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<document id="AC2CC8077C257D43B8F4E0EA6FB41960" ID-DOI="10.11646/zootaxa.3701.4.6" ID-GBIF-Dataset="dbf43e4d-705f-42a3-980d-244f665aab62" ID-ISSN="1175-5326" ID-Zenodo-Dep="219406" ID-ZooBank="D30C8478-F4DB-4820-8080-D760218207EC" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460668291003" checkinUser="plazi" docAuthor="Cruz Da Silva, Estevam L." docDate="2013" docId="1D04AF10E23DFFC1DAE3F9D7FA979632" docLanguage="en" docName="zt03701p470.pdf" docOrigin="Zootaxa 3701 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Enna rioja Silva, 2013, new species" docType="treatment" docVersion="7" lastPageNumber="470" masterDocId="E13DD768E23DFFC2DA74FFCAFFA4941E" masterDocTitle="A new Peruvian species of Enna (Araneae, Lycosoidea, Trechaleidae)" masterLastPageNumber="470" masterPageNumber="467" pageNumber="467" updateTime="1698347979324" updateUser="plazi">
<mods:mods id="1B54550C69BD15762217E0FA4B6BFFD2" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6B8B82D9A987A7808403CDAD544EB29C">
<mods:title id="E0BB1D2BBF8CD0BFD8E15EA16586E2AB">A new Peruvian species of Enna (Araneae, Lycosoidea, Trechaleidae)</mods:title>
</mods:titleInfo>
<mods:name id="2DAB2C37394D08E0FB5E2565488C1710" type="personal">
<mods:role id="E7C45321DEA1D285E4F2952C69C329ED">
<mods:roleTerm id="5D298F233D0A6F138EAE030DE2348BDF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="00D1A314B9BD4CC28E00003701C49924">Cruz Da Silva, Estevam L.</mods:namePart>
</mods:name>
<mods:typeOfResource id="8697A7674289CF14F474BCE4744F481F">text</mods:typeOfResource>
<mods:relatedItem id="55676DDBBC283B1109D18DD74648F644" type="host">
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<mods:title id="43A22622955594969EB4C18FBD70DE68">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="5C4445C329A204FFDB006EE616E147E6">
<mods:date id="124CCBB7F37C01CD1E17FE7A369BA384">2013</mods:date>
<mods:detail id="009839C6B863B0E6652EA96481121CAE" type="volume">
<mods:number id="8546B43283E21033C718E7BEEBAF43E4">3701</mods:number>
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<mods:detail id="D2131FAD8981B67AB69885CBC25047F1" type="issue">
<mods:number id="39D43AE197ED9465341E672F4B464693">4</mods:number>
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<mods:start id="31EA24D643BD12798CFE83F6F9FADE50">467</mods:start>
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</mods:extent>
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<mods:classification id="2CDBDF0499D17C60510D4068AC0F3B28">journal article</mods:classification>
<mods:identifier id="AE35923A72DDCA8355C353AAA84C9204" type="DOI">10.11646/zootaxa.3701.4.6</mods:identifier>
<mods:identifier id="CB10C515534E8A14187CD7D1120FBF2B" type="GBIF-Dataset">dbf43e4d-705f-42a3-980d-244f665aab62</mods:identifier>
<mods:identifier id="49708A8C1B6312BD0AE10A7B8F55DC7C" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="45B805F355E2C8A8A60041325946032F" type="Zenodo-Dep">219406</mods:identifier>
<mods:identifier id="8BCD793DB5C2A44CB728EDE8FD48F7D2" type="ZooBank">D30C8478-F4DB-4820-8080-D760218207EC</mods:identifier>
</mods:mods>
<treatment id="1D04AF10E23DFFC1DAE3F9D7FA979632" ID-DOI="http://doi.org/10.5281/zenodo.6159964" ID-GBIF-Taxon="119579023" ID-Zenodo-Dep="6159964" LSID="urn:lsid:plazi:treatment:1D04AF10E23DFFC1DAE3F9D7FA979632" httpUri="http://treatment.plazi.org/id/1D04AF10E23DFFC1DAE3F9D7FA979632" lastPageId="3" lastPageNumber="470" pageId="0" pageNumber="467">
<subSubSection id="DDB74D8DE23DFFC2DAE3F9D7FF579246" pageId="0" pageNumber="467" type="nomenclature">
<paragraph id="95121E06E23DFFC2DAE3F9D7FE159226" blockId="0.[151,433,1565,1624]" box="[151,433,1565,1592]" pageId="0" pageNumber="467">
