271 lines
34 KiB
XML
271 lines
34 KiB
XML
<document id="0721DF4F632F0613170DD223D103916F" ID-DOI="10.5281/zenodo.180540" ID-GBIF-Dataset="de2ad7f5-b005-43a6-bc79-c936647ff95f" ID-ISSN="1175-5326" ID-Zenodo-Dep="180540" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459809786369" checkinUser="plazi" docAuthor="Karaman, Ivo M. & Horvatović, Mladen" docDate="2008" docId="1C1B2230FFD8E271C1A30235FC78F83A" docLanguage="en" docName="zt01687p066.pdf" docOrigin="Zootaxa 1687" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Mladenoniscus belavodae" docType="treatment" docVersion="7" lastPageNumber="64" masterDocId="E0225A48FFD9E275C1340318FF94FFF2" masterDocTitle="Mladenoniscus belavodae n. g., n. sp., a troglobitic oniscid (Isopoda: Oniscidea: Trichoniscidae) from Macedonia" masterLastPageNumber="66" masterPageNumber="60" pageNumber="61" updateTime="1698220882663" updateUser="plazi">
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<mods:mods id="4193F1C3910AA161CAB58209B7F3C24B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="30F75FAB547A594E68C142CB86EB160F">
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<mods:title id="7C500A644169829E4A603210A1E03FF4">Mladenoniscus belavodae n. g., n. sp., a troglobitic oniscid (Isopoda: Oniscidea: Trichoniscidae) from Macedonia</mods:title>
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<mods:name id="B296BDF059F800D7E98095ED18D4579C" type="personal">
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<mods:role id="F7D2EA94B0EF76AAEEC60687818506D0">
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<mods:roleTerm id="0F21184FF1BDC2EAC72E8569F36F1517">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="DE6A38EA2764183091C8E27B6C351F68">Karaman, Ivo M.</mods:namePart>
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</mods:name>
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<mods:name id="89F305C01F0EDF9A0B521ACCD9B0106F" type="personal">
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<mods:role id="2F206686EE1C812E5AA4F19FA6383C36">
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<mods:roleTerm id="16FB76B476F31EE1462AE078EAE5D82B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F25E1F4642DD495067ED9F0D5ADEE6C0">Horvatović, Mladen</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="52E3630BAAB3B78D6AD4F05591F9943A">text</mods:typeOfResource>
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<mods:relatedItem id="1CDF6BB5133A15B339271620B0397059" type="host">
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<mods:titleInfo id="20161E4AF5501010498B48FD12F1E6EB">
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<mods:title id="5DFD7DCD8567DB0BE4E8D6408B98D789">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="C5946B1C10E99A3A54FD1278AB9932FE">
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<mods:date id="565161739375332A689722BA9BA81318">2008</mods:date>
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<mods:detail id="AC96A70C991C4AD79FABBB707C65D7E8" type="volume">
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<mods:number id="F3A409B93E0F66EC9EBD57A27D5773B8">1687</mods:number>
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<mods:start id="EF3773E41BA30D8CA8E56CAB10600D5B">60</mods:start>
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<mods:end id="810721497620DA467CE3EAAD8C4985E2">66</mods:end>
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<mods:classification id="AB5CD93849036255DA172F0F50905929">journal article</mods:classification>
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<mods:identifier id="4AA3B29E7841C072914C81BC8066E563" type="DOI">10.5281/zenodo.180540</mods:identifier>
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<mods:identifier id="ED6628870074B1BF6EE4FB7548C18116" type="GBIF-Dataset">de2ad7f5-b005-43a6-bc79-c936647ff95f</mods:identifier>
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<mods:identifier id="313AEA7FD1099E35BC084FEA31E30E0F" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="30EE40815C3B7EE7A00300DDF4820625" type="Zenodo-Dep">180540</mods:identifier>
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<treatment id="1C1B2230FFD8E271C1A30235FC78F83A" ID-DOI="http://doi.org/10.5281/zenodo.6227996" ID-GBIF-Taxon="119353538" ID-Zenodo-Dep="6227996" LSID="urn:lsid:plazi:treatment:1C1B2230FFD8E271C1A30235FC78F83A" httpUri="http://treatment.plazi.org/id/1C1B2230FFD8E271C1A30235FC78F83A" lastPageId="4" lastPageNumber="64" pageId="1" pageNumber="61">
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<subSubSection id="DCA8C0ADFFD8E274C1A30235FE8AFE98" pageId="1" pageNumber="61" type="nomenclature">
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<paragraph id="940D9326FFD8E274C1A30235FD77FEB5" blockId="1.[151,739,301,362]" box="[151,739,301,327]" pageId="1" pageNumber="61">
