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<document id="23EE5F4590B9EBB1316EB90DDFA494D7" ID-CLB-Dataset="305167" ID-DOI="10.5852/ejt.2024.959.2677" ID-GBIF-Dataset="bf132c8e-dbc7-4f3d-96e9-1a0b153e765a" ID-ISSN="2118-9773" ID-Zenodo-Dep="13945464" ID-ZooBank="30872C13-516F-42FA-AFA7-30ADC6BF1BAF" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1729166637250" checkinUser="plazi" docAuthor="Rosa, Cecilia Lozano De La, Cupello, Mario &amp; Vaz-De-Mello, Fernando Z." docDate="2024" docId="1D507666300C523CEF59FC0774FC03F8" docLanguage="en" docName="EJT.2024.959.1-272.pdf" docOrigin="European Journal of Taxonomy 959 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2677/12401" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Diabroctis cadmus" docType="treatment" docVersion="9" lastPageNumber="89" masterDocId="E1690E1E305A5264ED17FFA7770C013F" masterDocTitle="The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution" masterLastPageNumber="272" masterPageNumber="1" pageNumber="87" updateTime="1731534417997" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="E6A86209256A9CA5538F6F236613BEAA">
<mods:title id="1AE589945DA4BD185744FEBAFDB1E12B">The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution</mods:title>
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<mods:name id="9218E274383D59A40427255033969CED" type="personal">
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<mods:namePart id="63B216C13AD22D6CFD74D281F58FE378">Rosa, Cecilia Lozano De La</mods:namePart>
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<mods:affiliation id="4CECBAE31AA3201D136180AF9881E40D">Instituto Venezolano de Investigaciones Científicas, Centro de Estudios Botánicos y Agroforestales, Laboratorio de Ecología Espacial, Maracaibo, estado Zulia, Venezuela.</mods:affiliation>
<mods:nameIdentifier id="3774DEE77FA0D9DF024F4B3A11301CAB" type="email">lozanoceci@gmail.com</mods:nameIdentifier>
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<mods:roleTerm id="BAA422F18A5D13C3206655B61D3A4E07">Author</mods:roleTerm>
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<mods:namePart id="29C3B5709C1BF8E22049D7FE6021E1A9">Cupello, Mario</mods:namePart>
<mods:nameIdentifier id="EB5ADC8CD8E4679F4FEB96DDEE48E0CD" type="LSID">BDB03C18-7095-4EAA-8BDD-03CB4F79676D</mods:nameIdentifier>
<mods:affiliation id="EE0DC1E1D568C7164594AEA372FFC94F">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Curitiba, Paraná, Brazil. Currently at Department of Entomology, Texas A &amp; M University, College Station, TX, 77843, USA.</mods:affiliation>
<mods:nameIdentifier id="5D6F6EED48D1233C6D43F676075C2A72" type="email">mcupello@hotmail.com</mods:nameIdentifier>
</mods:name>
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<mods:namePart id="C650A3C8469D1CCCE8C55C75B190C57B">Vaz-De-Mello, Fernando Z.</mods:namePart>
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<mods:affiliation id="6E3D9AFA7A83B8583BB280689861C795">Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, Cuiabá, Mato Grosso, Brazil.</mods:affiliation>
<mods:nameIdentifier id="F5C4551CE9D0AF3E51E56DF6EC2592E3" type="email">vazdemello@gmail.com</mods:nameIdentifier>
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<mods:date id="AD7A49071A1B3C043072F5A15A95975C">2024</mods:date>
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<treatment id="1D507666300C523CEF59FC0774FC03F8" ID-DOI="http://doi.org/10.5281/zenodo.14154550" ID-GBIF-Taxon="246298087" ID-Zenodo-Dep="14154550" LSID="urn:lsid:plazi:treatment:1D507666300C523CEF59FC0774FC03F8" httpUri="http://treatment.plazi.org/id/1D507666300C523CEF59FC0774FC03F8" lastPageId="88" lastPageNumber="89" pageId="86" pageNumber="87">
<subSubSection id="DDE394FB300C5232EF59FC0774E90285" box="[590,997,928,955]" pageId="86" pageNumber="87" type="nomenclature">
<paragraph id="9546C770300C5232EF59FC0774E90285" blockId="86.[590,997,928,992]" box="[590,997,928,955]" pageId="86" pageNumber="87">
<heading id="CE0E701C300C5232EF59FC0774E90285" box="[590,997,928,955]" centered="true" fontSize="11" level="2" pageId="86" pageNumber="87" reason="2">
<taxonomicName id="52F9BCF3300C5232EF59FC0774E90285" ID-CoL="356T7" authority="(Harold, 1868)" baseAuthorityName="Harold" baseAuthorityYear="1868" box="[590,997,928,955]" class="Insecta" family="Scarabaeidae" genus="Diabroctis" kingdom="Animalia" order="Coleoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="cadmus">
<emphasis id="A78D1B62300C5232EF59FC07743C0284" bold="true" box="[590,816,928,955]" italics="true" pageId="86" pageNumber="87">Diabroctis cadmus</emphasis>
(Harold, 1868)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB300C5232EFF1FC62744302DF" box="[742,847,965,992]" pageId="86" pageNumber="87" type="description">
<paragraph id="9546C770300C5232EFF1FC62744302DF" blockId="86.[590,997,928,992]" box="[742,847,965,992]" pageId="86" pageNumber="87">
<figureCitation id="0DC2DBF5300C5232EFF1FC62744302DF" box="[742,847,965,992]" captionStart="Fig" captionStartId="87.[189,232,1963,1989]" captionTargetBox="[279,1302,278,1879]" captionTargetId="figure-12@87.[268,1320,265,1907]" captionTargetPageId="87" captionText="Fig. 35. Species distribution. A. Diabroctis cadmus (Harold, 1868). B. Diabroctis venezuelensis Martínez &amp; Clavijo, 1990. Green diamond = CEMT collection data; black circle = literature data." figureDoi="http://doi.org/10.5281/zenodo.13945559" httpUri="https://zenodo.org/record/13945559/files/figure.png" pageId="86" pageNumber="87">Fig. 35A</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB300C5232EDAAFBA076C5062A" pageId="86" pageNumber="87" type="reference_group">
<paragraph id="9546C770300C5232EDAAFBA07328057A" blockId="86.[189,1399,1031,1239]" pageId="86" pageNumber="87">
<treatmentCitationGroup id="B5E9E05E300C5232EDAAFBA07328057A" pageId="86" pageNumber="87">
<treatmentCitation id="1458E161300C5232EDAAFBA07569051E" author="Harold E." box="[189,613,1031,1057]" page="82" pageId="86" pageNumber="87" year="1868">
<taxonomicName id="52F9BCF3300C5232EDAAFBA07569051E" ID-CoL="4FSLT" authority="Harold, 1868 e: 82" authorityName="Harold" authorityPageNumber="82" authorityYear="1868" box="[189,613,1031,1057]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="cadmus">
<emphasis id="A78D1B62300C5232EDAAFBA0769C051E" box="[189,400,1031,1057]" italics="true" pageId="86" pageNumber="87">Phanaeus cadmus</emphasis>
<bibRefCitation id="F168BA81300C5232EC83FBA07569051E" author="Harold E." box="[404,613,1031,1057]" pageId="86" pageNumber="87" pagination="79 - 86" refId="ref123468" refString="Harold E. 1868 e. Diagnosen neuer Coprophagen. Coleopterologische Hefte 4: 79 - 86. Available from https: // www. biodiversitylibrary. org / page / 16069646 [accessed 16 Feb. 2024]." type="journal article" year="1868">Harold, 1868e: 82</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description). Type locality:
<collectingCountry id="EDEE87E0300C5232E903FBA073C8051E" box="[1044,1220,1031,1057]" name="French Guiana" pageId="86" pageNumber="87">French Guiana</collectingCountry>
. Name-bearing type:
