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<document id="D2F8031936D4CD7E0F713F3C71C3FB1B" ID-DOI="10.5281/zenodo.170590" ID-GBIF-Dataset="22b7f18d-d80f-41c9-a21b-7d4114bd504a" ID-ISSN="1175­5326" ID-Zenodo-Dep="170590" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459713953354" checkinUser="plazi" docAuthor="Hamilton, Steven W. &amp; Holzenthal, Ralph W." docDate="2005" docId="1D3D879BFFBBA14DFEEDFCE2B3C9124E" docLanguage="en" docName="zt00810.pdf" docOrigin="Zootaxa 810" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Polycentropus quadricuspidis Hamilton and Holzenthal, new species" docType="treatment" docVersion="9" lastPageNumber="9" masterDocId="E104FFE3FFBDA145FFE5FF96B1311170" masterDocTitle="Five new species of Polycentropodidae (Trichoptera) from Ecuador and Venezuela" masterLastPageNumber="14" masterPageNumber="1" pageNumber="7" updateTime="1698207473436" updateUser="plazi">
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<mods:title id="9EE9659FCF544ECAEE8E0AD8BF1DE088">Five new species of Polycentropodidae (Trichoptera) from Ecuador and Venezuela</mods:title>
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<mods:namePart id="869087F02022C41CDB3B8C9AE0BF8C48">Hamilton, Steven W.</mods:namePart>
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<treatment id="1D3D879BFFBBA14DFEEDFCE2B3C9124E" ID-DOI="http://doi.org/10.5281/zenodo.6266230" ID-GBIF-Taxon="119334120" ID-Zenodo-Dep="6266230" LSID="urn:lsid:plazi:treatment:1D3D879BFFBBA14DFEEDFCE2B3C9124E" httpUri="http://treatment.plazi.org/id/1D3D879BFFBBA14DFEEDFCE2B3C9124E" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7">
<subSubSection id="DD8E6506FFBBA143FEEDFCE2B07A12C1" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="952B368DFFBBA143FEEDFCE2B56012FE" blockId="6.[264,1105,884,945]" box="[264,1105,884,910]" pageId="6" pageNumber="7">
<heading id="CE6381E1FFBBA143FEEDFCE2B56012FE" bold="true" box="[264,1105,884,910]" fontSize="11" level="1" pageId="6" pageNumber="7" reason="1">
<emphasis id="A7E0EA9FFFBBA143FEEDFCE2B56012FE" bold="true" box="[264,1105,884,910]" pageId="6" pageNumber="7">
<taxonomicName id="52944D0EFFBBA143FEEDFCE2B28412FE" authority="Hamilton and Holzenthal" authorityName="Hamilton and Holzenthal" box="[264,949,884,910]" class="Insecta" family="Polycentropodidae" genus="Polycentropus" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="quadricuspidis" status="sp. nov.">
<emphasis id="A7E0EA9FFFBBA143FEEDFCE2B35F12FE" bold="true" box="[264,622,884,910]" italics="true" pageId="6" pageNumber="7">Polycentropus quadricuspidis</emphasis>
Hamilton and Holzenthal
</taxonomicName>
,
<taxonomicNameLabel id="BCD357E4FFBBA143FC27FCE2B56012FE" box="[962,1105,884,910]" pageId="6" pageNumber="7" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="952B368DFFBBA143FEEDFC01B07A12C1" blockId="6.[264,1105,884,945]" box="[264,331,919,945]" pageId="6" pageNumber="7">
<figureCitation id="0DAF2A08FFBBA143FEEDFC01B07A12C1" box="[264,331,919,945]" captionStart="FIGURE 3" captionStartId="7.[264,368,1789,1813]" captionTargetBox="[296,1291,291,1774]" captionTargetId="figure@7.[296,1292,291,1774]" captionTargetPageId="7" captionText="FIGURE 3. Polycentropus quadricuspidis, new species. Male genitalia: A—lateral; B—dorsal; C—ventral; D—inferior appendage, caudal; E—phallus, lateral; F—phallic sclerite, dorsal; Gphallobase, caudal; H—subphallic sclerite, caudal." httpUri="https://zenodo.org/record/170593/files/figure.png" pageId="6" pageNumber="7">Fig. 3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="DD8E6506FFBBA14DFEEDFC71B27B1306" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7" type="description">