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<heading id="CF45244AFFD8E274C1A30235FD77FEB5" bold="true" box="[151,739,301,327]" fontSize="11" level="1" pageId="1" pageNumber="61" reason="1">
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<emphasis id="A6C64F34FFD8E274C1A30235FD77FEB5" bold="true" box="[151,739,301,327]" pageId="1" pageNumber="61">
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||
<taxonomicName id="53B2E8A5FFD8E274C1A30235FE5BFEB5" ID-CoL="43S7V" box="[151,463,301,327]" class="Malacostraca" family="Trichoniscidae" genus="Mladenoniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="61" phylum="Arthropoda" rank="species" species="belavodae">
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<emphasis id="A6C64F34FFD8E274C1A30235FE5BFEB5" bold="true" box="[151,463,301,327]" italics="true" pageId="1" pageNumber="61">Mladenoniscus belavodae</emphasis>
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</taxonomicName>
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n. g., n. sp., Karaman
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="940D9326FFD8E274C1A30248FE8AFE98" blockId="1.[151,739,301,362]" box="[151,286,336,362]" pageId="1" pageNumber="61">
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||
(
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<figureCitation id="0C898FA3FFD8E274C1AB0248FE82FE98" box="[159,278,336,362]" captionStart-0="FIGURES 1 – 6" captionStart-1="FIGURES 7 – 11" captionStart-2="FIGURES 16 – 17" captionStartId-0="2.[151,269,1938,1962]" captionStartId-1="3.[151,269,1698,1722]" captionStartId-2="5.[151,269,698,722]" captionTargetBox-0="[151,1414,538,1924]" captionTargetBox-1="[159,1427,197,1661]" captionTargetBox-2="[151,1435,194,674]" captionTargetId-0="figure@2.[151,1416,500,1926]" captionTargetId-1="figure@3.[151,1436,194,1674]" captionTargetId-2="figure@5.[151,1436,194,674]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="5" captionText-0="FIGURES 1 – 6. Mladenoniscus belavodae n. g., n. sp., paratype female (2.46 mm). 1, Dorsal view of the body. 2, Lateral view of the body. 3, Cephalon and pereon tergites 1 – 3, dorsal view. 4, Cephalon, lateral view. 5, Cephalon, frontal view. 6, Anterior part of the animal, frontal view." captionText-1="FIGURES 7 – 11. Mladenoniscus belavodae n. g., n. sp. 7 – 10, Mouth parts of paratype female. 7, Endopodite of maxilla 1. 8, Exopodite of maxilla 1. 9, Maxilla 2. 10, Maxilliped. 11, Holotype, pereopod 7. Scale bar = 0.2 mm." captionText-2="FIGURES 16 – 17. Mladenoniscus belavodae n. g., n. sp. 16, Details of the integument ornamentic. 17, Paratype male, propodus and dactylus of pereopod 7." httpUri-0="https://zenodo.org/record/180541/files/figure.png" httpUri-1="https://zenodo.org/record/180542/files/figure.png" httpUri-2="https://zenodo.org/record/180543/files/figure.png" pageId="1" pageNumber="61">Figs 1–17</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="DCA8C0ADFFD8E274C1A30282FF4AFD56" pageId="1" pageNumber="61" type="materials_examined">
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<paragraph id="940D9326FFD8E274C1A30282FF4AFD56" blockId="1.[151,1437,410,2036]" pageId="1" pageNumber="61">
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<emphasis id="A6C64F34FFD8E274C1A30282FD83FE46" bold="true" box="[151,535,410,436]" pageId="1" pageNumber="61">
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Material examined.
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<typeStatus id="4B092D84FFD8E274C0AA0282FD86FE46" box="[414,530,410,436]" pageId="1" pageNumber="61" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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male, Bela voda cave (White water), Demir Kapija gorge (
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<collectingCountry id="ECA5D3B6FFD8E274C4330282FA04FE46" box="[1287,1424,410,436]" name="Macedonia" pageId="1" pageNumber="61">Macedonia</collectingCountry>
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), (UTM–FL08) 23. 0 2. 2007., leg. I. Karaman and M. Horvatović (Inv. No 920; collection hosted by the Department of Biology and Ecology—Novi Sad,
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<collectingCountry id="ECA5D3B6FFD8E274C3DF02F2FCAFFDF6" box="[747,827,490,516]" name="Serbia" pageId="1" pageNumber="61">Serbia</collectingCountry>
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).
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<typeStatus id="4B092D84FFD8E274C26402F2FC59FDF6" box="[848,973,490,516]" pageId="1" pageNumber="61" type="paratype">
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<emphasis id="A6C64F34FFD8E274C26402F2FC59FDF6" bold="true" box="[848,973,490,516]" pageId="1" pageNumber="61">Paratypes</emphasis>
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</typeStatus>
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.