<heading id="CE5AA96AE23DFFC2DAE3F9D7FE159226" bold="true" box="[151,433,1565,1592]" fontSize="11" level="1" pageId="0" pageNumber="467" reason="1">
<emphasis id="A7D9C214E23DFFC2DAE3F9D7FE159226" bold="true" box="[151,433,1565,1592]" pageId="0" pageNumber="467">
<taxonomicName id="52AD6585E23DFFC2DAE3F9D7FEBE9229" ID-CoL="39VKV" box="[151,282,1565,1591]" class="Arachnida" family="Trechaleidae" genus="Enna" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="467" phylum="Arthropoda" rank="species" species="rioja" status="sp. nov.">
<emphasis id="A7D9C214E23DFFC2DAE3F9D7FEBE9229" bold="true" box="[151,282,1565,1591]" italics="true" pageId="0" pageNumber="467">Enna rioja</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BCEA7F6FE23DFFC2DB55F9D4FE159226" box="[289,433,1566,1592]" pageId="0" pageNumber="467" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="95121E06E23DFFC2DAE3F98BFF579246" blockId="0.[151,433,1565,1624]" box="[151,243,1601,1624]" pageId="0" pageNumber="467">
<figureCitation id="0D960283E23DFFC2DAE3F98BFF579246" box="[151,243,1601,1624]" captionStart-0="FIGURES 1 – 6" captionStart-1="FIGURE 7" captionStartId-0="1.[151,269,1736,1759]" captionStartId-1="2.[151,255,1963,1986]" captionTargetBox-0="[151,1436,427,1712]" captionTargetBox-1="[252,1335,943,1938]" captionTargetId-0="figure@1.[151,1436,427,1712]" captionTargetId-1="figure@2.[252,1336,943,1938]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="FIGURES 1 – 6. Enna rioja sp. nov. Female habitus (1 dorsal, 2 lateral, 3 ventral, 4 frontal). Scale bar = 2 mm." captionText-1="FIGURE 7. Type locality of Enna rioja sp. nov. in Peru." httpUri-0="https://zenodo.org/record/219407/files/figure.png" httpUri-1="https://zenodo.org/record/219409/files/figure.png" pageId="0" pageNumber="467">Figs 1–7</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="DDB74D8DE23DFFC2DAE3F94CFAA892DE" pageId="0" pageNumber="467" type="materials_examined">
<paragraph id="95121E06E23DFFC2DAE3F94CFAA892DE" blockId="0.[151,1436,1670,2005]" pageId="0" pageNumber="467">
<emphasis id="A7D9C214E23DFFC2DAE3F94CFE1F9283" bold="true" box="[151,443,1670,1693]" pageId="0" pageNumber="467">
<typeStatus id="4A16A0A4E23DFFC2DAE3F94CFF6B9283" box="[151,207,1670,1693]" pageId="0" pageNumber="467">Type</typeStatus>
material:
<typeStatus id="4A16A0A4E23DFFC2DB3FF94CFE139283" box="[331,439,1670,1693]" pageId="0" pageNumber="467" type="holotype">Holotype</typeStatus>
:
</emphasis>
female from San Martin,
<taxonomicName id="52AD6585E23DFFC2D8AAF94CFCBF9283" box="[734,795,1670,1693]" class="Arachnida" family="Trechaleidae" genus="Enna" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="469" phylum="Arthropoda" rank="species" species="rioja" status="sp. nov.">Rioja</taxonomicName>
, Paitoja,
<collectingCountry id="EDBA5E96E23DFFC2D9F6F94CFC179283" box="[898,947,1670,1693]" name="Peru" pageId="0" pageNumber="467">Peru</collectingCountry>
[
<geoCoordinate id="F09978C1E23DFFC2D9B7F94CFBBB9283" box="[963,1055,1670,1693]" direction="south" orientation="latitude" pageId="0" pageNumber="467" precision="925" value="-6.05">06°03’S</geoCoordinate>
,
<geoCoordinate id="F09978C1E23DFFC2DE59F94CFB379283" box="[1069,1171,1670,1693]" direction="west" orientation="longitude" pageId="0" pageNumber="467" precision="925" value="-77.166664">77°10’W</geoCoordinate>
],
<date id="E11338C6E23DFFC2DEDCF94CFABE9283" box="[1192,1306,1670,1693]" pageId="0" pageNumber="467" value="2008-10-10">10.X.2008</date>
, C. Albujar (MUSM 505344).
<typeStatus id="4A16A0A4E23DFFC2DB2AF963FE1B92DE" box="[350,447,1705,1728]" pageId="0" pageNumber="467" type="paratype">Paratype</typeStatus>
: one female, same locality, date and collector as
<typeStatus id="4A16A0A4E23DFFC2D9B2F963FB8292DE" box="[966,1062,1705,1728]" pageId="0" pageNumber="467" type="holotype">holotype</typeStatus>
, deposited at FMNH.