<typeStatus id="4A4279D2300C5232EC3CFB8C769B057A" box="[299,407,1067,1093]" pageId="86" pageNumber="87" type="lectotype">lectotype</typeStatus>
(MNHN), designated by
<bibRefCitation id="F168BA81300C5232EFD6FB8C747E0579" author="Arnaud P." box="[705,882,1067,1094]" pageId="86" pageNumber="87" pagination="113 - 118" refId="ref109632" refString="Arnaud P. 1982. Liste des types de Phanaeini du Museum national d'Historie naturelle de Paris. Revue Francaise d'Entomologie 4 (3): 113 - 118." type="journal article" year="1982">Arnaud (1982)</bibRefCitation>
, not examined.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C770300C5232EDAAFBF7769C05E9" blockId="86.[189,1399,1031,1239]" pageId="86" pageNumber="87">
<treatmentCitationGroup id="B5E9E05E300C5232EDAAFBF7769C05E9" pageId="86" pageNumber="87">
<treatmentCitation id="1458E161300C5232EDAAFBF775BF0554" author="Ancey F. C." box="[189,691,1104,1131]" page="205" pageId="86" pageNumber="87" year="1880">
<taxonomicName id="52F9BCF3300C5232EDAAFBF775BF0554" ID-CoL="4FSN2" authority="Ancey, 1880: 205" authorityName="Ancey" authorityPageNumber="205" authorityYear="1880" box="[189,691,1104,1131]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="mimaeformis">
<emphasis id="A78D1B62300C5232EDAAFBF776D80555" box="[189,468,1104,1130]" italics="true" pageId="86" pageNumber="87">Phanaeus mimaeformis</emphasis>
<bibRefCitation id="F168BA81300C5232ECC9FBF775BF0554" author="Ancey F. C." box="[478,691,1104,1131]" pageId="86" pageNumber="87" pagination="205 - 206" refId="ref109388" refString="Ancey F. C. 1880. Description des Coleopteres nouveaux. Le Naturaliste 1 - 3: 205 - 206. Available from https: // www. biodiversitylibrary. org / page / 33928612 [accessed 14 Feb. 2024]." type="book chapter" year="1880">Ancey, 1880: 205</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description). Type locality: unknown; incorrectly stated as
<collectingCountry id="EDEE87E0300C5232EC41FBD376DA05B1" box="[342,470,1140,1166]" name="Costa Rica" pageId="86" pageNumber="87">Costa Rica</collectingCountry>
by
<bibRefCitation id="F168BA81300C5232EF16FBD375AE05B1" author="Ancey F. C." box="[513,674,1140,1166]" pageId="86" pageNumber="87" pagination="205 - 206" refId="ref109388" refString="Ancey F. C. 1880. Description des Coleopteres nouveaux. Le Naturaliste 1 - 3: 205 - 206. Available from https: // www. biodiversitylibrary. org / page / 33928612 [accessed 14 Feb. 2024]." type="book chapter" year="1880">Ancey (1880)</bibRefCitation>
based on mislabelled material, but the species does not occur there (
<bibRefCitation id="F168BA81300C5232EC24FB3F750E058D" author="Valois M. C. &amp; Harada L. &amp; Vaz-de-Mello F. Z. &amp; Silva F." box="[307,514,1176,1202]" pageId="86" pageNumber="87" pagination="347 - 374" refId="ref140212" refString="Valois M. C., Harada L., Vaz-de-Mello F. Z. &amp; Silva F. 2018. Synopsis of the genus Diabroctis Gistel, 1857 (Coleoptera: Scarabaeidae: Scarabaeinae) with a new species description. Insect Systematics &amp; Evolution 51: 347 - 374. https: // doi. org / 10.1163 / 1876312 X- 00002203" type="journal article" year="2018">
Valois
<emphasis id="A78D1B62300C5232EC94FB3E76B1058D" box="[387,445,1176,1202]" italics="true" pageId="86" pageNumber="87">et al.</emphasis>
2018
</bibRefCitation>
). Name-bearing type:
<typeStatus id="4A4279D2300C5232EE15FB3F7462058D" box="[770,878,1176,1202]" pageId="86" pageNumber="87" type="lectotype">lectotype</typeStatus>
(RBINS), designated by
<bibRefCitation id="F168BA81300C5232E986FB3F727E058D" author="Valois M. C. &amp; Harada L. &amp; Vaz-de-Mello F. Z. &amp; Silva F." box="[1169,1394,1176,1202]" pageId="86" pageNumber="87" pagination="347 - 374" refId="ref140212" refString="Valois M. C., Harada L., Vaz-de-Mello F. Z. &amp; Silva F. 2018. Synopsis of the genus Diabroctis Gistel, 1857 (Coleoptera: Scarabaeidae: Scarabaeinae) with a new species description. Insect Systematics &amp; Evolution 51: 347 - 374. https: // doi. org / 10.1163 / 1876312 X- 00002203" type="journal article" year="2018">
Valois
<emphasis id="A78D1B62300C5232E9F7FB3E7217058D" box="[1248,1307,1176,1202]" italics="true" pageId="86" pageNumber="87">et al.</emphasis>
(2018)
</bibRefCitation>
, not examined.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C770300C5232EDAAFB5875D4045D" blockId="86.[189,1399,1279,1813]" pageId="86" pageNumber="87">
<treatmentCitationGroup id="B5E9E05E300C5232EDAAFB5875D4045D" pageId="86" pageNumber="87">
<taxonomicName id="52F9BCF3300C5232EDAAFB5876960426" ID-CoL="4FSLT" authorityName="Harold" authorityYear="1868" box="[189,410,1279,1305]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="cadmus">
<emphasis id="A78D1B62300C5232EDAAFB5876960426" box="[189,410,1279,1305]" italics="true" pageId="86" pageNumber="87">Phanaeus cadmus</emphasis>
</taxonomicName>
<treatmentCitation id="1458E161300C5232ECD0FB5875CF0426" author="Harold E." box="[455,707,1279,1306]" page="1017" pageId="86" pageNumber="87" year="1869">
<bibRefCitation id="F168BA81300C5232ECD0FB5875CF0426" author="Harold E." box="[455,707,1279,1306]" pageId="86" pageNumber="87" refId="ref123629" refString="Harold E. 1869 d. Catalogus coleopterorum. Hucusque descriptorum, synonymicus et systematicus. Tom. IV. Scarabaeidae. E. H. Gummi, Munich. https: // doi. org / 10.5962 / bhl. title. 9089" type="book" year="1869">Harold 1869d: 1017</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="1458E161300C5232EE8EFB5873760426" author="Nevinson B. G." box="[921,1146,1279,1305]" page="2" pageId="86" pageNumber="87" year="1892">
<bibRefCitation id="F168BA81300C5232EE8EFB5873760426" author="Nevinson B. G." box="[921,1146,1279,1305]" pageId="86" pageNumber="87" refId="ref131974" refString="Nevinson B. G. 1892. Revised Synonymic List of Species in the Genera Phanӕus Macleay and Oxysternon Castelnau. Privately published, London. https: // doi. org / 10.5962 / bhl. title. 9416" type="book" year="1892">Nevinson 1892: 2</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="1458E161300C5232E813FAA776E20401" author="Heyne A. &amp; Taschenberg O." page="65" pageId="86" pageNumber="87" year="1908">
<bibRefCitation id="F168BA81300C5232E813FAA776E20401" author="Heyne A. &amp; Taschenberg O." pageId="86" pageNumber="87" refId="ref124102" refString="Heyne A. &amp; Taschenberg O. 1908. Die exotischen Kafer in Wort und Bild. J. F. Schreiber, Esslingen and Munich. https: // doi. org / 10.5962 / bhl. title. 9363" type="book" year="1908">Heyne &amp; Taschenberg 1908: 65</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="1458E161300C5232EF73FA8474680401" author="d'Olsoufieff G." box="[612,868,1315,1342]" page="15" pageId="86" pageNumber="87" year="1924">
<bibRefCitation id="F168BA81300C5232EF73FA8474680401" author="d'Olsoufieff G." box="[612,868,1315,1342]" pageId="86" pageNumber="87" pagination="4 - 172" refId="ref134049" refString="d'Olsoufieff G. 1924. Les Phanaeides, Famille Scarabaeidae. Tr. Coprini Insecta. Revue Illustree d'Entomologie 13: 4 - 172." type="journal article" year="1924">dOlsoufieff 1924: 15</bibRefCitation>
</treatmentCitation>
(comments).
