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<document id="B5BE65B63977242F6FF6221D9D8286A8" ID-DOI="10.5852/ejt.2024.959.2677" ID-ISSN="2118-9773" ID-Zenodo-Dep="13945464" ID-ZooBank="30872C13-516F-42FA-AFA7-30ADC6BF1BAF" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1729166637250" checkinUser="plazi" docAuthor="Rosa, Cecilia Lozano De La, Cupello, Mario &amp; Vaz-De-Mello, Fernando Z." docDate="2024" docId="1D5076663002523DEF4CF953732906E4" docLanguage="en" docName="EJT.2024.959.1-272.pdf" docOrigin="European Journal of Taxonomy 959 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2677/12401" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Dichotomius Hope 1838" docType="treatment" docVersion="6" lastPageNumber="90" masterDocId="E1690E1E305A5264ED17FFA7770C013F" masterDocTitle="The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution" masterLastPageNumber="272" masterPageNumber="1" pageNumber="89" updateTime="1731448046962" updateUser="carolina" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="82CB8FA0C05E76A76C117E70D880C78D">The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution</mods:title>
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<mods:namePart id="EBA91CDFACD39498F4F41C9095DD9A87">Rosa, Cecilia Lozano De La</mods:namePart>
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<mods:affiliation id="CD3082D31BE2DDBA1CBE254586CB226B">Instituto Venezolano de Investigaciones Científicas, Centro de Estudios Botánicos y Agroforestales, Laboratorio de Ecología Espacial, Maracaibo, estado Zulia, Venezuela.</mods:affiliation>
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<mods:namePart id="3468E9242806157289BC355DDD93FE2A">Cupello, Mario</mods:namePart>
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<mods:affiliation id="BB61A9682E931625119D994DD87BB0F6">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Curitiba, Paraná, Brazil. Currently at Department of Entomology, Texas A &amp; M University, College Station, TX, 77843, USA.</mods:affiliation>
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Genus
<taxonomicName id="52F9BCF33002523CEFBAF95374D50631" ID-CoL="44NT" authority="Hope, 1838" authorityName="Hope" authorityYear="1838" box="[685,985,1780,1807]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B623002523CEFBAF95374440630" bold="true" box="[685,840,1780,1807]" italics="true" pageId="88" pageNumber="89">Dichotomius</emphasis>
<bibRefCitation id="F168BA813002523CEE58F95274D50631" author="Hope F. W." box="[847,985,1780,1807]" pageId="88" pageNumber="89" pagination="312 - 326" refId="ref124432" refString="Hope F. W. 1838. Observations on the Lamellicorns of Olivier. The Entomological Magazine 5: 312 - 326. Available from https: // www. biodiversitylibrary. org / page / 8982116 [accessed 16 Feb. 2024]." type="journal article" year="1838">Hope, 1838</bibRefCitation>
</taxonomicName>
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<paragraph id="9546C7703002523CEDAAF89275DB064C" blockId="88.[189,1399,1845,2013]" pageId="88" pageNumber="89">
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<treatmentCitation id="1458E1613002523CEDAAF8927510066F" author="Hope F. W." box="[189,540,1845,1872]" page="321" pageId="88" pageNumber="89" year="1838">
<taxonomicName id="52F9BCF33002523CEDAAF8927510066F" ID-CoL="44NT" authority="Hope, 1838: 321" authorityName="Hope" authorityPageNumber="321" authorityYear="1838" box="[189,540,1845,1872]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B623002523CEDAAF892765F0670" box="[189,339,1845,1871]" italics="true" pageId="88" pageNumber="89">Dichotomius</emphasis>
<bibRefCitation id="F168BA813002523CEC4EF8917510066F" author="Hope F. W." box="[345,540,1846,1872]" pageId="88" pageNumber="89" pagination="312 - 326" refId="ref124432" refString="Hope F. W. 1838. Observations on the Lamellicorns of Olivier. The Entomological Magazine 5: 312 - 326. Available from https: // www. biodiversitylibrary. org / page / 8982116 [accessed 16 Feb. 2024]." type="journal article" year="1838">Hope, 1838: 321</bibRefCitation>
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(original description).
<typeStatus id="4A4279D23002523CEE3FF891746E066F" box="[808,866,1846,1872]" pageId="88" pageNumber="89">Type</typeStatus>
species:
<taxonomicName id="52F9BCF33002523CEEDCF8927241066F" ID-CoL="7YY4G" authority="Olivier, 1789" authorityName="Olivier" authorityYear="1789" box="[971,1357,1845,1872]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="boreus">
<emphasis id="A78D1B623002523CEEDCF89273A60670" box="[971,1194,1845,1871]" italics="true" pageId="88" pageNumber="89">Scarabaeus boreus</emphasis>
<bibRefCitation id="F168BA813002523CE9A7F8927241066F" author="Olivier G. A." box="[1200,1357,1845,1872]" pageId="88" pageNumber="89" refId="ref133896" refString="Olivier G. A. 1789. N ° 3. Scarabe. Scarabaeus. In: Olivier G. A. 1789. Entomologie, ou histoire naturelle des insectes, avec leurs caracteres generiques et specifiques, leur description, leur synonymie, et leur figure enluminee. Coleopteres. Tome Premier. Baudouin, Paris. [Publication date following Bousquet (2016).] https: // doi. org / 10.5962 / bhl. title. 61905" type="book" year="1789">Olivier, 1789</bibRefCitation>
</taxonomicName>
, by original designation; see
<bibRefCitation id="F168BA813002523CEF07F8FE75DE064C" author="Martinez A." box="[528,722,1881,1908]" pageId="88" pageNumber="89" pagination="138 - 142" refId="ref129742" refString="Martinez A. 1951. Notas coleopterologicas V. La invalidez del nombre generico Pinotus Erichson y dos nuevas sinonimias. Anales de la Sociedad Cientifica Argentina 152: 138 - 142." type="journal article" year="1951">Martínez (1951)</bibRefCitation>
.
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</paragraph>
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<treatmentCitationGroup id="B5E9E05E3002523DEDAAF8DA75CC0302" lastPageId="89" lastPageNumber="90" pageId="88" pageNumber="89">
<treatmentCitation id="1458E1613002523CEDAAF8DA752806A9" author="Erichson W. F." box="[189,548,1916,1942]" page="108" pageId="88" pageNumber="89" year="1847">
<taxonomicName id="52F9BCF33002523CEDAAF8DA752806A9" ID-CoL="7PS5K" authority="Erichson, 1847 a: 108" authorityName="Erichson" authorityPageNumber="108" authorityYear="1847" box="[189,548,1916,1942]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B623002523CEDAAF8DA761B06A9" box="[189,279,1917,1942]" italics="true" pageId="88" pageNumber="89">Pinotus</emphasis>
<bibRefCitation id="F168BA813002523CEC35F8DB752806A9" author="Erichson W. F." box="[290,548,1916,1942]" pageId="88" pageNumber="89" pagination="67 - 185" refId="ref117813" refString="Erichson W. F. 1847 a. Conspectus insectorum coleopterorum quae in Republica Peruana observata sunt. Archiv fur Naturgeschichte 13: 67 - 185. Available from https: // www. biodiversitylibrary. org / page / 7251438 [accessed 14 Feb. 2024]." type="journal article" year="1847">Erichson, 1847a: 108</bibRefCitation>
</taxonomicName>
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(original description).
