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<document id="C13CB5D1794E0AD5D1B25B93DA5911E6" ID-DOI="10.5852/ejt.2024.959.2677" ID-ISSN="2118-9773" ID-Zenodo-Dep="13945464" ID-ZooBank="30872C13-516F-42FA-AFA7-30ADC6BF1BAF" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1729166637250" checkinUser="plazi" docAuthor="Rosa, Cecilia Lozano De La, Cupello, Mario &amp; Vaz-De-Mello, Fernando Z." docDate="2024" docId="1D50766630005238ECF8F9C974520273" docLanguage="en" docName="EJT.2024.959.1-272.pdf" docOrigin="European Journal of Taxonomy 959 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2677/12401" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Dichotomius blancoi Nunes &amp; Vaz-de-Mello 2019" docType="treatment" docVersion="2" lastPageNumber="93" masterDocId="E1690E1E305A5264ED17FFA7770C013F" masterDocTitle="The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution" masterLastPageNumber="272" masterPageNumber="1" pageNumber="91" updateTime="1729169574746" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="8512996CB6234CF3BDB649056B6EB909">
<mods:title id="1DAE7A6019C5D24326965884B3E457FF">The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution</mods:title>
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<mods:name id="F21CB0C56BC8F2742337E531CA9FC765" type="personal">
<mods:role id="BB10D6BB04A78E9A497A508F6AF6BC57">
<mods:roleTerm id="BFB9F2AF4C0285405BE28E784E200876">Author</mods:roleTerm>
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<mods:namePart id="4F47F65983689733F985691BE4D7A47E">Rosa, Cecilia Lozano De La</mods:namePart>
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<mods:affiliation id="A4002BECDD13534393CAE1DF30054D04">Instituto Venezolano de Investigaciones Científicas, Centro de Estudios Botánicos y Agroforestales, Laboratorio de Ecología Espacial, Maracaibo, estado Zulia, Venezuela.</mods:affiliation>
<mods:nameIdentifier id="9662E9513A7BC9ABAB0803389F8E59BE" type="email">lozanoceci@gmail.com</mods:nameIdentifier>
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<mods:namePart id="F74C52FFE8143A9655689A0BE24AD163">Cupello, Mario</mods:namePart>
<mods:nameIdentifier id="764042090EEF49D5E6C27793E9761243" type="LSID">BDB03C18-7095-4EAA-8BDD-03CB4F79676D</mods:nameIdentifier>
<mods:affiliation id="BB5AB9B509EAD302548AB5B1FAE1FDB8">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Curitiba, Paraná, Brazil. Currently at Department of Entomology, Texas A &amp; M University, College Station, TX, 77843, USA.</mods:affiliation>
<mods:nameIdentifier id="6F6893CB0F5E009F3D3F619C683E4D79" type="email">mcupello@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="BD35974A2017F63AAED11B5AE2DAC424">Vaz-De-Mello, Fernando Z.</mods:namePart>
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<mods:affiliation id="FCB25DBB79B9E307946CA7EBAFA37566">Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, Cuiabá, Mato Grosso, Brazil.</mods:affiliation>
<mods:nameIdentifier id="D7064FDA0ACAAB1B70A6F75B24D15CC9" type="email">vazdemello@gmail.com</mods:nameIdentifier>
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<mods:date id="DB3585E601F22FF04D5C1D22D4B158E7">2024</mods:date>
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<subSubSection id="DDE394FB3000523EECF8F9C9734907B7" box="[495,1093,1646,1673]" pageId="90" pageNumber="91" type="nomenclature">
<paragraph id="9546C7703000523EECF8F9C9734907B7" blockId="90.[495,1093,1646,1705]" box="[495,1093,1646,1673]" pageId="90" pageNumber="91">