<emphasis id="A78D1B62300C5232E91FFA83732E0402" box="[1032,1058,1316,1341]" italics="true" pageId="86" pageNumber="87"></emphasis>
<treatmentCitation id="1458E161300C5232E93CFA8473E00402" author="Gillet J. J. E." box="[1067,1260,1315,1341]" page="81" pageId="86" pageNumber="87" year="1911">
<bibRefCitation id="F168BA81300C5232E93CFA8473E00402" author="Gillet J. J. E." box="[1067,1260,1315,1341]" pageId="86" pageNumber="87" refId="ref121235" refString="Gillet J. J. E. 1911 b. Coleopterorum Catalogus. Pars 38: Scarabaeidae: Coprinae I. W. Junk, Berlin. Available from https: // www. biodiversitylibrary. org / page / 48405679 [accesseed 16 Feb. 2024]." type="url" year="1911">Gillet 1911b: 81</bibRefCitation>
</treatmentCitation>
(catalogue, cited for
<collectingCountry id="EDEE87E0300C5232EC42FAEF76C0045D" box="[341,460,1352,1378]" name="Colombia" pageId="86" pageNumber="87">Colombia</collectingCountry>
, Guianas,
<collectingCountry id="EDEE87E0300C5232EF50FAEF75C0045D" box="[583,716,1352,1378]" name="Costa Rica" pageId="86" pageNumber="87">Costa Rica</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C770300C5232EDAAFACB72190495" blockId="86.[189,1399,1279,1813]" pageId="86" pageNumber="87">
<treatmentCitationGroup id="B5E9E05E300C5232EDAAFACB72190495" pageId="86" pageNumber="87">
<taxonomicName id="52F9BCF3300C5232EDAAFACB76DA04B9" ID-CoL="4FSN2" authorityName="Ancey" authorityYear="1880" box="[189,470,1388,1414]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="mimaeformis">
<emphasis id="A78D1B62300C5232EDAAFACB76DA04B9" box="[189,470,1388,1414]" italics="true" pageId="86" pageNumber="87">Phanaeus mimaeformis</emphasis>
</taxonomicName>
<treatmentCitation id="1458E161300C5232ECE9FACB75CB04B9" author="Bates H. W." box="[510,711,1388,1414]" page="387" pageId="86" pageNumber="87" year="1887">
<bibRefCitation id="F168BA81300C5232ECE9FACB75CB04B9" author="Bates H. W." box="[510,711,1388,1414]" pageId="86" pageNumber="87" refId="ref110759" refString="Bates H. W. 1887. Tribe Lamellicornia. In: Bates H. W. (1886 - 1889) Biologia Centrali-Americana. Insecta. Coleoptera. Vol. II. Part 2. Pectinicornia and Lamellicornia: 25 - 64. Taylor and Francis, London. https: // doi. org / 10.5962 / bhl. title. 730 [Publication date according to Lyal (2011) and Bousquet (2016).]" type="book" year="1887">Bates 1887: 387</bibRefCitation>
</treatmentCitation>
(cited for
<collectingCountry id="EDEE87E0300C5232EE46FACB74D004B9" box="[849,988,1388,1414]" name="Costa Rica" pageId="86" pageNumber="87">Costa Rica</collectingCountry>
).
<emphasis id="A78D1B62300C5232EEE3FACA730204B9" box="[1012,1038,1389,1414]" italics="true" pageId="86" pageNumber="87"></emphasis>
<treatmentCitation id="1458E161300C5232E90DFACB73FA04B9" author="Nevinson B. G." box="[1050,1270,1388,1414]" page="2" pageId="86" pageNumber="87" year="1892">
<bibRefCitation id="F168BA81300C5232E90DFACB73FA04B9" author="Nevinson B. G." box="[1050,1270,1388,1414]" pageId="86" pageNumber="87" refId="ref131974" refString="Nevinson B. G. 1892. Revised Synonymic List of Species in the Genera Phanӕus Macleay and Oxysternon Castelnau. Privately published, London. https: // doi. org / 10.5962 / bhl. title. 9416" type="book" year="1892">Nevinson 1892: 2</bibRefCitation>
</treatmentCitation>
(as junior synonym of
<taxonomicName id="52F9BCF3300C5232EC6AFA3676F70495" ID-CoL="4FSLT" authorityName="Harold" authorityYear="1868" box="[381,507,1424,1450]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="cadmus">
<emphasis id="A78D1B62300C5232EC6AFA3676F70495" box="[381,507,1424,1450]" italics="true" pageId="86" pageNumber="87">D. cadmus</emphasis>
</taxonomicName>
). —
<treatmentCitation id="1458E161300C5232EF2FFA3774390495" author="d'Olsoufieff G." box="[568,821,1424,1450]" page="63" pageId="86" pageNumber="87" year="1924">
<bibRefCitation id="F168BA81300C5232EF2FFA3774390495" author="d'Olsoufieff G." box="[568,821,1424,1450]" pageId="86" pageNumber="87" pagination="4 - 172" refId="ref134049" refString="d'Olsoufieff G. 1924. Les Phanaeides, Famille Scarabaeidae. Tr. Coprini Insecta. Revue Illustree d'Entomologie 13: 4 - 172." type="journal article" year="1924">dOlsoufieff 1924: 63</bibRefCitation>
</treatmentCitation>
(cited as junior synonym of
<taxonomicName id="52F9BCF3300C5232E990FA3672080495" ID-CoL="4FSLT" authorityName="Harold" authorityYear="1868" box="[1159,1284,1424,1450]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="cadmus">
<emphasis id="A78D1B62300C5232E990FA3672080495" box="[1159,1284,1424,1450]" italics="true" pageId="86" pageNumber="87">D. cadmus</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C770300C5232EDAAFA12766F0728" blockId="86.[189,1399,1279,1813]" pageId="86" pageNumber="87">
<treatmentCitationGroup id="B5E9E05E300C5232EDAAFA12766F0728" pageId="86" pageNumber="87">
<emphasis id="A78D1B62300C5232EDAAFA1276CF04F1" box="[189,451,1460,1486]" italics="true" pageId="86" pageNumber="87">
<taxonomicName id="52F9BCF3300C5232EDAAFA1276A004F1" box="[189,428,1460,1486]" class="Insecta" family="Scarabaeidae" genus="Taurocopris" kingdom="Animalia" order="Coleoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="cadmus">Taurocopris cadmus</taxonomicName>
</emphasis>
<treatmentCitation id="1458E161300C5232ECDCFA1375C704F0" author="d'Olsoufieff G." box="[459,715,1460,1487]" page="22" pageId="86" pageNumber="87" year="1924">
<bibRefCitation id="F168BA81300C5232ECDCFA1375C704F0" author="d'Olsoufieff G." box="[459,715,1460,1487]" pageId="86" pageNumber="87" pagination="4 - 172" refId="ref134049" refString="d'Olsoufieff G. 1924. Les Phanaeides, Famille Scarabaeidae. Tr. Coprini Insecta. Revue Illustree d'Entomologie 13: 4 - 172." type="journal article" year="1924">dOlsoufieff 1924: 22</bibRefCitation>
