treatments-xml/data/1D/28/6D/1D286D726C2A9F54FEB21C50F32CFD80.xml

<document id="DAE65B85D8FB12CA656F0638628AE4E1" ID-DOI="10.5281/zenodo.273585" ID-GBIF-Dataset="f2b78bf7-4c7a-4c18-9138-3961afe6a19c" ID-ISSN="1175­5326" ID-Zenodo-Dep="273585" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459790679642" checkinUser="plazi" docAuthor="Bray, Rodney A. &amp; Justine, Jean-Lou" docDate="2006" docId="1D286D726C2A9F54FEB21C50F32CFD80" docLanguage="en" docName="zt01326p044.pdf" docOrigin="Zootaxa 1326" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Hypocreadium toombo Bray &amp; Justine, 2006, n. sp." docType="treatment" docVersion="8" lastPageNumber="41" masterDocId="E111150A6C2B9F50FFBA1A5EF10CFFA6" masterDocTitle="Hypocreadium toombo n. sp. (Digenea: Lepocreadiidae) in the yellow­spotted triggerfish Pseudobalistes fuscus (Perciformes: Balistidae) and additional lepocreadiids parasitizing fishes from the waters off New Caledonia" masterLastPageNumber="44" masterPageNumber="37" pageNumber="38" updateTime="1698219176166" updateUser="plazi">
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<mods:title id="91FF21066D71D19F341EDB41C7E12317">Hypocreadium toombo n. sp. (Digenea: Lepocreadiidae) in the yellow­spotted triggerfish Pseudobalistes fuscus (Perciformes: Balistidae) and additional lepocreadiids parasitizing fishes from the waters off New Caledonia</mods:title>
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<mods:namePart id="AE0D44F04688B52AFD94F0A8D400DB50">Bray, Rodney A.</mods:namePart>
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<mods:namePart id="E07FF405D261D80CA511CA2A26E79E05">Justine, Jean-Lou</mods:namePart>
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<mods:relatedItem id="64B7CEA663765A8BC3C768AB762E98F5" type="host">
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<mods:date id="623F8714AB90246EC64A0241ECE73175">2006</mods:date>
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<treatment id="1D286D726C2A9F54FEB21C50F32CFD80" ID-DOI="http://doi.org/10.5281/zenodo.6255208" ID-GBIF-Taxon="119350331" ID-Zenodo-Dep="6255208" LSID="urn:lsid:zoobank.org:act:29022EC5-45D9-42B8-AAD9-57C944940841" httpUri="http://treatment.plazi.org/id/1D286D726C2A9F54FEB21C50F32CFD80" lastPageId="4" lastPageNumber="41" pageId="1" pageNumber="38">
<subSubSection id="DD9B8FEF6C2A9F51FEB21C50F086F907" pageId="1" pageNumber="38" type="nomenclature">
<paragraph id="953EDC646C2A9F51FEB21C50F3EBF98E" blockId="1.[264,743,1550,1576]" box="[264,743,1550,1576]" pageId="1" pageNumber="38">
<heading id="CE766B086C2A9F51FEB21C50F3EBF98E" bold="true" box="[264,743,1550,1576]" fontSize="11" level="1" pageId="1" pageNumber="38" reason="1">
<emphasis id="A7F500766C2A9F51FEB21C50F3EBF98E" bold="true" box="[264,743,1550,1576]" pageId="1" pageNumber="38">
<taxonomicName id="5281A7E76C2A9F51FEB21C50F311F98E" ID-CoL="6N3D7" box="[264,541,1550,1576]" class="Trematoda" family="Lepocreadiidae" genus="Hypocreadium" kingdom="Animalia" order="Plagiorchiida" pageId="1" pageNumber="38" phylum="Platyhelminthes" rank="species" species="toombo" status="sp. nov.">
<emphasis id="A7F500766C2A9F51FEB21C50F311F98E" bold="true" box="[264,541,1550,1576]" italics="true" pageId="1" pageNumber="38">Hypocreadium toombo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BCC6BD0D6C2A9F51FD991C50F36AF98E" box="[547,614,1550,1576]" pageId="1" pageNumber="38" rank="species">n. sp.</taxonomicNameLabel>
(