<heading id="CE0E701C3000523EECF8F9C9734907B7" box="[495,1093,1646,1673]" centered="true" fontSize="11" level="2" pageId="90" pageNumber="91" reason="2">
<taxonomicName id="52F9BCF33000523EECF8F9C9734907B7" ID-CoL="35N9P" authority="Nunes &amp; Vaz-de-Mello 2019" authorityName="Nunes &amp; Vaz-de-Mello" authorityYear="2019" box="[495,1093,1646,1673]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="blancoi">
<emphasis id="A78D1B623000523EECF8F9C975E707B6" bold="true" box="[495,747,1646,1673]" italics="true" pageId="90" pageNumber="91">Dichotomius blancoi</emphasis>
<bibRefCitation id="F168BA813000523EEFE5F9C8734907B7" author="Nunes R. V. &amp; Vaz-de-Mello F. Z." box="[754,1093,1646,1673]" pageId="90" pageNumber="91" pagination="2231 - 2351" refId="ref133607" refString="Nunes R. V. &amp; Vaz-de-Mello F. Z. 2019. Taxonomic revision of Dichotomius (Cephagonus) Luederwaldt 1929 and the taxonomic status of remaining Dichotomius Hope 1838 subgenera (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini). Journal of Natural History 53 (37 - 38): 2231 - 2351. https: // doi. org / 10.1080 / 00222933.2019.1692088" type="journal article" year="2019">Nunes &amp; Vaz-de-Mello 2019</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="DDE394FB3000523EEFF1F92874410796" box="[742,845,1679,1705]" pageId="90" pageNumber="91" type="description">
<paragraph id="9546C7703000523EEFF1F92874410796" blockId="90.[495,1093,1646,1705]" box="[742,845,1679,1705]" pageId="90" pageNumber="91">
<figureCitation id="0DC2DBF53000523EEFF1F92874410796" box="[742,845,1679,1705]" captionStart="Fig" captionStartId="91.[189,232,1963,1989]" captionTargetBox="[276,1304,278,1886]" captionTargetId="figure-12@91.[265,1322,265,1907]" captionTargetPageId="91" captionText="Fig. 36. Species distribution. A. Dichotomius agenor (Harold, 1869). B. Dichotomius blancoi Nunes &amp; Vaz-de-Mello, 2019. Green diamond = CEMT collection data; black circle = literature data." figureDoi="http://doi.org/10.5281/zenodo.13945565" httpUri="https://zenodo.org/record/13945565/files/figure.png" pageId="90" pageNumber="91">Fig. 36B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB3000523EEDAAF96876AF0614" pageId="90" pageNumber="91" type="reference_group">
<paragraph id="9546C7703000523EEDAAF96876AF0614" blockId="90.[189,1396,1742,1835]" pageId="90" pageNumber="91">
<treatmentCitationGroup id="B5E9E05E3000523EEDAAF96876AF0614" pageId="90" pageNumber="91">
<treatmentCitation id="1458E1613000523EEDAAF968744207D6" ID-DOI="http://doi.org/10.5281/zenodo.3671922" ID-Zenodo-Dep="3671922" author="Nunes R. V. &amp; Vaz-de-Mello F. Z." box="[189,846,1742,1769]" httpUri="http://treatment.plazi.org/id/C740D6092A37BB13FEFAE199E60828DE" page="2315" pageId="90" pageNumber="91" year="2019">
<taxonomicName id="52F9BCF33000523EEDAAF968744207D6" ID-CoL="35N9P" authority="Nunes &amp; Vaz-de-Mello, 2019: 2315" authorityName="Nunes" authorityYear="2019" box="[189,846,1742,1769]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="blancoi">
<emphasis id="A78D1B623000523EEDAAF96876A307D6" box="[189,431,1743,1769]" italics="true" pageId="90" pageNumber="91">Dichotomius blancoi</emphasis>