</paragraph>
</subSubSection>
<subSubSection id="DDB74D8DE23DFFC2DAB3F901FBB192FC" box="[199,1045,1739,1762]" pageId="0" pageNumber="467" type="etymology">
<paragraph id="95121E06E23DFFC2DAB3F901FBB192FC" blockId="0.[151,1436,1670,2005]" box="[199,1045,1739,1762]" pageId="0" pageNumber="467">
<emphasis id="A7D9C214E23DFFC2DAB3F901FEE392FC" bold="true" box="[199,327,1739,1762]" pageId="0" pageNumber="467">Etymology.</emphasis>
The specific name is a noun in apposition taken from
<typeStatus id="4A16A0A4E23DFFC2D9FEF901FC1C92FC" box="[906,952,1739,1762]" pageId="0" pageNumber="467">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="DDB74D8DE23DFFC3DAB3F924FEA69561" lastPageId="1" lastPageNumber="468" pageId="0" pageNumber="467" type="diagnosis">
<paragraph id="95121E06E23DFFC2DAB3F924FB44938E" blockId="0.[151,1436,1670,2005]" pageId="0" pageNumber="467">
<emphasis id="A7D9C214E23DFFC2DAB3F924FE98931B" bold="true" box="[199,316,1774,1797]" pageId="0" pageNumber="467">Diagnosis.</emphasis>
The female of
<taxonomicName id="52AD6585E23DFFC2DB90F924FD93931B" box="[484,567,1774,1797]" class="Arachnida" family="Trechaleidae" genus="Enna" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="467" phylum="Arthropoda" rank="species" species="rioja" status="sp. nov.">
<emphasis id="A7D9C214E23DFFC2DB90F924FD93931B" box="[484,567,1774,1797]" italics="true" pageId="0" pageNumber="467">E. rioja</emphasis>
</taxonomicName>
<emphasis id="A7D9C214E23DFFC2D834F924FD33931B" bold="true" box="[576,663,1774,1797]" pageId="0" pageNumber="467">
<taxonomicNameLabel id="BCEA7F6FE23DFFC2D834F924FD33931B" box="[576,663,1774,1797]" pageId="0" pageNumber="467" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
resembles those of
<taxonomicName id="52AD6585E23DFFC2D905F924FC6A931B" box="[881,974,1773,1797]" class="Arachnida" family="Trechaleidae" genus="Enna" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="467" phylum="Arthropoda" rank="species" species="baeza">
<emphasis id="A7D9C214E23DFFC2D905F924FC6A931B" box="[881,974,1773,1797]" italics="true" pageId="0" pageNumber="467">E. baeza</emphasis>
</taxonomicName>
(Silva
<emphasis id="A7D9C214E23DFFC2DE6AF924FBEA931B" box="[1054,1102,1773,1797]" italics="true" pageId="0" pageNumber="467">et al</emphasis>
., 2008: 93, figs 78, 79) by the small rounded projection on the head of spermathecae (HS) and
<taxonomicName id="52AD6585E23DFFC2D935F8DBFC1C9336" box="[833,952,1809,1832]" class="Arachnida" family="Trechaleidae" genus="Enna" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="467" phylum="Arthropoda" rank="species" species="osaensis">
<emphasis id="A7D9C214E23DFFC2D935F8DBFC1C9336" box="[833,952,1809,1832]" italics="true" pageId="0" pageNumber="467">E. osaensis</emphasis>
</taxonomicName>
(Silva
<emphasis id="A7D9C214E23DFFC2DE70F8DBFB969336" box="[1028,1074,1808,1832]" italics="true" pageId="0" pageNumber="467">et al</emphasis>
., 2012: 57, fig. 6) by the presence of a large accessory spermathecae (
<collectingCountry id="EDBA5E96E23DFFC2D87BF8F9FD979354" box="[527,563,1843,1866]" name="American Samoa" pageId="0" pageNumber="467">AS</collectingCountry>
) and by the shape of the projected scape (SC) of the middle field of the epigynum (MF), but can be distinguished from both species by the slightly projected lateral lobes (LL) (
<figureCitation id="0D960283E23DFFC2DE03F89CFB139373" box="[1143,1207,1878,1901]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,269,1736,1759]" captionTargetBox="[151,1436,427,1712]" captionTargetId="figure@1.[151,1436,427,1712]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Enna rioja sp. nov. Female habitus (1 dorsal, 2 lateral, 3 ventral, 4 frontal). Scale bar = 2 mm." httpUri="https://zenodo.org/record/219407/files/figure.png" pageId="0" pageNumber="467">Fig. 5</figureCitation>
) and by the large and conspicuous accessory spermathecae and by the elliptical shape of head of spermathecae (HS) (
<figureCitation id="0D960283E23DFFC2DEE5F8B3FB77938E" box="[1169,1235,1913,1936]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,269,1736,1759]" captionTargetBox="[151,1436,427,1712]" captionTargetId="figure@1.[151,1436,427,1712]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Enna rioja sp. nov. Female habitus (1 dorsal, 2 lateral, 3 ventral, 4 frontal). Scale bar = 2 mm." httpUri="https://zenodo.org/record/219407/files/figure.png" pageId="0" pageNumber="467">Fig. 6</figureCitation>
).