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species: uncertain. Usually claimed to be
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<taxonomicName id="52F9BCF33002523CEDFDF83875BD0686" ID-CoL="7YY4L" authority="Linnaeus, 1767" authorityName="Linnaeus" authorityYear="1767" box="[234,689,1951,1977]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="carolinus">
<emphasis id="A78D1B623002523CEDFDF83876E10686" box="[234,493,1951,1977]" italics="true" pageId="88" pageNumber="89">Scarabaeus carolinus</emphasis>
<bibRefCitation id="F168BA813002523CECEFF83875BD0686" author="Linnaeus C." box="[504,689,1951,1977]" pageId="88" pageNumber="89" refId="ref128194" refString="Linnaeus C. 1767. Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Editio duodecima, reformata. Tom. I. Pars II. Laurentii Salvii, Stockholm [Holmiae]. https: // doi. org / 10.5962 / bhl. title. 156772" type="book" year="1767">Linnaeus, 1767</bibRefCitation>
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(e.g.,
<treatmentCitation id="1458E1613002523CEE14F83874C70685" author="Lacordaire J. T." box="[771,971,1951,1978]" pageId="88" pageNumber="89" year="1855">
<bibRefCitation id="F168BA813002523CEE14F83874C70685" author="Lacordaire J. T." box="[771,971,1951,1978]" pageId="88" pageNumber="89" refId="ref126653" refString="Lacordaire J. T. 1855. Histoire naturelle des Insectes. Genera des coleopteres, ou expose methodique et critique de tous les genres proposes jusqu'ici dans cet ordre d'insectes. Tome troisieme. Contenant les familles des Pectinicornes et Lamellicornes. Librarie Encyclopedique de Roret, Paris. [Publication date following Bousquet (2016).] https: // doi. org / 10.5962 / bhl. title. 8864" type="journal volume" year="1855">Lacordaire 1855</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="1458E1613002523CEECCF83873B50686" author="Luederwaldt G." box="[987,1209,1951,1977]" pageId="88" pageNumber="89" year="1929">
<bibRefCitation id="F168BA813002523CEECCF83873B50686" author="Luederwaldt H." box="[987,1209,1951,1977]" pageId="88" pageNumber="89" pagination="603 - 775" refId="ref128687" refString="Luederwaldt H. 1929. As especies brasileiras do genero Pinotus, com algumas consideracoes tambem sobre outras especies. Revista do Museu Paulista 16: 603 - 775." type="journal article" year="1929">Luederwaldt 1929</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="1458E1613002523CE9DEF807769306E3" author="Nunes R. V. &amp; Vaz-de-Mello F." pageId="88" pageNumber="89" year="2019">
<bibRefCitation id="F168BA813002523CE9DEF807769306E3" author="Nunes R. V. &amp; Vaz-de-Mello F. Z." pageId="88" pageNumber="89" pagination="2231 - 2351" refId="ref133607" refString="Nunes R. V. &amp; Vaz-de-Mello F. Z. 2019. Taxonomic revision of Dichotomius (Cephagonus) Luederwaldt 1929 and the taxonomic status of remaining Dichotomius Hope 1838 subgenera (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini). Journal of Natural History 53 (37 - 38): 2231 - 2351. https: // doi. org / 10.1080 / 00222933.2019.1692088" type="journal article" year="2019">Nunes &amp; Vaz-de-Mello 2019</bibRefCitation>
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), but, as argued by
<treatmentCitation id="1458E1613002523CEF99F865745E06E2" author="Martinez A." box="[654,850,1986,2013]" pageId="88" pageNumber="89" year="1951">
<bibRefCitation id="F168BA813002523CEF99F865745E06E2" author="Martinez A." box="[654,850,1986,2013]" pageId="88" pageNumber="89" pagination="138 - 142" refId="ref129742" refString="Martinez A. 1951. Notas coleopterologicas V. La invalidez del nombre generico Pinotus Erichson y dos nuevas sinonimias. Anales de la Sociedad Cientifica Argentina 152: 138 - 142." type="journal article" year="1951">Martínez (1951)</bibRefCitation>
</treatmentCitation>
, this nominal species was not among the two expressly assigned to
<taxonomicName id="52F9BCF33003523DECEDFEAD7558001C" ID-CoL="7PS5K" authorityName="Erichson" authorityYear="1847" box="[506,596,266,291]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B623003523DECEDFEAD7558001C" box="[506,596,266,291]" italics="true" pageId="89" pageNumber="90">Pinotus</emphasis>
</taxonomicName>
by
<treatmentCitation id="1458E1613003523DEF9DFEAE746E001C" box="[650,866,265,291]" pageId="89" pageNumber="90">
<bibRefCitation id="F168BA813003523DEF9DFEAE7452001C" author="Erichson W. F." box="[650,862,265,291]" pageId="89" pageNumber="90" pagination="67 - 185" refId="ref117813" refString="Erichson W. F. 1847 a. Conspectus insectorum coleopterorum quae in Republica Peruana observata sunt. Archiv fur Naturgeschichte 13: 67 - 185. Available from https: // www. biodiversitylibrary. org / page / 7251438 [accessed 14 Feb. 2024]." type="journal article" year="1847">Erichson (1847a)</bibRefCitation>
.
</treatmentCitation>
<treatmentCitation id="1458E1613003523DEE79FEAE733F001B" author="Martinez A." box="[878,1075,265,292]" pageId="89" pageNumber="90" year="1951">
<bibRefCitation id="F168BA813003523DEE79FEAE733F001B" author="Martinez A." box="[878,1075,265,292]" pageId="89" pageNumber="90" pagination="138 - 142" refId="ref129742" refString="Martinez A. 1951. Notas coleopterologicas V. La invalidez del nombre generico Pinotus Erichson y dos nuevas sinonimias. Anales de la Sociedad Cientifica Argentina 152: 138 - 142." type="journal article" year="1951">Martínez (1951)</bibRefCitation>
</treatmentCitation>
stated that
<taxonomicName id="52F9BCF33003523DE9DEFEAD76910079" ID-CoL="7Y82L" authority="Erichson, 1847" authorityName="Erichson" authorityYear="1847" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="talaus">
<emphasis id="A78D1B623003523DE9DEFEAD727B001C" box="[1225,1399,265,291]" italics="true" pageId="89" pageNumber="90">Pinotus talaus</emphasis>
Erichson, 1847
</taxonomicName>
was the actual
<typeStatus id="4A4279D23003523DEF41FE8A75870078" box="[598,651,301,327]" pageId="89" pageNumber="90">type</typeStatus>
, and this was indeed one of the two species expressly assigned to the genus by
<treatmentCitation id="1458E1613003523DECB1FEE875780055" author="Erichson W. F." box="[422,628,335,362]" pageId="89" pageNumber="90" year="1847">
<bibRefCitation id="F168BA813003523DECB1FEE875780055" author="Erichson W. F." box="[422,628,335,362]" pageId="89" pageNumber="90" pagination="67 - 185" refId="ref117813" refString="Erichson W. F. 1847 a. Conspectus insectorum coleopterorum quae in Republica Peruana observata sunt. Archiv fur Naturgeschichte 13: 67 - 185. Available from https: // www. biodiversitylibrary. org / page / 7251438 [accessed 14 Feb. 2024]." type="journal article" year="1847">Erichson (1847a)</bibRefCitation>
</treatmentCitation>
. However, why Martínez believed that
<taxonomicName id="52F9BCF33003523DE956FEE873850056" ID-CoL="7Y82L" authorityName="Erichson" authorityYear="1847" box="[1089,1161,335,361]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="talaus">
<emphasis id="A78D1B623003523DE956FEE873850056" box="[1089,1161,335,361]" italics="true" pageId="89" pageNumber="90">talaus</emphasis>
</taxonomicName>
specifically was the
<typeStatus id="4A4279D23003523DEDFDFED4761100B2" box="[234,285,371,397]" pageId="89" pageNumber="90">type</typeStatus>
is unclear. Regardless, if no
<typeStatus id="4A4279D23003523DEF6EFED475A000B2" box="[633,684,371,397]" pageId="89" pageNumber="90">type</typeStatus>
species had been fixed prior to 1951,
<treatmentCitation id="1458E1613003523DE960FED4724700B2" author="Martinez A." box="[1143,1355,370,397]" pageId="89" pageNumber="90" year="1951">
<bibRefCitation id="F168BA813003523DE960FED4724700B2" author="Martinez A." box="[1143,1355,370,397]" pageId="89" pageNumber="90" pagination="138 - 142" refId="ref129742" refString="Martinez A. 1951. Notas coleopterologicas V. La invalidez del nombre generico Pinotus Erichson y dos nuevas sinonimias. Anales de la Sociedad Cientifica Argentina 152: 138 - 142." type="journal article" year="1951">Martínezs (1951)</bibRefCitation>
</treatmentCitation>
act should be interpreted as a valid subsequent designation of
<taxonomicName id="52F9BCF33003523DEEB0FE307301008F" ID-CoL="7Y82L" authorityName="Erichson" authorityYear="1847" box="[935,1037,406,432]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="talaus">
<emphasis id="A78D1B623003523DEEB0FE307301008F" box="[935,1037,406,432]" italics="true" pageId="89" pageNumber="90">P. talaus</emphasis>
</taxonomicName>
as the
<typeStatus id="4A4279D23003523DE975FE317399008F" box="[1122,1173,406,432]" pageId="89" pageNumber="90">type</typeStatus>
species of
<taxonomicName id="52F9BCF33003523DE809FE307274008F" ID-CoL="7PS5K" authorityName="Erichson" authorityYear="1847" box="[1310,1400,407,432]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B623003523DE809FE307274008F" box="[1310,1400,407,432]" italics="true" pageId="89" pageNumber="90">Pinotus</emphasis>
</taxonomicName>
under Art. 69.1.1 of the Code (
<bibRefCitation id="F168BA813003523DEF71FE1E75FA00EC" author="ICZN &amp; International Commission on Zoological Nomenclature" box="[614,758,441,467]" pageId="89" pageNumber="90" refId="ref125343" refString="ICZN [International Commission on Zoological Nomenclature] 1999. International Code of Zoological Nomenclature. Fourth edition adopted by the International Union of Biological Sciences. International Trust for Zoological Nomenclature, London." type="book" year="1999">ICZN 1999</bibRefCitation>
).
