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<document id="C18AFF8D26743C8872CFD0C07DAE3AA2" ID-DOI="10.5852/ejt.2024.959.2677" ID-ISSN="2118-9773" ID-Zenodo-Dep="13945464" ID-ZooBank="30872C13-516F-42FA-AFA7-30ADC6BF1BAF" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1729166637250" checkinUser="plazi" docAuthor="Rosa, Cecilia Lozano De La, Cupello, Mario &amp; Vaz-De-Mello, Fernando Z." docDate="2024" docId="1D5076663002523CECFDFD49732007F0" docLanguage="en" docName="EJT.2024.959.1-272.pdf" docOrigin="European Journal of Taxonomy 959 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2677/12401" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Diabroctis venezuelensis Martinez &amp; Clavijo 1990" docType="treatment" docVersion="6" lastPageNumber="89" masterDocId="E1690E1E305A5264ED17FFA7770C013F" masterDocTitle="The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution" masterLastPageNumber="272" masterPageNumber="1" pageNumber="89" updateTime="1731448046962" updateUser="carolina" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="8BB6B10BFB44FA45C577C1D2DC3D23CA">
<mods:title id="D0EEEBBE7C57BB85824DAF3E8A8A951F">The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution</mods:title>
</mods:titleInfo>
<mods:name id="6C9C892512B4E48D0EAAAEB1C7F49A9C" type="personal">
<mods:role id="767A7F91687F18C21A10104958EE22D5">
<mods:roleTerm id="09A104CB36DA8DB479880121380E2492">Author</mods:roleTerm>
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<mods:namePart id="A036ED84AD1ACF345791B189280150EE">Rosa, Cecilia Lozano De La</mods:namePart>
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<mods:affiliation id="C057F107F8B982293C952B328623E4F2">Instituto Venezolano de Investigaciones Científicas, Centro de Estudios Botánicos y Agroforestales, Laboratorio de Ecología Espacial, Maracaibo, estado Zulia, Venezuela.</mods:affiliation>
<mods:nameIdentifier id="277C3FB418D49BDC677B500DBA87AF7B" type="email">lozanoceci@gmail.com</mods:nameIdentifier>
</mods:name>
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<mods:namePart id="B80D34EEE4A673808B2E10FA911086FD">Cupello, Mario</mods:namePart>
<mods:nameIdentifier id="70EE1CCAE9A8FA26372EB17370844EE3" type="LSID">BDB03C18-7095-4EAA-8BDD-03CB4F79676D</mods:nameIdentifier>
<mods:affiliation id="8FE894C0785C61890FFFEEFF8A83A44E">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Curitiba, Paraná, Brazil. Currently at Department of Entomology, Texas A &amp; M University, College Station, TX, 77843, USA.</mods:affiliation>
<mods:nameIdentifier id="A830B211D082DEBC10F09F01DABCDF70" type="email">mcupello@hotmail.com</mods:nameIdentifier>
</mods:name>
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<mods:role id="0BE279C57241DCE559F01431020D907C">
<mods:roleTerm id="032E15842CA481A33AC168967BAA3045">Author</mods:roleTerm>
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<mods:namePart id="6A228CE55AE75F2FDF6B496589164E0F">Vaz-De-Mello, Fernando Z.</mods:namePart>
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<mods:affiliation id="9F3E34542EB7E9E007D659B202740E39">Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, Cuiabá, Mato Grosso, Brazil.</mods:affiliation>
<mods:nameIdentifier id="4CC21C93489F5C6258772BBA97E77C6C" type="email">vazdemello@gmail.com</mods:nameIdentifier>
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<mods:typeOfResource id="D9DEDF731A720E1F39569BF59E3A0B82">text</mods:typeOfResource>
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<mods:title id="7065D6923886023915903B3179714ACD">European Journal of Taxonomy</mods:title>
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<mods:date id="F9277204068422CB9C7A0DE633416F83">2024</mods:date>
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<mods:number id="0A3A8E3E313956B0618657DB2978B3E3">2024-10-11</mods:number>