</paragraph>
<paragraph id="95121E06E23DFFC3DAB3F851FEA69561" blockId="0.[151,1436,1670,2005]" lastBlockId="1.[151,1436,151,383]" lastPageId="1" lastPageNumber="468" pageId="0" pageNumber="467">
<emphasis id="A7D9C214E23DFFC2DAB3F851FEA793AC" bold="true" box="[199,259,1947,1970]" pageId="0" pageNumber="467">Note.</emphasis>
Ten species of
<taxonomicName id="52AD6585E23DFFC2DBDFF851FE4793AC" box="[427,483,1947,1970]" class="Arachnida" family="Trechaleidae" genus="Enna" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="467" phylum="Arthropoda" rank="genus">
<emphasis id="A7D9C214E23DFFC2DBDFF851FE4793AC" box="[427,483,1947,1970]" italics="true" pageId="0" pageNumber="467">Enna</emphasis>
</taxonomicName>
were described from
<collectingCountry id="EDBA5E96E23DFFC2D8BAF851FD5B93AC" box="[718,767,1947,1970]" name="Peru" pageId="0" pageNumber="467">Peru</collectingCountry>
and
<taxonomicName id="52AD6585E23DFFC2D940F851FB5293AC" authority="Silva, Lise &amp; Carico, 2008" authorityName="Silva, Lise &amp; Carico" authorityYear="2008" box="[820,1270,1946,1970]" class="Arachnida" family="Trechaleidae" genus="Enna" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="467" phylum="Arthropoda" rank="species" species="huanuco">
<emphasis id="A7D9C214E23DFFC2D940F851FC6B93AC" box="[820,975,1946,1970]" italics="true" pageId="0" pageNumber="467">Enna huanuco</emphasis>
Silva, Lise &amp; Carico, 2008
</taxonomicName>
is the only one with the female still unknown. In this paper, the female from San Martin,
<collectingCountry id="EDBA5E96E23DFFC2D9CDF874FC4E93CB" box="[953,1002,1982,2005]" name="Peru" pageId="0" pageNumber="467">Peru</collectingCountry>
is described as separate species and not grouped with
<taxonomicName id="52AD6585E23CFFC3DB40FF52FE1694B1" box="[308,434,151,175]" class="Arachnida" family="Trechaleidae" genus="Enna" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="468" phylum="Arthropoda" rank="species" species="huanuco">
<emphasis id="A7D9C214E23CFFC3DB40FF52FE1694B1" box="[308,434,151,175]" italics="true" pageId="1" pageNumber="468">E. huanuco</emphasis>
</taxonomicName>
, since the locations where they have been found are not close together (San Martin and
<taxonomicName id="52AD6585E23CFFC3DAE3FF70FF5E94CF" box="[151,250,186,209]" class="Arachnida" family="Trechaleidae" genus="Enna" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="468" phylum="Arthropoda" rank="species" species="huanuco">Huanuco</taxonomicName>
, respectively), distant about
<quantity id="5255B3E3E23CFFC3D843FF70FD2694CF" box="[567,642,186,209]" metricMagnitude="5" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="468" unit="km" value="400.0">400km</quantity>
apart and males and females have not been collected together from the same area. Both species (
<taxonomicName id="52AD6585E23CFFC3DBD3FF17FE5D94EA" box="[423,505,221,244]" class="Arachnida" family="Trechaleidae" genus="Enna" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="468" phylum="Arthropoda" rank="species" species="rioja" status="sp. nov.">
<emphasis id="A7D9C214E23CFFC3DBD3FF17FE5D94EA" box="[423,505,221,244]" italics="true" pageId="1" pageNumber="468">E. rioja</emphasis>
</taxonomicName>
<emphasis id="A7D9C214E23CFFC3D874FF17FDF294EA" bold="true" box="[512,598,221,244]" pageId="1" pageNumber="468">
<taxonomicNameLabel id="BCEA7F6FE23CFFC3D874FF17FDF294EA" box="[512,598,221,244]" pageId="1" pageNumber="468" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="52AD6585E23CFFC3D8FEFF17FB8894EA" authority="Silva, Lise &amp; Carico, 2008" authorityName="Silva, Lise &amp; Carico" authorityYear="2008" box="[650,1068,220,244]" class="Arachnida" family="Trechaleidae" genus="Enna" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="468" phylum="Arthropoda" rank="species" species="huanuco">
<emphasis id="A7D9C214E23CFFC3D8FEFF17FCA094EA" box="[650,772,220,244]" italics="true" pageId="1" pageNumber="468">E. huanuco</emphasis>
Silva, Lise &amp; Carico, 2008
</taxonomicName>
) also present some morphological differences: carapace colouration (