<treatmentCitation id="1458E1613003523DEE19FE1E732800EC" author="Bouchard P. &amp; Bousquet Y. &amp; Davies A. E. &amp; Cai C." box="[782,1060,441,468]" pageId="89" pageNumber="90" year="2024">
<bibRefCitation id="F168BA813003523DEE19FE1E732800EC" author="Bouchard P. &amp; Bousquet Y. &amp; Davies A. E. &amp; Cai C." box="[782,1060,441,468]" pageId="89" pageNumber="90" pagination="1 - 981" refId="ref112265" refString="Bouchard P., Bousquet Y., Davies A. E. &amp; Cai C. 2024. On the nomenclatural status of type genera in Coleoptera (Insecta). ZooKeys 1194: 1 - 981. https: // doi. org / 10.3897 / zookeys. 1194.106440" type="journal article" year="2024">
Bouchard
<emphasis id="A78D1B623003523DEE9DFE1D74CD00EC" box="[906,961,441,467]" italics="true" pageId="89" pageNumber="90">et al</emphasis>
. (2024)
</bibRefCitation>
</treatmentCitation>
, unaware of Martínezs act, recently designated the same nominal species as the
<typeStatus id="4A4279D23003523DEE70FE7A749000C8" box="[871,924,477,503]" pageId="89" pageNumber="90">type</typeStatus>
, but Martínezs designation would take precedence over theirs. To settle this issue, it must be determined whether no
<typeStatus id="4A4279D23003523DE992FDA773B40325" box="[1157,1208,512,538]" pageId="89" pageNumber="90">type</typeStatus>
species fixation indeed existed prior to
<treatmentCitation id="1458E1613003523DECEEFD8475CC0302" box="[505,704,546,573]" pageId="89" pageNumber="90">
<bibRefCitation id="F168BA813003523DECEEFD8475B00302" author="Martinez A." box="[505,700,546,573]" pageId="89" pageNumber="90" pagination="138 - 142" refId="ref129742" refString="Martinez A. 1951. Notas coleopterologicas V. La invalidez del nombre generico Pinotus Erichson y dos nuevas sinonimias. Anales de la Sociedad Cientifica Argentina 152: 138 - 142." type="journal article" year="1951">Martínez (1951)</bibRefCitation>
.
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C7703003523DEDAAFDE1754603BC" blockId="89.[189,1400,265,749]" pageId="89" pageNumber="90">
<treatmentCitationGroup id="B5E9E05E3003523DEDAAFDE1754603BC" pageId="89" pageNumber="90">
<treatmentCitation id="1458E1613003523DEDAAFDE17555035E" author="Haldeman S. S." box="[189,601,582,609]" page="125" pageId="89" pageNumber="90" year="1848">
<taxonomicName id="52F9BCF33003523DEDAAFDE17555035E" ID-CoL="7NR35" authority="Haldeman, 1848: 125" authorityName="Haldeman" authorityPageNumber="125" authorityYear="1848" box="[189,601,582,609]" class="Insecta" family="Scarabaeidae" genus="Brachycopris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B623003523DEDAAFDE17657035F" box="[189,347,582,608]" italics="true" pageId="89" pageNumber="90">Brachycopris</emphasis>
<bibRefCitation id="F168BA813003523DEC77FDE17555035E" author="Haldeman S. S." box="[352,601,582,609]" pageId="89" pageNumber="90" pagination="124 - 128" refId="ref122229" refString="Haldeman S. S. 1848. On several new genera and species of insects. Proceedings of the Academy of Natural Sciences of Philadelphia 3 (6): 124 - 128. Available from https: // www. biodiversitylibrary. org / page / 1659393 [accessed 16 Feb. 2024]." type="journal article" year="1848">Haldeman, 1848: 125</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description).
<typeStatus id="4A4279D23003523DEE76FDE17497035F" box="[865,923,582,608]" pageId="89" pageNumber="90">Type</typeStatus>
species:
<treatmentCitation id="1458E1613003523DE917FDE1762B03BC" author="Linnaeus C." pageId="89" pageNumber="90" year="1767">
<taxonomicName id="52F9BCF33003523DE917FDE1762B03BC" ID-CoL="7YY4L" authority="Linnaeus, 1767" authorityName="Linnaeus" authorityYear="1767" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="carolinus">
<emphasis id="A78D1B623003523DE917FDE173F0035F" box="[1024,1276,582,608]" italics="true" pageId="89" pageNumber="90">Scarabaeus carolinus</emphasis>
<bibRefCitation id="F168BA813003523DE816FDE1762B03BC" author="Linnaeus C." pageId="89" pageNumber="90" refId="ref128194" refString="Linnaeus C. 1767. Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Editio duodecima, reformata. Tom. I. Pars II. Laurentii Salvii, Stockholm [Holmiae]. https: // doi. org / 10.5962 / bhl. title. 156772" type="book" year="1767">Linnaeus, 1767</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, by original designation.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C7703003523DEDAAFD2B75FC03D2" blockId="89.[189,1400,265,749]" pageId="89" pageNumber="90">
<treatmentCitationGroup id="B5E9E05E3003523DEDAAFD2B75FC03D2" pageId="89" pageNumber="90">
<treatmentCitation id="1458E1613003523DEDAAFD2B75700399" author="Ferreira M. C." box="[189,636,652,678]" page="3" pageId="89" pageNumber="90" year="1970">
<taxonomicName id="52F9BCF33003523DEDAAFD2B75700399" ID-CoL="7PWGK" authority="Ferreira, 1970: 3" authorityName="Ferreira" authorityPageNumber="3" authorityYear="1970" box="[189,636,652,678]" class="Insecta" family="Scarabaeidae" genus="Pseudoheliocopris" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B623003523DEDAAFD2B769B0399" box="[189,407,652,678]" italics="true" pageId="89" pageNumber="90">Pseudoheliocopris</emphasis>
<bibRefCitation id="F168BA813003523DECB0FD2B75700399" author="Ferreira M. C." box="[423,636,652,678]" pageId="89" pageNumber="90" pagination="3 - 9" refId="ref119094" refString="Ferreira M. C. 1970. Contribuicao para o estudo dos escarabideos africanos. Mais um genero novo da tribo Coprini. Novos Taxa Entomologicos 76: 3 - 9." type="journal article" year="1970">Ferreira, 1970: 3</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description).