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<specimenCount id="82B458AFFFD8E274C2EA02F2FBA4FDF6" box="[990,1072,490,516]" pageId="1" pageNumber="61" type="male">1 male</specimenCount>
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<typeStatus id="4B092D84FFD8E274C50E02F2FB36FDF6" box="[1082,1186,490,516]" pageId="1" pageNumber="61" type="paratype">paratype</typeStatus>
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, ibid., 0 4. 0 4. 1998., leg. I. Karaman (Inv. No 1050),
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<specimenCount id="82B458AFFFD8E274C323010AFDF3FDDE" box="[535,615,530,556]" pageId="1" pageNumber="61" type="male">6 male</specimenCount>
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and
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<specimenCount id="82B458AFFFD8E274C397010AFC8CFDDE" box="[675,792,530,556]" pageId="1" pageNumber="61" type="female">14 female</specimenCount>
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<typeStatus id="4B092D84FFD8E274C214010AFC04FDDE" box="[800,912,530,556]" pageId="1" pageNumber="61" type="paratype">paratypes</typeStatus>
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and one juvenile; ibid., 23. 0 2. 2007. leg. I. Karaman and M. Horvatović (Inv. No 920; collection hosted by the Department of Biology and Ecology— Novi Sad,
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<collectingCountry id="ECA5D3B6FFD8E274C020017AFEF7FD8E" box="[276,355,610,636]" name="Serbia" pageId="1" pageNumber="61">Serbia</collectingCountry>
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;
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<specimenCount id="82B458AFFFD8E274C05A017AFE2AFD8E" box="[366,446,610,636]" pageId="1" pageNumber="61" type="male">1 male</specimenCount>
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and
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<specimenCount id="82B458AFFFD8E274C0C3017AFDC9FD8E" box="[503,605,610,636]" pageId="1" pageNumber="61" type="female">1 female</specimenCount>
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<typeStatus id="4B092D84FFD8E274C350017AFD40FD8E" box="[612,724,610,636]" pageId="1" pageNumber="61" type="paratype">paratypes</typeStatus>
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in Museo Zoologico "La Specola" dell'Universita, Florence,
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<collectingCountry id="ECA5D3B6FFD8E274C1A30192FF46FD56" box="[151,210,650,676]" name="Italy" pageId="1" pageNumber="61">Italy</collectingCountry>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="DCA8C0ADFFD8E274C1F201AAFBF4FCEE" pageId="1" pageNumber="61" type="diagnosis">
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<paragraph id="940D9326FFD8E274C1F201AAFBF4FCEE" blockId="1.[151,1437,410,2036]" pageId="1" pageNumber="61">
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<emphasis id="A6C64F34FFD8E274C1F201AAFED2FD3E" bold="true" box="[198,326,690,716]" pageId="1" pageNumber="61">Diagnosis.</emphasis>
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A blind, small species (up to
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<quantity id="534A3EC3FFD8E274C39101AAFC9AFD3E" box="[677,782,690,716]" metricMagnitude="-3" metricUnit="m" metricValue="2.69" pageId="1" pageNumber="61" unit="mm" value="2.69">2.69 mm</quantity>
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long), with slender body (four time as long as maximal width). Cephalon with three rows of tubercles; Pereon tergites with a row of 8 tubercles; 3. pleonal tergite with four tubercles; exopodite of male pleopod 1 oval, slightly concave terminally.
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</paragraph>
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</subSubSection>
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<subSubSection id="DCA8C0ADFFD8E271C1F20032FC78F83A" lastPageId="4" lastPageNumber="64" pageId="1" pageNumber="61" type="description">
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<paragraph id="940D9326FFD8E274C1F20032FCA1FC4E" blockId="1.[151,1437,410,2036]" pageId="1" pageNumber="61">
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<emphasis id="A6C64F34FFD8E274C1F20032FDA8FCB6" bold="true" box="[198,572,810,836]" pageId="1" pageNumber="61">
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Description of male
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<typeStatus id="4B092D84FFD8E274C0FD0032FDA3FCB6" box="[457,567,810,836]" pageId="1" pageNumber="61" type="holotype">holotype</typeStatus>
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.
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</emphasis>
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Total body length
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<quantity id="534A3EC3FFD8E274C2170032FC06FCB6" box="[803,914,810,836]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="1" pageNumber="61" unit="mm" value="2.3">2.30 mm</quantity>
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.
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<emphasis id="A6C64F34FFD8E274C2AB0032FB85FCB1" box="[927,1041,810,835]" italics="true" pageId="1" pageNumber="61">Cephalon</emphasis>
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(as on
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<figureCitation id="0C898FA3FFD8E274C55A0032FB43FCB6" box="[1134,1239,810,836]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,269,1938,1962]" captionTargetBox="[151,1414,538,1924]" captionTargetId="figure@2.[151,1416,500,1926]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Mladenoniscus belavodae n. g., n. sp., paratype female (2.46 mm). 1, Dorsal view of the body. 2, Lateral view of the body. 3, Cephalon and pereon tergites 1 – 3, dorsal view. 4, Cephalon, lateral view. 5, Cephalon, frontal view. 6, Anterior part of the animal, frontal view." httpUri="https://zenodo.org/record/180541/files/figure.png" pageId="1" pageNumber="61">Figs 3–5</figureCitation>
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) short and wide with three rows of tubercles; a row of six very small tubercles frontally, a row of six simple tubercles above them and a row of six big tubercles on the hind margin of the cephalon. Lateral pairs of the third row are doubled. Lateral lobes are short and wide, ventrally oriented.