<taxonomicName id="52AD6585E23CFFC3D873FECAFD259509" box="[519,641,255,279]" class="Arachnida" family="Trechaleidae" genus="Enna" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="468" phylum="Arthropoda" rank="species" species="huanuco">
<emphasis id="A7D9C214E23CFFC3D873FECAFD259509" box="[519,641,255,279]" italics="true" pageId="1" pageNumber="468">E. huanuco</emphasis>
</taxonomicName>
is dark brown with lateral brownish bands and
<taxonomicName id="52AD6585E23CFFC3DE08FECAFB6A9509" box="[1148,1230,256,279]" class="Arachnida" family="Trechaleidae" genus="Enna" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="468" phylum="Arthropoda" rank="species" species="rioja" status="sp. nov.">
<emphasis id="A7D9C214E23CFFC3DE08FECAFB6A9509" box="[1148,1230,256,279]" italics="true" pageId="1" pageNumber="468">E. rioja</emphasis>
</taxonomicName>
<emphasis id="A7D9C214E23CFFC3DEA0FECAFA8D9509" bold="true" box="[1236,1321,256,279]" pageId="1" pageNumber="468">
<taxonomicNameLabel id="BCEA7F6FE23CFFC3DEA0FECAFA8D9509" box="[1236,1321,256,279]" pageId="1" pageNumber="468" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has a dark brown carapace without bands) and carapace dimensions (most
<taxonomicName id="52AD6585E23CFFC3D914FEE8FC3C9527" box="[864,920,290,313]" class="Arachnida" family="Trechaleidae" genus="Enna" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="468" phylum="Arthropoda" rank="genus">
<emphasis id="A7D9C214E23CFFC3D914FEE8FC3C9527" box="[864,920,290,313]" italics="true" pageId="1" pageNumber="468">Enna</emphasis>
</taxonomicName>
males have a longer and wider carapace than females, but the female of
<taxonomicName id="52AD6585E23CFFC3DBB1FE8FFDBF9542" box="[453,539,325,348]" class="Arachnida" family="Trechaleidae" genus="Enna" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="468" phylum="Arthropoda" rank="species" species="rioja" status="sp. nov.">
<emphasis id="A7D9C214E23CFFC3DBB1FE8FFDBF9542" box="[453,539,325,348]" italics="true" pageId="1" pageNumber="468">E. rioja</emphasis>
</taxonomicName>
<emphasis id="A7D9C214E23CFFC3D851FE8FFDDA9542" bold="true" box="[549,638,325,348]" pageId="1" pageNumber="468">
<taxonomicNameLabel id="BCEA7F6FE23CFFC3D851FE8FFDDA9542" box="[549,638,325,348]" pageId="1" pageNumber="468" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has a longer and wider carapace compared to the male specimen of
<taxonomicName id="52AD6585E23CFFC3DFF1FE8FFF579561" class="Arachnida" family="Trechaleidae" genus="Enna" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="468" phylum="Arthropoda" rank="species" species="huanuco">
<emphasis id="A7D9C214E23CFFC3DFF1FE8FFF579561" italics="true" pageId="1" pageNumber="468">E. huanuco</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<caption id="C1D24E8EE23CFFC3DAE3F902FA9492C1" box="[151,1328,1736,1759]" httpUri="https://zenodo.org/record/219407/files/figure.png" pageId="1" pageNumber="468" targetBox="[151,1436,427,1712]" targetPageId="1">
<paragraph id="95121E06E23CFFC3DAE3F902FA9492C1" blockId="1.[151,1328,1736,1759]" box="[151,1328,1736,1759]" pageId="1" pageNumber="468">
<emphasis id="A7D9C214E23CFFC3DAE3F902FEE692C1" bold="true" box="[151,322,1736,1759]" pageId="1" pageNumber="468">FIGURES 1–6.</emphasis>
<taxonomicName id="52AD6585E23CFFC3DB3DF902FE1892C1" box="[329,444,1736,1759]" class="Arachnida" family="Trechaleidae" genus="Enna" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="468" phylum="Arthropoda" rank="species" species="rioja" status="sp. nov.">
<emphasis id="A7D9C214E23CFFC3DB3DF902FE1892C1" box="[329,444,1736,1759]" italics="true" pageId="1" pageNumber="468">Enna rioja</emphasis>
</taxonomicName>
<emphasis id="A7D9C214E23CFFC3DBB7F902FDBD92C1" bold="true" box="[451,537,1736,1759]" pageId="1" pageNumber="468">
<taxonomicNameLabel id="BCEA7F6FE23CFFC3DBB7F902FDBD92C1" box="[451,537,1736,1759]" pageId="1" pageNumber="468" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Female habitus (1 dorsal, 2 lateral, 3 ventral, 4 frontal). Scale bar = 2mm.