<typeStatus id="4A4279D23003523DEEB5FD2A74D00398" box="[930,988,653,679]" pageId="89" pageNumber="90">Type</typeStatus>
species:
<treatmentCitation id="1458E1613003523DE94FFD2B769A03F5" author="Ferreira M. C." pageId="89" pageNumber="90" year="1970">
<taxonomicName id="52F9BCF33003523DE94FFD2B769A03F5" ID-CoL="7YKYL" authority="Ferreira, 1970" authorityName="Ferreira" authorityYear="1970" class="Insecta" family="Scarabaeidae" genus="Pseudoheliocopris" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="freyi">
<emphasis id="A78D1B623003523DE94FFD2B727B0399" box="[1112,1399,652,678]" italics="true" pageId="89" pageNumber="90">Pseudoheliocopris freyi</emphasis>
<bibRefCitation id="F168BA813003523DEDFDFD08769A03F5" author="Ferreira M. C." box="[234,406,687,714]" pageId="89" pageNumber="90" pagination="3 - 9" refId="ref119094" refString="Ferreira M. C. 1970. Contribuicao para o estudo dos escarabideos africanos. Mais um genero novo da tribo Coprini. Novos Taxa Entomologicos 76: 3 - 9." type="journal article" year="1970">Ferreira, 1970</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, by original designation, not
<treatmentCitation id="1458E1613003523DEFECFD17733903F5" author="Ferreira M. C." box="[763,1077,687,714]" pageId="89" pageNumber="90" year="1970">
<taxonomicName id="52F9BCF33003523DEFECFD17733903F5" ID-CoL="7YKYK" authority="Ferreira, 1970" authorityName="Ferreira" authorityYear="1970" box="[763,1077,687,714]" class="Insecta" family="Scarabaeidae" genus="Pseudoheliocopris" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="cornutus">
<emphasis id="A78D1B623003523DEFECFD17748E03F6" box="[763,898,688,713]" italics="true" pageId="89" pageNumber="90">P. cornutus</emphasis>
<bibRefCitation id="F168BA813003523DEE9BFD08733903F5" author="Ferreira M. C." box="[908,1077,687,714]" pageId="89" pageNumber="90" pagination="3 - 9" refId="ref119094" refString="Ferreira M. C. 1970. Contribuicao para o estudo dos escarabideos africanos. Mais um genero novo da tribo Coprini. Novos Taxa Entomologicos 76: 3 - 9." type="journal article" year="1970">Ferreira, 1970</bibRefCitation>
</taxonomicName>
</treatmentCitation>
by original designation as informed by
<treatmentCitation id="1458E1613003523DEC95FD7475E103D2" author="Nunes L. G. O. A. &amp; Vaz-de-Mello F. Z." box="[386,749,722,749]" pageId="89" pageNumber="90" year="2019">
<bibRefCitation id="F168BA813003523DEC95FD7475E103D2" author="Nunes R. V. &amp; Vaz-de-Mello F. Z." box="[386,749,722,749]" pageId="89" pageNumber="90" pagination="2231 - 2351" refId="ref133607" refString="Nunes R. V. &amp; Vaz-de-Mello F. Z. 2019. Taxonomic revision of Dichotomius (Cephagonus) Luederwaldt 1929 and the taxonomic status of remaining Dichotomius Hope 1838 subgenera (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini). Journal of Natural History 53 (37 - 38): 2231 - 2351. https: // doi. org / 10.1080 / 00222933.2019.1692088" type="journal article" year="2019">Nunes &amp; Vaz-de-Mello (2019)</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C7703003523DEDAAFCB375580401" blockId="89.[189,1399,788,2011]" pageId="89" pageNumber="90">
<treatmentCitationGroup id="B5E9E05E3003523DEDAAFCB375580401" pageId="89" pageNumber="90">
<taxonomicName id="52F9BCF33003523DEDAAFCB3765F0211" ID-CoL="44NT" authorityName="Hope" authorityYear="1838" box="[189,339,788,814]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B623003523DEDAAFCB3765F0211" box="[189,339,788,814]" italics="true" pageId="89" pageNumber="90">Dichotomius</emphasis>
</taxonomicName>
<treatmentCitation id="1458E1613003523DEC64FCB3755E0210" author="Agassiz L." box="[371,594,788,815]" page="353" pageId="89" pageNumber="90" year="1846">
<bibRefCitation id="F168BA813003523DEC64FCB3755E0210" author="Agassiz L." box="[371,594,788,815]" pageId="89" pageNumber="90" refId="ref108975" refString="Agassiz L. 1846. Nomenclatoris zoologici index universalis, continens nomina systematica classium, ordinum, familiarum et generum animalium omnium, tam viventium quam fossilum, secundum ordinem alphabeticum unicum deposita, adjectis homonymus platarum, nec non variis adnotationibus et emendationibus. Jent and Gassman, Solothurn [Soloduri]. https: // doi. org / 10.5962 / bhl. title. 1819" type="book" year="1846">Agassiz 1846: 353</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="1458E1613003523DEE0FFCB373030211" author="Erichson W. F." box="[792,1039,788,814]" page="108" pageId="89" pageNumber="90" year="1847">
<bibRefCitation id="F168BA813003523DEE0FFCB373030211" author="Erichson W. F." box="[792,1039,788,814]" pageId="89" pageNumber="90" pagination="67 - 185" refId="ref117813" refString="Erichson W. F. 1847 a. Conspectus insectorum coleopterorum quae in Republica Peruana observata sunt. Archiv fur Naturgeschichte 13: 67 - 185. Available from https: // www. biodiversitylibrary. org / page / 7251438 [accessed 14 Feb. 2024]." type="journal article" year="1847">Erichson 1847a: 108</bibRefCitation>
</treatmentCitation>
(synonymy with
<taxonomicName id="52F9BCF33003523DE9F0FCB272320211" ID-CoL="7PS5K" authorityName="Erichson" authorityYear="1847" box="[1255,1342,789,814]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B623003523DE9F0FCB272320211" box="[1255,1342,789,814]" italics="true" pageId="89" pageNumber="90">Pinotus</emphasis>
</taxonomicName>
). —
<treatmentCitation id="1458E1613003523DEDFAFC9F76B8026E" author="Lucas R." box="[237,436,823,850]" page="237" pageId="89" pageNumber="90" year="1920">
<bibRefCitation id="F168BA813003523DEDFAFC9F76B8026E" author="Lucas R." box="[237,436,823,850]" pageId="89" pageNumber="90" refId="ref128561" refString="Lucas R. 1920. Catalogus alphabeticus generum et subgenerum Coleopterorum orbis terrarum totius (famil., trib., subtr., sect. incl.). Pars I. Nicolaische, Berlin." type="book" year="1920">Lucas 1920: 237</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF33003523DEFD4FC9F7417026E" ID-CoL="7PS5K" authorityName="Erichson" authorityYear="1847" box="[707,795,824,849]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B623003523DEFD4FC9F7417026E" box="[707,795,824,849]" italics="true" pageId="89" pageNumber="90">Pinotus</emphasis>
</taxonomicName>
). —
<treatmentCitation id="1458E1613003523DEE4CFC90737D026E" author="Blackwelder R. E." box="[859,1137,823,849]" page="206" pageId="89" pageNumber="90" year="1944">
<bibRefCitation id="F168BA813003523DEE4CFC90737D026E" author="Blackwelder R. E." box="[859,1137,823,849]" pageId="89" pageNumber="90" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 206</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF33003523DEDFBFCFC7648024B" ID-CoL="7PS5K" authorityName="Erichson" authorityYear="1847" box="[236,324,859,884]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B623003523DEDFBFCFC7648024B" box="[236,324,859,884]" italics="true" pageId="89" pageNumber="90">Pinotus</emphasis>
</taxonomicName>
). —
<treatmentCitation id="1458E1613003523DEC97FCFC756A024A" author="Martinez A." box="[384,614,858,885]" page="139" pageId="89" pageNumber="90" year="1951">
<bibRefCitation id="F168BA813003523DEC97FCFC756A024A" author="Martinez A." box="[384,614,858,885]" pageId="89" pageNumber="90" pagination="138 - 142" refId="ref129742" refString="Martinez A. 1951. Notas coleopterologicas V. La invalidez del nombre generico Pinotus Erichson y dos nuevas sinonimias. Anales de la Sociedad Cientifica Argentina 152: 138 - 142." type="journal article" year="1951">Martínez 1951: 139</bibRefCitation>
</treatmentCitation>
(comments);
<treatmentCitation id="1458E1613003523DEE10FCFD7462024A" author="Martinez A." box="[775,878,858,885]" page="80" pageId="89" pageNumber="90" year="1959">
<bibRefCitation id="F168BA813003523DEE10FCFD7462024A" author="Martinez A." box="[775,878,858,885]" pageId="89" pageNumber="90" pagination="1 - 126" refId="ref129799" refString="Martinez A. 1959. Catalogo de los Scarabaeidae Argentinos. Revista del Museo Argentino de Ciencias Naturales &quot; Bernardino Rivadavia &quot; 5: 1 - 126." type="journal article" year="1959">1959: 80</bibRefCitation>
</treatmentCitation>
(catalogue for