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</paragraph>
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<paragraph id="940D9326FFD8E274C1F200D2FB0AFBFE" blockId="1.[151,1437,410,2036]" pageId="1" pageNumber="61">
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<emphasis id="A6C64F34FFD8E274C1F200D2FEEEFC11" box="[198,378,970,995]" italics="true" pageId="1" pageNumber="61">Pereon tergites</emphasis>
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(as on
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<figureCitation id="0C898FA3FFD8E274C0E300D2FDCBFC16" box="[471,607,970,996]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,269,1938,1962]" captionTargetBox="[151,1414,538,1924]" captionTargetId="figure@2.[151,1416,500,1926]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Mladenoniscus belavodae n. g., n. sp., paratype female (2.46 mm). 1, Dorsal view of the body. 2, Lateral view of the body. 3, Cephalon and pereon tergites 1 – 3, dorsal view. 4, Cephalon, lateral view. 5, Cephalon, frontal view. 6, Anterior part of the animal, frontal view." httpUri="https://zenodo.org/record/180541/files/figure.png" pageId="1" pageNumber="61">Figs 1–3, 6</figureCitation>
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) convex, netlike ornamented (
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<figureCitation id="0C898FA3FFD8E274C2FB00D2FBB2FC16" box="[975,1062,970,996]" captionStart="FIGURES 16 – 17" captionStartId="5.[151,269,698,722]" captionTargetBox="[151,1435,194,674]" captionTargetId="figure@5.[151,1436,194,674]" captionTargetPageId="5" captionText="FIGURES 16 – 17. Mladenoniscus belavodae n. g., n. sp. 16, Details of the integument ornamentic. 17, Paratype male, propodus and dactylus of pereopod 7." httpUri="https://zenodo.org/record/180543/files/figure.png" pageId="1" pageNumber="61">Fig. 16</figureCitation>
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) with a row of eight tubercles. Medial pair of tubercles is single, sub lateral pair is tripled and lateral pairs are doubled.
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</paragraph>
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<paragraph id="940D9326FFD8E274C1F20702FF61FB76" blockId="1.[151,1437,410,2036]" pageId="1" pageNumber="61">
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<emphasis id="A6C64F34FFD8E274C1F20702FE9FFBC1" box="[198,267,1050,1075]" italics="true" pageId="1" pageNumber="61">Pleon</emphasis>
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(as on
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<figureCitation id="0C898FA3FFD8E274C0510702FE59FBC6" box="[357,461,1050,1076]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,269,1938,1962]" captionTargetBox="[151,1414,538,1924]" captionTargetId="figure@2.[151,1416,500,1926]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Mladenoniscus belavodae n. g., n. sp., paratype female (2.46 mm). 1, Dorsal view of the body. 2, Lateral view of the body. 3, Cephalon and pereon tergites 1 – 3, dorsal view. 4, Cephalon, lateral view. 5, Cephalon, frontal view. 6, Anterior part of the animal, frontal view." httpUri="https://zenodo.org/record/180541/files/figure.png" pageId="1" pageNumber="61">Figs 1–2</figureCitation>
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) convex; pleonal segments 3–5 with developed epimera. Pleonal segment 3. with four tubercles, medial pair more pronounced. Pleon 3. slightly netlike ornamented, other pleonal segments smooth.
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</paragraph>
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<paragraph id="940D9326FFD8E274C1F2078BFC54FB5E" blockId="1.[151,1437,410,2036]" box="[198,960,1170,1196]" pageId="1" pageNumber="61">
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<emphasis id="A6C64F34FFD8E274C1F2078BFED6FB5E" box="[198,322,1171,1196]" italics="true" pageId="1" pageNumber="61">Pleotelson</emphasis>
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(as on
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<figureCitation id="0C898FA3FFD8E274C0A3078AFE4EFB5E" box="[407,474,1170,1196]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,269,1938,1962]" captionTargetBox="[151,1414,538,1924]" captionTargetId="figure@2.[151,1416,500,1926]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Mladenoniscus belavodae n. g., n. sp., paratype female (2.46 mm). 1, Dorsal view of the body. 2, Lateral view of the body. 3, Cephalon and pereon tergites 1 – 3, dorsal view. 4, Cephalon, lateral view. 5, Cephalon, frontal view. 6, Anterior part of the animal, frontal view." httpUri="https://zenodo.org/record/180541/files/figure.png" pageId="1" pageNumber="61">Fig. 1</figureCitation>
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) smooth with widely rounded distal part.
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</paragraph>
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<paragraph id="940D9326FFD8E274C1F207A3FD6EFB0E" blockId="1.[151,1437,410,2036]" pageId="1" pageNumber="61">
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<taxonomicName id="53B2E8A5FFD8E274C1F207A3FEABFB26" box="[198,319,1211,1236]" class="Dothideomycetes" family="Euantennariaceae" genus="Antennula" higherTaxonomySource="GBIF" kingdom="Fungi" order="Capnodiales" pageId="1" pageNumber="61" phylum="Ascomycota" rank="genus">
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<emphasis id="A6C64F34FFD8E274C1F207A3FEABFB26" box="[198,319,1211,1236]" italics="true" pageId="1" pageNumber="61">Antennula</emphasis>
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</taxonomicName>
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(Fig. 12) bearing 3 aesthetascs and a spine like protrusion on terminal article latero-distally; terminal article elongated, 4 times longer than wide.
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</paragraph>
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<paragraph id="940D9326FFD8E274C1F20613FC73FAD6" blockId="1.[151,1437,410,2036]" box="[198,999,1290,1316]" pageId="1" pageNumber="61">
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<emphasis id="A6C64F34FFD8E274C1F20613FEBCFAD6" box="[198,296,1291,1316]" italics="true" pageId="1" pageNumber="61">Antenna</emphasis>
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(Fig. 13) with stout articles; flagellum longer than article 5.