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<mods:number id="89B7A26DF670BC3554FDB342A246493C">959</mods:number>
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<treatment id="1D5076663002523CECFDFD49732007F0" ID-DOI="http://doi.org/10.5281/zenodo.13945746" ID-Zenodo-Dep="13945746" LSID="urn:lsid:plazi:treatment:1D5076663002523CECFDFD49732007F0" httpUri="http://treatment.plazi.org/id/1D5076663002523CECFDFD49732007F0" lastPageNumber="89" pageId="88" pageNumber="89">
<subSubSection id="DDE394FB3002523CECFDFD4973460236" box="[490,1098,750,777]" pageId="88" pageNumber="89" type="nomenclature">
<paragraph id="9546C7703002523CECFDFD4973460236" blockId="88.[490,1098,750,813]" box="[490,1098,750,777]" pageId="88" pageNumber="89">
<heading id="CE0E701C3002523CECFDFD4973460236" box="[490,1098,750,777]" centered="true" fontSize="11" level="2" pageId="88" pageNumber="89" reason="2">
<taxonomicName id="52F9BCF33002523CECFDFD4973460236" ID-CoL="356TB" authority="Martinez &amp; Clavijo, 1990" authorityName="Martinez &amp; Clavijo" authorityYear="1990" box="[490,1098,750,777]" class="Insecta" family="Scarabaeidae" genus="Diabroctis" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="venezuelensis">
<emphasis id="A78D1B623002523CECFDFD4974180236" bold="true" box="[490,788,750,777]" italics="true" pageId="88" pageNumber="89">Diabroctis venezuelensis</emphasis>
<bibRefCitation id="F168BA813002523CEE0CFD4873460236" author="Martinez A. &amp; Martinez A." box="[795,1098,751,777]" pageId="88" pageNumber="89" pagination="159 - 163" refId="ref130132" refString="Martinez A. &amp; Martinez A. 1990. Dos nuevas especies de Ateuchus Weber (Col. Scarab. Ateuchina). Boletin de Entomologia Venezolana (N. S.) 5 (20): 159 - 163." type="journal article" year="1990">Martínez &amp; Clavijo, 1990</bibRefCitation>
</taxonomicName>
</heading>
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<subSubSection id="DDE394FB3002523CEFF1FCB574410212" box="[742,845,786,813]" pageId="88" pageNumber="89" type="description">
<paragraph id="9546C7703002523CEFF1FCB574410212" blockId="88.[490,1098,750,813]" box="[742,845,786,813]" pageId="88" pageNumber="89">
<figureCitation id="0DC2DBF53002523CEFF1FCB574410212" box="[742,845,786,813]" captionStart="Fig" captionStartId="87.[189,232,1963,1989]" captionTargetBox="[279,1302,278,1879]" captionTargetId="figure-12@87.[268,1320,265,1907]" captionTargetPageId="87" captionText="Fig. 35. Species distribution. A. Diabroctis cadmus (Harold, 1868). B. Diabroctis venezuelensis Martínez &amp; Clavijo, 1990. Green diamond = CEMT collection data; black circle = literature data." figureDoi="http://doi.org/10.5281/zenodo.13945559" httpUri="https://zenodo.org/record/13945559/files/figure.png" pageId="88" pageNumber="89">Fig. 35B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB3002523CEDAAFCF4753F05BD" pageId="88" pageNumber="89" type="reference_group">
<paragraph id="9546C7703002523CEDAAFCF476F1028B" blockId="88.[189,1399,851,948]" pageId="88" pageNumber="89">
<treatmentCitationGroup id="B5E9E05E3002523CEDAAFCF476F1028B" pageId="88" pageNumber="89">
<treatmentCitation id="1458E1613002523CEDAAFCF474B70251" author="Martinez A. &amp; Martinez A." box="[189,955,851,878]" page="147" pageId="88" pageNumber="89" year="1990">
<taxonomicName id="52F9BCF33002523CEDAAFCF474B70251" ID-CoL="356TB" authority="Martinez &amp; Clavijo, 1990: 147" authorityName="Martinez &amp; Clavijo" authorityPageNumber="147" authorityYear="1990" box="[189,955,851,878]" class="Insecta" family="Scarabaeidae" genus="Diabroctis" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="subSpecies" species="mimas" subSpecies="venezuelensis">
<emphasis id="A78D1B623002523CEDAAFCF475370252" box="[189,571,851,877]" italics="true" pageId="88" pageNumber="89">Diabroctis mimas venezuelensis</emphasis>