</paragraph>
</caption>
<subSubSection id="DDB74D8DE23CFFC1DAE3F8C7FCE49527" lastPageId="3" lastPageNumber="470" pageId="1" pageNumber="468" type="description">
<paragraph id="95121E06E23CFFC1DAE3F8C7FCE49527" blockId="1.[151,1437,1805,2036]" lastBlockId="3.[151,1436,152,348]" lastPageId="3" lastPageNumber="470" pageId="1" pageNumber="468">
<emphasis id="A7D9C214E23CFFC3DAE3F8C7FE86933A" bold="true" box="[151,290,1805,1828]" pageId="1" pageNumber="468">Description.</emphasis>
Female (
<typeStatus id="4A16A0A4E23CFFC3DBF1F8C7FE48933A" box="[389,492,1805,1828]" pageId="1" pageNumber="468" type="holotype">Holotype</typeStatus>
, San Martin,
<collectingCountry id="EDBA5E96E23CFFC3D80AF8C7FD15933A" box="[638,689,1805,1828]" name="Peru" pageId="1" pageNumber="468">Peru</collectingCountry>
, MUSM 505344). Total length 10.50. Carapace 5.58 long, 4.78 wide, dark brown, fovea marked (
<figureCitation id="0D960283E23CFFC3DBBCF8FAFD8A9359" box="[456,558,1840,1863]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,269,1736,1759]" captionTargetBox="[151,1436,427,1712]" captionTargetId="figure@1.[151,1436,427,1712]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Enna rioja sp. nov. Female habitus (1 dorsal, 2 lateral, 3 ventral, 4 frontal). Scale bar = 2 mm." httpUri="https://zenodo.org/record/219407/files/figure.png" pageId="1" pageNumber="468">Figs 1, 2</figureCitation>
). Clypeus dark brown, 0.46 high (
<figureCitation id="0D960283E23CFFC3D9DEF8FAFC4A9359" box="[938,1006,1840,1863]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,269,1736,1759]" captionTargetBox="[151,1436,427,1712]" captionTargetId="figure@1.[151,1436,427,1712]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Enna rioja sp. nov. Female habitus (1 dorsal, 2 lateral, 3 ventral, 4 frontal). Scale bar = 2 mm." httpUri="https://zenodo.org/record/219407/files/figure.png" pageId="1" pageNumber="468">Fig. 4</figureCitation>
). Anterior eye row straight, 1.51 wide; posterior 2.35 wide (
<figureCitation id="0D960283E23CFFC3DB03F899FE1E9374" box="[375,442,1875,1898]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,269,1736,1759]" captionTargetBox="[151,1436,427,1712]" captionTargetId="figure@1.[151,1436,427,1712]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Enna rioja sp. nov. Female habitus (1 dorsal, 2 lateral, 3 ventral, 4 frontal). Scale bar = 2 mm." httpUri="https://zenodo.org/record/219407/files/figure.png" pageId="1" pageNumber="468">Fig. 4</figureCitation>
). Eye diameters, interdistances, and median ocular quadrangle: AME 0.21, ALE 0.18, PME 0.31, PLE 0.18; AME-AME 0.18, AME-ALE 0.16, PME-PME 0.65, PME-PLE 0.49, OQA 0.65, OQP 1.20, OQH 0.71. Chelicerae reddish-brown, bristly (
<figureCitation id="0D960283E23CFFC3D866F852FDF293B1" box="[530,598,1944,1967]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,269,1736,1759]" captionTargetBox="[151,1436,427,1712]" captionTargetId="figure@1.[151,1436,427,1712]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Enna rioja sp. nov. Female habitus (1 dorsal, 2 lateral, 3 ventral, 4 frontal). Scale bar = 2 mm." httpUri="https://zenodo.org/record/219407/files/figure.png" pageId="1" pageNumber="468">Fig. 4</figureCitation>
); promargin and retromargin of fang furrow with three teeth equidistant and equal in size. Sternum yellow, with scattered setae (
<figureCitation id="0D960283E23CFFC3D8B8F871FCAB93CC" box="[716,783,1979,2002]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,269,1736,1759]" captionTargetBox="[151,1436,427,1712]" captionTargetId="figure@1.[151,1436,427,1712]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Enna rioja sp. nov. Female habitus (1 dorsal, 2 lateral, 3 ventral, 4 frontal). Scale bar = 2 mm." httpUri="https://zenodo.org/record/219407/files/figure.png" pageId="1" pageNumber="468">Fig. 3</figureCitation>
); 2.66 long, 2.52 wide. Labium dark brown, yellow distally (
<figureCitation id="0D960283E23CFFC3DAEBF817FF7A93EA" box="[159,222,2013,2036]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,269,1736,1759]" captionTargetBox="[151,1436,427,1712]" captionTargetId="figure@1.[151,1436,427,1712]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Enna rioja sp. nov. Female habitus (1 dorsal, 2 lateral, 3 ventral, 4 frontal). Scale bar = 2 mm." httpUri="https://zenodo.org/record/219407/files/figure.png" pageId="1" pageNumber="468">Fig. 3</figureCitation>
