A953010522EEC09FE12767A00F0" box="[286,374,437,464]" metricMagnitude="5" metricUnit="m" metricValue="1.35" pageId="74" pageNumber="75" unit="km" value="135.0">135 km</quantity>
S of
<location id="902691AB3010522EECA2FE12753F00F0" LSID="urn:lsid:plazi:treatment:1D5076663013522EECE9F9D3765D027D:902691AB3010522EECA2FE12753F00F0" box="[437,563,437,463]" country="Venezuela" name="El Dorado" pageId="74" pageNumber="75" stateProvince="Bolivar">El Dorado</location>
;
<date id="E147E1B03010522EEF29FE12742300F0" box="[574,815,437,463]" pageId="74" pageNumber="75" value="1986-07-20" valueMax="1986-08-07" valueMin="1986-07-20">
<collectingDate id="F10318583010522EEF29FE12742300F0" box="[574,815,437,463]" pageId="74" pageNumber="75" value="1986-07-20" valueMax="1986-08-07" valueMin="1986-07-20">20 Jul.‒7 Aug. 1986</collectingDate>
</date>
;
<collectorName id="380CA2A63010522EEE2CFE11748700F0" box="[827,907,437,464]" pageId="74" pageNumber="75">B. Gill</collectorName>
leg.; CEMT
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB3010522EEDAAFE51765D027D" pageId="74" pageNumber="75" type="distribution">
<paragraph id="9546C7703010522EEDAAFE51765A032F" blockId="74.[189,342,502,528]" box="[189,342,502,528]" pageId="74" pageNumber="75">
<heading id="CE0E701C3010522EEDAAFE51765A032F" bold="true" box="[189,342,502,528]" fontSize="11" level="3" pageId="74" pageNumber="75" reason="3">
<emphasis id="A78D1B623010522EEDAAFE51765A032F" bold="true" box="[189,342,502,528]" pageId="74" pageNumber="75">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="9546C7703010522EEDAAFD8276DE0300" blockId="74.[189,466,549,575]" box="[189,466,549,575]" pageId="74" pageNumber="75">
<collectingCountry id="EDEE87E03010522EEDAAFD82763A0300" box="[189,310,549,575]" name="Venezuela" pageId="74" pageNumber="75">Venezuela</collectingCountry>
and
<collectingCountry id="EDEE87E03010522EEC78FD8276C20300" box="[367,462,549,575]" name="Guyana" pageId="74" pageNumber="75">Guyana</collectingCountry>
.
</paragraph>
<paragraph id="9546C7703010522EEDAAFDC176E103BE" blockId="74.[189,493,614,641]" box="[189,493,614,641]" pageId="74" pageNumber="75">
<heading id="CE0E701C3010522EEDAAFDC176E103BE" bold="true" box="[189,493,614,641]" fontSize="11" level="3" pageId="74" pageNumber="75" reason="3">
<emphasis id="A78D1B623010522EEDAAFDC176E103BE" bold="true" box="[189,493,614,641]" pageId="74" pageNumber="75">
Subregions of
<collectingCountry id="EDEE87E03010522EEC78FDC076E103BE" box="[367,493,615,641]" name="Venezuela" pageId="74" pageNumber="75">Venezuela</collectingCountry>
</emphasis>
</heading>
</paragraph>
<paragraph id="9546C7703010522EEDAAFD3272430390" blockId="74.[189,1359,661,687]" box="[189,1359,661,687]" pageId="74" pageNumber="75">System of low mountains and hills Imataca-Cuyuní of Northeast Guiana Shield, and Guiana Shield.</paragraph>
<paragraph id="9546C7703010522EEDAAFD7176A803CE" blockId="74.[189,420,726,753]" box="[189,420,726,753]" pageId="74" pageNumber="75">
<heading id="CE0E701C3010522EEDAAFD7176A803CE" bold="true" box="[189,420,726,753]" fontSize="11" level="3" pageId="74" pageNumber="75" reason="3">
<emphasis id="A78D1B623010522EEDAAFD7176A803CE" bold="true" box="[189,420,726,753]" pageId="74" pageNumber="75">Literature records</emphasis>
</heading>
</paragraph>
<paragraph id="9546C7703010522EEDAAFCA2765D027D" blockId="74.[189,1398,773,834]" pageId="74" pageNumber="75">
Halffter &amp; Martínez, 1976: 40 (
<collectingCountry id="EDEE87E03010522EEF52FCA275C80220" box="[581,708,773,799]" name="Venezuela" pageId="74" pageNumber="75">Venezuela</collectingCountry>
:
<collectingRegion id="573D09923010522EEFC4FCA2743C0220" box="[723,816,773,799]" country="Venezuela" name="Bolivar" pageId="74" pageNumber="75">Bolívar</collectingRegion>
: Meseta de Sarisariñama). —
<bibRefCitation id="F168BA813010522EE9B0FCA2727A0220" author="Krajcik M." box="[1191,1398,773,799]" pageId="74" pageNumber="75" pagination="1 - 278" refId="ref126632" refString="Krajcik M. 2012. Checklist of the World Scarabaeoidea. Animma. X Supplement 5: 1 - 278." type="journal article" year="2012">Krajcik 2012: 88</bibRefCitation>
(
<collectingCountry id="EDEE87E03010522EEDD2FC8F7649027D" box="[197,325,808,834]" name="Venezuela" pageId="74" pageNumber="75">Venezuela</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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