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</paragraph>
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<paragraph id="940D9326FFD8E274C1F2062BFF4CFA6E" blockId="1.[151,1437,410,2036]" pageId="1" pageNumber="61">
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<emphasis id="A6C64F34FFD8E274C1F2062BFEEDFABE" box="[198,377,1331,1356]" italics="true" pageId="1" pageNumber="61">Right mandible</emphasis>
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with one penicil; pars incisiva with three teeth; lacinia mobilis finely thorny in apical widened part; pars molaris with a short reduced penicil anteriorly on medial margin, grinding surfaces with 6 furrows.
|
||
</paragraph>
|
||
<paragraph id="940D9326FFD8E274C1F206B3FC32FA1E" blockId="1.[151,1437,410,2036]" pageId="1" pageNumber="61">
|
||
<emphasis id="A6C64F34FFD8E274C1F206B3FEFDFA36" box="[198,361,1451,1476]" italics="true" pageId="1" pageNumber="61">Left mandible</emphasis>
|
||
with three penicils; pars incisiva lateralis with four teeth; pars incisiva medialis with three teeth; pars molaris oval in shape, grinding surfaces with 5 furrows.
|
||
</paragraph>
|
||
<paragraph id="940D9326FFD8E274C1F206E3FE6CF996" blockId="1.[151,1437,410,2036]" pageId="1" pageNumber="61">
|
||
<emphasis id="A6C64F34FFD8E274C1F206E3FEADF9E6" box="[198,313,1531,1556]" italics="true" pageId="1" pageNumber="61">Maxilla 1</emphasis>
|
||
exopodite (as on
|
||
<figureCitation id="0C898FA3FFD8E274C32606E2FDCDF9E6" box="[530,601,1530,1556]" captionStart="FIGURES 7 – 11" captionStartId="3.[151,269,1698,1722]" captionTargetBox="[159,1427,197,1661]" captionTargetId="figure@3.[151,1436,194,1674]" captionTargetPageId="3" captionText="FIGURES 7 – 11. Mladenoniscus belavodae n. g., n. sp. 7 – 10, Mouth parts of paratype female. 7, Endopodite of maxilla 1. 8, Exopodite of maxilla 1. 9, Maxilla 2. 10, Maxilliped. 11, Holotype, pereopod 7. Scale bar = 0.2 mm." httpUri="https://zenodo.org/record/180542/files/figure.png" pageId="1" pageNumber="61">Fig. 8</figureCitation>
|
||
) with 3 strong outer teeth, 7 inner tooth, two of them very short; one elongated pectinate seta terminally; endopodite (as on
|
||
<figureCitation id="0C898FA3FFD8E274C219053AFCE1F9CE" box="[813,885,1570,1596]" captionStart="FIGURES 7 – 11" captionStartId="3.[151,269,1698,1722]" captionTargetBox="[159,1427,197,1661]" captionTargetId="figure@3.[151,1436,194,1674]" captionTargetPageId="3" captionText="FIGURES 7 – 11. Mladenoniscus belavodae n. g., n. sp. 7 – 10, Mouth parts of paratype female. 7, Endopodite of maxilla 1. 8, Exopodite of maxilla 1. 9, Maxilla 2. 10, Maxilliped. 11, Holotype, pereopod 7. Scale bar = 0.2 mm." httpUri="https://zenodo.org/record/180542/files/figure.png" pageId="1" pageNumber="61">Fig. 7</figureCitation>
|
||
) with three penicils, terminal one very short, inner one stout and elongated.
|
||
</paragraph>
|
||
<paragraph id="940D9326FFD8E274C1F2056BFF4DF92E" blockId="1.[151,1437,410,2036]" pageId="1" pageNumber="61">
|
||
<emphasis id="A6C64F34FFD8E274C1F2056BFEACF97E" box="[198,312,1651,1676]" italics="true" pageId="1" pageNumber="61">Maxilla 2</emphasis>
|
||
(as on
|
||
<figureCitation id="0C898FA3FFD8E274C0A4056AFE42F97E" box="[400,470,1650,1676]" captionStart="FIGURES 7 – 11" captionStartId="3.[151,269,1698,1722]" captionTargetBox="[159,1427,197,1661]" captionTargetId="figure@3.[151,1436,194,1674]" captionTargetPageId="3" captionText="FIGURES 7 – 11. Mladenoniscus belavodae n. g., n. sp. 7 – 10, Mouth parts of paratype female. 7, Endopodite of maxilla 1. 8, Exopodite of maxilla 1. 9, Maxilla 2. 10, Maxilliped. 11, Holotype, pereopod 7. Scale bar = 0.2 mm." httpUri="https://zenodo.org/record/180542/files/figure.png" pageId="1" pageNumber="61">Fig. 9</figureCitation>
|
||
) outer lobe with 2 strong rodlike setae on its tip; inner lobe with a row of rodlike setae on its tip and medial edge; inner side of both lobes with brush like surface of dense, very tiny elongated setae.