<collectingCountry id="EDEE87E03003523DE90AFCFD739B024B" box="[1053,1175,858,884]" name="Argentina" pageId="89" pageNumber="90">Argentina</collectingCountry>
). —
<treatmentCitation id="1458E1613003523DE9DAFCFD760B02A7" author="Pereira F. S." page="57" pageId="89" pageNumber="90" year="1954">
<bibRefCitation id="F168BA813003523DE9DAFCFD760B02A7" author="Pereira F. S." pageId="89" pageNumber="90" pagination="53 - 78" refId="ref135304" refString="Pereira F. S. 1954 a. O genero Scatonomus Er. (Coleoptera, Scarabaeidae). Revista Brasileira de Entomologia 1: 53 - 78." type="journal article" year="1954">Pereira 1954a: 57</bibRefCitation>
</treatmentCitation>
(key). —
<treatmentCitation id="1458E1613003523DEC6BFCD9752902A7" author="Roze J. A." box="[380,549,893,920]" page="44" pageId="89" pageNumber="90" year="1955">
<bibRefCitation id="F168BA813003523DEC6BFCD9752902A7" author="Roze J. A." box="[380,549,893,920]" pageId="89" pageNumber="90" pagination="39 - 63" refId="ref137914" refString="Roze J. A. 1955. Lista preliminar de la familia Scarabaeidae sensu lato de Venezuela. Boletin del Museo de Ciencias Naturales 1 (1): 39 - 63." type="journal article" year="1955">Roze 1955: 44</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E03003523DEFDBFCD9744602A7" box="[716,842,894,920]" name="Venezuela" pageId="89" pageNumber="90">Venezuela</collectingCountry>
). —
<treatmentCitation id="1458E1613003523DEE97FCD973E302A7" author="Halffter G. &amp; Matthews E. G." box="[896,1263,893,920]" page="257" pageId="89" pageNumber="90" year="1966">
<bibRefCitation id="F168BA813003523DEE97FCD973E302A7" author="Halffter G. &amp; Matthews E. G." box="[896,1263,893,920]" pageId="89" pageNumber="90" pagination="1 - 312" refId="ref122681" refString="Halffter G. &amp; Matthews E. G. 1966. The natural history of dung beetles of the subfamily Scarabaeinae (Coleoptera: Scarabaeidae). Folia Entomologica Mexicana 12 - 14: 1 - 312." type="book chapter" year="1966">Halffter &amp; Matthews 1966: 257</bibRefCitation>
</treatmentCitation>
(catalogue, distribution). —
<treatmentCitation id="1458E1613003523DECA1FC0674110283" author="Vulcano M. A. &amp; Pereira F. S." box="[438,797,929,956]" page="577" pageId="89" pageNumber="90" year="1967">
<bibRefCitation id="F168BA813003523DECA1FC0674110283" author="Vulcano M. A. &amp; Pereira F. S." box="[438,797,929,956]" pageId="89" pageNumber="90" pagination="533 - 603" refId="ref141365" refString="Vulcano M. A. &amp; Pereira F. S. 1967. Sinopse dos Passalidae e Scarabaeidae s. str. da regiao amazonica (Insecta, Coleoptera). Atas do Simposio sobre a Biota Amazonica 5: 533 - 603." type="journal article" year="1967">Vulcano &amp; Pereira 1967: 577</bibRefCitation>
</treatmentCitation>
(key). —
<treatmentCitation id="1458E1613003523DEEB6FC0673F90284" author="Howden H. F. &amp; Young O. P." box="[929,1269,929,955]" page="13" pageId="89" pageNumber="90" year="1981">
<bibRefCitation id="F168BA813003523DEEB6FC0673F90284" author="Howden H. F. &amp; Young O. P." box="[929,1269,929,955]" pageId="89" pageNumber="90" pagination="1 - 204" refId="ref124824" refString="Howden H. F. &amp; Young O. P. 1981. Panamanian Scarabaeinae: taxonomy, distribution and habits (Coleoptera, Scarabaeidae). Contributions of the American Entomological Institute 18 (1): 1 - 204." type="journal article" year="1981">Howden &amp; Young 1981: 13</bibRefCitation>
</treatmentCitation>
, 123 (key, redescription). —
<treatmentCitation id="1458E1613003523DECDEFC63744B02E1" author="Halffter G. &amp; Edmonds W. D." box="[457,839,964,990]" page="137" pageId="89" pageNumber="90" year="1982">
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Medina
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2001: 138
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Ratcliffe
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2002: 49
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Hamel-Leigue
<emphasis id="A78D1B623003523DEE1FFBF574490554" box="[776,837,1105,1131]" italics="true" pageId="89" pageNumber="90">et al.</emphasis>
2006: 15
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Vaz-de-Mello
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2011a: 20
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Carvajal
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2011: 129
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Ferrer-Paris
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2013: 96
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Chamorro
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2018: 77
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Pardo-Díaz
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2019: 14
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Cassenote
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2020: 1
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Nieto
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2020: 136
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species designation). —
<treatmentCitation id="1458E1613003523DE99FFAE0727A045E" author="Harold E." box="[1160,1398,1351,1377]" page="124" pageId="89" pageNumber="90" year="1869">
<bibRefCitation id="F168BA813003523DE99FFAE0727A045E" author="Harold E." box="[1160,1398,1351,1377]" pageId="89" pageNumber="90" pagination="123 - 144" refId="ref123559" refString="Harold E. 1869 b. Revision des especes qui rentrent dans le genre Pinotus Erichs. L'Abeille 6: 123 - 144." type="journal article" year="1869">Harold 1869b: 124</bibRefCitation>
</treatmentCitation>
(redescription);
<treatmentCitation id="1458E1613003523DECB4FACD753804BA" author="Harold E." box="[419,564,1386,1413]" page="1009" pageId="89" pageNumber="90" year="1869">
<bibRefCitation id="F168BA813003523DECB4FACD753804BA" author="Harold E." box="[419,564,1386,1413]" firstAuthor="d" pageId="89" pageNumber="90" refId="ref123629" refString="Harold E. 1869 d. Catalogus coleopterorum. Hucusque descriptorum, synonymicus et systematicus. Tom. IV. Scarabaeidae. E. H. Gummi, Munich. https: // doi. org / 10.5962 / bhl. title. 9089" type="book" year="1869">1869d: 1009</bibRefCitation>
</treatmentCitation>
(catalogue).—
<treatmentCitation id="1458E1613003523DEFFEFACD74AB04BA" author="Bruch C." box="[745,935,1386,1413]" page="187" pageId="89" pageNumber="90" year="1911">
<bibRefCitation id="F168BA813003523DEFFEFACD74AB04BA" author="Bruch C." box="[745,935,1386,1413]" pageId="89" pageNumber="90" pagination="143 - 225" refId="ref112509" refString="Bruch C. 1911. Catalogo sistematico de los coleopteros de la Republica Argentina. Revista del Museo de La Plata 17: 143 - 225. https: // doi. org / 10.5962 / bhl. title. 8799" type="journal article" year="1911">Bruch 1911: 187</bibRefCitation>
</treatmentCitation>
(list).
<emphasis id="A78D1B623003523DEEFCFACC730904BB" box="[1003,1029,1387,1412]" italics="true" pageId="89" pageNumber="90"></emphasis>
<treatmentCitation id="1458E1613003523DE91BFACD73C904BA" author="Gillet J. J. E." box="[1036,1221,1386,1413]" page="59" pageId="89" pageNumber="90" year="1911">
<bibRefCitation id="F168BA813003523DE91BFACD73C904BA" author="Gillet J. J. E." box="[1036,1221,1386,1413]" pageId="89" pageNumber="90" refId="ref121235" refString="Gillet J. J. E. 1911 b. Coleopterorum Catalogus. Pars 38: Scarabaeidae: Coprinae I. W. Junk, Berlin. Available from https: // www. biodiversitylibrary. org / page / 48405679 [accesseed 16 Feb. 2024]." type="url" year="1911">Gillet 1911b: 59</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="1458E1613003523DEDF0FA2976A70497" author="Lucas R." box="[231,427,1421,1448]" page="514" pageId="89" pageNumber="90" year="1920">
<bibRefCitation id="F168BA813003523DEDF0FA2976A70497" author="Lucas R." box="[231,427,1421,1448]" pageId="89" pageNumber="90" refId="ref128561" refString="Lucas R. 1920. Catalogus alphabeticus generum et subgenerum Coleopterorum orbis terrarum totius (famil., trib., subtr., sect. incl.). Pars I. Nicolaische, Berlin." type="book" year="1920">Lucas 1920: 514</bibRefCitation>
</treatmentCitation>
(catalogue, distribution). —