<bibRefCitation id="F168BA813002523CEF52FCF374B70251" author="Martinez A. &amp; Martinez A." box="[581,955,851,878]" pageId="88" pageNumber="89" pagination="159 - 163" refId="ref130132" refString="Martinez A. &amp; Martinez A. 1990. Dos nuevas especies de Ateuchus Weber (Col. Scarab. Ateuchina). Boletin de Entomologia Venezolana (N. S.) 5 (20): 159 - 163." type="journal article" year="1990">Martínez &amp; Clavijo, 1990: 147</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description). Type locality:
<collectingCountry id="EDEE87E03002523CEDFDFCD0766B02AE" box="[234,359,887,913]" name="Venezuela" pageId="88" pageNumber="89">Venezuela</collectingCountry>
:
<collectingRegion id="573D09923002523CEC66FCD076F402AE" box="[369,504,887,913]" country="Venezuela" name="Portuguesa" pageId="88" pageNumber="89">Portuguesa</collectingRegion>
:Araure. Name-bearing type:
<typeStatus id="4A4279D23002523CEE5CFCD074BE02AE" box="[843,946,887,913]" pageId="88" pageNumber="89" type="holotype">holotype</typeStatus>
(MIZA) (
<bibRefCitation id="F168BA813002523CE934FCD073FE02AE" author="Valois M. C. &amp; Harada L. &amp; Vaz-de-Mello F. Z. &amp; Silva F." box="[1059,1266,887,913]" pageId="88" pageNumber="89" pagination="347 - 374" refId="ref140212" refString="Valois M. C., Harada L., Vaz-de-Mello F. Z. &amp; Silva F. 2018. Synopsis of the genus Diabroctis Gistel, 1857 (Coleoptera: Scarabaeidae: Scarabaeinae) with a new species description. Insect Systematics &amp; Evolution 51: 347 - 374. https: // doi. org / 10.1163 / 1876312 X- 00002203" type="journal article" year="2018">
Valois
<emphasis id="A78D1B623002523CE964FCD073A002AE" box="[1139,1196,887,913]" italics="true" pageId="88" pageNumber="89">et al.</emphasis>
2018
</bibRefCitation>
), examined from photos by FZVM.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C7703002523CEDAAFC7C73260503" blockId="88.[189,1399,987,1154]" pageId="88" pageNumber="89">
<treatmentCitationGroup id="B5E9E05E3002523CEDAAFC7C73260503" pageId="88" pageNumber="89">
<taxonomicName id="52F9BCF33002523CEDAAFC7C753F02CA" ID-CoL="356TB" authorityName="Martinez &amp; Clavijo" authorityYear="1990" box="[189,563,987,1013]" class="Insecta" family="Scarabaeidae" genus="Diabroctis" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="subSpecies" species="mimas" subSpecies="venezuelensis">
<emphasis id="A78D1B623002523CEDAAFC7C753F02CA" box="[189,563,987,1013]" italics="true" pageId="88" pageNumber="89">Diabroctis mimas venezuelensis</emphasis>
</taxonomicName>
–
<treatmentCitation id="1458E1613002523CEF51FC7C741502CA" author="Arnaud P." box="[582,793,987,1013]" page="18" pageId="88" pageNumber="89" year="2002">
<bibRefCitation id="F168BA813002523CEF51FC7C741502CA" author="Arnaud P." box="[582,793,987,1013]" pageId="88" pageNumber="89" refId="ref109736" refString="Arnaud P. 2002 b. Les Coleopteres du Monde, Vol. 28. Phanaeini. Dendropaemon, Tetramereia, Homalotarsus, Megatharsis, Diabroctis, Coprophanaeus, Oxysternon, Phanaeus, Sulcophanaeus. Hillside Books, Canterbury, England." type="book" year="2002">Arnaud 2002b: 18</bibRefCitation>
</treatmentCitation>
(cited as
<taxonomicName id="52F9BCF33002523CEE9FFC7C76870526" authority="Martinez &amp; Clavijo, 1990" authorityName="Martinez &amp; Clavijo" authorityYear="1990" class="Insecta" family="Scarabaeidae" genus="Diabroctis" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="subSpecies" species="mimas" subSpecies="venezuelina">
<emphasis id="A78D1B623002523CEE9FFC7C73E402CA" box="[904,1256,987,1013]" italics="true" pageId="88" pageNumber="89">Diabroctis mimas venezuelina</emphasis>