|
||
</paragraph>
|
||
<paragraph id="940D9326FFD8E274C1F205F3FD74F856" blockId="1.[151,1437,410,2036]" pageId="1" pageNumber="61">
|
||
<emphasis id="A6C64F34FFD8E274C1F205F3FED1F8F6" box="[198,325,1771,1796]" italics="true" pageId="1" pageNumber="61">Maxilliped</emphasis>
|
||
(as on
|
||
<figureCitation id="0C898FA3FFD8E274C0AE05F2FE79F8F6" box="[410,493,1770,1796]" captionStart="FIGURES 7 – 11" captionStartId="3.[151,269,1698,1722]" captionTargetBox="[159,1427,197,1661]" captionTargetId="figure@3.[151,1436,194,1674]" captionTargetPageId="3" captionText="FIGURES 7 – 11. Mladenoniscus belavodae n. g., n. sp. 7 – 10, Mouth parts of paratype female. 7, Endopodite of maxilla 1. 8, Exopodite of maxilla 1. 9, Maxilla 2. 10, Maxilliped. 11, Holotype, pereopod 7. Scale bar = 0.2 mm." httpUri="https://zenodo.org/record/180542/files/figure.png" pageId="1" pageNumber="61">Fig. 10</figureCitation>
|
||
) endopodite with widened latero-distal margin, terminally protruding; margin of protrusion with rows of remarquable elongated tiny setae; palp terminally and on medial edge bearing strong setae; outer edge with two low protrusions, each with 1 strong and 1 simple setae; endite distally truncate, with a stout penicil near medial margin and a spine like protrusion of latero-distal margin, a large seta subapically and a short spine apically near inner margin.
|
||
</paragraph>
|
||
<paragraph id="940D9326FFD8E274C1F204ABFCC5F806" blockId="1.[151,1437,410,2036]" pageId="1" pageNumber="61">
|
||
<emphasis id="A6C64F34FFD8E274C1F204ABFEE3F83E" box="[198,375,1971,1996]" italics="true" pageId="1" pageNumber="61">Pereopods 1–7</emphasis>
|
||
similar in shape, with stout articles; carpus and merus as long as wide. pereopod 7 (
|
||
<figureCitation id="0C898FA3FFD8E274C45C04AAFF21F806" captionStart="FIGURES 7 – 11" captionStartId="3.[151,269,1698,1722]" captionTargetBox="[159,1427,197,1661]" captionTargetId="figure@3.[151,1436,194,1674]" captionTargetPageId="3" captionText="FIGURES 7 – 11. Mladenoniscus belavodae n. g., n. sp. 7 – 10, Mouth parts of paratype female. 7, Endopodite of maxilla 1. 8, Exopodite of maxilla 1. 9, Maxilla 2. 10, Maxilliped. 11, Holotype, pereopod 7. Scale bar = 0.2 mm." httpUri="https://zenodo.org/record/180542/files/figure.png" pageId="1" pageNumber="61">Figs 11</figureCitation>
|
||
,
|
||
<figureCitation id="0C898FA3FFD8E274C1F604C2FF74F806" box="[194,224,2010,2036]" captionStart="FIGURES 16 – 17" captionStartId="5.[151,269,698,722]" captionTargetBox="[151,1435,194,674]" captionTargetId="figure@5.[151,1436,194,674]" captionTargetPageId="5" captionText="FIGURES 16 – 17. Mladenoniscus belavodae n. g., n. sp. 16, Details of the integument ornamentic. 17, Paratype male, propodus and dactylus of pereopod 7." httpUri="https://zenodo.org/record/180543/files/figure.png" pageId="1" pageNumber="61">17</figureCitation>
|
||
) without noticeable secondary sexual characteristics.
|
||
</paragraph>
|
||
<paragraph id="940D9326FFDBE277C1F20380FD0EFF40" blockId="2.[151,1436,152,458]" box="[198,666,152,178]" pageId="2" pageNumber="62">
|
||
<emphasis id="A6C64F34FFDBE277C1F20380FEECFF43" box="[198,376,152,177]" italics="true" pageId="2" pageNumber="62">Genital papilla</emphasis>
|
||
(Fig. 14) simple, setose.
|
||
</paragraph>
|
||
<paragraph id="940D9326FFDBE277C1F203D8FA9DFEF0" blockId="2.[151,1436,152,458]" pageId="2" pageNumber="62">
|
||
<emphasis id="A6C64F34FFDBE277C1F203D8FEABFF2B" box="[198,319,192,217]" italics="true" pageId="2" pageNumber="62">Pleopod 1</emphasis>
|
||
(Fig. 14) endopodite biarticulated, terminal article setose, longer than basal. Basal article with longitudinal carina medio-ventrally; exopodite oval, as long as wide, slightly concave terminally.
|
||
</paragraph>
|
||
<paragraph id="940D9326FFDBE277C1F20208FEA2FE88" blockId="2.[151,1436,152,458]" pageId="2" pageNumber="62">
|
||
<emphasis id="A6C64F34FFDBE277C1F20208FEABFEDB" box="[198,319,272,297]" italics="true" pageId="2" pageNumber="62">Pleopod 2</emphasis>
|
||
(Fig. 15). Endopodite stout, triarticulated; distal article spiniform, medial article twice as long as basal article. Trapezoidal exopodite wider than long; distal margin almost straight with a spine like setae paramedially.
|
||
</paragraph>
|
||
<paragraph id="940D9326FFDBE277C1F20290FC5BFE38" blockId="2.[151,1436,152,458]" pageId="2" pageNumber="62">
|
||
<emphasis id="A6C64F34FFDBE277C1F20290FEB4FE53" box="[198,288,392,417]" italics="true" pageId="2" pageNumber="62">Uropod</emphasis>
|
||
basis dilated posteriorly; exopodite and endopodite conical, articulated almost at the same level and with a tuft of setae at the apex; endopodite shorter than exopodite.