<figureCitation id="0DBAC0E16C2A9F51FDCE1C50F3D2F98E" box="[628,734,1550,1576]" captionStart="FIGURES 1 – 2" captionStartId="2.[266,385,1727,1751]" captionTargetBox="[444,1162,344,1675]" captionTargetId="figure@2.[368,1226,320,1688]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Hypocreadium toombo n. sp. 1. Ventral view of holotype, uterus in bold outline. 2. Terminal genitalia. Abbreviations: Ej, Ejaculatory duct; Ep, Excretory pore; Esv, External seminal vesicle; Isv, Internal seminal vesicle; Mt, Metraterm; Ov, Ovary; Pp, Pars prostatica; T, testis; Ut, Uterus. Scale bars: 1, 1 mm; 2, 200 μm." httpUri="https://zenodo.org/record/273586/files/figure.png" pageId="1" pageNumber="38">Figs 1–2</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="953EDC646C2A9F51FEB21C00F086F907" blockId="1.[264,1323,1630,1857]" pageId="1" pageNumber="38">
<emphasis id="A7F500766C2A9F51FEB21C00F087F9DE" bold="true" box="[264,395,1630,1656]" pageId="1" pageNumber="38">Type­host:</emphasis>
<taxonomicName id="5281A7E76C2A9F51FE1B1C01F2D8F9DF" authority="Bloch &amp; Schneider" authorityName="Bloch &amp; Schneider" box="[417,980,1631,1657]" class="Actinopterygii" family="Balistidae" genus="Pseudobalistes" kingdom="Animalia" order="Tetraodontiformes" pageId="1" pageNumber="38" phylum="Chordata" rank="species" species="fuscus">
<emphasis id="A7F500766C2A9F51FE1B1C01F3A2F9DE" box="[417,686,1631,1656]" italics="true" pageId="1" pageNumber="38">Pseudobalistes fuscus</emphasis>
(Bloch &amp; Schneider)
</taxonomicName>
, 
<taxonomicName id="5281A7E76C2A9F51FC561C01F568F9DF" box="[1004,1124,1631,1657]" class="Actinopterygii" family="Balistidae" kingdom="Animalia" order="Tetraodontiformes" pageId="1" pageNumber="38" phylum="Chordata" rank="family">Balistidae</taxonomicName>
, yellow­spotted triggerfish.
</paragraph>
</subSubSection>
<subSubSection id="DD9B8FEF6C2A9F53FE821CF0F52DFE08" lastPageId="3" lastPageNumber="40" pageId="1" pageNumber="38" type="description">
<paragraph id="953EDC646C2A9F51FE821CF0F0E9F96F" blockId="1.[264,1323,1630,1857]" box="[312,485,1710,1737]" pageId="1" pageNumber="38">
<emphasis id="A7F500766C2A9F51FE821CF0F07DF96E" bold="true" box="[312,369,1710,1736]" pageId="1" pageNumber="38">Site:</emphasis>
Intestine.
</paragraph>
<paragraph id="953EDC646C2A9F51FE821C88F5E3F957" blockId="1.[264,1323,1630,1857]" box="[312,1263,1750,1777]" pageId="1" pageNumber="38">
<emphasis id="A7F500766C2A9F51FE821C88F0EFF956" bold="true" box="[312,483,1750,1776]" pageId="1" pageNumber="38">Type­locality:</emphasis>
Récif 
<taxonomicName id="5281A7E76C2A9F51FD881C89F39BF957" box="[562,663,1751,1777]" class="Trematoda" family="Lepocreadiidae" genus="Hypocreadium" kingdom="Animalia" order="Plagiorchiida" pageId="1" pageNumber="43" phylum="Platyhelminthes" rank="species" species="toombo" status="sp. nov.">Toombo</taxonomicName>
, 
<collectingCountry id="ED969CF46C2A9F51FD181C89F254F957" box="[674,856,1751,1777]" name="New Caledonia" pageId="1" pageNumber="38">New Caledonia</collectingCountry>
(
<geoCoordinate id="F0B5BAA36C2A9F51FCD21C89F2C0F957" box="[872,972,1751,1777]" direction="south" orientation="latitude" pageId="1" pageNumber="38" precision="925" value="-22.433332">22°26’S</geoCoordinate>
, 
<geoCoordinate id="F0B5BAA36C2A9F51FC621C89F540F957" box="[984,1100,1751,1777]" direction="east" orientation="longitude" pageId="1" pageNumber="38" precision="925" value="166.55">166°33’E</geoCoordinate>
, 
<date id="E13FFAA46C2A9F51FBE21C89F5EEF957" box="[1112,1250,1751,1777]" pageId="1" pageNumber="38" value="2005-12-13">13/12/2005</date>
).