<bibRefCitation id="F168BA813002523CE9F9FC7C76870526" author="Martinez A. &amp; Martinez A." pageId="88" pageNumber="89" pagination="159 - 163" refId="ref130132" refString="Martinez A. &amp; Martinez A. 1990. Dos nuevas especies de Ateuchus Weber (Col. Scarab. Ateuchina). Boletin de Entomologia Venezolana (N. S.) 5 (20): 159 - 163." type="journal article" year="1990">Martínez &amp; Clavijo, 1990</bibRefCitation>
</taxonomicName>
, cited as synonym of
<taxonomicName id="52F9BCF33002523CEF90FC59745B0527" ID-CoL="356T8" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[647,855,1022,1048]" class="Insecta" family="Scarabaeidae" genus="Diabroctis" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="mimas">
<emphasis id="A78D1B623002523CEF90FC59745B0527" box="[647,855,1022,1048]" italics="true" pageId="88" pageNumber="89">Diabroctis mimas</emphasis>
</taxonomicName>
). —
<treatmentCitation id="1458E1613002523CEE84FC58735A0526" author="Gamez J." box="[915,1110,1023,1049]" page="48" pageId="88" pageNumber="89" year="2004">
<bibRefCitation id="F168BA813002523CEE84FC58735A0526" author="Gamez J." box="[915,1110,1023,1049]" pageId="88" pageNumber="89" pagination="43 - 60" refId="ref119830" refString="Gamez J. 2004 (&quot; 2002 &quot;). Phanaeini (Coleoptera: Scarabaeinae) de la Cordillera de los Andes, depresion de Maracaibo y Llanos de Venezuela. Memoria de la Fundacion La Salle de Ciencias Naturales 158: 43 - 60." type="journal article" year="2004">Gámez 2004: 48</bibRefCitation>
</treatmentCitation>
, 51–52, 58 (key, list). —
<treatmentCitation id="1458E1613002523CEDFDFB8575480503" author="Hamel-Leigue A. &amp; Herzog S. K. &amp; Mann D. J. &amp; Larsen T. H. &amp; Gill B. D. &amp; Edmonds W. D. &amp; Spector S." box="[234,580,1058,1084]" page="60" pageId="88" pageNumber="89" year="2009">
<bibRefCitation id="F168BA813002523CEDFDFB8575480503" author="Hamel-Leigue A. &amp; Herzog S. K. &amp; Mann D. J. &amp; Larsen T. H. &amp; Gill B. D. &amp; Edmonds W. D. &amp; Spector S." box="[234,580,1058,1084]" pageId="88" pageNumber="89" pagination="43 - 95" refId="ref122950" refString="Hamel-Leigue A., Herzog S. K., Mann D. J., Larsen T. H., Gill B. D., Edmonds W. D., Spector S. 2009. Distribucion e historia natural de escarabajos coprofagos de la tribu Phanaeini (Coleoptera: Scarabaeidae: Scarabaeinae) en Bolivia. Kempffiana 5: 43 - 95." type="journal article" year="2009">
Hamel-Leigue
<emphasis id="A78D1B623002523CEC8CFB8576DA0503" box="[411,470,1058,1084]" italics="true" pageId="88" pageNumber="89">et al.</emphasis>
2009: 60
</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF33002523CEE5FFB8573150504" ID-CoL="356T8" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[840,1049,1058,1084]" class="Insecta" family="Scarabaeidae" genus="Diabroctis" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="mimas">
<emphasis id="A78D1B623002523CEE5FFB8573150504" box="[840,1049,1058,1084]" italics="true" pageId="88" pageNumber="89">Diabroctis mimas</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C7703002523CEDAAFBE2753F05BD" blockId="88.[189,1399,987,1154]" pageId="88" pageNumber="89">
<treatmentCitationGroup id="B5E9E05E3002523CEDAAFBE2753F05BD" pageId="88" pageNumber="89">
<taxonomicName id="52F9BCF33002523CEDAAFBE276ED0560" ID-CoL="356TB" authorityName="Martinez &amp; Clavijo" authorityYear="1990" box="[189,481,1093,1119]" class="Insecta" family="Scarabaeidae" genus="Diabroctis" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="venezuelensis">
<emphasis id="A78D1B623002523CEDAAFBE276ED0560" box="[189,481,1093,1119]" italics="true" pageId="88" pageNumber="89">Diabroctis venezuelensis</emphasis>
</taxonomicName>
–
<treatmentCitation id="1458E1613002523CEF17FBE27407055F" author="Valois M. C. &amp; Harada L. &amp; Vaz-de-Mello F. Z. &amp; Silva F." box="[512,779,1093,1120]" page="357" pageId="88" pageNumber="89" year="2018">