<bibRefCitation id="F168BA813000523EECA3F968744207D6" author="Nunes R. V. &amp; Vaz-de-Mello F. Z." box="[436,846,1742,1769]" pageId="90" pageNumber="91" pagination="2231 - 2351" refId="ref133607" refString="Nunes R. V. &amp; Vaz-de-Mello F. Z. 2019. Taxonomic revision of Dichotomius (Cephagonus) Luederwaldt 1929 and the taxonomic status of remaining Dichotomius Hope 1838 subgenera (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini). Journal of Natural History 53 (37 - 38): 2231 - 2351. https: // doi. org / 10.1080 / 00222933.2019.1692088" type="journal article" year="2019">Nunes &amp; Vaz-de-Mello, 2019: 2315</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description).Type locality:
<collectingCountry id="EDEE87E03000523EE9E4F968727C07D6" box="[1267,1392,1743,1769]" name="Venezuela" pageId="90" pageNumber="91">Venezuela</collectingCountry>
:
<collectingRegion id="573D09923000523EEDF0F95776490635" box="[231,325,1776,1802]" country="Venezuela" name="Tachira" pageId="90" pageNumber="91">Táchira</collectingRegion>
: Parque Nacional Natural Paramillo. Name-bearing type:
<typeStatus id="4A4279D23000523EEEE1F95773510635" box="[1014,1117,1776,1802]" pageId="90" pageNumber="91" type="holotype">holotype</typeStatus>
(CEMT), examined by CL and FZVM.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB30005238EDAAF8F774B300C9" lastPageId="92" lastPageNumber="93" pageId="90" pageNumber="91" type="materials_examined">
<paragraph id="9546C7703000523EEDAAF8F776A60655" blockId="90.[189,426,1872,1898]" box="[189,426,1872,1898]" pageId="90" pageNumber="91">
<heading id="CE0E701C3000523EEDAAF8F776A60655" bold="true" box="[189,426,1872,1898]" fontSize="11" level="3" pageId="90" pageNumber="91" reason="3">
<emphasis id="A78D1B623000523EEDAAF8F776A60655" bold="true" box="[189,426,1872,1898]" pageId="90" pageNumber="91">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="9546C7703000523EEDFDF8DA765006A8" blockId="90.[189,1399,1917,2009]" box="[234,348,1917,1943]" pageId="90" pageNumber="91">
<emphasis id="A78D1B623000523EEDFDF8DA765006A8" bold="true" box="[234,348,1917,1943]" pageId="90" pageNumber="91">
<typeStatus id="4A4279D23000523EEDFDF8DA765006A8" box="[234,348,1917,1943]" pageId="90" pageNumber="91" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="9546C7703000523EEDAAF839761806E6" blockId="90.[189,1399,1917,2009]" pageId="90" pageNumber="91">
<materialsCitation id="2591CD2D3000523EEDAAF839761806E6" collectingDate="1993-05" collectorName="Leg. J. Blanco" country="Venezuela" location="Tachira" pageId="90" pageNumber="91" specimenCount="1" specimenCount-male="1" stateProvince="Tachira" typeStatus="holotype">
<collectingCountry id="EDEE87E03000523EEDAAF83976670687" box="[189,363,1950,1976]" name="Venezuela" pageId="90" pageNumber="91">VENEZUELA</collectingCountry>
‒
<emphasis id="A78D1B623000523EEC9EF83A76E00687" bold="true" box="[393,492,1949,1976]" pageId="90" pageNumber="91">
<collectingRegion id="573D09923000523EEC9EF83A76E00687" box="[393,492,1949,1976]" country="Venezuela" name="Tachira" pageId="90" pageNumber="91">Táchira</collectingRegion>
</emphasis>
•
<specimenCount id="83FF0CF93000523EEF1FF83975340687" box="[520,568,1950,1976]" count="1" pageId="90" pageNumber="91" type="male">1 ♂</specimenCount>
; “Parque Paramillo/ edo. Táchira- Vzla/
<date id="E147E1B03000523EE90AF83973AF0687" box="[1053,1187,1950,1976]" pageId="90" pageNumber="91" value="1993-05">
<collectingDate id="F10318583000523EE90AF83973AF0687" box="[1053,1187,1950,1976]" pageId="90" pageNumber="91" value="1993-05">Mayo 1993</collectingDate>
</date>
/
<collectorName id="380CA2A63000523EE9A2F839726B0688" box="[1205,1383,1949,1976]" pageId="90" pageNumber="91">Leg. J. Blanco</collectorName>
”; CEMT.