</treatmentCitation>
, 62, 140 (key, comments, distribution). —
<treatmentCitation id="1458E161300C5232E9D0FA1277F704CC" author="Janssens A." page="2" pageId="86" pageNumber="87" year="1940">
<bibRefCitation id="F168BA81300C5232E9D0FA1277F704CC" author="Janssens A." pageId="86" pageNumber="87" refId="ref125561" refString="Janssens A. 1940. Contribution a l'etude des coleopteres lamellicornes coprophages. Bulletin du Musee royal d'Histoire naturelle de Belgique 16 (34): [1] - 7." type="journal volume" year="1940">Janssens 1940: 2</bibRefCitation>
</treatmentCitation>
, 4 (cited, key). —
<treatmentCitation id="1458E161300C5232ECC0FA7E75E404CC" author="Blackwelder R. E." box="[471,744,1497,1523]" page="208" pageId="86" pageNumber="87" year="1944">
<bibRefCitation id="F168BA81300C5232ECC0FA7E75E404CC" author="Blackwelder R. E." box="[471,744,1497,1523]" pageId="86" pageNumber="87" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 208</bibRefCitation>
</treatmentCitation>
(list, cited for
<collectingCountry id="EDEE87E0300C5232EE81FA7E731704CC" box="[918,1051,1497,1523]" name="Costa Rica" pageId="86" pageNumber="87">Costa Rica</collectingCountry>
). —
<treatmentCitation id="1458E161300C5232E945FA7E761A0727" author="Vulcano M. A. &amp; Pereira F. S." page="567" pageId="86" pageNumber="87" year="1967">
<bibRefCitation id="F168BA81300C5232E945FA7E761A0727" author="Vulcano M. A. &amp; Pereira F. S." pageId="86" pageNumber="87" pagination="533 - 603" refId="ref141365" refString="Vulcano M. A. &amp; Pereira F. S. 1967. Sinopse dos Passalidae e Scarabaeidae s. str. da regiao amazonica (Insecta, Coleoptera). Atas do Simposio sobre a Biota Amazonica 5: 533 - 603." type="journal article" year="1967">Vulcano &amp; Pereira 1967: 567</bibRefCitation>
</treatmentCitation>
(key).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C770300C5232EDAAF98676C5062A" blockId="86.[189,1399,1279,1813]" pageId="86" pageNumber="87">
<treatmentCitationGroup id="B5E9E05E300C5232EDAAF98676C5062A" pageId="86" pageNumber="87">
<emphasis id="A78D1B62300C5232EDAAF98676B50704" box="[189,441,1569,1595]" italics="true" pageId="86" pageNumber="87">
<taxonomicName id="52F9BCF3300C5232EDAAF98676930704" ID-CoL="356T7" baseAuthorityName="Harold" baseAuthorityYear="1868" box="[189,415,1569,1595]" class="Insecta" family="Scarabaeidae" genus="Diabroctis" kingdom="Animalia" order="Coleoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="cadmus">Diabroctis cadmus</taxonomicName>
</emphasis>
<treatmentCitation id="1458E161300C5232ECD2F98675B50704" author="Edmonds W. D." box="[453,697,1569,1595]" page="821" pageId="86" pageNumber="87" year="1972">
<bibRefCitation id="F168BA81300C5232ECD2F98675B50704" author="Edmonds W. D." box="[453,697,1569,1595]" firstAuthor="Edmonds" journalOrPublisher="The University of Kansas Science Bulletin" pageId="86" pageNumber="87" pagination="731 - 874" part="49" refId="ref117282" refString="Edmonds W. D. 1972. Comparative skeletal morphology, systematics and evolution of the Phanaeine dung beetles (Coleoptera: Scarabaeidae). The University of Kansas Science Bulletin 49: 731 - 874." title="Comparative skeletal morphology, systematics and evolution of the Phanaeine dung beetles (Coleoptera: Scarabaeidae)" type="journal article" year="1972">Edmonds 1972: 821</bibRefCitation>
</treatmentCitation>
(cited). —
<treatmentCitation id="1458E161300C5232EE5BF98673250703" author="Arnaud P." box="[844,1065,1569,1596]" page="115" pageId="86" pageNumber="87" year="1982">
<bibRefCitation id="F168BA81300C5232EE5BF98673250703" author="Arnaud P." box="[844,1065,1569,1596]" pageId="86" pageNumber="87" pagination="113 - 118" refId="ref109632" refString="Arnaud P. 1982. Liste des types de Phanaeini du Museum national d'Historie naturelle de Paris. Revue Francaise d'Entomologie 4 (3): 113 - 118." type="journal article" year="1982">Arnaud 1982: 115</bibRefCitation>
</treatmentCitation>
(list of
<typeStatus id="4A4279D2300C5232E985F98573DF0703" box="[1170,1235,1570,1596]" pageId="86" pageNumber="87">types</typeStatus>
, MNHN). —
<treatmentCitation id="1458E161300C5232EDFDF9E17540075F" author="Martinez A. &amp; Martinez A." box="[234,588,1605,1632]" page="151" pageId="86" pageNumber="87" year="1990">
<bibRefCitation id="F168BA81300C5232EDFDF9E17540075F" author="Martinez A. &amp; Martinez A." box="[234,588,1605,1632]" pageId="86" pageNumber="87" pagination="159 - 163" refId="ref130132" refString="Martinez A. &amp; Martinez A. 1990. Dos nuevas especies de Ateuchus Weber (Col. Scarab. Ateuchina). Boletin de Entomologia Venezolana (N. S.) 5 (20): 159 - 163." type="journal article" year="1990">Martínez &amp; Clavijo 1990: 151</bibRefCitation>
</treatmentCitation>
(key, comments). —
<treatmentCitation id="1458E161300C5232EE5EF9E27328075F" author="Escobar F." box="[841,1060,1605,1632]" page="208" pageId="86" pageNumber="87" year="2000">
<bibRefCitation id="F168BA81300C5232EE5EF9E27328075F" author="Escobar F." box="[841,1060,1605,1632]" pageId="86" pageNumber="87" pagination="197 - 210" refId="ref118039" refString="Escobar F. 2000. Diversidad y distribution de los escarabajos del estiercol (Coleoptera: Scarabaeidae: Scarabaeinae) de Colombia. In: Martin-Piera F., Morrone J. J. &amp; Melic A. (eds.) Hacia un proyecto CYTED para el Inventario y Estimacion de la Diversidad Entomologica en Iberoamerica: PRIBES- 2000: 197 - 210. m 3 m: Monografias Tercer Milenio, vol. 1. Sociedad Entomologica Aragonesa (SEA), Zaragoza." type="book chapter" year="2000">Escobar 2000: 208</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E0300C5232E9DAF9E2724A0760" box="[1229,1350,1605,1631]" name="Colombia" pageId="86" pageNumber="87">Colombia</collectingCountry>
). —
<treatmentCitation id="1458E161300C5232EDFDF9CD751A07BB" author="Gamez J. &amp; Mora E." box="[234,534,1642,1668]" page="17" pageId="86" pageNumber="87" year="2000">
<bibRefCitation id="F168BA81300C5232EDFDF9CD751A07BB" author="Gamez J. &amp; Mora E." box="[234,534,1642,1668]" pageId="86" pageNumber="87" pagination="3 - 16" refId="ref120147" refString="Gamez J. &amp; Mora E. 2000. Aspectos autoecologicos y distribucion local de Phanaeus (Phanaeus) prasinus Harold, 1868 (Coleoptera: Scarabaeidae: Scarabaeinae) en Venezuela. Memorias de la Fundacion La Salle de Ciencias Naturales 60 (153): 3 - 16." type="journal article" year="2000">Gámez &amp; Mora 2000: 17</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="1458E161300C5232EF9BF9CD745B07BA" author="Vitolo A. L." box="[652,855,1642,1669]" page="595" pageId="86" pageNumber="87" year="2000">
<bibRefCitation id="F168BA81300C5232EF9BF9CD745B07BA" author="Vitolo A. L." box="[652,855,1642,1669]" pageId="86" pageNumber="87" pagination="591 - 601" refId="ref141219" refString="Vitolo A. L. 2000. Clave para la identificacion de los generos y especies Phanaeinas de Colombia. Revista de la Academia Colombiana de Ciencias Exactas Fisicas y Naturales 24 (93): 591 - 601." type="journal article" year="2000">Vítolo 2000: 595</bibRefCitation>
</treatmentCitation>
(key);
<treatmentCitation id="1458E161300C5232EEB8F9CD732A07BB" author="Vitolo A." box="[943,1062,1642,1668]" page="279" pageId="86" pageNumber="87" year="2004">
<bibRefCitation id="F168BA81300C5232EEB8F9CD732A07BB" author="Vitolo A." box="[943,1062,1642,1668]" pageId="86" pageNumber="87" pagination="277 - 318" refId="ref141259" refString="Vitolo A. 2004. Escarabajos estercoleros de la tribu Phanaeini de Colombia (Coleoptera: Scarabaeoidea: Scarabaeidae). In: Fernandez F., Andrade M. G. &amp; Amat G. D. (eds) Insectos de Colombia 3: 277 - 318. Facultad de Ciencias, Universidad Nacional de Colombia, Bogota." type="book chapter" year="2004">2004: 279</bibRefCitation>
</treatmentCitation>
(diagnosis, distribution). —
<treatmentCitation id="1458E161300C5232EDFBF92976B50797" author="Arnaud P." box="[236,441,1678,1704]" page="18" pageId="86" pageNumber="87" year="2002">
<bibRefCitation id="F168BA81300C5232EDFBF92976B50797" author="Arnaud P." box="[236,441,1678,1704]" pageId="86" pageNumber="87" refId="ref109736" refString="Arnaud P. 2002 b. Les Coleopteres du Monde, Vol. 28. Phanaeini. Dendropaemon, Tetramereia, Homalotarsus, Megatharsis, Diabroctis, Coprophanaeus, Oxysternon, Phanaeus, Sulcophanaeus. Hillside Books, Canterbury, England." type="book" year="2002">Arnaud2002b:18</bibRefCitation>
</treatmentCitation>
(key, distribution).—
<treatmentCitation id="1458E161300C5232EFA1F929747F0797" author="Gamez J." box="[694,883,1678,1704]" page="48" pageId="86" pageNumber="87" year="2004">
<bibRefCitation id="F168BA81300C5232EFA1F929747F0797" author="Gamez J." box="[694,883,1678,1704]" pageId="86" pageNumber="87" pagination="43 - 60" refId="ref119830" refString="Gamez J. 2004 (&quot; 2002 &quot;). Phanaeini (Coleoptera: Scarabaeinae) de la Cordillera de los Andes, depresion de Maracaibo y Llanos de Venezuela. Memoria de la Fundacion La Salle de Ciencias Naturales 158: 43 - 60." type="journal article" year="2004">Gámez 2004:48</bibRefCitation>
</treatmentCitation>
, 51, 52,58 (list, distribution, key).—
<treatmentCitation id="1458E161300C5232E80FF92976AD07F3" author="Noriega J. A. &amp; Solis C. &amp; Escobar F. &amp; Realpe E." page="83" pageId="86" pageNumber="87" year="2007">
<bibRefCitation id="F168BA81300C5232E80FF92976AD07F3" author="Noriega J. A. &amp; Solis C. &amp; Escobar F. &amp; Realpe E." pageId="86" pageNumber="87" pagination="77 - 86" refId="ref132574" refString="Noriega J. A., Solis C., Escobar F. &amp; Realpe E. 2007. Escarabajos coprofagos (Coleoptera: Scarabaeidae) de la provincia de la Sierra Nevada de Santa Marta. Biota Colombiana 8 (1): 77 - 86." type="journal article" year="2007">
Noriega
<emphasis id="A78D1B62300C5232EDF0F914762507F3" box="[231,297,1714,1740]" italics="true" pageId="86" pageNumber="87">et al.</emphasis>
2007: 83
</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="1458E161300C5232EF37F915743007F3" author="Giraldo C. &amp; Montoya S. &amp; Escobar F." box="[544,828,1714,1740]" page="22" pageId="86" pageNumber="87" year="2018">
<bibRefCitation id="F168BA81300C5232EF37F915743007F3" author="Giraldo C. &amp; Montoya S. &amp; Escobar F." box="[544,828,1714,1740]" pageId="86" pageNumber="87" refId="ref121556" refString="Giraldo C., Montoya S. &amp; Escobar F. 2018. Escarabajos del Estiercol en Paisajes Ganaderos de Colombia. Fundacion CIPAV, Cali, Colombia." type="book" year="2018">
Giraldo
<emphasis id="A78D1B62300C5232EF92F91475C907F3" box="[645,709,1714,1740]" italics="true" pageId="86" pageNumber="87">et al.</emphasis>
2018: 22
</bibRefCitation>
</treatmentCitation>
(guide). —
<treatmentCitation id="1458E161300C5232EECFF91573F907F2" author="Valois M. C. &amp; Harada L. &amp; Vaz-de-Mello F. Z. &amp; Silva F." box="[984,1269,1714,1741]" page="351" pageId="86" pageNumber="87" year="2018">
<bibRefCitation id="F168BA81300C5232EECFF91573F907F2" author="Valois M. C. &amp; Harada L. &amp; Vaz-de-Mello F. Z. &amp; Silva F." box="[984,1269,1714,1741]" pageId="86" pageNumber="87" pagination="347 - 374" refId="ref140212" refString="Valois M. C., Harada L., Vaz-de-Mello F. Z. &amp; Silva F. 2018. Synopsis of the genus Diabroctis Gistel, 1857 (Coleoptera: Scarabaeidae: Scarabaeinae) with a new species description. Insect Systematics &amp; Evolution 51: 347 - 374. https: // doi. org / 10.1163 / 1876312 X- 00002203" type="journal article" year="2018">
Valois
<emphasis id="A78D1B62300C5232E93AF914736007F3" box="[1069,1132,1714,1740]" italics="true" pageId="86" pageNumber="87">et al.</emphasis>
2018: 351
</bibRefCitation>
</treatmentCitation>
, 365 (key, diagnosis). —
<treatmentCitation id="1458E161300C5232EC8BF971742B07CE" ID-DOI="http://doi.org/10.5281/zenodo.3683843" ID-Zenodo-Dep="3683843" author="Hielkema A. J. &amp; Hielkema M. A." box="[412,807,1750,1777]" httpUri="http://treatment.plazi.org/id/03D7FF1CFFAEFF8BFF48FC5D330EFD69" page="100" pageId="86" pageNumber="87" year="2019">
<bibRefCitation id="F168BA81300C5232EC8BF971742B07CE" author="Hielkema A. J. &amp; Hielkema M. A." box="[412,807,1750,1777]" pageId="86" pageNumber="87" pagination="1 - 306" refId="ref124225" refString="Hielkema A. J. &amp; Hielkema M. A. 2019. An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas. Insecta Mundi 732 (732): 1 - 306." type="journal article" year="2019">Hielkema &amp; Hielkema 2019:100</bibRefCitation>
</treatmentCitation>
, 292 (catalogue for the Guianas). —
<treatmentCitation id="1458E161300C5232E9F9F971766C062A" author="Nieto M. J. A. &amp; Giraldo E. C. &amp; Quevedo V. C. J. &amp; Chara J. &amp; Medina A. C." page="136" pageId="86" pageNumber="87" year="2020">
<bibRefCitation id="F168BA81300C5232E9F9F971766C062A" author="Nieto M. J. A. &amp; Giraldo E. C. &amp; Quevedo V. C. J. &amp; Chara J. &amp; Medina A. C." pageId="86" pageNumber="87" pagination="134 - 141" refId="ref132145" refString="Nieto M. J. A., Giraldo E. C., Quevedo V. C. J., Chara J. &amp; Medina A. C. 2020. Escarabajos estercoleros asociados a sistemas de ganaderia sostenible en diferentes regiones de Colombia. Biota Colombiana 21: 134 - 141. https: // doi. org / 10.21068 / c 2020. v 21 n 02 a 09" type="journal article" year="2020">
Nieto
<emphasis id="A78D1B62300C5232E82EF970727B07CF" box="[1337,1399,1750,1776]" italics="true" pageId="86" pageNumber="87">et al.</emphasis>
2020: 136
</bibRefCitation>
</treatmentCitation>
(report).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB300C5232EDAAF89A766B0694" pageId="86" pageNumber="87" type="materials_examined">
<paragraph id="9546C770300C5232EDAAF89A76A60668" blockId="86.[189,426,1853,1879]" box="[189,426,1853,1879]" pageId="86" pageNumber="87">
<heading id="CE0E701C300C5232EDAAF89A76A60668" bold="true" box="[189,426,1853,1879]" fontSize="11" level="3" pageId="86" pageNumber="87" reason="3">
<emphasis id="A78D1B62300C5232EDAAF89A76A60668" bold="true" box="[189,426,1853,1879]" pageId="86" pageNumber="87">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="9546C770300C5232EDAAF8CA766B0694" blockId="86.[189,1399,1901,1963]" pageId="86" pageNumber="87">
<materialsCitation id="2591CD2D300C5232EDAAF8CA766E0694" ID-GBIF-Occurrence="4979307467" collectedFrom="faeces" collectingDate="2006-07-20" collectionCode="CEMT" collectorName="curso NM 2006" country="Venezuela" location="Rosario de Perija" pageId="86" pageNumber="87" specimenCount="1" stateProvince="Zulia">
<collectingCountry id="EDEE87E0300C5232EDAAF8CA766706B8" box="[189,363,1901,1927]" name="Venezuela" pageId="86" pageNumber="87">VENEZUELA</collectingCountry>
<emphasis id="A78D1B62300C5232EC9BF8CA76C306B8" bold="true" box="[396,463,1901,1927]" pageId="86" pageNumber="87">
<collectingRegion id="573D0992300C5232EC9BF8CA76C306B8" box="[396,463,1901,1927]" country="Venezuela" name="Zulia" pageId="86" pageNumber="87">Zulia</collectingRegion>
</emphasis>
<specimenCount id="83FF0CF9300C5232ECFAF8CA754906B8" box="[493,581,1901,1927]" count="1" pageId="86" pageNumber="87" type="generic">1 spec.</specimenCount>
;
<location id="902691AB300C5232EF43F8CA742A06B8" LSID="urn:lsid:plazi:treatment:1D507666300C523CEF59FC0774FC03F8:902691AB300C5232EF43F8CA742A06B8" box="[596,806,1901,1927]" country="Venezuela" name="Rosario de Perija" pageId="86" pageNumber="87" stateProvince="Zulia">Rosario de Perijá</location>
;
<date id="E147E1B0300C5232EE22F8CA74C206B8" box="[821,974,1901,1927]" pageId="86" pageNumber="87" value="2006-07-20">
<collectingDate id="F1031858300C5232EE22F8CA74C206B8" box="[821,974,1901,1927]" pageId="86" pageNumber="87" value="2006-07-20">20 Jul. 2006</collectingDate>
</date>
;
<collectorName id="380CA2A6300C5232EECAF8CA739C06B8" box="[989,1168,1901,1927]" pageId="86" pageNumber="87">curso NM2006</collectorName>
leg.;
<collectedFrom id="26E5848D300C5232E9C0F8CA722F06B8" box="[1239,1315,1901,1927]" pageId="86" pageNumber="87">faeces</collectedFrom>
, 13h 00:00;
<collectionCode id="F3E85FB5300C5232EC18F836766E0694" box="[271,354,1937,1963]" pageId="86" pageNumber="87">CEMT</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="C18697F8300D5233EDAAF80C723806D7" ID-DOI="http://doi.org/10.5281/zenodo.13945559" ID-Zenodo-Dep="13945559" httpUri="https://zenodo.org/record/13945559/files/figure.png" pageId="87" pageNumber="88" startId="87.[189,232,1963,1989]" targetBox="[279,1302,278,1879]" targetPageId="87">
<paragraph id="9546C770300D5233EDAAF80C723806D7" blockId="87.[189,1399,1963,2025]" pageId="87" pageNumber="88">
<emphasis id="A78D1B62300D5233EDAAF80C762E06FA" bold="true" box="[189,290,1963,1989]" pageId="87" pageNumber="88">Fig. 35.</emphasis>
Species distribution.
<emphasis id="A78D1B62300D5233EF20F80C754006FA" bold="true" box="[567,588,1963,1989]" pageId="87" pageNumber="88">A</emphasis>
.
<taxonomicName id="52F9BCF3300D5233EF75F80C730306FA" authority="(Harold, 1868)" baseAuthorityName="Harold" baseAuthorityYear="1868" box="[610,1039,1963,1989]" class="Insecta" family="Scarabaeidae" genus="Diabroctis" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="cadmus">
<emphasis id="A78D1B62300D5233EF75F80C744B06FA" box="[610,839,1963,1989]" italics="true" pageId="87" pageNumber="88">Diabroctis cadmus</emphasis>
(Harold, 1868)
</taxonomicName>
.
<emphasis id="A78D1B62300D5233E935F80C733A06FA" bold="true" box="[1058,1078,1963,1989]" pageId="87" pageNumber="88">B</emphasis>
.
<taxonomicName id="52F9BCF3300D5233E95BF80C76E306D7" authority="Martinez &amp; Clavijo, 1990" authorityName="Martinez &amp; Clavijo" authorityYear="1990" class="Insecta" family="Scarabaeidae" genus="Diabroctis" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="venezuelensis">
<emphasis id="A78D1B62300D5233E95BF80C727B06FA" box="[1100,1399,1963,1989]" italics="true" pageId="87" pageNumber="88">Diabroctis venezuelensis</emphasis>
<bibRefCitation id="F168BA81300D5233EDAAF86976E306D7" author="Martinez A. &amp; Martinez A." box="[189,495,1998,2024]" pageId="87" pageNumber="88" pagination="159 - 163" refId="ref130132" refString="Martinez A. &amp; Martinez A. 1990. Dos nuevas especies de Ateuchus Weber (Col. Scarab. Ateuchina). Boletin de Entomologia Venezolana (N. S.) 5 (20): 159 - 163." type="journal article" year="1990">Martínez &amp; Clavijo, 1990</bibRefCitation>
</taxonomicName>
. Green diamond = CEMT collection data; black circle = literature data.
</paragraph>
</caption>
<subSubSection id="DDE394FB3002523CEDAAFEAF730200FB" pageId="88" pageNumber="89" type="distribution">
<paragraph id="9546C7703002523CEDAAFEAF765A001D" blockId="88.[189,342,264,290]" box="[189,342,264,290]" pageId="88" pageNumber="89">
<heading id="CE0E701C3002523CEDAAFEAF765A001D" bold="true" box="[189,342,264,290]" fontSize="11" level="3" pageId="88" pageNumber="89" reason="3">
<emphasis id="A78D1B623002523CEDAAFEAF765A001D" bold="true" box="[189,342,264,290]" pageId="88" pageNumber="89">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="9546C7703002523CEDAAFE9F76E6006D" blockId="88.[189,490,312,338]" box="[189,490,312,338]" pageId="88" pageNumber="89">
<collectingCountry id="EDEE87E03002523CEDAAFE9F763D006D" box="[189,305,312,338]" name="Colombia" pageId="88" pageNumber="89">Colombia</collectingCountry>
and
<collectingCountry id="EDEE87E03002523CEC7EFE9F76EA006D" box="[361,486,312,338]" name="Venezuela" pageId="88" pageNumber="89">Venezuela</collectingCountry>
.
</paragraph>
<paragraph id="9546C7703002523CEDABFEDD76E100AA" blockId="88.[188,493,378,405]" box="[188,493,378,405]" pageId="88" pageNumber="89">
<heading id="CE0E701C3002523CEDABFEDD76E100AA" bold="true" box="[188,493,378,405]" fontSize="11" level="3" pageId="88" pageNumber="89" reason="3">
<emphasis id="A78D1B623002523CEDABFEDD76E100AA" bold="true" box="[188,493,378,405]" pageId="88" pageNumber="89">
Subregions of
<collectingCountry id="EDEE87E03002523CEC78FEDC76E100AA" box="[367,493,379,405]" name="Venezuela" pageId="88" pageNumber="89">Venezuela</collectingCountry>
</emphasis>
</heading>
</paragraph>
<paragraph id="9546C7703002523CEDAAFE0D730200FB" blockId="88.[189,1038,426,452]" box="[189,1038,426,452]" pageId="88" pageNumber="89">Maracaibo Depression, and System of hills and low sierras Lara-Falcón.</paragraph>
</subSubSection>
<subSubSection id="DDE394FB3002523CEDAAFE4B74FC03F8" pageId="88" pageNumber="89" type="reference_group">
<paragraph id="9546C7703002523CEDAAFE4B74FC03F8" blockId="88.[189,420,492,519]" lastBlockId="88.[189,1399,540,712]" pageId="88" pageNumber="89">
<heading id="CE0E701C3002523CEDAAFE4B76A80338" bold="true" box="[189,420,492,519]" fontSize="11" level="3" pageId="88" pageNumber="89" reason="3">
<emphasis id="A78D1B623002523CEDAAFE4B76A80338" bold="true" box="[189,420,492,519]" pageId="88" pageNumber="89">Literature records</emphasis>
</heading>
<bibRefCitation id="F168BA813002523CEDAAFDBB76C60309" author="d'Olsoufieff G." box="[189,458,540,566]" pageId="88" pageNumber="89" pagination="4 - 172" refId="ref134049" refString="d'Olsoufieff G. 1924. Les Phanaeides, Famille Scarabaeidae. Tr. Coprini Insecta. Revue Illustree d'Entomologie 13: 4 - 172." type="journal article" year="1924">dOlsoufieff 1924: 62</bibRefCitation>
, 140 (
<collectingCountry id="EDEE87E03002523CEF09FDBB75920309" box="[542,670,540,566]" name="Venezuela" pageId="88" pageNumber="89">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA813002523CEFF5FDBA74960308" author="Roze J. A." box="[738,922,540,567]" pageId="88" pageNumber="89" pagination="39 - 63" refId="ref137914" refString="Roze J. A. 1955. Lista preliminar de la familia Scarabaeidae sensu lato de Venezuela. Boletin del Museo de Ciencias Naturales 1 (1): 39 - 63." type="journal article" year="1955">Roze 1955: 45</bibRefCitation>
(
<collectingCountry id="EDEE87E03002523CEEA7FDBB73230309" box="[944,1071,540,566]" name="Venezuela" pageId="88" pageNumber="89">Venezuela</collectingCountry>
:
<collectingRegion id="573D09923002523CE957FDBB73980309" box="[1088,1172,540,566]" country="Venezuela" name="Falcon" pageId="88" pageNumber="89">Falcón</collectingRegion>
: Buena Vista). —
<bibRefCitation id="F168BA813002523CEDAAFDE6751C0364" author="Martinez A. &amp; Martinez A." box="[189,528,576,603]" pageId="88" pageNumber="89" pagination="159 - 163" refId="ref130132" refString="Martinez A. &amp; Martinez A. 1990. Dos nuevas especies de Ateuchus Weber (Col. Scarab. Ateuchina). Boletin de Entomologia Venezolana (N. S.) 5 (20): 159 - 163." type="journal article" year="1990">Martínez &amp; Clavijo 1990: 5</bibRefCitation>
(
<collectingCountry id="EDEE87E03002523CEF35FDE775AD0365" box="[546,673,576,602]" name="Venezuela" pageId="88" pageNumber="89">Venezuela</collectingCountry>
:
<collectingRegion id="573D09923002523CEFB8FDE775FD0365" box="[687,753,576,602]" country="Venezuela" name="Zulia" pageId="88" pageNumber="89">Zulia</collectingRegion>
: Maracaibo, El Guaco). —
<bibRefCitation id="F168BA813002523CE954FDE6727A0364" author="Gamez J. &amp; Mora E." box="[1091,1398,577,603]" pageId="88" pageNumber="89" pagination="3 - 16" refId="ref120147" refString="Gamez J. &amp; Mora E. 2000. Aspectos autoecologicos y distribucion local de Phanaeus (Phanaeus) prasinus Harold, 1868 (Coleoptera: Scarabaeidae: Scarabaeinae) en Venezuela. Memorias de la Fundacion La Salle de Ciencias Naturales 60 (153): 3 - 16." type="journal article" year="2000">Gámez &amp; Mora 2000: 17</bibRefCitation>
(
<collectingCountry id="EDEE87E03002523CEDD2FDC276490340" box="[197,325,613,639]" name="Venezuela" pageId="88" pageNumber="89">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA813002523CEC6DFDC275420340" author="Arnaud P." box="[378,590,613,639]" pageId="88" pageNumber="89" refId="ref109736" refString="Arnaud P. 2002 b. Les Coleopteres du Monde, Vol. 28. Phanaeini. Dendropaemon, Tetramereia, Homalotarsus, Megatharsis, Diabroctis, Coprophanaeus, Oxysternon, Phanaeus, Sulcophanaeus. Hillside Books, Canterbury, England." type="book" year="2002">Arnaud 2002b: 18</bibRefCitation>
(
<collectingCountry id="EDEE87E03002523CEF4AFDC275D00340" box="[605,732,613,639]" name="Venezuela" pageId="88" pageNumber="89">Venezuela</collectingCountry>
:
<collectingRegion id="573D09923002523CEFF0FDC274260340" box="[743,810,613,639]" country="Venezuela" name="Zulia" pageId="88" pageNumber="89">Zulia</collectingRegion>
). —
<bibRefCitation id="F168BA813002523CEE76FDC2732E03BF" author="Gamez J." box="[865,1058,613,640]" pageId="88" pageNumber="89" pagination="43 - 60" refId="ref119830" refString="Gamez J. 2004 (&quot; 2002 &quot;). Phanaeini (Coleoptera: Scarabaeinae) de la Cordillera de los Andes, depresion de Maracaibo y Llanos de Venezuela. Memoria de la Fundacion La Salle de Ciencias Naturales 158: 43 - 60." type="journal article" year="2004">Gámez 2004: 58</bibRefCitation>
(
<collectingCountry id="EDEE87E03002523CE926FDC273BC0340" box="[1073,1200,613,639]" name="Venezuela" pageId="88" pageNumber="89">Venezuela</collectingCountry>
:
<collectingRegion id="573D09923002523CE9ADFDC273F00340" box="[1210,1276,613,639]" country="Venezuela" name="Zulia" pageId="88" pageNumber="89">Zulia</collectingRegion>
: Mara and El Guaco). —
<bibRefCitation id="F168BA813002523CEC7EFD2E753E039C" author="Vitolo A." box="[361,562,649,675]" pageId="88" pageNumber="89" pagination="277 - 318" refId="ref141259" refString="Vitolo A. 2004. Escarabajos estercoleros de la tribu Phanaeini de Colombia (Coleoptera: Scarabaeoidea: Scarabaeidae). In: Fernandez F., Andrade M. G. &amp; Amat G. D. (eds) Insectos de Colombia 3: 277 - 318. Facultad de Ciencias, Universidad Nacional de Colombia, Bogota." type="book chapter" year="2004">Vítolo 2004: 279</bibRefCitation>
(
<collectingCountry id="EDEE87E03002523CEF55FD2E75CD039C" box="[578,705,649,675]" name="Venezuela" pageId="88" pageNumber="89">Venezuela</collectingCountry>
:
<collectingRegion id="573D09923002523CEFDAFD2E741C039C" box="[717,784,649,675]" country="Venezuela" name="Zulia" pageId="88" pageNumber="89">Zulia</collectingRegion>
). —
<bibRefCitation id="F168BA813002523CEE5EFD2E739A039C" author="Ferrer-Paris J. &amp; Sanchez-Mercado A. &amp; Rodriguez J. P." box="[841,1174,649,676]" pageId="88" pageNumber="89" pagination="89 - 110" refId="ref119203" refString="Ferrer-Paris J. R, Sanchez-Mercado A. &amp; Rodriguez J. P. 2013. Optimizacion del muestreo de invertebrados tropicales: un ejemplo con escarabajos coprofagos (Coleoptera: Scarabaeinae) en Venezuela. Revista de Biologia Tropical 61 (1): 89 - 110. https: // doi. org / 10.15517 / rbt. v 61 i 1.10941" type="journal article" year="2013">
Ferrer-Paris
<emphasis id="A78D1B623002523CEECAFD2D731C039C" box="[989,1040,649,675]" italics="true" pageId="88" pageNumber="89">et al</emphasis>
. 2013: 109
</bibRefCitation>
(
<collectingCountry id="EDEE87E03002523CE9B1FD2E7229039C" box="[1190,1317,649,675]" name="Venezuela" pageId="88" pageNumber="89">Venezuela</collectingCountry>
:
<collectingRegion id="573D09923002523CE827FD2E727E039C" box="[1328,1394,649,675]" country="Venezuela" name="Zulia" pageId="88" pageNumber="89">Zulia</collectingRegion>
: Rosario de Perijá). —
<bibRefCitation id="F168BA813002523CECD3FD0A75C303F7" author="Valois M. C. &amp; Harada L. &amp; Vaz-de-Mello F. Z. &amp; Silva F." box="[452,719,685,712]" pageId="88" pageNumber="89" pagination="347 - 374" refId="ref140212" refString="Valois M. C., Harada L., Vaz-de-Mello F. Z. &amp; Silva F. 2018. Synopsis of the genus Diabroctis Gistel, 1857 (Coleoptera: Scarabaeidae: Scarabaeinae) with a new species description. Insect Systematics &amp; Evolution 51: 347 - 374. https: // doi. org / 10.1163 / 1876312 X- 00002203" type="journal article" year="2018">
Valois
<emphasis id="A78D1B623002523CEF03FD09754403F8" box="[532,584,685,711]" italics="true" pageId="88" pageNumber="89">et al</emphasis>
. 2018: 350
</bibRefCitation>
, 365 (
<collectingCountry id="EDEE87E03002523CEE01FD0A749903F8" box="[790,917,685,711]" name="Venezuela" pageId="88" pageNumber="89">Venezuela</collectingCountry>
:
<collectingRegion id="573D09923002523CEEB7FD0A74EF03F8" box="[928,995,685,711]" country="Venezuela" name="Zulia" pageId="88" pageNumber="89">Zulia</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>