<bibRefCitation id="F168BA813002523CEF17FBE27407055F" author="Valois M. C. &amp; Harada L. &amp; Vaz-de-Mello F. Z. &amp; Silva F." box="[512,779,1093,1120]" pageId="88" pageNumber="89" pagination="347 - 374" refId="ref140212" refString="Valois M. C., Harada L., Vaz-de-Mello F. Z. &amp; Silva F. 2018. Synopsis of the genus Diabroctis Gistel, 1857 (Coleoptera: Scarabaeidae: Scarabaeinae) with a new species description. Insect Systematics &amp; Evolution 51: 347 - 374. https: // doi. org / 10.1163 / 1876312 X- 00002203" type="journal article" year="2018">
Valois
<emphasis id="A78D1B623002523CEF46FBE175800560" box="[593,652,1093,1119]" italics="true" pageId="88" pageNumber="89">et al.</emphasis>
2018: 357
</bibRefCitation>
</treatmentCitation>
(revision). —
<treatmentCitation id="1458E1613002523CEEAEFBE272320560" ID-DOI="http://doi.org/10.5281/zenodo.3683833" ID-Zenodo-Dep="3683833" author="Hielkema A. J. &amp; Hielkema M. A." box="[953,1342,1093,1119]" httpUri="http://treatment.plazi.org/id/03D7FF1CFFADFF8BFF48FE00344FFA23" page="100" pageId="88" pageNumber="89" year="2019">
<bibRefCitation id="F168BA813002523CEEAEFBE272320560" author="Hielkema A. J. &amp; Hielkema M. A." box="[953,1342,1093,1119]" pageId="88" pageNumber="89" pagination="1 - 306" refId="ref124225" refString="Hielkema A. J. &amp; Hielkema M. A. 2019. An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas. Insecta Mundi 732 (732): 1 - 306." type="journal article" year="2019">Hielkema &amp; Hielkema 2019:100</bibRefCitation>
</treatmentCitation>
, 292 (catalogue for the Guianas).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB3002523CEDAAFB0E720D042A" pageId="88" pageNumber="89" type="materials_examined">
<paragraph id="9546C7703002523CEDAAFB0E76A605FC" blockId="88.[189,426,1193,1219]" box="[189,426,1193,1219]" pageId="88" pageNumber="89">
<heading id="CE0E701C3002523CEDAAFB0E76A605FC" bold="true" box="[189,426,1193,1219]" fontSize="11" level="3" pageId="88" pageNumber="89" reason="3">
<emphasis id="A78D1B623002523CEDAAFB0E76A605FC" bold="true" box="[189,426,1193,1219]" pageId="88" pageNumber="89">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="9546C7703002523CEDAAFB7F720D042A" blockId="88.[189,1399,1239,1302]" pageId="88" pageNumber="89">
<materialsCitation id="2591CD2D3002523CEDAAFB7F725405CD" box="[189,1368,1239,1267]" collectingDate="2004-05-15" collectionCode="CEMT" country="Venezuela" elevation="100" location="Rio Paraguaz" pageId="88" pageNumber="89" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Bolivar">
<collectingCountry id="EDEE87E03002523CEDAAFB7F766705CD" box="[189,363,1240,1266]" name="Venezuela" pageId="88" pageNumber="89">VENEZUELA</collectingCountry>
‒
<emphasis id="A78D1B623002523CEC86FB7076E105CD" bold="true" box="[401,493,1239,1266]" pageId="88" pageNumber="89">
<collectingRegion id="573D09923002523CEC86FB7076E105CD" box="[401,493,1239,1266]" country="Venezuela" name="Bolivar" pageId="88" pageNumber="89">Bolívar</collectingRegion>
</emphasis>
•
<specimenCount id="83FF0CF93002523CEF07FB7F754F05CC" box="[528,579,1240,1267]" count="1" pageId="88" pageNumber="89" type="male">1 ♂</specimenCount>
,
<specimenCount id="83FF0CF93002523CEF42FB7F759105CC" box="[597,669,1240,1267]" count="2" pageId="88" pageNumber="89" type="female">2 ♀♀</specimenCount>
; near
<location id="902691AB3002523CEFF8FB7F749805CC" LSID="urn:lsid:plazi:treatment:1D5076663002523CECFDFD49732007F0:902691AB3002523CEFF8FB7F749805CC" box="[751,916,1240,1267]" country="Venezuela" name="Rio Paraguaz" pageId="88" pageNumber="89" stateProvince="Bolivar">Río Paraguaz</location>
,
<elevation id="1ED420433002523CEEB2FB7F733B05CD" box="[933,1079,1240,1267]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="88" pageNumber="89" unit="m" value="100.0">
<quantity id="52016A953002523CEEB2FB7F74F805CC" box="[933,1012,1240,1267]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="88" pageNumber="89" unit="m" value="100.0">100 m</quantity>
a.s.l.
</elevation>
,
<date id="E147E1B03002523CE95DFB7F73F805CD" box="[1098,1268,1240,1267]" pageId="88" pageNumber="89" value="2004-05-15">
<collectingDate id="F10318583002523CE95DFB7F73F805CD" box="[1098,1268,1240,1267]" pageId="88" pageNumber="89" value="2004-05-15">15 May 2004</collectingDate>
</date>
;
<collectionCode id="F3E85FB53002523CE812FB7F725405CD" box="[1285,1368,1240,1266]" pageId="88" pageNumber="89">CEMT</collectionCode>
</materialsCitation>
. ‒
<materialsCitation id="2591CD2D3002523CEDAAFB5C7591042A" box="[189,669,1275,1302]" collectionCode="CEMT" collectorName="Araure" country="Venezuela" location="Araure" pageId="88" pageNumber="89" specimenCount="1" specimenCount-male="1" stateProvince="Portuguesa">
<emphasis id="A78D1B623002523CEDAAFB5C7647042A" bold="true" box="[189,331,1275,1301]" pageId="88" pageNumber="89">
<collectingRegion id="573D09923002523CEDAAFB5C7647042A" box="[189,331,1275,1301]" country="Venezuela" name="Portuguesa" pageId="88" pageNumber="89">Portuguesa</collectingRegion>
</emphasis>
•
<specimenCount id="83FF0CF93002523CEC73FB5C769F0429" box="[356,403,1275,1302]" count="1" pageId="88" pageNumber="89" type="male">1 ♂</specimenCount>
;
<location id="902691AB3002523CEC88FB5C76FC042A" LSID="urn:lsid:plazi:treatment:1D5076663002523CECFDFD49732007F0:902691AB3002523CEC88FB5C76FC042A" box="[415,496,1275,1301]" country="Venezuela" name="Araure" pageId="88" pageNumber="89" stateProvince="Portuguesa">Araure</location>
; 1978;
<collectionCode id="F3E85FB53002523CEF5DFB5C7591042A" box="[586,669,1275,1301]" pageId="88" pageNumber="89">CEMT</collectionCode>
</materialsCitation>
. ‒
<materialsCitation id="2591CD2D3002523CEFA9FB5C73F1042A" box="[702,1277,1275,1302]" collectingDate="1978-05-15" collectionCode="CEMT" country="Venezuela" location="Tinaquillo" pageId="88" pageNumber="89" specimenCount="1" specimenCount-female="1" stateProvince="Cojedes">
<emphasis id="A78D1B623002523CEFA9FB5C742D042A" bold="true" box="[702,801,1275,1301]" pageId="88" pageNumber="89">
<collectingRegion id="573D09923002523CEFA9FB5C742D042A" box="[702,801,1275,1301]" country="Venezuela" name="Cojedes" pageId="88" pageNumber="89">Cojedes</collectingRegion>
</emphasis>
•
<specimenCount id="83FF0CF93002523CEE2CFB5C74640429" box="[827,872,1275,1302]" count="1" pageId="88" pageNumber="89" type="female">1 ♀</specimenCount>
;
<location id="902691AB3002523CEE62FB5C74FE042A" LSID="urn:lsid:plazi:treatment:1D5076663002523CECFDFD49732007F0:902691AB3002523CEE62FB5C74FE042A" box="[885,1010,1275,1301]" country="Venezuela" name="Tinaquillo" pageId="88" pageNumber="89" stateProvince="Cojedes">Tinaquillo</location>
;
<date id="E147E1B03002523CEEE9FB5C7392042A" box="[1022,1182,1275,1302]" pageId="88" pageNumber="89" value="1978-05-15">
<collectingDate id="F10318583002523CEEE9FB5C7392042A" box="[1022,1182,1275,1302]" pageId="88" pageNumber="89" value="1978-05-15">15 May 1978</collectingDate>
</date>
;
<collectionCode id="F3E85FB53002523CE9BDFB5C73F1042A" box="[1194,1277,1275,1301]" pageId="88" pageNumber="89">CEMT</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB3002523CEDAAFA9B76DE0726" pageId="88" pageNumber="89" type="distribution">
<paragraph id="9546C7703002523CEDAAFA9B765A0469" blockId="88.[189,342,1340,1366]" box="[189,342,1340,1366]" pageId="88" pageNumber="89">
<heading id="CE0E701C3002523CEDAAFA9B765A0469" bold="true" box="[189,342,1340,1366]" fontSize="11" level="3" pageId="88" pageNumber="89" reason="3">
<emphasis id="A78D1B623002523CEDAAFA9B765A0469" bold="true" box="[189,342,1340,1366]" pageId="88" pageNumber="89">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="9546C7703002523CEDAAFACC76B004BA" blockId="88.[189,444,1387,1413]" box="[189,444,1387,1413]" pageId="88" pageNumber="89">
<collectingCountry id="EDEE87E03002523CEDAAFACC763A04BA" box="[189,310,1387,1413]" name="Venezuela" pageId="88" pageNumber="89">Venezuela</collectingCountry>
(endemic).
</paragraph>
<paragraph id="9546C7703002523CEDAAFA0B76E104F8" blockId="88.[189,493,1452,1479]" box="[189,493,1452,1479]" pageId="88" pageNumber="89">
<heading id="CE0E701C3002523CEDAAFA0B76E104F8" bold="true" box="[189,493,1452,1479]" fontSize="11" level="3" pageId="88" pageNumber="89" reason="3">
<emphasis id="A78D1B623002523CEDAAFA0B76E104F8" bold="true" box="[189,493,1452,1479]" pageId="88" pageNumber="89">
Subregions of
<collectingCountry id="EDEE87E03002523CEC78FA0A76E104F8" box="[367,493,1453,1479]" name="Venezuela" pageId="88" pageNumber="89">Venezuela</collectingCountry>
</emphasis>
</heading>
</paragraph>
<paragraph id="9546C7703002523CEDAAFA7C76DE0726" blockId="88.[188,1398,1499,1561]" pageId="88" pageNumber="89">
Plains, System of hills and low piedmont mountains of the Guiana Shield, Andes mountains, and Central
<collectingRegion id="573D09923002523CEDABFA5877F30726" box="[188,255,1535,1561]" country="New Zealand" name="West Coast" pageId="88" pageNumber="89">Coast</collectingRegion>
Mountain Range.
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB3002523CEDAAF9E7732007F0" pageId="88" pageNumber="89" type="reference_group">
<paragraph id="9546C7703002523CEDAAF9E776A80765" blockId="88.[189,420,1600,1626]" box="[189,420,1600,1626]" pageId="88" pageNumber="89">
<heading id="CE0E701C3002523CEDAAF9E776A80765" bold="true" box="[189,420,1600,1626]" fontSize="11" level="3" pageId="88" pageNumber="89" reason="3">
<emphasis id="A78D1B623002523CEDAAF9E776A80765" bold="true" box="[189,420,1600,1626]" pageId="88" pageNumber="89">Literature records</emphasis>
</heading>
</paragraph>
<paragraph id="9546C7703002523CEDAAF9C9732007F0" blockId="88.[189,1399,1646,1744]" pageId="88" pageNumber="89">
<bibRefCitation id="F168BA813002523CEDAAF9C9753A07B7" author="Martinez A. &amp; Martinez A." box="[189,566,1646,1672]" pageId="88" pageNumber="89" pagination="159 - 163" refId="ref130132" refString="Martinez A. &amp; Martinez A. 1990. Dos nuevas especies de Ateuchus Weber (Col. Scarab. Ateuchina). Boletin de Entomologia Venezolana (N. S.) 5 (20): 159 - 163." type="journal article" year="1990">Martínez &amp; Clavijo 1990: 147</bibRefCitation>
(
<collectingCountry id="EDEE87E03002523CEF5CF9C975C607B7" box="[587,714,1646,1672]" name="Venezuela" pageId="88" pageNumber="89">Venezuela</collectingCountry>
:
<collectingRegion id="573D09923002523CEFCDF9C9743007B7" box="[730,828,1646,1672]" country="Venezuela" name="Cojedes" pageId="88" pageNumber="89">Cojedes</collectingRegion>
;
<collectingRegion id="573D09923002523CEE59F9C874D907B6" box="[846,981,1647,1673]" country="Venezuela" name="Portuguesa" pageId="88" pageNumber="89">Portuguesa</collectingRegion>
: Araure;
<collectingRegion id="573D09923002523CE95DF9C8738C07B6" box="[1098,1152,1647,1673]" country="Venezuela" name="Lara" pageId="88" pageNumber="89">Lara</collectingRegion>
[actually,
<collectingRegion id="573D09923002523CE813F9C8726007B6" box="[1284,1388,1647,1673]" country="Venezuela" name="Yaracuy" pageId="88" pageNumber="89">Yaracuy</collectingRegion>
]: Sabana de Parra). —
<bibRefCitation id="F168BA813002523CECD4F93675920793" author="Arnaud P." box="[451,670,1681,1708]" pageId="88" pageNumber="89" refId="ref109736" refString="Arnaud P. 2002 b. Les Coleopteres du Monde, Vol. 28. Phanaeini. Dendropaemon, Tetramereia, Homalotarsus, Megatharsis, Diabroctis, Coprophanaeus, Oxysternon, Phanaeus, Sulcophanaeus. Hillside Books, Canterbury, England." type="book" year="2002">Arnaud 2002b: 18</bibRefCitation>
(
<collectingCountry id="EDEE87E03002523CEFA6F936743D0794" box="[689,817,1681,1707]" name="Venezuela" pageId="88" pageNumber="89">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA813002523CEE78F935733B0793" author="Gamez J." box="[879,1079,1682,1708]" pageId="88" pageNumber="89" pagination="43 - 60" refId="ref119830" refString="Gamez J. 2004 (&quot; 2002 &quot;). Phanaeini (Coleoptera: Scarabaeinae) de la Cordillera de los Andes, depresion de Maracaibo y Llanos de Venezuela. Memoria de la Fundacion La Salle de Ciencias Naturales 158: 43 - 60." type="journal article" year="2004">Gámez 2004: 48</bibRefCitation>
(subregion of Llanos). —
<bibRefCitation id="F168BA813002523CEDAAF912750D07EF" author="Valois M. C. &amp; Harada L. &amp; Vaz-de-Mello F. Z. &amp; Silva F." box="[189,513,1717,1744]" pageId="88" pageNumber="89" pagination="347 - 374" refId="ref140212" refString="Valois M. C., Harada L., Vaz-de-Mello F. Z. &amp; Silva F. 2018. Synopsis of the genus Diabroctis Gistel, 1857 (Coleoptera: Scarabaeidae: Scarabaeinae) with a new species description. Insect Systematics &amp; Evolution 51: 347 - 374. https: // doi. org / 10.1163 / 1876312 X- 00002203" type="journal article" year="2018">
Valois
<emphasis id="A78D1B623002523CEC19F911764D07F0" box="[270,321,1717,1743]" italics="true" pageId="88" pageNumber="89">et al</emphasis>
. 2018: 357–358
</bibRefCitation>
(
<collectingCountry id="EDEE87E03002523CEF07F912758307F0" box="[528,655,1717,1743]" name="Venezuela" pageId="88" pageNumber="89">Venezuela</collectingCountry>
:
<collectingRegion id="573D09923002523CEF8DF91275F007F0" box="[666,764,1717,1743]" country="Venezuela" name="Cojedes" pageId="88" pageNumber="89">Cojedes</collectingRegion>
,
<collectingRegion id="573D09923002523CEE10F912745307F0" box="[775,863,1717,1743]" country="Venezuela" name="Bolivar" pageId="88" pageNumber="89">Bolívar</collectingRegion>
and
<collectingRegion id="573D09923002523CEE8FF912732C07F0" box="[920,1056,1717,1743]" country="Venezuela" name="Portuguesa" pageId="88" pageNumber="89">Portuguesa</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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