</paragraph>
<paragraph id="953EDC646C2A9F51FE821CA0F36BF8E7" blockId="1.[264,1323,1630,1857]" pageId="1" pageNumber="38">
<emphasis id="A7F500766C2A9F51FE821CA0F30BF8BE" bold="true" box="[312,519,1790,1816]" pageId="1" pageNumber="38">Type­specimens:</emphasis>
<typeStatus id="4A3A62C66C2A9F51FDA11CA1F384F8BF" box="[539,648,1791,1817]" pageId="1" pageNumber="38" type="holotype">Holotype</typeStatus>
MNHN JNC1680–D1; 
<typeStatus id="4A3A62C66C2A9F51FC7E1CA1F53BF8BF" box="[964,1079,1791,1817]" pageId="1" pageNumber="38" type="paratype">paratypes</typeStatus>
, MNHN JNC1680 D2–5, BMNH 2006.8.23.1–5.
</paragraph>
<caption id="C1FE8CEC6C299F52FEB01CE1F3B8F898" httpUri="https://zenodo.org/record/273586/files/figure.png" pageId="2" pageNumber="39" targetBox="[444,1162,344,1675]" targetPageId="2">
<paragraph id="953EDC646C299F52FEB01CE1F3B8F898" blockId="2.[264,1322,1727,1855]" pageId="2" pageNumber="39">
<emphasis id="A7F500766C299F52FEB01CE1F0A3F971" bold="true" box="[266,431,1727,1751]" pageId="2" pageNumber="39">FIGURES 1–2</emphasis>
. 
<taxonomicName id="5281A7E76C299F52FE061C9EF3BCF971" box="[444,688,1728,1751]" class="Trematoda" family="Lepocreadiidae" genus="Hypocreadium" kingdom="Animalia" order="Plagiorchiida" pageId="2" pageNumber="39" phylum="Platyhelminthes" rank="species" species="toombo" status="sp. nov.">
<emphasis id="A7F500766C299F52FE061C9EF3BCF971" box="[444,688,1728,1751]" italics="true" pageId="2" pageNumber="39">Hypocreadium toombo</emphasis>
</taxonomicName>
<emphasis id="A7F500766C299F52FD0C1CE1F207F971" bold="true" box="[694,779,1727,1751]" pageId="2" pageNumber="39">
<taxonomicNameLabel id="BCC6BD0D6C299F52FD0C1CE1F3FDF971" box="[694,753,1727,1751]" pageId="2" pageNumber="39" rank="species">n. sp.</taxonomicNameLabel>
1.
</emphasis>
Ventral view of holotype, uterus in bold outline. 
<emphasis id="A7F500766C299F52FAAC1CE1F426F971" bold="true" box="[1302,1322,1727,1751]" pageId="2" pageNumber="39">2.</emphasis>
Terminal genitalia. Abbreviations: Ej, Ejaculatory duct; Ep, Excretory pore; Esv, External seminal vesicle; Isv, Internal seminal vesicle; Mt, Metraterm; Ov, Ovary; Pp, Pars prostatica; T, testis; Ut, Uterus. 
<emphasis id="A7F500766C299F52FEE61D76F0C0F899" box="[348,460,1832,1855]" italics="true" pageId="2" pageNumber="39">Scale bars</emphasis>
: 1, 1 mm; 2, 200 μm.
</paragraph>
</caption>
<paragraph id="953EDC646C289F53FE821B45F5E0FEF8" blockId="3.[264,1325,283,1870]" pageId="3" pageNumber="40">
<emphasis id="A7F500766C289F53FE821B45F316FE93" bold="true" box="[312,538,283,309]" pageId="3" pageNumber="40">Note on the host:</emphasis>
the specimen studied was 
<quantity id="527971816C289F53FCD11B42F2B5FE90" box="[875,953,284,310]" metricMagnitude="-1" metricUnit="m" metricValue="5.5" pageId="3" pageNumber="40" unit="cm" value="55.0">55 cm</quantity>
in total length, the maximum recorded for this species (
<bibRefCitation id="F110A1956C289F53FD8F1B1AF3DBFEF8" author="Randall" box="[565,727,324,350]" pageId="3" pageNumber="44" refString="Randall, J. E. (2005) Reef and Shore Fishes of the South Pacific. New Caledonia to Tahiti and the Pitcairn Islands. University of Hawai'i Press, Honolulu, 707 pp." type="book" year="2005">Randall 2005</bibRefCitation>
; 
<bibRefCitation id="F110A1956C289F53FD591B1AF2ECFEF8" author="Froese" box="[739,992,324,350]" pageId="3" pageNumber="44" refString="Froese, R. &amp; Pauly, D. (2006) FishBase: World Wide Web electronic publication. Available on http: // www. fishbase. org." type="book" year="2006">Froese &amp; Pauly 2006</bibRefCitation>
), and 
<quantity id="527971816C289F53FB9C1B1AF563FEF8" box="[1062,1135,324,350]" metricMagnitude="0" metricUnit="kg" metricValue="4.7" pageId="3" pageNumber="40" unit="kg" value="4.7">4.7 kg</quantity>
in weight.