<treatmentCitation id="1458E1613003523DEE15FA2974FB0497" author="Luederwaldt H." box="[770,1015,1422,1448]" page="8" pageId="89" pageNumber="90" year="1929">
<bibRefCitation id="F168BA813003523DEE15FA2974FB0497" author="Luederwaldt H." box="[770,1015,1422,1448]" pageId="89" pageNumber="90" pagination="603 - 775" refId="ref128687" refString="Luederwaldt H. 1929. As especies brasileiras do genero Pinotus, com algumas consideracoes tambem sobre outras especies. Revista do Museu Paulista 16: 603 - 775." type="journal article" year="1929">Luederwaldt 1929: 8</bibRefCitation>
</treatmentCitation>
, 10 (redescription, key);
<treatmentCitation id="1458E1613003523DE832FA29761A04F4" author="Luederwaldt H." page="369" pageId="89" pageNumber="90" year="1931">
<bibRefCitation id="F168BA813003523DE832FA29761A04F4" author="Luederwaldt H." firstAuthor="b" pageId="89" pageNumber="90" pagination="297 - 304" refId="ref128781" refString="Luederwaldt H. 1931 b. Novos subsidios para o conhecimento do genero Pinotus. Revista de Entomologia 1 (3): 297 - 304." type="journal article" year="1931">1931b: 369</bibRefCitation>
</treatmentCitation>
(key). —
<treatmentCitation id="1458E1613003523DEC84FA16756704F4" author="Paulian R." box="[403,619,1457,1483]" page="234" pageId="89" pageNumber="90" year="1938">
<bibRefCitation id="F168BA813003523DEC84FA16756704F4" author="Paulian R." box="[403,619,1457,1483]" pageId="89" pageNumber="90" pagination="213 - 296" refId="ref134801" refString="Paulian R. 1938. Contribution a l'etude des Canthonides americans [Coleopt. Lamellic.]. Annales de la Societe entomologique de France 107: 213 - 296." type="journal article" year="1938">Paulian 1938: 234</bibRefCitation>
</treatmentCitation>
(key). —
<treatmentCitation id="1458E1613003523DEFF0FA16731604F4" author="Pessoa S. B. &amp; Lane F." box="[743,1050,1457,1483]" page="437" pageId="89" pageNumber="90" year="1941">
<bibRefCitation id="F168BA813003523DEFF0FA16731604F4" author="Pessoa S. B. &amp; Lane F." box="[743,1050,1457,1483]" pageId="89" pageNumber="90" pagination="389 - 504" refId="ref135859" refString="Pessoa S. B. &amp; Lane F. 1941. Coleopteros necrofagos de interesse medico-legal: ensaio monografico sobre a familia Scarabaeidae de S. Paulo e regioes vizinhas. Arquivos de Zoologia do Estado de Sao Paulo 2: 389 - 504. https: // doi. org / 10.11606 / issn. 2176 - 7793.19412389 - 504" type="journal article" year="1941">Pessôa &amp; Lane 1941: 437</bibRefCitation>
</treatmentCitation>
(key). —
<treatmentCitation id="1458E1613003523DE980FA16761404D1" author="Blackwelder R. E." page="206" pageId="89" pageNumber="90" year="1944">
<bibRefCitation id="F168BA813003523DE980FA16761404D1" author="Blackwelder R. E." pageId="89" pageNumber="90" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 206</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="1458E1613003523DEC8CFA73758204D1" author="Martinez A." box="[411,654,1492,1519]" page="139" pageId="89" pageNumber="90" year="1951">
<bibRefCitation id="F168BA813003523DEC8CFA73758204D1" author="Martinez A." box="[411,654,1492,1519]" pageId="89" pageNumber="90" pagination="138 - 142" refId="ref129742" refString="Martinez A. 1951. Notas coleopterologicas V. La invalidez del nombre generico Pinotus Erichson y dos nuevas sinonimias. Anales de la Sociedad Cientifica Argentina 152: 138 - 142." type="journal article" year="1951">Martínez 1951: 139</bibRefCitation>
</treatmentCitation>
(synonym of
<taxonomicName id="52F9BCF33003523DEE56FA7374DB04D1" ID-CoL="44NT" authorityName="Hope" authorityYear="1838" box="[833,983,1492,1518]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B623003523DEE56FA7374DB04D1" box="[833,983,1492,1518]" italics="true" pageId="89" pageNumber="90">Dichotomius</emphasis>
</taxonomicName>
);
<treatmentCitation id="1458E1613003523DEEE1FA73736F04D1" author="Martinez A." box="[1014,1123,1492,1519]" page="80" pageId="89" pageNumber="90" year="1959">
<bibRefCitation id="F168BA813003523DEEE1FA73736F04D1" author="Martinez A." box="[1014,1123,1492,1519]" pageId="89" pageNumber="90" pagination="1 - 126" refId="ref129799" refString="Martinez A. 1959. Catalogo de los Scarabaeidae Argentinos. Revista del Museo Argentino de Ciencias Naturales &quot; Bernardino Rivadavia &quot; 5: 1 - 126." type="journal article" year="1959">1959: 80</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF33003523DEDFDFA50768C072E" ID-CoL="44NT" authorityName="Hope" authorityYear="1838" box="[234,384,1527,1553]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B623003523DEDFDFA50768C072E" box="[234,384,1527,1553]" italics="true" pageId="89" pageNumber="90">Dichotomius</emphasis>
</taxonomicName>
). —
<treatmentCitation id="1458E1613003523DECACFA507424072D" author="Halffter G. &amp; Matthews E. G." box="[443,808,1527,1554]" page="257" pageId="89" pageNumber="90" year="1966">
<bibRefCitation id="F168BA813003523DECACFA507424072D" author="Halffter G. &amp; Matthews E. G." box="[443,808,1527,1554]" pageId="89" pageNumber="90" pagination="1 - 312" refId="ref122681" refString="Halffter G. &amp; Matthews E. G. 1966. The natural history of dung beetles of the subfamily Scarabaeinae (Coleoptera: Scarabaeidae). Folia Entomologica Mexicana 12 - 14: 1 - 312." type="book chapter" year="1966">Halffter &amp; Matthews 1966: 257</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF33003523DE933FA5073B6072E" ID-CoL="44NT" authorityName="Hope" authorityYear="1838" box="[1060,1210,1527,1553]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B623003523DE933FA5073B6072E" box="[1060,1210,1527,1553]" italics="true" pageId="89" pageNumber="90">Dichotomius</emphasis>
</taxonomicName>
). —
<treatmentCitation id="1458E1613003523DE9E1FA5076BA070A" author="Howden H. F. &amp; Young O. P." page="123" pageId="89" pageNumber="90" year="1981">
<bibRefCitation id="F168BA813003523DE9E1FA5076BA070A" author="Howden H. F. &amp; Young O. P." pageId="89" pageNumber="90" pagination="1 - 204" refId="ref124824" refString="Howden H. F. &amp; Young O. P. 1981. Panamanian Scarabaeinae: taxonomy, distribution and habits (Coleoptera, Scarabaeidae). Contributions of the American Entomological Institute 18 (1): 1 - 204." type="journal article" year="1981">Howden &amp; Young 1981: 123</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF33003523DEFD4F9BD7454070B" ID-CoL="44NT" authorityName="Hope" authorityYear="1838" box="[707,856,1562,1588]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B623003523DEFD4F9BD7454070B" box="[707,856,1562,1588]" italics="true" pageId="89" pageNumber="90">Dichotomius</emphasis>
</taxonomicName>
). —
<treatmentCitation id="1458E1613003523DEE80F9BD7206070A" author="Halffter G. &amp; Edmonds W. D." box="[919,1290,1562,1589]" page="137" pageId="89" pageNumber="90" year="1982">
<bibRefCitation id="F168BA813003523DEE80F9BD7206070A" author="Halffter G. &amp; Edmonds W. D." box="[919,1290,1562,1589]" pageId="89" pageNumber="90" refId="ref122441" refString="Halffter G. &amp; Edmonds W. D. 1982. The Nesting Behavior of Dung Beetles (Scarabaeinae). An Ecological and Evolutive Approach. Instituto de Ecologia, Mexico, D. F." type="book" year="1982">Halffter &amp; Edmonds 1982: 137</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF33003523DEC6BF999751E0767" ID-CoL="44NT" authorityName="Hope" authorityYear="1838" box="[380,530,1598,1624]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B623003523DEC6BF999751E0767" box="[380,530,1598,1624]" italics="true" pageId="89" pageNumber="90">Dichotomius</emphasis>
</taxonomicName>
). —
<treatmentCitation id="1458E1613003523DEF58F99974A40767" author="Kohlmann B. &amp; Solis A." box="[591,936,1598,1624]" page="344" pageId="89" pageNumber="90" year="1997">
<bibRefCitation id="F168BA813003523DEF58F99974A40767" author="Kohlmann B. &amp; Solis A." box="[591,936,1598,1624]" pageId="89" pageNumber="90" pagination="343 - 382" refId="ref126101" refString="Kohlmann B. &amp; Solis A. 1997. El genero Dichotomius en Costa Rica. Giornale Italiano di Entomologia 8: 343 - 382." type="journal article" year="1997">Kohlmann &amp; Solís 1997: 344</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF33003523DE9BCF999724D0767" ID-CoL="44NT" authorityName="Hope" authorityYear="1838" box="[1195,1345,1598,1624]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B623003523DE9BCF999724D0767" box="[1195,1345,1598,1624]" italics="true" pageId="89" pageNumber="90">Dichotomius</emphasis>