|
||
</paragraph>
|
||
<caption id="C0CDC3AEFFDBE277C1A3048AFD37F81C" httpUri="https://zenodo.org/record/180541/files/figure.png" pageId="2" pageNumber="62" targetBox="[151,1414,538,1924]" targetPageId="2">
|
||
<paragraph id="940D9326FFDBE277C1A3048AFD37F81C" blockId="2.[151,1436,1938,2030]" pageId="2" pageNumber="62">
|
||
<emphasis id="A6C64F34FFDBE277C1A3048AFED7F858" bold="true" box="[151,323,1938,1962]" pageId="2" pageNumber="62">FIGURES 1–6.</emphasis>
|
||
<taxonomicName id="53B2E8A5FFDBE277C07F048AFDF6F85B" box="[331,610,1938,1961]" class="Malacostraca" family="Trichoniscidae" genus="Mladenoniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="62" phylum="Arthropoda" rank="species" species="belavodae">
|
||
<emphasis id="A6C64F34FFDBE277C07F048AFDF6F85B" box="[331,610,1938,1961]" italics="true" pageId="2" pageNumber="62">Mladenoniscus belavodae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A6C64F34FFDBE277C35D048AFD71F858" bold="true" box="[617,741,1938,1962]" pageId="2" pageNumber="62">n. g., n. sp.</emphasis>
|
||
, paratype female (2.46 mm). 1, Dorsal view of the body. 2, Lateral view of the body. 3, Cephalon and pereon tergites 1–3, dorsal view. 4, Cephalon, lateral view. 5, Cephalon, frontal view. 6, Anterior part of the animal, frontal view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="C0CDC3AEFFDAE276C1A305BAFB6FF929" httpUri="https://zenodo.org/record/180542/files/figure.png" pageId="3" pageNumber="63" targetBox="[159,1427,197,1661]" targetPageId="3">
|
||
<paragraph id="940D9326FFDAE276C1A305BAFB6FF929" blockId="3.[151,1436,1698,1755]" pageId="3" pageNumber="63">
|
||
<emphasis id="A6C64F34FFDAE276C1A305BAFEDBF948" bold="true" box="[151,335,1698,1722]" pageId="3" pageNumber="63">FIGURES 7–11.</emphasis>
|
||
<taxonomicName id="53B2E8A5FFDAE276C06105BAFDFFF94B" box="[341,619,1698,1721]" class="Malacostraca" family="Trichoniscidae" genus="Mladenoniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="63" phylum="Arthropoda" rank="species" species="belavodae">
|
||
<emphasis id="A6C64F34FFDAE276C06105BAFDFFF94B" box="[341,619,1698,1721]" italics="true" pageId="3" pageNumber="63">Mladenoniscus belavodae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A6C64F34FFDAE276C34605BAFD7FF948" bold="true" box="[626,747,1698,1722]" pageId="3" pageNumber="63">n. g., n. sp.</emphasis>
|
||
7–10, Mouth parts of paratype female. 7, Endopodite of maxilla 1. 8, Exopodite of maxilla 1. 9, Maxilla 2. 10, Maxilliped. 11, Holotype, pereopod 7. Scale bar = 0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="940D9326FFDAE276C1F20417FC4CF8DB" blockId="3.[151,1436,1807,2033]" box="[198,984,1807,1833]" pageId="3" pageNumber="63">Females differ from males only in the structure of pleopod 1 and 2.</paragraph>
|
||
<paragraph id="940D9326FFDAE271C1F2042FFCB3FF28" blockId="3.[151,1436,1807,2033]" lastBlockId="4.[151,1436,152,218]" lastPageId="4" lastPageNumber="64" pageId="3" pageNumber="63">
|
||
<emphasis id="A6C64F34FFDAE276C1F2042FFED5F8A3" bold="true" box="[198,321,1847,1873]" pageId="3" pageNumber="63">Variation.</emphasis>
|
||
Range of measurements in body length of males
|
||
<quantity id="534A3EC3FFDAE276C2B3042FFBA6F8A3" box="[903,1074,1847,1873]" metricMagnitude="-3" metricUnit="m" metricValue="2.07" metricValueMax="2.3" metricValueMin="1.84" pageId="3" pageNumber="63" unit="mm" value="2.07" valueMax="2.3" valueMin="1.84">1.84–2.30 mm</quantity>
|
||
and females
|
||
<quantity id="534A3EC3FFDAE276C5F9042FFAEEF8A3" box="[1229,1402,1847,1873]" metricMagnitude="-3" metricUnit="m" metricValue="2.21" metricValueMax="2.56" metricValueMin="1.86" pageId="3" pageNumber="63" unit="mm" value="2.21" valueMax="2.56" valueMin="1.86">1.86–2.56 mm</quantity>
|
||
.