), 1.06 long, 0.93 wide. Legs light brown with dark brown annuli on femora (
<figureCitation id="0D960283E23CFFC3DE7AF817FBC893EA" box="[1038,1132,2013,2036]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,269,1736,1759]" captionTargetBox="[151,1436,427,1712]" captionTargetId="figure@1.[151,1436,427,1712]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Enna rioja sp. nov. Female habitus (1 dorsal, 2 lateral, 3 ventral, 4 frontal). Scale bar = 2 mm." httpUri="https://zenodo.org/record/219407/files/figure.png" pageId="1" pageNumber="468">Figs 1, 2</figureCitation>
), relative length: IV-I-II-III, I—femur 6.25/ tibia-patella 8.64/ metatarsus 6.65/ tarsus 2.66/ total 24.20; II—6.38/ 8.37/ 6.11/ 2.55/ 23.41; III—5.18/ 5.45/ 4.12/ 2.12/ 16.87; IV—6.65/ 7.98/ 7.31/ 2.92/ 24.85. Ventral pairs of macrosetae on tibiae:
<date id="E11338C6E23EFFC1DED8FF70FB7794CF" box="[1196,1235,186,209]" bridgedPair="-" pageId="3" pageNumber="470">I–4</date>
;
<date id="E11338C6E23EFFC1DEABFF70FAAA94CF" box="[1247,1294,186,209]" bridgedPair="-" pageId="3" pageNumber="470">II–3</date>
;
<date id="E11338C6E23EFFC1DF6EFF70FAF794CF" box="[1306,1363,186,209]" bridgedPair="-" pageId="3" pageNumber="470">III–3</date>
;
<date id="E11338C6E23EFFC1DF2AFF70FA3894CF" box="[1374,1436,186,209]" bridgedPair="-" pageId="3" pageNumber="470">IV–3.</date>
Abdomen 4.92 long, gray with two pairs of sigillae (
<figureCitation id="0D960283E23EFFC1D89EFF17FC8994EA" box="[746,813,221,244]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,269,1736,1759]" captionTargetBox="[151,1436,427,1712]" captionTargetId="figure@1.[151,1436,427,1712]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Enna rioja sp. nov. Female habitus (1 dorsal, 2 lateral, 3 ventral, 4 frontal). Scale bar = 2 mm." httpUri="https://zenodo.org/record/219407/files/figure.png" pageId="3" pageNumber="470">Fig. 1</figureCitation>
). Venter light brown, bristly (
<figureCitation id="0D960283E23EFFC1DE0BFF17FB6094EA" box="[1151,1220,221,244]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,269,1736,1759]" captionTargetBox="[151,1436,427,1712]" captionTargetId="figure@1.[151,1436,427,1712]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Enna rioja sp. nov. Female habitus (1 dorsal, 2 lateral, 3 ventral, 4 frontal). Scale bar = 2 mm." httpUri="https://zenodo.org/record/219407/files/figure.png" pageId="3" pageNumber="470">Fig. 3</figureCitation>
). Epigynum with a projected scape (SC) (
<figureCitation id="0D960283E23EFFC1DBF3FECAFE6D9509" box="[391,457,256,279]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,269,1736,1759]" captionTargetBox="[151,1436,427,1712]" captionTargetId="figure@1.[151,1436,427,1712]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Enna rioja sp. nov. Female habitus (1 dorsal, 2 lateral, 3 ventral, 4 frontal). Scale bar = 2 mm." httpUri="https://zenodo.org/record/219407/files/figure.png" pageId="3" pageNumber="470">Fig. 5</figureCitation>
). Head of spermathecae (HS) rounded and with a rounded whitish projection (
<figureCitation id="0D960283E23EFFC1DF6FFECAFAF99509" box="[1307,1373,256,279]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,269,1736,1759]" captionTargetBox="[151,1436,427,1712]" captionTargetId="figure@1.[151,1436,427,1712]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Enna rioja sp. nov. Female habitus (1 dorsal, 2 lateral, 3 ventral, 4 frontal). Scale bar = 2 mm." httpUri="https://zenodo.org/record/219407/files/figure.png" pageId="3" pageNumber="470">Fig. 6</figureCitation>
, HS). Accessory spermathecae conspicuous and rounded (
<figureCitation id="0D960283E23EFFC1D8B6FEE8FCA79527" box="[706,771,290,313]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,269,1736,1759]" captionTargetBox="[151,1436,427,1712]" captionTargetId="figure@1.[151,1436,427,1712]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Enna rioja sp. nov. Female habitus (1 dorsal, 2 lateral, 3 ventral, 4 frontal). Scale bar = 2 mm." httpUri="https://zenodo.org/record/219407/files/figure.png" pageId="3" pageNumber="470">Fig. 6</figureCitation>
,
<collectingCountry id="EDBA5E96E23EFFC1D97AFEE8FC909527" box="[782,820,290,313]" name="American Samoa" pageId="3" pageNumber="470">AS</collectingCountry>
).
</paragraph>
</subSubSection>
<caption id="C1D24E8EE23FFFC0DAE3FCF5FDD89767" httpUri="https://zenodo.org/record/219408/files/figure.png" pageId="2" pageNumber="469" targetBox="[151,1434,194,805]" targetPageId="2">
<paragraph id="95121E06E23FFFC0DAE3FCF5FDD89767" blockId="2.[151,1436,831,889]" pageId="2" pageNumber="469">
<emphasis id="A7D9C214E23FFFC0DAE3FCF5FEEB9748" bold="true" box="[151,335,831,854]" pageId="2" pageNumber="469">FIGURES 5, 6.</emphasis>
<taxonomicName id="52AD6585E23FFFC0DB29FC8AFE739749" box="[349,471,832,855]" class="Arachnida" family="Trechaleidae" genus="Enna" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="469" phylum="Arthropoda" rank="species" species="rioja" status="sp. nov.">
<emphasis id="A7D9C214E23FFFC0DB29FC8AFE739749" box="[349,471,832,855]" italics="true" pageId="2" pageNumber="469">Enna rioja</emphasis>
</taxonomicName>
<emphasis id="A7D9C214E23FFFC0DB90FCF5FDE49748" bold="true" box="[484,576,831,854]" pageId="2" pageNumber="469">
<taxonomicNameLabel id="BCEA7F6FE23FFFC0DB90FCF5FDE49748" box="[484,576,831,854]" pageId="2" pageNumber="469" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Female epigynum (5 ventral, 6 dorsal). (AS, accessory spermathecae; FD, fertilization duct; HS, head of spermathecae).
</paragraph>
</caption>
<caption id="C1D24E8EE23FFFC0DAE3F861FD5C93DC" box="[151,760,1963,1986]" httpUri="https://zenodo.org/record/219409/files/figure.png" pageId="2" pageNumber="469" targetBox="[252,1335,943,1938]" targetPageId="2">
<paragraph id="95121E06E23FFFC0DAE3F861FD5C93DC" blockId="2.[151,760,1963,1986]" box="[151,760,1963,1986]" pageId="2" pageNumber="469">
<emphasis id="A7D9C214E23FFFC0DAE3F861FEBD93DC" bold="true" box="[151,281,1963,1986]" pageId="2" pageNumber="469">FIGURE 7.</emphasis>
Type locality of
<taxonomicName id="52AD6585E23FFFC0DBBBF861FDE693DC" box="[463,578,1963,1986]" class="Arachnida" family="Trechaleidae" genus="Enna" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="469" phylum="Arthropoda" rank="species" species="rioja" status="sp. nov.">
<emphasis id="A7D9C214E23FFFC0DBBBF861FDE693DC" box="[463,578,1963,1986]" italics="true" pageId="2" pageNumber="469">Enna rioja</emphasis>
</taxonomicName>
<emphasis id="A7D9C214E23FFFC0D83CF861FD3A93DC" bold="true" box="[584,670,1963,1986]" pageId="2" pageNumber="469">
<taxonomicNameLabel id="BCEA7F6FE23FFFC0D83CF861FD3A93DC" box="[584,670,1963,1986]" pageId="2" pageNumber="469" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in Peru.
</paragraph>
</caption>
<subSubSection id="DDB74D8DE23EFFC1DAB3FE8FFA979632" pageId="3" pageNumber="470" type="distribution">
<paragraph id="95121E06E23EFFC1DAB3FE8FFCD39542" blockId="3.[151,1436,152,348]" box="[199,887,325,348]" pageId="3" pageNumber="470">
<emphasis id="A7D9C214E23EFFC1DAB3FE8FFEFD9542" bold="true" box="[199,345,325,348]" pageId="3" pageNumber="470">Distribution.</emphasis>
Known only from the
<typeStatus id="4A16A0A4E23EFFC1D838FE8FFDDE9542" box="[588,634,325,348]" pageId="3" pageNumber="470">type</typeStatus>
locality (
<collectingCountry id="EDBA5E96E23EFFC1D8ABFE8FFCB79542" box="[735,787,325,348]" name="Peru" pageId="3" pageNumber="470">Peru</collectingCountry>
) (
<figureCitation id="0D960283E23EFFC1D95DFE8FFCCD9542" box="[809,873,325,348]" captionStart="FIGURE 7" captionStartId="2.[151,255,1963,1986]" captionTargetBox="[252,1335,943,1938]" captionTargetId="figure@2.[252,1336,943,1938]" captionTargetPageId="2" captionText="FIGURE 7. Type locality of Enna rioja sp. nov. in Peru." httpUri="https://zenodo.org/record/219409/files/figure.png" pageId="3" pageNumber="470">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="95121E06E23EFFC1DAE3FE67FECC95DA" blockId="3.[151,360,429,452]" box="[151,360,429,452]" pageId="3" pageNumber="470">
<emphasis id="A7D9C214E23EFFC1DAE3FE67FECC95DA" bold="true" box="[151,360,429,452]" pageId="3" pageNumber="470">Acknowledgments</emphasis>
</paragraph>
<paragraph id="95121E06E23EFFC1DAE3FE38FA979632" blockId="3.[151,1436,498,556]" pageId="3" pageNumber="470">I wish to thank Diana Silva-Dávila (MUSM) for the loan of the material. The images were taken in Petra Sierwald’s lab at the Field Museum, Robin Delapena (FMNH) graciously helped obtaining the photos of the female genitalia.</paragraph>
</subSubSection>
</treatment>
</document>
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