</taxonomicName>
). —
<treatmentCitation id="1458E1613003523DEDFDF9C676CA0743" author="Ratcliffe B. C." box="[234,454,1633,1660]" page="15" pageId="89" pageNumber="90" year="2002">
<bibRefCitation id="F168BA813003523DEDFDF9C676CA0743" author="Ratcliffe B. C." box="[234,454,1633,1660]" firstAuthor="Ratcliffe" pageId="89" pageNumber="90" pagination="1 - 48" refId="ref136799" refString="Ratcliffe B. C. 2002. A checklist of the Scarabaeoidea of Panama. Zootaxa 32: 1 - 48. https: // doi. org / 10.11646 / zootaxa. 32.1.1" type="journal article" year="2002">Ratcliffe 2002: 15</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF33003523DEFC9F9C674780744" ID-CoL="44NT" authorityName="Hope" authorityYear="1838" box="[734,884,1633,1659]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B623003523DEFC9F9C674780744" box="[734,884,1633,1659]" italics="true" pageId="89" pageNumber="90">Dichotomius</emphasis>
</taxonomicName>
). —
<treatmentCitation id="1458E1613003523DEEAEF9C672090744" author="Solis A. &amp; Kohlmann B." box="[953,1285,1633,1659]" page="6" pageId="89" pageNumber="90" year="2012">
<bibRefCitation id="F168BA813003523DEEAEF9C672090744" author="Solis A. &amp; Kohlmann B." box="[953,1285,1633,1659]" pageId="89" pageNumber="90" pagination="1 - 32" refId="ref139201" refString="Solis A. &amp; Kohlmann B. 2012. Checklist and distribution atlas of the Scarabaeinae of Costa Rica. Zootaxa 3482: 1 - 32. https: // doi. org / 10.11646 / zootaxa. 3482.1.1" type="journal article" year="2012">Solís &amp; Kohlmann 2012: 6</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF33003523DEC93F923751607A1" ID-CoL="44NT" authorityName="Hope" authorityYear="1838" box="[388,538,1668,1694]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B623003523DEC93F923751607A1" box="[388,538,1668,1694]" italics="true" pageId="89" pageNumber="90">Dichotomius</emphasis>
</taxonomicName>
). —
<treatmentCitation id="1458E1613003523DEF49F923736707A0" author="Sarmiento-Garces R. &amp; Amat-Garcia G." box="[606,1131,1668,1695]" page="25" pageId="89" pageNumber="90" year="2014">
<bibRefCitation id="F168BA813003523DEF49F923736707A0" author="Sarmiento-Garces R. &amp; Amat-Garcia G." box="[606,1131,1668,1695]" pageId="89" pageNumber="90" refId="ref138148" refString="Sarmiento-Garces R. &amp; Amat-Garcia G. 2014. Escarabajos del Genero Dichotomius Hope 1838 (Scarabaeidae: Scarabaeinae) en Colombia. Fauna de Colombia, Monografia n ° 4, Universidad Nacional de Colombia, Bogota. https: // doi. org / 10.18257 / raccefyn. 33 (127). 2009.2364" type="book" year="2014">Sarmiento-Garcés &amp; Amat-García 2014: 25</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF33003523DEDFDF900768C07FE" ID-CoL="44NT" authorityName="Hope" authorityYear="1838" box="[234,384,1703,1729]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B623003523DEDFDF900768C07FE" box="[234,384,1703,1729]" italics="true" pageId="89" pageNumber="90">Dichotomius</emphasis>
</taxonomicName>
). —
<treatmentCitation id="1458E1613003523DECAAF90F746007FE" author="Nunes R. V. &amp; Vaz-de-Mello F. Z." box="[445,876,1703,1730]" page="37" pageId="89" pageNumber="90" year="2019">
<bibRefCitation id="F168BA813003523DECAAF90F746007FE" author="Nunes R. V. &amp; Vaz-de-Mello F. Z." box="[445,876,1703,1730]" pageId="89" pageNumber="90" pagination="2231 - 2351" refId="ref133607" refString="Nunes R. V. &amp; Vaz-de-Mello F. Z. 2019. Taxonomic revision of Dichotomius (Cephagonus) Luederwaldt 1929 and the taxonomic status of remaining Dichotomius Hope 1838 subgenera (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini). Journal of Natural History 53 (37 - 38): 2231 - 2351. https: // doi. org / 10.1080 / 00222933.2019.1692088" type="journal article" year="2019">Nunes &amp; Vaz-de-Mello 2019: 3738</bibRefCitation>
</treatmentCitation>
, 22312351 (revision).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C7703003523DEDAAF96D74BA0687" blockId="89.[189,1399,788,2011]" pageId="89" pageNumber="90">
<treatmentCitationGroup id="B5E9E05E3003523DEDAAF96D74BA0687" pageId="89" pageNumber="90">
<taxonomicName id="52F9BCF33003523DEDAAF96D765707DB" ID-CoL="7NR35" authorityName="Haldeman" authorityYear="1848" box="[189,347,1738,1764]" class="Insecta" family="Scarabaeidae" genus="Brachycopris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B623003523DEDAAF96D765707DB" box="[189,347,1738,1764]" italics="true" pageId="89" pageNumber="90">Brachycopris</emphasis>
</taxonomicName>
<treatmentCitation id="1458E1613003523DEC94F96D757707DA" author="Harold E." box="[387,635,1738,1765]" page="1009" pageId="89" pageNumber="90" year="1869">
<bibRefCitation id="F168BA813003523DEC94F96D757707DA" author="Harold E." box="[387,635,1738,1765]" pageId="89" pageNumber="90" refId="ref123629" refString="Harold E. 1869 d. Catalogus coleopterorum. Hucusque descriptorum, synonymicus et systematicus. Tom. IV. Scarabaeidae. E. H. Gummi, Munich. https: // doi. org / 10.5962 / bhl. title. 9089" type="book" year="1869">Harold 1869d: 1009</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF33003523DEE8EF96C74FF07DB" ID-CoL="7PS5K" authorityName="Erichson" authorityYear="1847" box="[921,1011,1739,1764]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B623003523DEE8EF96C74FF07DB" box="[921,1011,1739,1764]" italics="true" pageId="89" pageNumber="90">Pinotus</emphasis>
</taxonomicName>
).
<emphasis id="A78D1B623003523DE906F96C732707DB" box="[1041,1067,1739,1764]" italics="true" pageId="89" pageNumber="90"></emphasis>
<treatmentCitation id="1458E1613003523DE92FF96D720D07DA" author="Gillet J. J. E." box="[1080,1281,1738,1765]" page="59" pageId="89" pageNumber="90" year="1911">
<bibRefCitation id="F168BA813003523DE92FF96D720D07DA" author="Gillet J. J. E." box="[1080,1281,1738,1765]" pageId="89" pageNumber="90" refId="ref121235" refString="Gillet J. J. E. 1911 b. Coleopterorum Catalogus. Pars 38: Scarabaeidae: Coprinae I. W. Junk, Berlin. Available from https: // www. biodiversitylibrary. org / page / 48405679 [accesseed 16 Feb. 2024]." type="url" year="1911">Gillet 1911b: 59</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF33003523DEC68F94976D40638" ID-CoL="7PS5K" authorityName="Erichson" authorityYear="1847" box="[383,472,1774,1799]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B623003523DEC68F94976D40638" box="[383,472,1774,1799]" italics="true" pageId="89" pageNumber="90">Pinotus</emphasis>
</taxonomicName>
). —
<treatmentCitation id="1458E1613003523DEF01F94975D70638" author="Lucas R." box="[534,731,1773,1800]" page="146" pageId="89" pageNumber="90" year="1920">
<bibRefCitation id="F168BA813003523DEF01F94975D70638" author="Lucas R." box="[534,731,1773,1800]" pageId="89" pageNumber="90" refId="ref128561" refString="Lucas R. 1920. Catalogus alphabeticus generum et subgenerum Coleopterorum orbis terrarum totius (famil., trib., subtr., sect. incl.). Pars I. Nicolaische, Berlin." type="book" year="1920">Lucas 1920: 146</bibRefCitation>
</treatmentCitation>
(synonym of
<taxonomicName id="52F9BCF33003523DEE96F94974D70638" ID-CoL="7PS5K" authorityName="Erichson" authorityYear="1847" box="[897,987,1774,1799]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B623003523DEE96F94974D70638" box="[897,987,1774,1799]" italics="true" pageId="89" pageNumber="90">Pinotus</emphasis>
</taxonomicName>
). —
<treatmentCitation id="1458E1613003523DE90EF94972200638" author="Blackwelder R. E." box="[1049,1324,1773,1800]" page="206" pageId="89" pageNumber="90" year="1944">
<bibRefCitation id="F168BA813003523DE90EF94972200638" author="Blackwelder R. E." box="[1049,1324,1773,1800]" pageId="89" pageNumber="90" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 206</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF33003523DEC8FF8B576FE0614" ID-CoL="7PS5K" authorityName="Erichson" authorityYear="1847" box="[408,498,1810,1835]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B623003523DEC8FF8B576FE0614" box="[408,498,1810,1835]" italics="true" pageId="89" pageNumber="90">Pinotus</emphasis>
</taxonomicName>
). —
<treatmentCitation id="1458E1613003523DEF3AF8B6741E0614" author="Martinez A." box="[557,786,1809,1836]" page="139" pageId="89" pageNumber="90" year="1951">
<bibRefCitation id="F168BA813003523DEF3AF8B6741E0614" author="Martinez A." box="[557,786,1809,1836]" pageId="89" pageNumber="90" pagination="138 - 142" refId="ref129742" refString="Martinez A. 1951. Notas coleopterologicas V. La invalidez del nombre generico Pinotus Erichson y dos nuevas sinonimias. Anales de la Sociedad Cientifica Argentina 152: 138 - 142." type="journal article" year="1951">Martínez 1951: 139</bibRefCitation>
</treatmentCitation>
(comments, synonym of
<taxonomicName id="52F9BCF33003523DE921F8B5739C0614" ID-CoL="7PS5K" authorityName="Erichson" authorityYear="1847" box="[1078,1168,1810,1835]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B623003523DE921F8B5739C0614" box="[1078,1168,1810,1835]" italics="true" pageId="89" pageNumber="90">Pinotus</emphasis>
</taxonomicName>
);
<treatmentCitation id="1458E1613003523DE9BEF8B672030614" author="Martinez A." box="[1193,1295,1809,1836]" page="80" pageId="89" pageNumber="90" year="1959">
<bibRefCitation id="F168BA813003523DE9BEF8B672030614" author="Martinez A." box="[1193,1295,1809,1836]" pageId="89" pageNumber="90" pagination="1 - 126" refId="ref129799" refString="Martinez A. 1959. Catalogo de los Scarabaeidae Argentinos. Revista del Museo Argentino de Ciencias Naturales &quot; Bernardino Rivadavia &quot; 5: 1 - 126." type="journal article" year="1959">1959: 80</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF33003523DEC6BF893751E0671" ID-CoL="44NT" authorityName="Hope" authorityYear="1838" box="[380,530,1844,1870]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B623003523DEC6BF893751E0671" box="[380,530,1844,1870]" italics="true" pageId="89" pageNumber="90">Dichotomius</emphasis>
</taxonomicName>
). —
<treatmentCitation id="1458E1613003523DEF58F89374A40671" author="Kohlmann B. &amp; Solis A." box="[591,936,1844,1870]" page="344" pageId="89" pageNumber="90" year="1997">
<bibRefCitation id="F168BA813003523DEF58F89374A40671" author="Kohlmann B. &amp; Solis A." box="[591,936,1844,1870]" pageId="89" pageNumber="90" pagination="343 - 382" refId="ref126101" refString="Kohlmann B. &amp; Solis A. 1997. El genero Dichotomius en Costa Rica. Giornale Italiano di Entomologia 8: 343 - 382." type="journal article" year="1997">Kohlmann &amp; Solís 1997: 344</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF33003523DE9BCF893724D0671" ID-CoL="44NT" authorityName="Hope" authorityYear="1838" box="[1195,1345,1844,1870]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B623003523DE9BCF893724D0671" box="[1195,1345,1844,1870]" italics="true" pageId="89" pageNumber="90">Dichotomius</emphasis>
</taxonomicName>
). —
<treatmentCitation id="1458E1613003523DEDFDF8F076CA064D" author="Ratcliffe B. C." box="[234,454,1879,1906]" page="15" pageId="89" pageNumber="90" year="2002">
<bibRefCitation id="F168BA813003523DEDFDF8F076CA064D" author="Ratcliffe B. C." box="[234,454,1879,1906]" firstAuthor="Ratcliffe" pageId="89" pageNumber="90" pagination="1 - 48" refId="ref136799" refString="Ratcliffe B. C. 2002. A checklist of the Scarabaeoidea of Panama. Zootaxa 32: 1 - 48. https: // doi. org / 10.11646 / zootaxa. 32.1.1" type="journal article" year="2002">Ratcliffe 2002: 15</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF33003523DEFC9F8F07478064E" ID-CoL="44NT" authorityName="Hope" authorityYear="1838" box="[734,884,1879,1905]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B623003523DEFC9F8F07478064E" box="[734,884,1879,1905]" italics="true" pageId="89" pageNumber="90">Dichotomius</emphasis>
</taxonomicName>
). —
<treatmentCitation id="1458E1613003523DEEAEF8F07209064E" author="Solis A. &amp; Kohlmann B." box="[953,1285,1879,1905]" page="6" pageId="89" pageNumber="90" year="2012">
<bibRefCitation id="F168BA813003523DEEAEF8F07209064E" author="Solis A. &amp; Kohlmann B." box="[953,1285,1879,1905]" pageId="89" pageNumber="90" pagination="1 - 32" refId="ref139201" refString="Solis A. &amp; Kohlmann B. 2012. Checklist and distribution atlas of the Scarabaeinae of Costa Rica. Zootaxa 3482: 1 - 32. https: // doi. org / 10.11646 / zootaxa. 3482.1.1" type="journal article" year="2012">Solís &amp; Kohlmann 2012: 6</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF33003523DEC93F8DD751606AB" ID-CoL="44NT" authorityName="Hope" authorityYear="1838" box="[388,538,1914,1940]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B623003523DEC93F8DD751606AB" box="[388,538,1914,1940]" italics="true" pageId="89" pageNumber="90">Dichotomius</emphasis>
</taxonomicName>
). —
<treatmentCitation id="1458E1613003523DEF49F8DD736706AA" author="Sarmiento-Garces R. &amp; Amat-Garcia G." box="[606,1131,1914,1941]" page="25" pageId="89" pageNumber="90" year="2014">
<bibRefCitation id="F168BA813003523DEF49F8DD736706AA" author="Sarmiento-Garces R. &amp; Amat-Garcia G." box="[606,1131,1914,1941]" pageId="89" pageNumber="90" refId="ref138148" refString="Sarmiento-Garces R. &amp; Amat-Garcia G. 2014. Escarabajos del Genero Dichotomius Hope 1838 (Scarabaeidae: Scarabaeinae) en Colombia. Fauna de Colombia, Monografia n ° 4, Universidad Nacional de Colombia, Bogota. https: // doi. org / 10.18257 / raccefyn. 33 (127). 2009.2364" type="book" year="2014">Sarmiento-Garcés &amp; Amat-García 2014: 25</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF33003523DEDFDF839768C0687" ID-CoL="44NT" authorityName="Hope" authorityYear="1838" box="[234,384,1950,1976]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B623003523DEDFDF839768C0687" box="[234,384,1950,1976]" italics="true" pageId="89" pageNumber="90">Dichotomius</emphasis>
</taxonomicName>
). —
<treatmentCitation id="1458E1613003523DECAAF83974570687" author="Nunes R. V. &amp; Vaz-de-Mello F. Z." box="[445,859,1950,1976]" page="2239" pageId="89" pageNumber="90" year="2019">
<bibRefCitation id="F168BA813003523DECAAF83974570687" author="Nunes R. V. &amp; Vaz-de-Mello F. Z." box="[445,859,1950,1976]" pageId="89" pageNumber="90" pagination="2231 - 2351" refId="ref133607" refString="Nunes R. V. &amp; Vaz-de-Mello F. Z. 2019. Taxonomic revision of Dichotomius (Cephagonus) Luederwaldt 1929 and the taxonomic status of remaining Dichotomius Hope 1838 subgenera (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini). Journal of Natural History 53 (37 - 38): 2231 - 2351. https: // doi. org / 10.1080 / 00222933.2019.1692088" type="journal article" year="2019">Nunes &amp; Vaz-de-Mello 2019: 2239</bibRefCitation>
</treatmentCitation>
(cited).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C7703003523DEDAAF866732906E4" blockId="89.[189,1399,788,2011]" box="[189,1061,1985,2011]" pageId="89" pageNumber="90">
<treatmentCitationGroup id="B5E9E05E3003523DEDAAF866732906E4" box="[189,1061,1985,2011]" pageId="89" pageNumber="90">
<taxonomicName id="52F9BCF33003523DEDAAF866769B06E4" ID-CoL="7PWGK" authorityName="Ferreira" authorityYear="1970" box="[189,407,1985,2011]" class="Insecta" family="Scarabaeidae" genus="Pseudoheliocopris" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B623003523DEDAAF866769B06E4" box="[189,407,1985,2011]" italics="true" pageId="89" pageNumber="90">Pseudoheliocopris</emphasis>
</taxonomicName>
<treatmentCitation id="1458E1613003523DECA2F86675BC06E4" author="Cambefort Y. " box="[437,688,1985,2011]" page="270" pageId="89" pageNumber="90" year="1976">
<bibRefCitation id="F168BA813003523DECA2F86675BC06E4" box="[437,688,1985,2011]" firstAuthor="Cambefort" pageId="89" pageNumber="90" year="1976">Cambefort 1976: 270</bibRefCitation>
</treatmentCitation>
(synonymy with
<taxonomicName id="52F9BCF33003523DEE69F866731806E4" ID-CoL="44NT" authorityName="Hope" authorityYear="1838" box="[894,1044,1985,2011]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B623003523DEE69F866731806E4" box="[894,1044,1985,2011]" italics="true" pageId="89" pageNumber="90">Dichotomius</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
</treatment>
</document>