</materialsCitation>
</paragraph>
<caption id="C18697F83001523FEDAAF80C73FC06D7" ID-DOI="http://doi.org/10.5281/zenodo.13945565" ID-Zenodo-Dep="13945565" httpUri="https://zenodo.org/record/13945565/files/figure.png" pageId="91" pageNumber="92" startId="91.[189,232,1963,1989]" targetBox="[276,1304,278,1886]" targetPageId="91">
<paragraph id="9546C7703001523FEDAAF80C73FC06D7" blockId="91.[189,1398,1963,2025]" pageId="91" pageNumber="92">
<emphasis id="A78D1B623001523FEDAAF80C761606FA" bold="true" box="[189,282,1963,1989]" pageId="91" pageNumber="92">Fig. 36.</emphasis>
Species distribution.
<emphasis id="A78D1B623001523FEF00F80C752006FA" bold="true" box="[535,556,1963,1989]" pageId="91" pageNumber="92">A</emphasis>
.
<taxonomicName id="52F9BCF33001523FEF2CF80C74E906FA" authority="(Harold, 1869)" baseAuthorityName="Harold" baseAuthorityYear="1869" box="[571,997,1963,1989]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="agenor">
<emphasis id="A78D1B623001523FEF2CF80C742706FA" box="[571,811,1963,1989]" italics="true" pageId="91" pageNumber="92">Dichotomius agenor</emphasis>
(Harold, 1869)
</taxonomicName>
.
<emphasis id="A78D1B623001523FEEF8F80C730F06FA" bold="true" box="[1007,1027,1963,1989]" pageId="91" pageNumber="92">B</emphasis>
.
<taxonomicName id="52F9BCF33001523FE906F80C76A706D7" authority="Nunes &amp; Vaz-de-Mello, 2019" authorityName="Nunes &amp; Vaz-de-Mello" authorityYear="2019" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="blancoi">
<emphasis id="A78D1B623001523FE906F80C720A06FA" box="[1041,1286,1963,1989]" italics="true" pageId="91" pageNumber="92">Dichotomius blancoi</emphasis>
<bibRefCitation id="F168BA813001523FE81AF80B76A706D7" author="Nunes R. V. &amp; Vaz-de-Mello F. Z." pageId="91" pageNumber="92" pagination="2231 - 2351" refId="ref133607" refString="Nunes R. V. &amp; Vaz-de-Mello F. Z. 2019. Taxonomic revision of Dichotomius (Cephagonus) Luederwaldt 1929 and the taxonomic status of remaining Dichotomius Hope 1838 subgenera (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini). Journal of Natural History 53 (37 - 38): 2231 - 2351. https: // doi. org / 10.1080 / 00222933.2019.1692088" type="journal article" year="2019">Nunes &amp; Vaz-de-Mello, 2019</bibRefCitation>
</taxonomicName>
. Green diamond = CEMT collection data; black circle = literature data.
</paragraph>
</caption>
<paragraph id="9546C77030065238EDFDFEAE766B001C" blockId="92.[189,1035,265,327]" box="[234,359,265,291]" pageId="92" pageNumber="93">
<emphasis id="A78D1B6230065238EDFDFEAE766B001C" bold="true" box="[234,359,265,291]" pageId="92" pageNumber="93">
<typeStatus id="4A4279D230065238EDFDFEAE766B001C" box="[234,359,265,291]" pageId="92" pageNumber="93" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="9546C77030065238EDAAFE8A73070078" blockId="92.[189,1035,265,327]" box="[189,1035,300,327]" pageId="92" pageNumber="93">
<materialsCitation id="2591CD2D30065238EDAAFE8A730B0078" box="[189,1031,300,327]" collectingDate="1993-05" collectorName="Leg. J. Blanco" country="Venezuela" location="Tachira" pageId="92" pageNumber="93" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Tachira" typeStatus="paratype">
<collectingCountry id="EDEE87E030065238EDAAFE8A76670078" box="[189,363,301,327]" name="Venezuela" pageId="92" pageNumber="93">VENEZUELA</collectingCountry>
‒
<emphasis id="A78D1B6230065238EC91FE8B76E50078" bold="true" box="[390,489,300,327]" pageId="92" pageNumber="93">
<collectingRegion id="573D099230065238EC91FE8B76E50078" box="[390,489,300,327]" country="Venezuela" name="Tachira" pageId="92" pageNumber="93">Táchira</collectingRegion>
</emphasis>
•
<specimenCount id="83FF0CF930065238EF15FE8A753C0078" box="[514,560,301,327]" count="1" pageId="92" pageNumber="93" type="male">1 ♂</specimenCount>
,
<specimenCount id="83FF0CF930065238EF2AFE8A75660078" box="[573,618,301,327]" count="1" pageId="92" pageNumber="93" type="female">1 ♀</specimenCount>
; same data as for holotype; CEMT
</materialsCitation>
.
</paragraph>
<paragraph id="9546C77030065238EDFDFEC876ED00B6" blockId="92.[189,1399,367,503]" box="[234,481,367,393]" pageId="92" pageNumber="93">
<emphasis id="A78D1B6230065238EDFDFEC876ED00B6" bold="true" box="[234,481,367,393]" pageId="92" pageNumber="93">Additional material</emphasis>
</paragraph>
<paragraph id="9546C77030065238EDAAFE3374B300C9" blockId="92.[189,1399,367,503]" pageId="92" pageNumber="93">
<collectingCountry id="EDEE87E030065238EDAAFE3376670091" box="[189,363,404,430]" name="Venezuela" pageId="92" pageNumber="93">VENEZUELA</collectingCountry>
‒
<emphasis id="A78D1B6230065238EC9AFE3476E00092" bold="true" box="[397,492,403,429]" pageId="92" pageNumber="93">
<collectingRegion id="573D099230065238EC9AFE3476E00092" box="[397,492,403,429]" country="Venezuela" name="Aragua" pageId="92" pageNumber="93">Aragua</collectingRegion>
</emphasis>
• 3 specs; Mario Briceño Iragorry, Parque Nacional Henri Pittier, Estación Biológica Rancho Grande, way to Pico Periquito;
<geoCoordinate id="F0CDA1B730065238EE30FE1174CD00ED" box="[807,961,438,466]" degrees="10" direction="north" minutes="20" orientation="latitude" pageId="92" pageNumber="93" precision="15" seconds="51" value="10.3475">10°20′51″ N</geoCoordinate>
,
<geoCoordinate id="F0CDA1B730065238EEC6FE11737C00ED" box="[977,1136,438,466]" degrees="67" direction="west" minutes="41" orientation="longitude" pageId="92" pageNumber="93" precision="15" seconds="17" value="-67.68806">67°41′17″ W</geoCoordinate>
;
<quantity id="52016A9530065238E997FE1F73DD00EC" box="[1152,1233,440,467]" metricMagnitude="3" metricUnit="m" metricValue="1.145" pageId="92" pageNumber="93" unit="m" value="1145.0">1145m</quantity>
a.s.l.; human faeces;
<date id="E147E1B030065238EC01FE7B76E800C9" box="[278,484,476,502]" pageId="92" pageNumber="93" value="2010-08-18" valueMax="2010-08-20" valueMin="2010-08-18">18‒20 Aug. 2010</date>
; R. Acconcia and J. Gámez leg.; CEMT.
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB30065238EDAAFDB974520273" pageId="92" pageNumber="93" type="distribution">
<paragraph id="9546C77030065238EDAAFDB9765A0307" blockId="92.[189,342,542,568]" box="[189,342,542,568]" pageId="92" pageNumber="93">
<heading id="CE0E701C30065238EDAAFDB9765A0307" bold="true" box="[189,342,542,568]" fontSize="11" level="3" pageId="92" pageNumber="93" reason="3">
<emphasis id="A78D1B6230065238EDAAFDB9765A0307" bold="true" box="[189,342,542,568]" pageId="92" pageNumber="93">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="9546C77030065238EDAAFDE976B20357" blockId="92.[189,446,590,616]" box="[189,446,590,616]" pageId="92" pageNumber="93">
<collectingCountry id="EDEE87E030065238EDAAFDE9763A0357" box="[189,310,590,616]" name="Venezuela" pageId="92" pageNumber="93">Venezuela</collectingCountry>
and
<collectingCountry id="EDEE87E030065238EC78FDE976B60357" box="[367,442,590,616]" name="Brazil" pageId="92" pageNumber="93">Brazil</collectingCountry>
.
</paragraph>
<paragraph id="9546C77030065238EDAAFD3776E10394" blockId="92.[189,493,656,683]" box="[189,493,656,683]" pageId="92" pageNumber="93">
<heading id="CE0E701C30065238EDAAFD3776E10394" bold="true" box="[189,493,656,683]" fontSize="11" level="3" pageId="92" pageNumber="93" reason="3">
<emphasis id="A78D1B6230065238EDAAFD3776E10394" bold="true" box="[189,493,656,683]" pageId="92" pageNumber="93">
Subregions of
<collectingCountry id="EDEE87E030065238EC78FD3676E10394" box="[367,493,657,683]" name="Venezuela" pageId="92" pageNumber="93">Venezuela</collectingCountry>
</emphasis>
</heading>
</paragraph>
<paragraph id="9546C77030065238EDAAFD67743203E4" blockId="92.[189,830,704,731]" box="[189,830,704,731]" pageId="92" pageNumber="93">
Andes mountains, and Central
<collectingRegion id="573D099230065238EF3FFD67756703E5" box="[552,619,704,730]" country="New Zealand" name="West Coast" pageId="92" pageNumber="93">Coast</collectingRegion>
Mountain Range.
</paragraph>
<paragraph id="9546C77030065238EDAAFCA576A80222" blockId="92.[189,420,770,797]" box="[189,420,770,797]" pageId="92" pageNumber="93">
<heading id="CE0E701C30065238EDAAFCA576A80222" bold="true" box="[189,420,770,797]" fontSize="11" level="3" pageId="92" pageNumber="93" reason="3">
<emphasis id="A78D1B6230065238EDAAFCA576A80222" bold="true" box="[189,420,770,797]" pageId="92" pageNumber="93">Literature records</emphasis>
</heading>
</paragraph>
<paragraph id="9546C77030065238EDAAFC9474520273" blockId="92.[189,862,818,845]" box="[189,862,818,845]" pageId="92" pageNumber="93">
<bibRefCitation id="F168BA8130065238EDAAFC9475570272" author="Nunes R. V. &amp; Vaz-de-Mello F. Z." box="[189,603,818,845]" pageId="92" pageNumber="93" pagination="2231 - 2351" refId="ref133607" refString="Nunes R. V. &amp; Vaz-de-Mello F. Z. 2019. Taxonomic revision of Dichotomius (Cephagonus) Luederwaldt 1929 and the taxonomic status of remaining Dichotomius Hope 1838 subgenera (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini). Journal of Natural History 53 (37 - 38): 2231 - 2351. https: // doi. org / 10.1080 / 00222933.2019.1692088" type="journal article" year="2019">Nunes &amp; Vaz-de-Mello 2019: 2315</bibRefCitation>
(
<collectingCountry id="EDEE87E030065238EF7DFC9575E50273" box="[618,745,818,844]" name="Venezuela" pageId="92" pageNumber="93">Venezuela</collectingCountry>
:
<collectingRegion id="573D099230065238EFE4FC95745E0273" box="[755,850,818,844]" country="Venezuela" name="Tachira" pageId="92" pageNumber="93">Táchira</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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