</paragraph>
<paragraph id="953EDC646C289F53FE821B35F52DFE08" blockId="3.[264,1325,283,1870]" pageId="3" pageNumber="40">
<emphasis id="A7F500766C289F53FE821B35F0C6FE23" bold="true" box="[312,458,363,389]" pageId="3" pageNumber="40">Prevalence:</emphasis>
50%, 1/2, intensity 20; a second, smaller (
<quantity id="527971816C289F53FC7B1B32F503FE20" box="[961,1039,364,390]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="40" unit="cm" value="30.0">30 cm</quantity>
, 
<quantity id="527971816C289F53FBA11B32F579FE20" box="[1051,1141,364,390]" metricMagnitude="-1" metricUnit="kg" metricValue="8.0" pageId="3" pageNumber="40" unit="kg" value="0.8">0.80 kg</quantity>
) fish, caught at Ilôt Signal, off Nouméa, 
<date id="E13FFAA46C289F53FD971BCAF3BAFE08" box="[557,694,404,430]" pageId="3" pageNumber="40" value="2006-05-30">30/05/2006</date>
, did not harbour this digenean.
</paragraph>
</subSubSection>
<subSubSection id="DD9B8FEF6C289F53FE821BE5F402FE70" box="[312,1294,443,470]" pageId="3" pageNumber="40" type="etymology">
<paragraph id="953EDC646C289F53FE821BE5F402FE70" blockId="3.[264,1325,283,1870]" box="[312,1294,443,470]" pageId="3" pageNumber="40">
<emphasis id="A7F500766C289F53FE821BE5F0C4FE73" bold="true" box="[312,456,443,469]" pageId="3" pageNumber="40">Etymology:</emphasis>
This species is named after the reef from where the host was collected.
</paragraph>
</subSubSection>
<subSubSection id="DD9B8FEF6C289F54FE821BBDF32CFD80" lastPageId="4" lastPageNumber="41" pageId="3" pageNumber="40" type="description">
<paragraph id="953EDC646C289F53FE821BBDF282FC10" blockId="3.[264,1325,283,1870]" pageId="3" pageNumber="40">
<emphasis id="A7F500766C289F53FE821BBDF0DEFE5B" bold="true" box="[312,466,483,509]" pageId="3" pageNumber="40">Description:</emphasis>
Based on 
<specimenCount id="838717ED6C289F53FDF71BBAF3FCFE58" box="[589,752,484,510]" pageId="3" pageNumber="40" type="generic">10 specimens</specimenCount>
. Body almost circular, slightly longer than wide, flattened, distinct anterior notch present, 1,572–2,310 × 1,446–2,160 (2,026 × 1,863); width 89.3–97.4 (92.0) % of body­length (
<figureCitation id="0DBAC0E16C289F53FCBA186AF269FDE8" box="[768,869,564,590]" captionStart="FIGURES 1 – 2" captionStartId="2.[266,385,1727,1751]" captionTargetBox="[444,1162,344,1675]" captionTargetId="figure@2.[368,1226,320,1688]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Hypocreadium toombo n. sp. 1. Ventral view of holotype, uterus in bold outline. 2. Terminal genitalia. Abbreviations: Ej, Ejaculatory duct; Ep, Excretory pore; Esv, External seminal vesicle; Isv, Internal seminal vesicle; Mt, Metraterm; Ov, Ovary; Pp, Pars prostatica; T, testis; Ut, Uterus. Scale bars: 1, 1 mm; 2, 200 μm." httpUri="https://zenodo.org/record/273586/files/figure.png" pageId="3" pageNumber="40">Figure 1</figureCitation>
). No tegumental spines seen. Pre­oral lobe distinct, 27–54 (36) long. Oral sucker subglobular, with ventral aperture, 124–173 × 138–188 (149 × 167). Ventral sucker rounded, equatorial, 178–264 × 188–260 (230 × 228). Sucker­length ratio 1:1.32–1.78 (1.54); sucker­width ratio 1:1.14–1.49 (1.36). Forebody 651–983 (860) long, 40.7–44.5 (42.4) % of body­length. Prepharynx not seen. Pharynx broadly oval, 85–131 × 94–134 (109 × 117). Pharynx/oral sucker ratio 1:1.37–1.53 (1.43). Oesophagus distinct, 84–157 (111). Intestinal bifurcation in midforebody, 308–463 (402) from ventral sucker. Caeca narrow, irregularly arcuate around gonads, converge posteriorly, then turn anteriorly, terminating just posterior to testes; nearest approach to posterior extremity 282–556 (368).
</paragraph>
<paragraph id="953EDC646C289F53FE82199AF59BFAC8" blockId="3.[264,1325,283,1870]" pageId="3" pageNumber="40">
Testes 2, irregularly oval, symmetrical, 234–362 × 211–345 (316 × 272), in anterior hindbody, separated by 165–260 (220). Post­testicular region 529–782 (663) long, 28.7–35.3 (32.7) % of body­length. External seminal vesicle bi­ or tripartite, at level of ventral sucker, 145–389 × 71–145 (230 × 112). Cirrus­sac large, 417–632 × 195–251 (541 × 222), claviform, reaching from point dextro­lateral to ventral sucker diagonally across posterior forebody, surrounded by gland­cells; length 23.9–29.6 (26.7) % of body­length. Internal seminal vesicle oval, in proximal end of cirrus­sac. Pars prostatica large, vesicular, folded, lined with large anuclear cell­like bodies (
<figureCitation id="0DBAC0E16C289F53FC5A1E82F546FB50" box="[992,1098,1244,1270]" captionStart="FIGURES 1 – 2" captionStartId="2.[266,385,1727,1751]" captionTargetBox="[444,1162,344,1675]" captionTargetId="figure@2.[368,1226,320,1688]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Hypocreadium toombo n. sp. 1. Ventral view of holotype, uterus in bold outline. 2. Terminal genitalia. Abbreviations: Ej, Ejaculatory duct; Ep, Excretory pore; Esv, External seminal vesicle; Isv, Internal seminal vesicle; Mt, Metraterm; Ov, Ovary; Pp, Pars prostatica; T, testis; Ut, Uterus. Scale bars: 1, 1 mm; 2, 200 μm." httpUri="https://zenodo.org/record/273586/files/figure.png" pageId="3" pageNumber="40">Figure 2</figureCitation>
). Ejaculatory duct muscular, long, complexly folded, lined with tessellated surface of distinct protruding bosses, becoming larger distally. Genital atrium distinct. Genital pore just sinistral to intestinal bifurcation or just post­bifurcal, 247–399 (337) from ventral sucker.
</paragraph>
<paragraph id="953EDC646C289F54FE821F22F268FE08" blockId="3.[264,1325,283,1870]" lastBlockId="4.[264,1323,284,550]" lastPageId="4" lastPageNumber="41" pageId="3" pageNumber="40">Ovary more or less oval, intertesticular, 172–257 × 130–196 (214 × 162), usually contiguous with sinistral testis, but separated by up to 38, separated by uterus from dextral testis, 12–92 (36) from ventral sucker. Post­ovarian region 528–820 (675) long, 30.9–35.5 (33.3) % of body­length. Seminal receptacle large, saccular, between sinistral testis and ventral sucker. Laurer’s canal opens dorsally to ovary or between ovary and closest testis. Mehlis’ gland anterior to ovary. Uterus passes between ovary and dextral testis into postovarian region, then anteriorly, narrowing dorsally to ventral sucker, then widens to form metraterm. Metraterm passes anteriorly, with thick muscular wall and sheath of glandcells, often dilated with eggs, 442–708 (587) long. Post­uterine distance 407–613 (506), 22.9–26.5 (25.0) % of body­length. Eggs tanned, operculate, 63–79 × 33–39 (70 × 36). Vitellarium follicular; follicles numerous, surround gonads, well separated from bodymargins, contiguous or nearly so pre­bifurcally, contiguous or not immediately postbifurcally where follicles may form continuous narrow band dorsal to terminal genitalia, contiguous posteriorly, lateral, dorsal and median to caeca but not ventral; previtelline distance 194–287 (256), 8.97–14.9 (12.7) % of body­length; post­vitelline distance 157–252 (208), 7.72–11.4 (10.3) % of body­length.</paragraph>
<paragraph id="953EDC646C2F9F54FE821BE2F32CFD80" blockId="4.[264,1323,284,550]" pageId="4" pageNumber="41">Excretory pore dorsal to posterior extension of uterus, between caeca. Extent of vesicle obscured by eggs, arms of excretory system reach into forebody laterally to level of oesophagus or pharynx.</paragraph>
</subSubSection>
</treatment>
</document>