<paragraph id="952B368DFFBBA143FEEDFC71B30F1779" blockId="6.[264,1323,999,1905]" pageId="6" pageNumber="7">
From
<collectingCountry id="ED83761DFFBBA143FEB4FC71B0851571" box="[337,436,999,1025]" name="Ecuador" pageId="6" pageNumber="7">Ecuador</collectingCountry>
,
<taxonomicName id="52944D0EFFBBA143FE27FC71B3BE1570" box="[450,655,999,1024]" class="Insecta" family="Polycentropodidae" genus="Polycentropus" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="quadricuspidis" status="sp. nov.">
<emphasis id="A7E0EA9FFFBBA143FE27FC71B3BE1570" box="[450,655,999,1024]" italics="true" pageId="6" pageNumber="7">P. quadricuspidis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BCD357E4FFBBA143FD7DFC71B3E81571" box="[664,729,999,1025]" pageId="6" pageNumber="7" rank="species">n. sp.</taxonomicNameLabel>
is placed in the
<taxonomicName id="52944D0EFFBBA143FC47FC71B51B1570" box="[930,1066,999,1024]" class="Insecta" family="Polycentropodidae" genus="Polycentropus" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="joergenseni">
<emphasis id="A7E0EA9FFFBBA143FC47FC71B51B1570" box="[930,1066,999,1024]" italics="true" pageId="6" pageNumber="7">joergenseni</emphasis>
</taxonomicName>
complex and is most closely related to
<taxonomicName id="52944D0EFFBBA143FE32FB99B34A1558" box="[471,635,1039,1064]" class="Insecta" family="Polycentropodidae" genus="Polycentropus" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="joergenseni">
<emphasis id="A7E0EA9FFFBBA143FE32FB99B34A1558" box="[471,635,1039,1064]" italics="true" pageId="6" pageNumber="7">P. joergenseni</emphasis>
</taxonomicName>
and
<taxonomicName id="52944D0EFFBBA143FD56FB99B2411558" box="[691,880,1039,1064]" class="Insecta" family="Polycentropodidae" genus="Polycentropus" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="longispinosus">
<emphasis id="A7E0EA9FFFBBA143FD56FB99B2411558" box="[691,880,1039,1064]" italics="true" pageId="6" pageNumber="7">P. longispinosus</emphasis>
</taxonomicName>
. These 3 species share 4 synapomorphies not found in other species of the
<taxonomicName id="52944D0EFFBBA143FD29FBA1B2651520" box="[716,852,1079,1104]" class="Insecta" family="Polycentropodidae" genus="Polycentropus" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="joergenseni">
<emphasis id="A7E0EA9FFFBBA143FD29FBA1B2651520" box="[716,852,1079,1104]" italics="true" pageId="6" pageNumber="7">joergenseni</emphasis>
</taxonomicName>
complex: (1) a ventral sclerotized band in the phallic membranes (
<figureCitation id="0DAF2A08FFBBA143FDA2FBC9B3BC1509" box="[583,653,1119,1145]" captionStart="FIGURE 3" captionStartId="7.[264,368,1789,1813]" captionTargetBox="[296,1291,291,1774]" captionTargetId="figure@7.[296,1292,291,1774]" captionTargetPageId="7" captionText="FIGURE 3. Polycentropus quadricuspidis, new species. Male genitalia: A—lateral; B—dorsal; C—ventral; D—inferior appendage, caudal; E—phallus, lateral; F—phallic sclerite, dorsal; Gphallobase, caudal; H—subphallic sclerite, caudal." httpUri="https://zenodo.org/record/170593/files/figure.png" pageId="6" pageNumber="7">Fig. 3</figureCitation>
E), (2) a pair of ventral sclerotized bumps in the phallic membrane apical to the ventral band (
<figureCitation id="0DAF2A08FFBBA143FD2FFB11B22115D1" box="[714,784,1159,1185]" captionStart="FIGURE 3" captionStartId="7.[264,368,1789,1813]" captionTargetBox="[296,1291,291,1774]" captionTargetId="figure@7.[296,1292,291,1774]" captionTargetPageId="7" captionText="FIGURE 3. Polycentropus quadricuspidis, new species. Male genitalia: A—lateral; B—dorsal; C—ventral; D—inferior appendage, caudal; E—phallus, lateral; F—phallic sclerite, dorsal; Gphallobase, caudal; H—subphallic sclerite, caudal." httpUri="https://zenodo.org/record/170593/files/figure.png" pageId="6" pageNumber="7">Fig. 3</figureCitation>
E), (3) an apicoventral process on the phallobase that is reduced to a pair of ventral points (
<figureCitation id="0DAF2A08FFBBA143FCC8FB39B24015B9" box="[813,881,1199,1225]" captionStart="FIGURE 3" captionStartId="7.[264,368,1789,1813]" captionTargetBox="[296,1291,291,1774]" captionTargetId="figure@7.[296,1292,291,1774]" captionTargetPageId="7" captionText="FIGURE 3. Polycentropus quadricuspidis, new species. Male genitalia: A—lateral; B—dorsal; C—ventral; D—inferior appendage, caudal; E—phallus, lateral; F—phallic sclerite, dorsal; Gphallobase, caudal; H—subphallic sclerite, caudal." httpUri="https://zenodo.org/record/170593/files/figure.png" pageId="6" pageNumber="7">Fig. 3</figureCitation>
E &amp; G), and (4) somewhat broadened and flattened inferior appendages (
<figureCitation id="0DAF2A08FFBBA143FD41FB41B3DB1581" box="[676,746,1239,1265]" captionStart="FIGURE 3" captionStartId="7.[264,368,1789,1813]" captionTargetBox="[296,1291,291,1774]" captionTargetId="figure@7.[296,1292,291,1774]" captionTargetPageId="7" captionText="FIGURE 3. Polycentropus quadricuspidis, new species. Male genitalia: A—lateral; B—dorsal; C—ventral; D—inferior appendage, caudal; E—phallus, lateral; F—phallic sclerite, dorsal; Gphallobase, caudal; H—subphallic sclerite, caudal." httpUri="https://zenodo.org/record/170593/files/figure.png" pageId="6" pageNumber="7">Fig. 3</figureCitation>
A). The inferior appendage of
<taxonomicName id="52944D0EFFBBA143FBBBFB41B41B1580" box="[1118,1322,1239,1264]" class="Insecta" family="Polycentropodidae" genus="Polycentropus" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="quadricuspidis">
<emphasis id="A7E0EA9FFFBBA143FBBBFB41B41B1580" box="[1118,1322,1239,1264]" italics="true" pageId="6" pageNumber="7">P. quadricuspidis</emphasis>
</taxonomicName>
(
<figureCitation id="0DAF2A08FFBBA143FEF4FB69B0651469" box="[273,340,1279,1305]" captionStart="FIGURE 3" captionStartId="7.[264,368,1789,1813]" captionTargetBox="[296,1291,291,1774]" captionTargetId="figure@7.[296,1292,291,1774]" captionTargetPageId="7" captionText="FIGURE 3. Polycentropus quadricuspidis, new species. Male genitalia: A—lateral; B—dorsal; C—ventral; D—inferior appendage, caudal; E—phallus, lateral; F—phallic sclerite, dorsal; Gphallobase, caudal; H—subphallic sclerite, caudal." httpUri="https://zenodo.org/record/170593/files/figure.png" pageId="6" pageNumber="7">Fig. 3</figureCitation>
A &amp; C) is shorter than that of
<taxonomicName id="52944D0EFFBBA143FD52FB69B2441468" box="[695,885,1279,1304]" class="Insecta" family="Polycentropodidae" genus="Polycentropus" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="longispinosus">
<emphasis id="A7E0EA9FFFBBA143FD52FB69B2441468" box="[695,885,1279,1304]" italics="true" pageId="6" pageNumber="7">P. longispinosus</emphasis>
</taxonomicName>
and resembles that of
<taxonomicName id="52944D0EFFBBA143FB9BFB69B4121468" box="[1150,1315,1279,1304]" class="Insecta" family="Polycentropodidae" genus="Polycentropus" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="joergenseni">
<emphasis id="A7E0EA9FFFBBA143FB9BFB69B4121468" box="[1150,1315,1279,1304]" italics="true" pageId="6" pageNumber="7">P. joergenseni</emphasis>
</taxonomicName>
. The intermediate appendage (
<figureCitation id="0DAF2A08FFBBA143FD82FAB1B39D1431" box="[615,684,1319,1345]" captionStart="FIGURE 3" captionStartId="7.[264,368,1789,1813]" captionTargetBox="[296,1291,291,1774]" captionTargetId="figure@7.[296,1292,291,1774]" captionTargetPageId="7" captionText="FIGURE 3. Polycentropus quadricuspidis, new species. Male genitalia: A—lateral; B—dorsal; C—ventral; D—inferior appendage, caudal; E—phallus, lateral; F—phallic sclerite, dorsal; Gphallobase, caudal; H—subphallic sclerite, caudal." httpUri="https://zenodo.org/record/170593/files/figure.png" pageId="6" pageNumber="7">Fig. 3</figureCitation>
A &amp; B) is intermediate between that of
<taxonomicName id="52944D0EFFBBA143FB62FAB1B01C1418" class="Insecta" family="Polycentropodidae" genus="Polycentropus" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="longispinosus">
<emphasis id="A7E0EA9FFFBBA143FB62FAB1B01C1418" italics="true" pageId="6" pageNumber="7">P. longispinosus</emphasis>
</taxonomicName>
and
<taxonomicName id="52944D0EFFBBA143FE82FAD9B33B1418" box="[359,522,1359,1384]" class="Insecta" family="Polycentropodidae" genus="Polycentropus" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="joergenseni">
<emphasis id="A7E0EA9FFFBBA143FE82FAD9B33B1418" box="[359,522,1359,1384]" italics="true" pageId="6" pageNumber="7">P. joergenseni</emphasis>
</taxonomicName>
in both length and thickness.
<taxonomicName id="52944D0EFFBBA143FC97FAD9B5FE1418" box="[882,1231,1359,1384]" class="Insecta" family="Polycentropodidae" genus="Polycentropus" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="quadricuspidis">
<emphasis id="A7E0EA9FFFBBA143FC97FAD9B5FE1418" box="[882,1231,1359,1384]" italics="true" pageId="6" pageNumber="7">Polycentropus quadricuspidis</emphasis>
</taxonomicName>
is most easily distinguished from these sister species by the 2 pairs of apicoventral points on the phallobase (
<figureCitation id="0DAF2A08FFBBA143FE7CFA09B0D014C9" box="[409,481,1439,1465]" captionStart="FIGURE 3" captionStartId="7.[264,368,1789,1813]" captionTargetBox="[296,1291,291,1774]" captionTargetId="figure@7.[296,1292,291,1774]" captionTargetPageId="7" captionText="FIGURE 3. Polycentropus quadricuspidis, new species. Male genitalia: A—lateral; B—dorsal; C—ventral; D—inferior appendage, caudal; E—phallus, lateral; F—phallic sclerite, dorsal; Gphallobase, caudal; H—subphallic sclerite, caudal." httpUri="https://zenodo.org/record/170593/files/figure.png" pageId="6" pageNumber="7">Fig. 3</figureCitation>
G), the more prominent, spicule­covered pair of ventral bumps in the phallic membranes (
<figureCitation id="0DAF2A08FFBBA143FE13FA51B30A1491" box="[502,571,1479,1505]" captionStart="FIGURE 3" captionStartId="7.[264,368,1789,1813]" captionTargetBox="[296,1291,291,1774]" captionTargetId="figure@7.[296,1292,291,1774]" captionTargetPageId="7" captionText="FIGURE 3. Polycentropus quadricuspidis, new species. Male genitalia: A—lateral; B—dorsal; C—ventral; D—inferior appendage, caudal; E—phallus, lateral; F—phallic sclerite, dorsal; Gphallobase, caudal; H—subphallic sclerite, caudal." httpUri="https://zenodo.org/record/170593/files/figure.png" pageId="6" pageNumber="7">Fig. 3</figureCitation>
E), and the slight dorsal concavity of the lateral face of the inferior appendages (
<figureCitation id="0DAF2A08FFBBA143FE37FA79B3261779" box="[466,535,1519,1545]" captionStart="FIGURE 3" captionStartId="7.[264,368,1789,1813]" captionTargetBox="[296,1291,291,1774]" captionTargetId="figure@7.[296,1292,291,1774]" captionTargetPageId="7" captionText="FIGURE 3. Polycentropus quadricuspidis, new species. Male genitalia: A—lateral; B—dorsal; C—ventral; D—inferior appendage, caudal; E—phallus, lateral; F—phallic sclerite, dorsal; Gphallobase, caudal; H—subphallic sclerite, caudal." httpUri="https://zenodo.org/record/170593/files/figure.png" pageId="6" pageNumber="7">Fig. 3</figureCitation>
A).
</paragraph>
<paragraph id="952B368DFFBBA143FEDDF981B52817D9" blockId="6.[264,1323,999,1905]" pageId="6" pageNumber="7">Adult. Length of forewing: male 9.0 mm (n=2), female 8.0 mm (n=1). Body stramineous, legs stramineous, dorsum of head and thorax brown, clothed with long, dark setae; general vestiture of forewing with fine, brown to dark brown setae, with many small, scattered areas of golden setae, base of forewing with long, erect setae.</paragraph>
<paragraph id="952B368DFFBBA14DFEDDF921B27B1306" blockId="6.[264,1323,999,1905]" lastBlockId="8.[264,1323,284,830]" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7">
Male. Genitalia as in
<figureCitation id="0DAF2A08FFBBA143FDA4F921B3BA17A1" box="[577,651,1719,1745]" captionStart="FIGURE 3" captionStartId="7.[264,368,1789,1813]" captionTargetBox="[296,1291,291,1774]" captionTargetId="figure@7.[296,1292,291,1774]" captionTargetPageId="7" captionText="FIGURE 3. Polycentropus quadricuspidis, new species. Male genitalia: A—lateral; B—dorsal; C—ventral; D—inferior appendage, caudal; E—phallus, lateral; F—phallic sclerite, dorsal; Gphallobase, caudal; H—subphallic sclerite, caudal." httpUri="https://zenodo.org/record/170593/files/figure.png" pageId="6" pageNumber="7">Fig. 3</figureCitation>
. Sternum IX in lateral view subquadrate, posterolateral corner slightly produced, in ventral view slightly trapezoidal; anterior margin concave, posterior margin slightly concave. Tergum IX and X membranous. Intermediate appendage sinuate, apex curved medially, long, exceeding preanal appendage by more than 1/2 its length; basal region simple, not expanded; apex acuminate, without apical setae. Preanal appendage short, densely setose, rounded, broad basally; mesoventral process absent. Inferior appendage densely setose, short, subtriangular, wide dorsoventrally, subequal to length; dorsolateral flange broadly rounded; ventrally with dorsally­directed medial point, hidden in lateral view; in ventral view, inferior appendage quadrate; medial point prominent, acute; in caudal view, inferior appendage broadly oval, medial point rounded. Phallobase moderately elongate; apicoventral projection short, broad, with two pairs of points; phallic membranes without spines, but with ventromesal pair of conical, spiculate lobes and dorsal and ventral sclerotized bands; phallic sclerite elongate, apex upturned, spatulate. Subphallic sclerite U­shaped in caudal view.
</paragraph>
</subSubSection>
<caption id="C1EB6605FFBAA142FEEDF96BB2151612" httpUri="https://zenodo.org/record/170593/files/figure.png" pageId="7" pageNumber="8" targetBox="[296,1291,291,1774]" targetPageId="7">
<paragraph id="952B368DFFBAA142FEEDF96BB2151612" blockId="7.[264,1323,1789,1890]" pageId="7" pageNumber="8">
<emphasis id="A7E0EA9FFFBAA142FEEDF96BB0BD1665" bold="true" box="[264,396,1789,1813]" pageId="7" pageNumber="8">FIGURE 3.</emphasis>
<taxonomicName id="52944D0EFFBAA142FE79F968B3ED1665" box="[412,732,1790,1813]" class="Insecta" family="Polycentropodidae" genus="Polycentropus" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="quadricuspidis" status="sp. nov.">
<emphasis id="A7E0EA9FFFBAA142FE79F968B3ED1665" box="[412,732,1790,1813]" italics="true" pageId="7" pageNumber="8">Polycentropus quadricuspidis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="BCD357E4FFBAA142FD0EF96BB2411664" box="[747,880,1789,1812]" pageId="7" pageNumber="8" rank="species">new species</taxonomicNameLabel>
. Male genitalia: A—lateral; B—dorsal; C—ventral; D—inferior appendage, caudal; E—phallus, lateral; F—phallic sclerite, dorsal; Gphallobase, caudal; H—subphallic sclerite, caudal.
</paragraph>
</caption>
<subSubSection id="DD8E6506FFB5A14DFEDDFD12B3C9124E" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="952B368DFFB5A14DFEDDFD12B39D13B6" blockId="8.[264,1323,284,830]" pageId="8" pageNumber="9">
<emphasis id="A7E0EA9FFFB5A14DFEDDFD12B28413EE" bold="true" box="[312,949,644,670]" pageId="8" pageNumber="9">
<typeStatus id="4A2F882FFFB5A14DFEDDFD12B09B13EE" box="[312,426,644,670]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
male:
<collectingCountry id="ED83761DFFB5A14DFDEEFD12B39513EE" box="[523,676,644,670]" name="Ecuador" pageId="8" pageNumber="9">ECUADOR</collectingCountry>
: Zamora­Chinchipe:
</emphasis>
<quantity id="526C9B68FFB5A14DFC23FD12B52313EE" box="[966,1042,644,670]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="9" unit="km" value="30.0">30 km</quantity>
E Loja, el.
<quantity id="526C9B68FFB5A14DFB47FD12B5CE13EE" box="[1186,1279,644,670]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="9" unit="m" value="2000.0">2000 m</quantity>
,
<date id="E12A104DFFB5A14DFAE8FD12B0B913B6" pageId="8" pageNumber="9" value="1990-09-23">23 Sept. 1990</date>
, O.S. Flint, Jr. (
<collectionCode id="F385AE48FFB5A14DFDA5FD3AB39013B6" box="[576,673,684,710]" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="9">NMNH</collectionCode>
).
</paragraph>
<paragraph id="952B368DFFB5A14DFEDDFD42B51E139E" blockId="8.[264,1323,284,830]" box="[312,1071,724,750]" pageId="8" pageNumber="9">
<emphasis id="A7E0EA9FFFB5A14DFEDDFD42B08E139E" bold="true" box="[312,447,724,750]" pageId="8" pageNumber="9">
<typeStatus id="4A2F882FFFB5A14DFEDDFD42B08B139E" box="[312,442,724,750]" pageId="8" pageNumber="9" type="paratype">Paratypes</typeStatus>
:
</emphasis>
Same data as holotype—
<specimenCount id="8392FD04FFB5A14DFD13FD42B273139E" box="[758,834,724,750]" pageId="8" pageNumber="9" type="male">1 male</specimenCount>
,
<specimenCount id="8392FD04FFB5A14DFCABFD42B285139E" box="[846,948,724,750]" pageId="8" pageNumber="9" type="female">1 female</specimenCount>
(
<collectionCode id="F385AE48FFB5A14DFC26FD42B512139E" box="[963,1059,724,750]" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="9">NMNH</collectionCode>
).
</paragraph>
<paragraph id="952B368DFFB5A14DFEDDFD6AB3C9124E" blockId="8.[264,1323,284,830]" pageId="8" pageNumber="9">
Etymology. From the Latin words
<emphasis id="A7E0EA9FFFB5A14DFD36FD6AB2061265" box="[723,823,764,789]" italics="true" pageId="8" pageNumber="9">quattuor</emphasis>
, four, and
<emphasis id="A7E0EA9FFFB5A14DFC51FD6AB2CC1265" box="[948,1021,764,789]" italics="true" pageId="8" pageNumber="9">cuspis</emphasis>
, point, for the four points on the apicoventral area of the phallobase.
</paragraph>
</subSubSection>
</treatment>
</document>