|
||
<emphasis id="A6C64F34FFDAE276C1F20447FEABF88B" bold="true" box="[198,319,1887,1913]" pageId="3" pageNumber="63">Remarks.</emphasis>
|
||
In
|
||
<date id="E00CB5E6FFDAE276C0530447FD83F88B" box="[359,535,1887,1913]" pageId="3" pageNumber="63" value="2007-02">February 2007</date>
|
||
, a few specimens of
|
||
<taxonomicName id="53B2E8A5FFDAE276C2390478FC22F88B" box="[781,950,1888,1913]" class="Malacostraca" family="Trichoniscidae" genus="Alpioniscus" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="63" phylum="Arthropoda" rank="species" species="vardarensis">
|
||
<emphasis id="A6C64F34FFDAE276C2390478FC22F88B" box="[781,950,1888,1913]" italics="true" pageId="3" pageNumber="63">A. vardarensis</emphasis>
|
||
</taxonomicName>
|
||
, originally described from the Bela Voda Cave, were found in mesovoid shallow substratum (MSS) above a fountain in the Demir Kapija gorge, together with specimens of some
|
||
<taxonomicName id="53B2E8A5FFDAE276C31204A8FD7EF83B" box="[550,746,1968,1993]" class="Malacostraca" family="Trichoniscidae" genus="Haplophthalmus" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="63" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A6C64F34FFDAE276C31204A8FD7EF83B" box="[550,746,1968,1993]" italics="true" pageId="3" pageNumber="63">Haplophthalmus</emphasis>
|
||
</taxonomicName>
|
||
species. No specimens of
|
||
<taxonomicName id="53B2E8A5FFDAE276C51304A8FB5CF83B" box="[1063,1224,1968,1993]" class="Malacostraca" family="Trichoniscidae" genus="Mladenoniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="63" phylum="Arthropoda" rank="species" species="belavodae">
|
||
<emphasis id="A6C64F34FFDAE276C51304A8FB5CF83B" box="[1063,1224,1968,1993]" italics="true" pageId="3" pageNumber="63">M. belavodae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A6C64F34FFDAE276C5E504B7FACFF83B" bold="true" box="[1233,1371,1967,1993]" pageId="3" pageNumber="63">n. g., n. sp.</emphasis>
|
||
were found in the same location. We consider that
|
||
<taxonomicName id="53B2E8A5FFDAE276C38F04C0FCC8F803" box="[699,860,2008,2033]" class="Malacostraca" family="Trichoniscidae" genus="Mladenoniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="63" phylum="Arthropoda" rank="species" species="belavodae">
|
||
<emphasis id="A6C64F34FFDAE276C38F04C0FCC8F803" box="[699,860,2008,2033]" italics="true" pageId="3" pageNumber="63">M. belavodae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A6C64F34FFDAE276C25204CFFC67F803" bold="true" box="[870,1011,2007,2033]" pageId="3" pageNumber="63">n. g., n. sp.</emphasis>
|
||
could be regarded as eutroglobitic, since we found it only in the deeper parts of the cave with persistently increased humidity, while
|
||
<taxonomicName id="53B2E8A5FFDDE271C43C0380FF22FF2B" class="Malacostraca" family="Trichoniscidae" genus="Alpioniscus" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="64" phylum="Arthropoda" rank="species" species="vardarensis">
|
||
<emphasis id="A6C64F34FFDDE271C43C0380FF22FF2B" italics="true" pageId="4" pageNumber="64">A. vardarensis</emphasis>
|
||
</taxonomicName>
|
||
shows somewhat higher tolerance against this factor.
|
||
</paragraph>
|
||
<paragraph id="940D9326FFDDE271C1A30496FC78F83A" blockId="4.[151,1436,1934,1992]" pageId="4" pageNumber="64">
|
||
<emphasis id="A6C64F34FFDDE271C1A30496FECAF854" bold="true" box="[151,350,1934,1958]" pageId="4" pageNumber="64">FIGURES 12–15.</emphasis>
|
||
<taxonomicName id="53B2E8A5FFDDE271C0520497FDE9F854" box="[358,637,1935,1958]" class="Malacostraca" family="Trichoniscidae" genus="Mladenoniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="64" phylum="Arthropoda" rank="species" species="belavodae">
|
||
<emphasis id="A6C64F34FFDDE271C0520497FDE9F854" box="[358,637,1935,1958]" italics="true" pageId="4" pageNumber="64">Mladenoniscus belavodae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A6C64F34FFDDE271C3B10496FC97F854" bold="true" box="[645,771,1934,1958]" pageId="4" pageNumber="64">n. g., n. sp.</emphasis>
|
||
12–13,
|
||
<typeStatus id="4B092D84FFDDE271C2680496FC2DF857" box="[860,953,1934,1957]" pageId="4" pageNumber="64" type="paratype">Paratype</typeStatus>
|
||
female. 12,
|
||
<taxonomicName id="53B2E8A5FFDDE271C5760496FB20F857" box="[1090,1204,1934,1957]" class="Dothideomycetes" family="Euantennariaceae" genus="Antennula" higherTaxonomySource="GBIF" kingdom="Fungi" order="Capnodiales" pageId="4" pageNumber="61" phylum="Ascomycota" rank="genus">Antennula</taxonomicName>
|
||
; 13. Antenna, 14–15.
|
||
<typeStatus id="4B092D84FFDDE271C1A304A9FF6AF83A" box="[151,254,1969,1992]" pageId="4" pageNumber="64" type="holotype">Holotype</typeStatus>
|
||
. 14, Pleopod 1 and genital papilla. 15, Pleopod 2. Scale bar =
|
||
<quantity id="534A3EC3FFDDE271C2A704A9FC7CF83A" box="[915,1000,1969,1992]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="64" unit="mm" value="0.1">0.1 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |