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<document id="039CB656259CE48805C2CE6C16D09729" ID-DOI="10.5281/zenodo.188493" ID-GBIF-Dataset="c0ed2eaa-3e81-4a19-8a61-0cdfead50fd4" ID-ISSN="1175-5326" ID-Zenodo-Dep="188493" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461094063333" checkinUser="plazi" docAuthor="Zamanpoore, Mehrdad, Poeckl, Manfred, Grabowski, Michal &amp; Schiemer, Friedrich" docDate="2009" docId="1C19C76EFFBD8A19FF3C23999553FC44" docLanguage="en" docName="zt02136p039.pdf" docOrigin="Zootaxa 2136" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Gammarus zagrosensis Zamanpoore, Poeckl, Grabowski &amp; Schiemer, 2009, n. sp." docType="treatment" docUuid="F5D7A8DC-2EE4-47C1-994D-278B84C054E6" docUuidSource="ZooBank" docVersion="9" lastPageNumber="31" masterDocId="E020BF16FFBF8A13FFAB2053976DFFD7" masterDocTitle="Two new sympatric species of freshwater Gammarus (Crustacea: Amphipoda) from Southern Zagros Region, Iran" masterLastPageNumber="39" masterPageNumber="21" pageNumber="23" updateTime="1698588735859" updateUser="plazi">
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<mods:title id="123E9A0337D9504560C64D5C3994BE69">Two new sympatric species of freshwater Gammarus (Crustacea: Amphipoda) from Southern Zagros Region, Iran</mods:title>
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<treatment id="1C19C76EFFBD8A19FF3C23999553FC44" ID-DOI="http://doi.org/10.5281/zenodo.3503236" ID-GBIF-Taxon="119619966" ID-Zenodo-Dep="3503236" LSID="urn:lsid:zoobank.org:act:F5D7A8DC-2EE4-47C1-994D-278B84C054E6" httpUri="http://treatment.plazi.org/id/1C19C76EFFBD8A19FF3C23999553FC44" lastPageId="10" lastPageNumber="31" pageId="2" pageNumber="23">
<subSubSection id="DCAA25F3FFBD8A11FF3C23999675FBD0" pageId="2" pageNumber="23" type="nomenclature">
<paragraph id="940F7678FFBD8A11FF3C23999694FC33" blockId="2.[151,505,970,1031]" box="[151,505,970,996]" pageId="2" pageNumber="23">
<heading id="CF47C114FFBD8A11FF3C23999694FC33" bold="true" box="[151,505,970,996]" fontSize="11" level="1" pageId="2" pageNumber="23" reason="1">
<emphasis id="A6C4AA6AFFBD8A11FF3C23999694FC33" bold="true" box="[151,505,970,996]" pageId="2" pageNumber="23">
<taxonomicName id="53B00DFBFFBD8A11FF3C239996C2FC33" ID-CoL="3F8N7" box="[151,431,970,996]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="23" phylum="Arthropoda" rank="species" species="zagrosensis" status="sp. nov.">
<emphasis id="A6C4AA6AFFBD8A11FF3C239996C2FC33" bold="true" box="[151,431,970,996]" italics="true" pageId="2" pageNumber="23">Gammarus zagrosensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BDF71711FFBD8A11FE1C23999694FC33" box="[439,505,970,996]" pageId="2" pageNumber="23" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="940F7678FFBD8A11FF3C23BE9675FBD0" blockId="2.[151,505,970,1031]" box="[151,280,1005,1031]" pageId="2" pageNumber="23">
(
<figureCitation id="0C8B6AFDFFBD8A11FF3423BE9662FBD0" box="[159,271,1005,1031]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStartId-0="3.[151,254,1942,1966]" captionStartId-1="5.[151,254,2006,2030]" captionStartId-2="7.[151,257,1972,1996]" captionStartId-3="8.[151,254,1109,1133]" captionTargetBox-0="[214,1370,193,1913]" captionTargetBox-1="[191,1404,191,1980]" captionTargetBox-2="[212,1368,193,1938]" captionTargetBox-3="[172,1410,191,1075]" captionTargetId-0="figure@3.[194,1399,165,1937]" captionTargetId-1="figure@5.[185,1409,182,1982]" captionTargetId-2="figure@7.[192,1397,185,1957]" captionTargetId-3="figure@8.[151,1436,178,1085]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-2="7" captionTargetPageId-3="8" captionText-0="FIGURE 2. Gammarus zagrosensis n. sp., 3, 17.4 mm. A (HC), B (A 1), and C (A 2) on scale i; D (Mn) and E (MnPl) on scale ii, F (exopodite of Maxilipede), G (left palp of maxilla 1), and H (right palp of maxilla 1) on scale iii, and I (T) on scale iv." captionText-1="FIGURE 3. Gammarus zagrosensis n. sp., 3, 17.4 mm. A (G 1) and B (G 2) on scale i, C (P 3) and D (P 4) on scale ii." captionText-2="FIGURE 4. Gammarus zagrosensis n. sp., 3, 17.4 mm. A (P 5), B (P 6), and C (P 7), Ci and Cii (bases of P 7 in two paratypes) on scale i; D (US) and E (EP) on scale ii; F (U 3) on scale iii." captionText-3="FIGURE 5. Gammarus zagrosensis n. sp., Ƥ, 12 mm. A (basis of P 5), B (basis of P 6), C (basis of P 7) and E (oostegite 1) on scale i; D (EP) on scale ii." httpUri-0="https://zenodo.org/record/188495/files/figure.png" httpUri-1="https://zenodo.org/record/188496/files/figure.png" httpUri-2="https://zenodo.org/record/188497/files/figure.png" httpUri-3="https://zenodo.org/record/188498/files/figure.png" pageId="2" pageNumber="23">Figs. 25</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="DCAA25F3FFBD8A11FF3C2469961DFA29" pageId="2" pageNumber="23" type="materials_examined">
<paragraph id="940F7678FFBD8A11FF3C2469961DFA29" blockId="2.[151,1437,1082,2036]" pageId="2" pageNumber="23">
<emphasis id="A6C4AA6AFFBD8A11FF3C246996FEFB83" bold="true" box="[151,403,1082,1108]" pageId="2" pageNumber="23">Material examined.</emphasis>
Many males and females. Male
<typeStatus id="4B0BC8DAFFBD8A11FC8A246994E3FB83" box="[801,910,1082,1108]" pageId="2" pageNumber="23" type="holotype">holotype</typeStatus>
, body length
<quantity id="5348DB9DFFBD8A11FB98246993CEFB83" box="[1075,1187,1082,1108]" metricMagnitude="-2" metricUnit="m" metricValue="1.7399999999999998" pageId="2" pageNumber="23" unit="mm" value="17.4">17.4 mm</quantity>
,
<date id="E00E50B8FFBD8A11FB0424699271FB83" box="[1199,1308,1082,1108]" pageId="2" pageNumber="23" value="2007-02">February</date>
&amp;
<date id="E00E50B8FFBD8A11FAEE246997B4FBAC" pageId="2" pageNumber="23" value="2007-08">August 2007</date>
, coll. M. Zamanpoore.
<emphasis id="A6C4AA6AFFBD8A11FDA9243295C2FBAD" box="[514,687,1121,1146]" italics="true" pageId="2" pageNumber="23">Locus typicus</emphasis>
: Vezge-Morad spring (locality 2),
<quantity id="5348DB9DFFBD8A11FBC1243293C7FBAC" box="[1130,1194,1121,1147]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="23" unit="km" value="2.0">2 km</quantity>
S of Sepidan,
<collectingCountry id="ECA736E8FFBD8A11FACC243292F0FBAC" box="[1383,1437,1121,1147]" name="Iran" pageId="2" pageNumber="23">Fars</collectingCountry>
Province,
<collectingCountry id="ECA736E8FFBD8A11FEB824DB9629FB75" box="[275,324,1160,1186]" name="Iran" pageId="2" pageNumber="23">Iran</collectingCountry>
(
<geoCoordinate id="F18410BFFFBD8A11FEFD24DB96AFFB75" box="[342,450,1160,1186]" direction="north" orientation="latitude" pageId="2" pageNumber="23" precision="925" value="30.216667">30°13´N</geoCoordinate>
,
<geoCoordinate id="F18410BFFFBD8A11FE7B24DB9555FB75" box="[464,568,1160,1186]" direction="east" orientation="longitude" pageId="2" pageNumber="23" precision="925" value="51.966667">51°58´E</geoCoordinate>
), Altitude
<quantity id="5348DB9DFFBD8A11FD1724DB9471FB75" box="[700,796,1160,1186]" metricMagnitude="3" metricUnit="m" metricValue="2.19" pageId="2" pageNumber="23" unit="m" value="2190.0">2190 m</quantity>
.
<typeStatus id="4B0BC8DAFFBD8A11FC8224DB94CEFB75" box="[809,931,1160,1186]" pageId="2" pageNumber="23" type="paratype">Paratypes</typeStatus>
, many males and females; same date and locality. Additional samples: Khani Varg spring (locality 3),
<quantity id="5348DB9DFFBD8A11FCCA24FD94F1FB1F" box="[865,924,1198,1224]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="23" unit="km" value="4.0">4 km</quantity>
S of Sepidan (
<geoCoordinate id="F18410BFFFBD8A11FBE724FD93D9FB1F" box="[1100,1204,1198,1224]" direction="north" orientation="latitude" pageId="2" pageNumber="23" precision="925" value="30.2">30°12´N</geoCoordinate>
,
<geoCoordinate id="F18410BFFFBD8A11FB6A24FD924BFB1F" box="[1217,1318,1198,1224]" direction="east" orientation="longitude" pageId="2" pageNumber="23" precision="925" value="51.983334">51°59´E</geoCoordinate>
), Altitude
<quantity id="5348DB9DFFBD8A11FF3C2486979EFB38" box="[151,243,1237,1263]" metricMagnitude="3" metricUnit="m" metricValue="2.053" pageId="2" pageNumber="23" unit="m" value="2053.0">2053 m</quantity>
; Baraghan spring (locality 4),
<quantity id="5348DB9DFFBD8A11FDF0248695FAFB38" box="[603,663,1237,1263]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="23" unit="km" value="5.0">5 km</quantity>
SE of Sepidan (
<geoCoordinate id="F18410BFFFBD8A11FCFC248694ADFB38" box="[855,960,1237,1263]" direction="north" orientation="latitude" pageId="2" pageNumber="23" precision="925" value="30.25">30°15´N</geoCoordinate>
,
<geoCoordinate id="F18410BFFFBD8A11FC672486935CFB38" box="[972,1073,1237,1263]" direction="east" orientation="longitude" pageId="2" pageNumber="23" precision="925" value="51.983334">51°59´E</geoCoordinate>
), Altitude
<quantity id="5348DB9DFFBD8A11FB0524869266FB38" box="[1198,1291,1237,1263]" metricMagnitude="3" metricUnit="m" metricValue="2.2119999999999997" pageId="2" pageNumber="23" unit="m" value="2212.0">2212 m</quantity>
; Shesh-Peer (spring brook) (locality 5),
<quantity id="5348DB9DFFBD8A11FE5B24AF9552FAC1" box="[496,575,1276,1302]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="23" unit="km" value="10.0">10 km</quantity>
SE of Sepidan (
<geoCoordinate id="F18410BFFFBD8A11FCA624AF9411FAC1" box="[781,892,1276,1302]" direction="north" orientation="latitude" pageId="2" pageNumber="23" precision="925" value="30.25">30°15´N</geoCoordinate>
,
<geoCoordinate id="F18410BFFFBD8A11FC2124AF9498FAC1" box="[906,1013,1276,1302]" direction="east" orientation="longitude" pageId="2" pageNumber="23" precision="925" value="52.05">52°03´E</geoCoordinate>
), Altitude
<quantity id="5348DB9DFFBD8A11FBD724AF93B3FAC1" box="[1148,1246,1276,1302]" metricMagnitude="3" metricUnit="m" metricValue="2.35" pageId="2" pageNumber="23" unit="m" value="2350.0">2350 m</quantity>
. All specimens including
<typeStatus id="4B0BC8DAFFBD8A11FEB325719622FAEB" box="[280,335,1314,1340]" pageId="2" pageNumber="23">type</typeStatus>
series are kept in the amphipod collection of the Museum of
<collectingCountry id="ECA736E8FFBD8A11FBC3257193F2FAEB" box="[1128,1183,1314,1340]" name="Iran" pageId="2" pageNumber="23">Fars</collectingCountry>
Research Centre of Agriculture and Natural Resources (FARSAGRES), Shiraz,
<collectingCountry id="ECA736E8FFBD8A11FCDD251A94C6FAB4" box="[886,939,1353,1379]" name="Iran" pageId="2" pageNumber="23">Iran</collectingCountry>
.
<typeStatus id="4B0BC8DAFFBD8A11FC1C251A9345FAB4" box="[951,1064,1353,1379]" pageId="2" pageNumber="23" type="holotype">Holotype</typeStatus>
is deposited in the Zoological Museum of Amsterdam (
<collectionCode id="F2A1EEBDFFBD8A11FE7F25239573FA5D" box="[468,542,1392,1418]" httpUri="http://grbio.org/cool/a65z-wv6r" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="2" pageNumber="23">ZMA</collectionCode>
, Amph. 206052, 2009). Descriptions were based on the
<typeStatus id="4B0BC8DAFFBD8A11FB442523920FFA5D" box="[1263,1378,1392,1418]" pageId="2" pageNumber="23" type="holotype">holotype</typeStatus>
, and compared to 5 winter and 5 summer
<typeStatus id="4B0BC8DAFFBD8A11FDD825C59583FA67" box="[627,750,1430,1456]" pageId="2" pageNumber="23" type="paratype">paratypes</typeStatus>
from the
<typeStatus id="4B0BC8DAFFBD8A11FCC425C594CBFA67" box="[879,934,1430,1456]" pageId="2" pageNumber="23">type</typeStatus>
locality for variations.
<typeStatus id="4B0BC8DAFFBD8A11FB7125C5923BFA67" box="[1242,1366,1430,1456]" pageId="2" pageNumber="23" type="paratype">Paratypes</typeStatus>
were completely dissected for staining with Lignin Pink and mounted with Euparal (Carl Roth GmbH &amp; Co) on microscope slides.
</paragraph>
</subSubSection>
<subSubSection id="DCAA25F3FFBD8A11FF6D265993DCF94F" pageId="2" pageNumber="23" type="diagnosis">
<paragraph id="940F7678FFBD8A11FF6D265993DCF94F" blockId="2.[151,1437,1082,2036]" pageId="2" pageNumber="23">
<emphasis id="A6C4AA6AFFBD8A11FF6D26599652F9F3" bold="true" box="[198,319,1546,1572]" pageId="2" pageNumber="23">Diagnosis</emphasis>
. Medium-sized to large species. In general, this species can be distinguished from other adjacent species by a combination of the following characters: (1) extra-ordinary long setae on inferior margin of EP2 and 3, (2) endopodite of U3 about 3/4th of exopodite, (3) existence of some setae on postero-distal corner of P6 and P7, and (4) antenna 2 with short setae on flagellum and bearing calceoli.
</paragraph>
</subSubSection>
<subSubSection id="DCAA25F3FFBD8A11FF6D26F6969EF8DB" pageId="2" pageNumber="23" type="etymology">
<paragraph id="940F7678FFBD8A11FF6D26F6969EF8DB" blockId="2.[151,1437,1082,2036]" pageId="2" pageNumber="23">
<emphasis id="A6C4AA6AFFBD8A11FF6D26F69626F968" bold="true" box="[198,331,1701,1727]" pageId="2" pageNumber="23">Etymology</emphasis>
. The specific name
<taxonomicName id="53B00DFBFFBD8A11FD9526F695ABF969" box="[574,710,1701,1726]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="23" phylum="Arthropoda" rank="species" species="zagrosensis">
<emphasis id="A6C4AA6AFFBD8A11FD9526F695ABF969" box="[574,710,1701,1726]" italics="true" pageId="2" pageNumber="23">zagrosensis</emphasis>
</taxonomicName>
is an adjective made from the name of the Zagros Mountain Chains, located along the eastern edge of the Iranian Plateau, an area with a high biodiversity. The
<typeStatus id="4B0BC8DAFFBD8A11FAC3269F92F0F931" box="[1384,1437,1740,1766]" pageId="2" pageNumber="23">type</typeStatus>
locality is in its southern part.
</paragraph>
</subSubSection>
<subSubSection id="DCAA25F3FFBD8A15FF6D274B9327F806" lastPageId="6" lastPageNumber="27" pageId="2" pageNumber="23" type="description">
<paragraph id="940F7678FFBD8A11FF6D274B9213F857" blockId="2.[151,1437,1082,2036]" pageId="2" pageNumber="23">
<emphasis id="A6C4AA6AFFBD8A11FF6D274B96C7F8E4" bold="true" box="[198,426,1816,1843]" pageId="2" pageNumber="23">
Description.
<emphasis id="A6C4AA6AFFBD8A11FEC7274A96C7F8E4" bold="true" box="[364,426,1817,1843]" italics="true" pageId="2" pageNumber="23">Male</emphasis>
</emphasis>
. Maximum body length (based on 100 individuals)
<quantity id="5348DB9DFFBD8A11FB90274A93FAF8E4" box="[1083,1175,1817,1843]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="23" unit="mm" value="22.0">22 mm</quantity>
. Body robust; lateral cephalic lobes more or less rounded, anterior lower part of head extended forward; eyes elliptical to slightly reniform, short (less than diameter of first segment of antenna 1), less than twice as long as wide (
<figureCitation id="0C8B6AFDFFBD8A11FAB927359234F857" box="[1298,1369,1894,1920]" captionStart="FIGURE 2" captionStartId="3.[151,254,1942,1966]" captionTargetBox="[214,1370,193,1913]" captionTargetId="figure@3.[194,1399,165,1937]" captionTargetPageId="3" captionText="FIGURE 2. Gammarus zagrosensis n. sp., 3, 17.4 mm. A (HC), B (A 1), and C (A 2) on scale i; D (Mn) and E (MnPl) on scale ii, F (exopodite of Maxilipede), G (left palp of maxilla 1), and H (right palp of maxilla 1) on scale iii, and I (T) on scale iv." httpUri="https://zenodo.org/record/188495/files/figure.png" pageId="2" pageNumber="23">Fig. 2</figureCitation>
A).
</paragraph>
<paragraph id="940F7678FFBD8A11FF6E27DE940CF823" blockId="2.[151,1437,1082,2036]" pageId="2" pageNumber="23">
<emphasis id="A6C4AA6AFFBD8A11FF6E27DE963EF870" bold="true" box="[197,339,1933,1959]" pageId="2" pageNumber="23">Antenna 1:</emphasis>
About one-third of total body length, third peduncle segment shorter than first and second, about half length of first. Main and accessory flagellums with 2832 and 34 segments, respectively, with very short simple setae in groups of 5 on each side (
<figureCitation id="0C8B6AFDFFBD8A11FD5C27899453F823" box="[759,830,2010,2036]" captionStart="FIGURE 2" captionStartId="3.[151,254,1942,1966]" captionTargetBox="[214,1370,193,1913]" captionTargetId="figure@3.[194,1399,165,1937]" captionTargetPageId="3" captionText="FIGURE 2. Gammarus zagrosensis n. sp., 3, 17.4 mm. A (HC), B (A 1), and C (A 2) on scale i; D (Mn) and E (MnPl) on scale ii, F (exopodite of Maxilipede), G (left palp of maxilla 1), and H (right palp of maxilla 1) on scale iii, and I (T) on scale iv." httpUri="https://zenodo.org/record/188495/files/figure.png" pageId="2" pageNumber="23">Fig. 2</figureCitation>
B).
</paragraph>
<caption id="C0CF26F0FFBC8A10FF3C27C59780F824" httpUri="https://zenodo.org/record/188495/files/figure.png" pageId="3" pageNumber="24" targetBox="[214,1370,193,1913]" targetPageId="3">
<paragraph id="940F7678FFBC8A10FF3C27C59780F824" blockId="3.[151,1436,1942,2035]" pageId="3" pageNumber="24">
<emphasis id="A6C4AA6AFFBC8A10FF3C27C59675F879" bold="true" box="[151,280,1942,1966]" pageId="3" pageNumber="24">FIGURE 2.</emphasis>
<taxonomicName id="53B00DFBFFBC8A10FEB527C5957BF87A" box="[286,534,1942,1965]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="24" phylum="Arthropoda" rank="species" species="zagrosensis" status="sp. nov.">
<emphasis id="A6C4AA6AFFBC8A10FEB527C5957BF87A" box="[286,534,1942,1965]" italics="true" pageId="3" pageNumber="24">Gammarus zagrosensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BDF71711FFBC8A10FDB627C59535F87A" box="[541,600,1942,1966]" pageId="3" pageNumber="24" rank="species">
<emphasis id="A6C4AA6AFFBC8A10FDB627C5953CF879" bold="true" box="[541,593,1942,1966]" pageId="3" pageNumber="24">n. sp</emphasis>
.
</taxonomicNameLabel>
, 3, 17.4 mm. A (HC), B (A1), and C (A2) on scale i; D (Mn) and E (MnPl) on scale ii, F (exopodite of Maxilipede), G (left palp of maxilla 1), and H (right palp of maxilla 1) on scale iii, and I (T) on scale iv.
</paragraph>
</caption>
<paragraph id="940F7678FFBB8A17FF6E20C49608FEA4" blockId="4.[151,1437,151,2034]" pageId="4" pageNumber="25">
<emphasis id="A6C4AA6AFFBB8A17FF6E20C4963AFF66" bold="true" box="[197,343,151,177]" pageId="4" pageNumber="25">Antenna 2:</emphasis>
Gland cones rather slender, not reaching distal end of third peduncle segment; peduncle segments 4 and 5 of almost equal length, armed with tufts of setae of the same length or slightly longer than diameter of segments; flagellum with 1117 segments which are not swollen or compressed; groups of setae near to the end of each flagellar segment at both outer and inner surfaces. A row of setae implanted submarginal to the distal end of each segment at the inner surface, never forming a flag-like brush; calceoli present (
<figureCitation id="0C8B6AFDFFBB8A17FF57210A962FFEA4" box="[252,322,345,371]" captionStart="FIGURE 2" captionStartId="3.[151,254,1942,1966]" captionTargetBox="[214,1370,193,1913]" captionTargetId="figure@3.[194,1399,165,1937]" captionTargetPageId="3" captionText="FIGURE 2. Gammarus zagrosensis n. sp., 3, 17.4 mm. A (HC), B (A 1), and C (A 2) on scale i; D (Mn) and E (MnPl) on scale ii, F (exopodite of Maxilipede), G (left palp of maxilla 1), and H (right palp of maxilla 1) on scale iii, and I (T) on scale iv." httpUri="https://zenodo.org/record/188495/files/figure.png" pageId="4" pageNumber="25">Fig. 2</figureCitation>
C).
</paragraph>
<paragraph id="940F7678FFBB8A17FF6E212C968CFE17" blockId="4.[151,1437,151,2034]" pageId="4" pageNumber="25">
<emphasis id="A6C4AA6AFFBB8A17FF6E212C9625FE4E" bold="true" box="[197,328,383,409]" pageId="4" pageNumber="25">Mandible:</emphasis>
Incisor processes,
<emphasis id="A6C4AA6AFFBB8A17FD8521D3958DFE4E" box="[558,736,384,409]" italics="true" pageId="4" pageNumber="25">lacinia mobilis</emphasis>
and ridged molar process well developed, a plumose long spine row present (
<figureCitation id="0C8B6AFDFFBB8A17FEDE21F596D1FE17" box="[373,444,422,448]" captionStart="FIGURE 2" captionStartId="3.[151,254,1942,1966]" captionTargetBox="[214,1370,193,1913]" captionTargetId="figure@3.[194,1399,165,1937]" captionTargetPageId="3" captionText="FIGURE 2. Gammarus zagrosensis n. sp., 3, 17.4 mm. A (HC), B (A 1), and C (A 2) on scale i; D (Mn) and E (MnPl) on scale ii, F (exopodite of Maxilipede), G (left palp of maxilla 1), and H (right palp of maxilla 1) on scale iii, and I (T) on scale iv." httpUri="https://zenodo.org/record/188495/files/figure.png" pageId="4" pageNumber="25">Fig. 2</figureCitation>
D).
</paragraph>
<paragraph id="940F7678FFBB8A17FF6E219E9460FD8C" blockId="4.[151,1437,151,2034]" pageId="4" pageNumber="25">
<emphasis id="A6C4AA6AFFBB8A17FF6E219E96EAFE30" bold="true" box="[197,391,461,487]" pageId="4" pageNumber="25">Mandible palp:</emphasis>
First segment not armed; second segment with ventral setae, 45 proximal setae shorter than the width of the segment and 710 closely placed distal setae longer than the width of the segment; inferior margin of the third segment armed with a comb-like row of 2529 D-setae, 34 long E-setae, two groups of B-setae and one group of A-setae (
<figureCitation id="0C8B6AFDFFBB8A17FD0E22129581FD8C" box="[677,748,577,603]" captionStart="FIGURE 2" captionStartId="3.[151,254,1942,1966]" captionTargetBox="[214,1370,193,1913]" captionTargetId="figure@3.[194,1399,165,1937]" captionTargetPageId="3" captionText="FIGURE 2. Gammarus zagrosensis n. sp., 3, 17.4 mm. A (HC), B (A 1), and C (A 2) on scale i; D (Mn) and E (MnPl) on scale ii, F (exopodite of Maxilipede), G (left palp of maxilla 1), and H (right palp of maxilla 1) on scale iii, and I (T) on scale iv." httpUri="https://zenodo.org/record/188495/files/figure.png" pageId="4" pageNumber="25">Fig. 2</figureCitation>
E).
</paragraph>
<paragraph id="940F7678FFBB8A17FF6E22349567FD7F" blockId="4.[151,1437,151,2034]" pageId="4" pageNumber="25">
<emphasis id="A6C4AA6AFFBB8A17FF6E22349658FD56" bold="true" box="[197,309,615,641]" pageId="4" pageNumber="25">Labium:</emphasis>
Simple, with a group of short setae on the apices of the inner lobe, and a group of very fine bristles on inner and outer lobe.
</paragraph>
<paragraph id="940F7678FFBB8A17FF6E22E696DFFCBD" blockId="4.[151,1437,151,2034]" pageId="4" pageNumber="25">
<emphasis id="A6C4AA6AFFBB8A17FF6E22E6962BFD18" bold="true" box="[197,326,693,719]" pageId="4" pageNumber="25">Maxilla 1:</emphasis>
Inner basal lobe with long finely plumose setae; outer lobe with stout serrate spines; left palp at its end has an armature of 4 median spines accompanied by a short seta, one longer separate subapical spine on its outer margin and 3 smaller spines on distal inner angle (
<figureCitation id="0C8B6AFDFFBB8A17FCD2235194ADFCCB" box="[889,960,770,796]" captionStart="FIGURE 2" captionStartId="3.[151,254,1942,1966]" captionTargetBox="[214,1370,193,1913]" captionTargetId="figure@3.[194,1399,165,1937]" captionTargetPageId="3" captionText="FIGURE 2. Gammarus zagrosensis n. sp., 3, 17.4 mm. A (HC), B (A 1), and C (A 2) on scale i; D (Mn) and E (MnPl) on scale ii, F (exopodite of Maxilipede), G (left palp of maxilla 1), and H (right palp of maxilla 1) on scale iii, and I (T) on scale iv." httpUri="https://zenodo.org/record/188495/files/figure.png" pageId="4" pageNumber="25">Fig. 2</figureCitation>
G), right palp with five robust tooth-like spines at its end, one longer separate subapical spine on its outer margin opposed by one smaller spine, anterodistally (
<figureCitation id="0C8B6AFDFFBB8A17FEED230396E0FCBD" box="[326,397,848,874]" captionStart="FIGURE 2" captionStartId="3.[151,254,1942,1966]" captionTargetBox="[214,1370,193,1913]" captionTargetId="figure@3.[194,1399,165,1937]" captionTargetPageId="3" captionText="FIGURE 2. Gammarus zagrosensis n. sp., 3, 17.4 mm. A (HC), B (A 1), and C (A 2) on scale i; D (Mn) and E (MnPl) on scale ii, F (exopodite of Maxilipede), G (left palp of maxilla 1), and H (right palp of maxilla 1) on scale iii, and I (T) on scale iv." httpUri="https://zenodo.org/record/188495/files/figure.png" pageId="4" pageNumber="25">Fig. 2</figureCitation>
H).
</paragraph>
<paragraph id="940F7678FFBB8A17FF6E2325967AFB85" blockId="4.[151,1437,151,2034]" pageId="4" pageNumber="25">
<emphasis id="A6C4AA6AFFBB8A17FF6E23259603FC47" bold="true" box="[197,366,886,912]" pageId="4" pageNumber="25">Maxillipedes:</emphasis>
Exopodite with a row of three distal strong teeth, accompanied by three short thick setae; a row of setae on distal margin becomes plumose from the middle towards the inferior side, where it joins about 10 very long regularly placed plumose setae at the inferior edge (
<figureCitation id="0C8B6AFDFFBB8A17FC21239794BDFC09" box="[906,976,964,990]" captionStart="FIGURE 2" captionStartId="3.[151,254,1942,1966]" captionTargetBox="[214,1370,193,1913]" captionTargetId="figure@3.[194,1399,165,1937]" captionTargetPageId="3" captionText="FIGURE 2. Gammarus zagrosensis n. sp., 3, 17.4 mm. A (HC), B (A 1), and C (A 2) on scale i; D (Mn) and E (MnPl) on scale ii, F (exopodite of Maxilipede), G (left palp of maxilla 1), and H (right palp of maxilla 1) on scale iii, and I (T) on scale iv." httpUri="https://zenodo.org/record/188495/files/figure.png" pageId="4" pageNumber="25">Fig. 2</figureCitation>
F); endopodite with a row of long distal plumose setae which gradually shorten and lose plumosity towards the interior, joining to a row of inferior strong spine-teeth gradually shorten proximally and finally merging to a row of long setae. The palp is well developed.
</paragraph>
<paragraph id="940F7678FFBB8A17FF6E240D9327FA50" blockId="4.[151,1437,151,2034]" pageId="4" pageNumber="25">
<emphasis id="A6C4AA6AFFBB8A17FF6E240D9619FBAF" bold="true" box="[197,372,1118,1144]" pageId="4" pageNumber="25">Gnathopod 1:</emphasis>
Coxal plate distally not or slightly wider than proximally, rounded corners with a setae at the postero-ventral corner and three to four setae at antero-ventral corner, one fine setae at 1/4th of the postero-ventral surface of the plate; basis is expanded at 1/3rd, gradually becomes narrower toward the base of the segment; carpus and merus with groups of setae which are plumose at posterior margin of carpus; propodus pyriform, palm oblique with one medial palmar spine and several palmar angle spines, five groups of small spines at posterior palmar margin, a group of four setae very close to medial palmar spine, two groups of setae at the propodal anterior margin, a row of five groups of short setae on propodal surface, a row of three groups of small spines (2 each) on posterior sab-margin, dactylus long (
<figureCitation id="0C8B6AFDFFBB8A17FC75253E934EFA50" box="[990,1059,1389,1415]" captionStart="FIGURE 3" captionStartId="5.[151,254,2006,2030]" captionTargetBox="[191,1404,191,1980]" captionTargetId="figure@5.[185,1409,182,1982]" captionTargetPageId="5" captionText="FIGURE 3. Gammarus zagrosensis n. sp., 3, 17.4 mm. A (G 1) and B (G 2) on scale i, C (P 3) and D (P 4) on scale ii." httpUri="https://zenodo.org/record/188496/files/figure.png" pageId="4" pageNumber="25">Fig. 3</figureCitation>
A).
</paragraph>
<paragraph id="940F7678FFBB8A17FF6E25C096A7F9F5" blockId="4.[151,1437,151,2034]" pageId="4" pageNumber="25">
<emphasis id="A6C4AA6AFFBB8A17FF6E25C09618FA7A" bold="true" box="[197,373,1427,1453]" pageId="4" pageNumber="25">Gnathopod 2:</emphasis>
Distal end of coxal plate finely narrower than proximal; propodus is less elongated than propodus of GP1; posterior propodal margin is convex; a group of setae close to the medial palmar spine, two groups of setae at propodal anterior margin; palm is oblique having a robust medial palmar spine and palmar angle spines (3) (
<figureCitation id="0C8B6AFDFFBB8A17FECB265B96CBF9F5" box="[352,422,1544,1570]" captionStart="FIGURE 3" captionStartId="5.[151,254,2006,2030]" captionTargetBox="[191,1404,191,1980]" captionTargetId="figure@5.[185,1409,182,1982]" captionTargetPageId="5" captionText="FIGURE 3. Gammarus zagrosensis n. sp., 3, 17.4 mm. A (G 1) and B (G 2) on scale i, C (P 3) and D (P 4) on scale ii." httpUri="https://zenodo.org/record/188496/files/figure.png" pageId="4" pageNumber="25">Fig. 3</figureCitation>
B).
</paragraph>
<paragraph id="940F7678FFBB8A17FF6E267D9262F880" blockId="4.[151,1437,151,2034]" pageId="4" pageNumber="25">
<emphasis id="A6C4AA6AFFBB8A17FF6E267D9631F99F" bold="true" box="[197,348,1582,1608]" pageId="4" pageNumber="25">Pereopod 3:</emphasis>
Coxal plate rounded distally, concave posteriorly, convex at anterior margin, with 34 very short setae implanted at antero-distal corner, one at postero-distal corner, and a longer setae on outer surface of the plate. Basis-carpus bear groups of setae at posterior margin, about 1 to 1.5 times as long as the diameter of the segments in basis and ischium; those of merus and carpus are in dense groups and 23 times longer than the diameter of the segments; anterior margin of merus with two groups of few short setae and a shorter spine, and another group of long setae and a spine at anterior tip; carpus has one spine among each of its first three groups of setae, and a long spine and a group of longer setae are implanted on both its anterior and posterior tip; posterior margin of propodus with 5 groups of one small spine and some long setae (
<figureCitation id="0C8B6AFDFFBB8A17FB0E276E9387F880" box="[1189,1258,1853,1879]" captionStart="FIGURE 3" captionStartId="5.[151,254,2006,2030]" captionTargetBox="[191,1404,191,1980]" captionTargetId="figure@5.[185,1409,182,1982]" captionTargetPageId="5" captionText="FIGURE 3. Gammarus zagrosensis n. sp., 3, 17.4 mm. A (G 1) and B (G 2) on scale i, C (P 3) and D (P 4) on scale ii." httpUri="https://zenodo.org/record/188496/files/figure.png" pageId="4" pageNumber="25">Fig. 3</figureCitation>
C).
</paragraph>
<paragraph id="940F7678FFBB8A17FF6E27309267F825" blockId="4.[151,1437,151,2034]" pageId="4" pageNumber="25">
<emphasis id="A6C4AA6AFFBB8A17FF6E27309636F8AA" bold="true" box="[197,347,1891,1917]" pageId="4" pageNumber="25">Pereopod 4:</emphasis>
Coxal plate with five small setae implanted at antero-distal margin and eight at postero-distal margin; segments similar to pereopod 3, but setae are shorter and the number of setae and groups is lower. Anterior margin of merus with just one group of short setae and one spine, one long spine implanted at anterior tip among a group of setae; carpus has two spines among most groups of setae (
<figureCitation id="0C8B6AFDFFBB8A17FB34278B9389F825" box="[1183,1252,2008,2034]" captionStart="FIGURE 3" captionStartId="5.[151,254,2006,2030]" captionTargetBox="[191,1404,191,1980]" captionTargetId="figure@5.[185,1409,182,1982]" captionTargetPageId="5" captionText="FIGURE 3. Gammarus zagrosensis n. sp., 3, 17.4 mm. A (G 1) and B (G 2) on scale i, C (P 3) and D (P 4) on scale ii." httpUri="https://zenodo.org/record/188496/files/figure.png" pageId="4" pageNumber="25">Fig. 3</figureCitation>
D).
</paragraph>
<caption id="C0CF26F0FFBA8A16FF3C27859219F83A" box="[151,1396,2006,2030]" httpUri="https://zenodo.org/record/188496/files/figure.png" pageId="5" pageNumber="26" targetBox="[191,1404,191,1980]" targetPageId="5">
<paragraph id="940F7678FFBA8A16FF3C27859219F83A" blockId="5.[151,1396,2006,2030]" box="[151,1396,2006,2030]" pageId="5" pageNumber="26">
<emphasis id="A6C4AA6AFFBA8A16FF3C27859675F839" bold="true" box="[151,280,2006,2030]" pageId="5" pageNumber="26">FIGURE 3.</emphasis>
<taxonomicName id="53B00DFBFFBA8A16FEB427859575F83A" box="[287,536,2006,2029]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="26" phylum="Arthropoda" rank="species" species="zagrosensis" status="sp. nov.">
<emphasis id="A6C4AA6AFFBA8A16FEB427859575F83A" box="[287,536,2006,2029]" italics="true" pageId="5" pageNumber="26">Gammarus zagrosensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BDF71711FFBA8A16FDB527859537F83A" box="[542,602,2006,2030]" pageId="5" pageNumber="26" rank="species">
<emphasis id="A6C4AA6AFFBA8A16FDB527859539F839" bold="true" box="[542,596,2006,2030]" pageId="5" pageNumber="26">n. sp</emphasis>
.
</taxonomicNameLabel>
, 3, 17.4 mm. A (G1) and B (G2) on scale i, C (P3) and D (P4) on scale ii.
</paragraph>
</caption>
<paragraph id="940F7678FFB98A15FF6E20C494BAFE9B" blockId="6.[151,1437,151,2001]" pageId="6" pageNumber="27">
<emphasis id="A6C4AA6AFFB98A15FF6E20C49631FF66" bold="true" box="[197,348,151,177]" pageId="6" pageNumber="27">Pereopod 5:</emphasis>
Basis with a sub-rectangular shape, posterior margin slightly concave (variation: straight to slightly convex in some individuals), postero-distal lobe well developed, posterior margin with 14 very short setae, anterior basis with one to two groups of 3 marginal setae, 3-4 spines mixed with a fine seta, one seta at sub-margin of antero-proximal part of basis; merus and carpus with small spines and setae (as long as spines), propodus having 6 transverse rows of spines (233332) (
<figureCitation id="0C8B6AFDFFB98A15FCC0216194DFFE9B" box="[875,946,306,332]" captionStart="FIGURE 4" captionStartId="7.[151,257,1972,1996]" captionTargetBox="[212,1368,193,1938]" captionTargetId="figure@7.[192,1397,185,1957]" captionTargetPageId="7" captionText="FIGURE 4. Gammarus zagrosensis n. sp., 3, 17.4 mm. A (P 5), B (P 6), and C (P 7), Ci and Cii (bases of P 7 in two paratypes) on scale i; D (US) and E (EP) on scale ii; F (U 3) on scale iii." httpUri="https://zenodo.org/record/188497/files/figure.png" pageId="6" pageNumber="27">Fig. 4</figureCitation>
A).
</paragraph>
<paragraph id="940F7678FFB98A15FF6E210B97A7FE17" blockId="6.[151,1437,151,2001]" pageId="6" pageNumber="27">
<emphasis id="A6C4AA6AFFB98A15FF6E210B9633FEA5" bold="true" box="[197,350,344,370]" pageId="6" pageNumber="27">Pereopod 6:</emphasis>
Similar to pereopod 5, but longer (1.2 times longer than P5); basis more slender and more concave at postero-distal, 13 setae at the inner surface of the postero-distal corner of protruding lobe (
<figureCitation id="0C8B6AFDFFB98A15FAC721D397C5FE17" captionStart="FIGURE 4" captionStartId="7.[151,257,1972,1996]" captionTargetBox="[212,1368,193,1938]" captionTargetId="figure@7.[192,1397,185,1957]" captionTargetPageId="7" captionText="FIGURE 4. Gammarus zagrosensis n. sp., 3, 17.4 mm. A (P 5), B (P 6), and C (P 7), Ci and Cii (bases of P 7 in two paratypes) on scale i; D (US) and E (EP) on scale ii; F (U 3) on scale iii." httpUri="https://zenodo.org/record/188497/files/figure.png" pageId="6" pageNumber="27">Fig. 4</figureCitation>
B).
</paragraph>
<paragraph id="940F7678FFB98A15FF6E219E9564FD18" blockId="6.[151,1437,151,2001]" pageId="6" pageNumber="27">
<emphasis id="A6C4AA6AFFB98A15FF6E219E9631FE30" bold="true" box="[197,348,461,487]" pageId="6" pageNumber="27">Pereopod 7:</emphasis>
Basis wider proximally; postero-distal protruding lobe less developed than P6, set with few long setae at postero-inferior corner, 18 very short setae on posterior margin, starting from the base of the basis; anterior margin of merus and carpus with spines and longer setae; merus with a long spine and some short setae at the middle of posterior margin; carpus with one long and one smaller spine at two-third of posterior margin; propodus with spines and setae which are as long as spines, 7 transverse rows of spines on anterior margin of propodus (3333332); 2 longer spines at posterior tip of propodus intermixed with a group of longer setae (
<figureCitation id="0C8B6AFDFFB98A15FE3422E6968BFD18" box="[415,486,693,719]" captionStart="FIGURE 4" captionStartId="7.[151,257,1972,1996]" captionTargetBox="[212,1368,193,1938]" captionTargetId="figure@7.[192,1397,185,1957]" captionTargetPageId="7" captionText="FIGURE 4. Gammarus zagrosensis n. sp., 3, 17.4 mm. A (P 5), B (P 6), and C (P 7), Ci and Cii (bases of P 7 in two paratypes) on scale i; D (US) and E (EP) on scale ii; F (U 3) on scale iii." httpUri="https://zenodo.org/record/188497/files/figure.png" pageId="6" pageNumber="27">Fig. 4</figureCitation>
C).
</paragraph>
<paragraph id="940F7678FFB98A15FF6E228895AEFC47" blockId="6.[151,1437,151,2001]" pageId="6" pageNumber="27">
<emphasis id="A6C4AA6AFFB98A15FF6E22889625FD22" bold="true" box="[197,328,731,757]" pageId="6" pageNumber="27">Uropod 3:</emphasis>
Endopodite more than two-thirds (0.71) of the exopodite; setae on both rami are more than twice as long as the width of the part of the segment they are implanted on, many setae on outer margin of both exopodite and endopodite are plumose, a group of 1 spine and 5 setae on two-thirds of the basis and a number of setae and short to long robust spines implanted on the apex; long spines at intervals on outer margins of endopodite and exopodite (
<figureCitation id="0C8B6AFDFFB98A15FDF1232595F2FC47" box="[602,671,886,912]" captionStart="FIGURE 3" captionStartId="5.[151,254,2006,2030]" captionTargetBox="[191,1404,191,1980]" captionTargetId="figure@5.[185,1409,182,1982]" captionTargetPageId="5" captionText="FIGURE 3. Gammarus zagrosensis n. sp., 3, 17.4 mm. A (G 1) and B (G 2) on scale i, C (P 3) and D (P 4) on scale ii." httpUri="https://zenodo.org/record/188496/files/figure.png" pageId="6" pageNumber="27">Fig. 3</figureCitation>
E).
</paragraph>
<paragraph id="940F7678FFB98A15FF6E23CE9575FB85" blockId="6.[151,1437,151,2001]" pageId="6" pageNumber="27">
<emphasis id="A6C4AA6AFFB98A15FF6E23CE9625FC60" bold="true" box="[197,328,925,951]" pageId="6" pageNumber="27">Epimeres:</emphasis>
Posterior-inferior corner of the first epimeral plate rounded with a small pointed node, 68 long setae on antero-distal corner; EP2 sub-rectangular with rounded anterior margin, EP3 moderately pointed; 47 extraordinary long setae on distal sub-marginal area of 2nd and 3rd plate rarely mixed with spine; a row of setules on posterior margin of all 3 plates; dorsal surface of metasomites posteriorly covered with very short setules (
<figureCitation id="0C8B6AFDFFB98A15FE04246B969BFB85" box="[431,502,1080,1106]" captionStart="FIGURE 4" captionStartId="7.[151,257,1972,1996]" captionTargetBox="[212,1368,193,1938]" captionTargetId="figure@7.[192,1397,185,1957]" captionTargetPageId="7" captionText="FIGURE 4. Gammarus zagrosensis n. sp., 3, 17.4 mm. A (P 5), B (P 6), and C (P 7), Ci and Cii (bases of P 7 in two paratypes) on scale i; D (US) and E (EP) on scale ii; F (U 3) on scale iii." httpUri="https://zenodo.org/record/188497/files/figure.png" pageId="6" pageNumber="27">Fig. 4</figureCitation>
E).
</paragraph>
<paragraph id="940F7678FFB98A15FF6E240D9792FB11" blockId="6.[151,1437,151,2001]" pageId="6" pageNumber="27">
<emphasis id="A6C4AA6AFFB98A15FF6E240D9671FBAF" bold="true" box="[197,284,1118,1144]" pageId="6" pageNumber="27">Telson:</emphasis>
Rather elongated, length of the lobes about twice their widest width; two to three long and robust spines and some 57 long setae on distal margin; four groups of 26 setae at the dorsal surface of the lobes (
<figureCitation id="0C8B6AFDFFB98A15FF3424FF978BFB11" box="[159,230,1196,1222]" captionStart="FIGURE 2" captionStartId="3.[151,254,1942,1966]" captionTargetBox="[214,1370,193,1913]" captionTargetId="figure@3.[194,1399,165,1937]" captionTargetPageId="3" captionText="FIGURE 2. Gammarus zagrosensis n. sp., 3, 17.4 mm. A (HC), B (A 1), and C (A 2) on scale i; D (Mn) and E (MnPl) on scale ii, F (exopodite of Maxilipede), G (left palp of maxilla 1), and H (right palp of maxilla 1) on scale iii, and I (T) on scale iv." httpUri="https://zenodo.org/record/188495/files/figure.png" pageId="6" pageNumber="27">
Fig.
<date id="E00E50B8FFB98A15FF7D24FF978BFB11" box="[214,230,1196,1222]" pageId="6" pageNumber="27">2</date>
</figureCitation>
I).
</paragraph>
<paragraph id="940F7678FFB98A15FF6E2481931BFAC4" blockId="6.[151,1437,151,2001]" pageId="6" pageNumber="27">
<emphasis id="A6C4AA6AFFB98A15FF6E24819631FB3B" bold="true" box="[197,348,1234,1260]" pageId="6" pageNumber="27">Urosomites:</emphasis>
Moderate elevations on postero-dorsal of urosome segments, dorsal armature consists of 12 lateral spines and 2 mid-dorsal spines, all combined with some short setae (
<figureCitation id="0C8B6AFDFFB98A15FBA124AA933CFAC4" box="[1034,1105,1273,1299]" captionStart="FIGURE 4" captionStartId="7.[151,257,1972,1996]" captionTargetBox="[212,1368,193,1938]" captionTargetId="figure@7.[192,1397,185,1957]" captionTargetPageId="7" captionText="FIGURE 4. Gammarus zagrosensis n. sp., 3, 17.4 mm. A (P 5), B (P 6), and C (P 7), Ci and Cii (bases of P 7 in two paratypes) on scale i; D (US) and E (EP) on scale ii; F (U 3) on scale iii." httpUri="https://zenodo.org/record/188497/files/figure.png" pageId="6" pageNumber="27">Fig. 4</figureCitation>
D).
</paragraph>
<paragraph id="940F7678FFB98A15FF6D254C9645FAED" blockId="6.[151,1437,151,2001]" box="[198,296,1311,1338]" pageId="6" pageNumber="27">
<emphasis id="A6C4AA6AFFB98A15FF6D254C964CFAEE" bold="true" box="[198,289,1311,1337]" italics="true" pageId="6" pageNumber="27">Female</emphasis>
.
</paragraph>
<paragraph id="940F7678FFB98A15FF6D25159652FAB7" blockId="6.[151,1437,151,2001]" box="[198,319,1350,1376]" pageId="6" pageNumber="27">Female is:</paragraph>
<paragraph id="940F7678FFB98A15FF3C253E957BFA50" blockId="6.[151,1437,151,2001]" box="[151,534,1389,1415]" pageId="6" pageNumber="27">
- Smaller than male (
<quantity id="5348DB9DFFB98A15FE00253E9568FA50" box="[427,517,1389,1415]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="27" unit="mm" value="12.0">12 mm</quantity>
.).
</paragraph>
<paragraph id="940F7678FFB98A15FF3C25C792F1FA79" blockId="6.[151,1437,151,2001]" box="[151,1436,1428,1454]" pageId="6" pageNumber="27">- Palm in propodus of gnathopod 1 is less oblique; medial palmar spine absent, one long palmar angle</paragraph>
<paragraph id="940F7678FFB98A15FF6C25E992F1FA2C" blockId="6.[151,1437,151,2001]" pageId="6" pageNumber="27">spine, with 23 spines at posterior palmar margin, a group of setae at the medial palmar region. - Propodus of gnathopod 2 sub-rectangular in shape, and lack the medial palmar spine; 2 palmar angle</paragraph>
<paragraph id="940F7678FFB98A15FF6C265B92F6F99F" blockId="6.[151,1437,151,2001]" pageId="6" pageNumber="27">spines, a group of setae at medial palmar region, no spines on posterior palmar margin. - Propodus of gnathopod 1 and 2 is smaller in respect to the other segments, and the whole appendage</paragraph>
<paragraph id="940F7678FFB98A15FF6C260696A5F9B8" blockId="6.[151,1437,151,2001]" box="[199,456,1621,1647]" pageId="6" pageNumber="27">smaller than in males.</paragraph>
<paragraph id="940F7678FFB98A15FF3C262F933AF941" blockId="6.[151,1437,151,2001]" box="[151,1111,1660,1686]" pageId="6" pageNumber="27">- Proximal 3/4th of gnathopod 1 and 2 bases are not wider than their distal part.</paragraph>
<paragraph id="940F7678FFB98A15FF3C26F192F1F96B" blockId="6.[151,1437,151,2001]" box="[151,1436,1698,1724]" pageId="6" pageNumber="27">- Basis of pereopods 57 less elongated than in males, rather distinct convex angles at the middle of</paragraph>
<paragraph id="940F7678FFB98A15FF6C269A957AF934" blockId="6.[151,1437,151,2001]" box="[199,535,1737,1763]" pageId="6" pageNumber="27">
posterior edges (
<figureCitation id="0C8B6AFDFFB98A15FE22269A96A0F934" box="[393,461,1737,1763]" captionStart="FIGURE 5" captionStartId="8.[151,254,1109,1133]" captionTargetBox="[172,1410,191,1075]" captionTargetId="figure@8.[151,1436,178,1085]" captionTargetPageId="8" captionText="FIGURE 5. Gammarus zagrosensis n. sp., Ƥ, 12 mm. A (basis of P 5), B (basis of P 6), C (basis of P 7) and E (oostegite 1) on scale i; D (EP) on scale ii." httpUri="https://zenodo.org/record/188498/files/figure.png" pageId="6" pageNumber="27">Fig. 5</figureCitation>
AC).
</paragraph>
<paragraph id="940F7678FFB98A15FF3C26A392F1F8DD" blockId="6.[151,1437,151,2001]" box="[151,1436,1776,1802]" pageId="6" pageNumber="27">
- Epimeral plates of the females do not show any significant variation in respect to those of the males (
<figureCitation id="0C8B6AFDFFB98A15FAC726A392F1F8DD" box="[1388,1436,1776,1802]" captionStart="FIGURE 5" captionStartId="8.[151,254,1109,1133]" captionTargetBox="[172,1410,191,1075]" captionTargetId="figure@8.[151,1436,178,1085]" captionTargetPageId="8" captionText="FIGURE 5. Gammarus zagrosensis n. sp., Ƥ, 12 mm. A (basis of P 5), B (basis of P 6), C (basis of P 7) and E (oostegite 1) on scale i; D (EP) on scale ii." httpUri="https://zenodo.org/record/188498/files/figure.png" pageId="6" pageNumber="27">Fig.</figureCitation>
</paragraph>
<paragraph id="940F7678FFB98A15FF6C274595D6F8E7" blockId="6.[151,1437,151,2001]" box="[199,699,1814,1840]" pageId="6" pageNumber="27">5D). Oostegite 1 is illustrated in figure 5E.</paragraph>
<paragraph id="940F7678FFB98A15FF6E276E9327F806" blockId="6.[151,1437,151,2001]" pageId="6" pageNumber="27">
<emphasis id="A6C4AA6AFFB98A15FF6E276E9566F880" bold="true" box="[197,523,1853,1879]" pageId="6" pageNumber="27">Cuticular ultrastructure:</emphasis>
HC cuticular polygons with both linear and scattered
<typeStatus id="4B0BC8DAFFB98A15FB47276E9247F880" box="[1260,1322,1853,1879]" pageId="6" pageNumber="27">types</typeStatus>
of pore arrangements (
<figureCitation id="0C8B6AFDFFB98A15FEED273796E3F8A9" box="[326,398,1892,1918]" captionStart="FIGURE 6" captionStartId="8.[151,254,1670,1694]" captionTargetBox="[157,1431,1221,1646]" captionTargetId="figure@8.[151,1436,1221,1646]" captionTargetPageId="8" captionText="FIGURE 6. SEM photographs of head region in Gammarus zagrosensis n. sp. (Right), and G. sepidannus n. sp. (Left)." httpUri="https://zenodo.org/record/188499/files/figure.png" pageId="6" pageNumber="27">Fig. 6</figureCitation>
, left); lines with a distance of 0.561.48 (Mean = 0.97, SD = 0.20); density of pores in line-type (L-type) polygons 6280 pores per 25 µm2 (Mean = 71, SD = 5), while scattered-type (S-type) polygons with higher density of 75244 pores per
<quantity id="5348DB9DFFB98A15FD4E27E4944CF806" box="[741,801,1975,2001]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="6" pageNumber="27" unit="m" value="25.0">25 m</quantity>
2 (Mean = 163, SD = 52).
</paragraph>
</subSubSection>
<caption id="C0CF26F0FFB88A14FF3C27E794FEF839" httpUri="https://zenodo.org/record/188497/files/figure.png" pageId="7" pageNumber="28" targetBox="[212,1368,193,1938]" targetPageId="7">
<paragraph id="940F7678FFB88A14FF3C27E794FEF839" blockId="7.[151,1436,1972,2030]" pageId="7" pageNumber="28">
<emphasis id="A6C4AA6AFFB88A14FF3C27E79673F81B" bold="true" box="[151,286,1972,1996]" pageId="7" pageNumber="28">FIGURE 4.</emphasis>
<taxonomicName id="53B00DFBFFB88A14FE8E27E6954BF81B" box="[293,550,1973,1996]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="7" pageNumber="28" phylum="Arthropoda" rank="species" species="zagrosensis" status="sp. nov.">
<emphasis id="A6C4AA6AFFB88A14FE8E27E6954BF81B" box="[293,550,1973,1996]" italics="true" pageId="7" pageNumber="28">Gammarus zagrosensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BDF71711FFB88A14FD8527E79501F81B" box="[558,620,1972,1996]" pageId="7" pageNumber="28" rank="species">
<emphasis id="A6C4AA6AFFB88A14FD8527E7950BF81B" bold="true" box="[558,614,1972,1996]" pageId="7" pageNumber="28">n. sp</emphasis>
.
</taxonomicNameLabel>
, 3, 17.4 mm. A (P5), B (P6), and C (P7), Ci and Cii (bases of P7 in two paratypes) on scale i; D (US) and E (EP) on scale ii; F (U3) on scale iii.
</paragraph>
</caption>
<caption id="C0CF26F0FFB78A1BFF3C24069686FB58" httpUri="https://zenodo.org/record/188498/files/figure.png" pageId="8" pageNumber="29" targetBox="[172,1410,191,1075]" targetPageId="8">
<paragraph id="940F7678FFB78A1BFF3C24069686FB58" blockId="8.[151,1436,1109,1167]" pageId="8" pageNumber="29">
<emphasis id="A6C4AA6AFFB78A1BFF3C24069674FBBA" bold="true" box="[151,281,1109,1133]" pageId="8" pageNumber="29">FIGURE 5.</emphasis>
<taxonomicName id="53B00DFBFFB78A1BFE8B24059574FBBA" box="[288,537,1110,1133]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="29" phylum="Arthropoda" rank="species" species="zagrosensis" status="sp. nov.">
<emphasis id="A6C4AA6AFFB78A1BFE8B24059574FBBA" box="[288,537,1110,1133]" italics="true" pageId="8" pageNumber="29">Gammarus zagrosensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BDF71711FFB78A1BFD8A24069530FBBB" box="[545,605,1109,1133]" pageId="8" pageNumber="29" rank="species">
<emphasis id="A6C4AA6AFFB78A1BFD8A2406953BFBBA" bold="true" box="[545,598,1109,1133]" pageId="8" pageNumber="29">n. sp</emphasis>
.
</taxonomicNameLabel>
, Ƥ, 12 mm. A (basis of P5), B (basis of P6), C (basis of P7) and E (oostegite 1) on scale i; D (EP) on scale ii.
</paragraph>
</caption>
<caption id="C0CF26F0FFB78A1BFF3C26D592FDF94A" box="[151,1424,1670,1694]" httpUri="https://zenodo.org/record/188499/files/figure.png" pageId="8" pageNumber="29" targetBox="[157,1431,1221,1646]" targetPageId="8">
<paragraph id="940F7678FFB78A1BFF3C26D592FDF94A" blockId="8.[151,1424,1670,1694]" box="[151,1424,1670,1694]" pageId="8" pageNumber="29">
<emphasis id="A6C4AA6AFFB78A1BFF3C26D59674F949" bold="true" box="[151,281,1670,1694]" pageId="8" pageNumber="29">FIGURE 6.</emphasis>
SEM photographs of head region in
<taxonomicName id="53B00DFBFFB78A1BFD0826D494F6F949" box="[675,923,1671,1694]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="29" phylum="Arthropoda" rank="species" species="zagrosensis" status="sp. nov.">
<emphasis id="A6C4AA6AFFB78A1BFD0826D494F6F949" box="[675,923,1671,1694]" italics="true" pageId="8" pageNumber="29">Gammarus zagrosensis</emphasis>
</taxonomicName>
<emphasis id="A6C4AA6AFFB78A1BFC0926D594B3F949" bold="true" box="[930,990,1670,1694]" pageId="8" pageNumber="29">
<taxonomicNameLabel id="BDF71711FFB78A1BFC0926D594B3F949" box="[930,990,1670,1694]" pageId="8" pageNumber="29" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(Right), and
<taxonomicName id="53B00DFBFFB78A1BFBC726D4926FF949" box="[1132,1282,1671,1694]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="29" phylum="Arthropoda" rank="species" species="sepidannus" status="sp. nov.">
<emphasis id="A6C4AA6AFFB78A1BFBC726D4926FF949" box="[1132,1282,1671,1694]" italics="true" pageId="8" pageNumber="29">G. sepidannus</emphasis>
</taxonomicName>
<emphasis id="A6C4AA6AFFB78A1BFAA326D59229F949" bold="true" box="[1288,1348,1670,1694]" pageId="8" pageNumber="29">
<taxonomicNameLabel id="BDF71711FFB78A1BFAA326D59229F949" box="[1288,1348,1670,1694]" pageId="8" pageNumber="29" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(Left).
</paragraph>
</caption>
<subSubSection id="DCAA25F3FFB78A1AFF6D2698951CFEA4" lastPageId="9" lastPageNumber="30" pageId="8" pageNumber="29" type="discussion">
<paragraph id="940F7678FFB78A1BFF6D26989210F8DB" blockId="8.[151,1437,1739,2036]" pageId="8" pageNumber="29">
<emphasis id="A6C4AA6AFFB78A1BFF6D2698963CF932" bold="true" box="[198,337,1739,1765]" pageId="8" pageNumber="29">Discussion.</emphasis>
<taxonomicName id="53B00DFBFFB78A1BFECA269F9516F932" box="[353,635,1740,1765]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="29" phylum="Arthropoda" rank="species" species="zagrosensis">
<emphasis id="A6C4AA6AFFB78A1BFECA269F9516F932" box="[353,635,1740,1765]" italics="true" pageId="8" pageNumber="29">Gammarus zagrosensis</emphasis>
</taxonomicName>
shows minute similarities with some of the well known taxa. Comparisons with these morphologically closest species, and species from adjacent regions are given here.
</paragraph>
<paragraph id="940F7678FFB78A1BFF6E274A94F1F823" blockId="8.[151,1437,1739,2036]" pageId="8" pageNumber="29">
This species has some similarities with
<taxonomicName id="53B00DFBFFB78A1BFD3D274A948BF8E4" authority="G. O. Sars, 1863" authorityName="G. O. Sars" authorityYear="1863" box="[662,998,1817,1843]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="29" phylum="Arthropoda" rank="species" species="lacustris">
<emphasis id="A6C4AA6AFFB78A1BFD3D274A9476F8E5" box="[662,795,1817,1842]" italics="true" pageId="8" pageNumber="29">G. lacustris</emphasis>
G. O. Sars, 1863
</taxonomicName>
, a widely distributed species, e.g. the relatively short A1, shape and armature of U3, and of telson. On the other hand, the new species differs from
<taxonomicName id="53B00DFBFFB78A1BFF3C27349676F857" box="[151,283,1895,1920]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="29" phylum="Arthropoda" rank="species" species="lacustris">
<emphasis id="A6C4AA6AFFB78A1BFF3C27349676F857" box="[151,283,1895,1920]" italics="true" pageId="8" pageNumber="29">G. lacustris</emphasis>
</taxonomicName>
by its elliptical eye, much denser and longer setae on P34; existence of setae on merus and carpus of P7 (as long as/longer than twice the length of spines) and the posterior-inferior corner setae of P7 basis, having dorsal elevation on epimeres, non-sharp EP2, long setae on EP2 and EP3, in addition to long setae on the tip of telson, which can clearly differentiate these two species.
</paragraph>
<paragraph id="940F7678FFB68A1AFF6E20CB951CFEA4" blockId="9.[151,1437,152,2034]" pageId="9" pageNumber="30">
<emphasis id="A6C4AA6AFFB68A1AFF6E20CB9673FF66" box="[197,286,152,177]" italics="true" pageId="9" pageNumber="30">G. ro u x i</emphasis>
Pinkster &amp; Goedmakers, 1975 is the only species having long setae on EP 2 and 3, in addition to possession of setules on metasomites' dorsal surfaces, but has many other features which separate it strongly from this species, among them are: much shorter A1 and A2, short and very few setae on U3 and telson, less pointed EP3, longer setae on A2, longer setae on P57 segments, very different shape of P6 and P7 basis and having many setae on their interior surface. It is also worthy to add the point that
<emphasis id="A6C4AA6AFFB68A1AFBCF216193D3FE9C" box="[1124,1214,306,331]" italics="true" pageId="9" pageNumber="30">G. ro u x i</emphasis>
has been reported from North Africa, far distant from
<collectingCountry id="ECA736E8FFB68A1AFD91210A9500FEA4" box="[570,621,345,371]" name="Iran" pageId="9" pageNumber="30">Iran</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="DCAA25F3FFB68A1AFF6E21D395B9F872" pageId="9" pageNumber="30" type="reference_group">
<paragraph id="940F7678FFB68A1AFF6E21D3931EFDE3" blockId="9.[151,1437,152,2034]" pageId="9" pageNumber="30">
<taxonomicName id="53B00DFBFFB68A1AFF6E21D396C9FE4E" ID-CoL="3F8J9" box="[197,420,384,409]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="30" phylum="Arthropoda" rank="species" species="pseudosyriacus">
<emphasis id="A6C4AA6AFFB68A1AFF6E21D396C9FE4E" box="[197,420,384,409]" italics="true" pageId="9" pageNumber="30">G. pseudosyriacus</emphasis>
</taxonomicName>
<bibRefCitation id="F0210B89FFB68A1AFE0621D39597FE4D" author="Karaman" box="[429,762,384,410]" pageId="9" pageNumber="39" refString="Karaman, G. S. &amp; Pinkster, S. (1977) Fresh water Gammarus species from Europe, North Africa and adjacent regions of Asia (Crustacea, Amphipoda), part 1. Gammarus pulex - group and related species. Bijdragen Tot Der Dierkunde. 47 (1), 1 - 97." type="journal article" year="1977">Karaman &amp; Pinkster, 1977</bibRefCitation>
, which has been reported by
<bibRefCitation id="F0210B89FFB68A1AFBCD21D39223FE4D" box="[1126,1358,384,410]" pageId="9" pageNumber="39" refString="Stock, J. H., Mirzajani, A. R., Vonk, R., Naderi, S. &amp; Kiabi, B. H. (1998) Limnic and brackish water Amphipoda (Crustacea) from Iran. Beaufortia, 48 (9), 173 - 234." type="journal article">
Stock
<emphasis id="A6C4AA6AFFB68A1AFB1F21D39386FE4E" box="[1204,1259,384,409]" italics="true" pageId="9" pageNumber="30">et al</emphasis>
. (1998)
</bibRefCitation>
in the adjacent western province, has larger eyes, less setose A1, shorter setae on A2, a longer antennal gland cone (reaches to the tip of the third peduncle segment), less spines on posterior propodus margin of GP1, no setae on P57 merus and carpus, and the posterior-inferior corner setae, shorter setae on both endo- and exopodite of U3, and more pointed EP2 without long setae on inferior margin of EP1 and EP2.
</paragraph>
<paragraph id="940F7678FFB68A1AFF6E221293BFFD18" blockId="9.[151,1437,152,2034]" pageId="9" pageNumber="30">
<taxonomicName id="53B00DFBFFB68A1AFF6E2212950BFD8C" ID-CoL="6JWXF" authority="Stock et al., 1998" authorityName="Stock et al." authorityYear="1998" box="[197,614,577,603]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="30" phylum="Arthropoda" rank="species" species="crinicaudatus">
<emphasis id="A6C4AA6AFFB68A1AFF6E221296E4FD8D" box="[197,393,577,602]" italics="true" pageId="9" pageNumber="30">G. crinicaudatus</emphasis>
<bibRefCitation id="F0210B89FFB68A1AFE3A2212950BFD8C" author="Stock" box="[401,614,577,603]" pageId="9" pageNumber="39" refString="Stock, J. H., Mirzajani, A. R., Vonk, R., Naderi, S. &amp; Kiabi, B. H. (1998) Limnic and brackish water Amphipoda (Crustacea) from Iran. Beaufortia, 48 (9), 173 - 234." type="journal article" year="1998">
Stock
<emphasis id="A6C4AA6AFFB68A1AFE772212957DFD8D" box="[476,528,577,602]" italics="true" pageId="9" pageNumber="30">et al</emphasis>
., 1998
</bibRefCitation>
</taxonomicName>
, which was firstly reported and described from inside
<collectingCountry id="ECA736E8FFB68A1AFB5B2212924EFD8C" box="[1264,1315,577,603]" name="Iran" pageId="9" pageNumber="30">Fars</collectingCountry>
Province, has a shorter antennal gland cone, longer and more groups of setae on A2, merus and carpus of P57, and also on dorsal of epimeres, fewer groups of setae on GP1 propodus surface, additional accessory spines near the palmar angle spine, more numerous long setae on telson, and no long setae on EP2 and EP3.
</paragraph>
<paragraph id="940F7678FFB68A1AFF6E228F9597FC47" blockId="9.[151,1437,152,2034]" pageId="9" pageNumber="30">
<taxonomicName id="53B00DFBFFB68A1AFF6E228F9466FD21" ID-CoL="3F895" authority="Khalaji-Pirbalouty &amp; Sari, 2004" authorityName="Khalaji-Pirbalouty &amp; Sari" authorityYear="2004" box="[197,779,732,758]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="30" phylum="Arthropoda" rank="species" species="bakhteyaricus">
<emphasis id="A6C4AA6AFFB68A1AFF6E228F96E4FD22" box="[197,393,732,757]" italics="true" pageId="9" pageNumber="30">G. bakhteyaricus</emphasis>
<bibRefCitation id="F0210B89FFB68A1AFE3A228F9466FD21" author="Khalaji-Pirbalouty" box="[401,779,732,758]" pageId="9" pageNumber="39" refString="Khalaji-Pirbalouty, V. &amp; Sari, A. (2004) Biogeography of amphipods (Crustacea: Amphipoda: Gammaridae) from the central Zagros Mountains, Iran, with descriptions of two new species. Journal of Natural History, 38, 2425 - 2445." type="journal article" year="2004">Khalaji-Pirbalouty &amp; Sari, 2004</bibRefCitation>
</taxonomicName>
reported and described from the neighbouring province at North-West of the location has shorter setae on A2, and shorter setae and spines on telson. It has two rows of setae and a row of small spines on the palm face of GP1 and two rows of setae, in addition of small accessory spines, near the palmar angle spine in GP2, and a much sharper EP2. P57 without setae on merus and carpus, P7 basis has a higher length/width ratio.
</paragraph>
<paragraph id="940F7678FFB68A1AFF6E23CE94B5FB48" blockId="9.[151,1437,152,2034]" pageId="9" pageNumber="30">
<taxonomicName id="53B00DFBFFB68A1AFF6E23CE9475FC60" ID-CoL="3F8FC" authority="Khalaji-Pirbalouty &amp; Sari, 2004" authorityName="Khalaji-Pirbalouty &amp; Sari" authorityYear="2004" box="[197,792,925,951]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="30" phylum="Arthropoda" rank="species" species="lordeganensis">
<emphasis id="A6C4AA6AFFB68A1AFF6E23CE96E0FC61" box="[197,397,925,950]" italics="true" pageId="9" pageNumber="30">G. lordeganensis</emphasis>
<bibRefCitation id="F0210B89FFB68A1AFE3E23CE9475FC60" author="Khalaji-Pirbalouty" box="[405,792,925,951]" pageId="9" pageNumber="39" refString="Khalaji-Pirbalouty, V. &amp; Sari, A. (2004) Biogeography of amphipods (Crustacea: Amphipoda: Gammaridae) from the central Zagros Mountains, Iran, with descriptions of two new species. Journal of Natural History, 38, 2425 - 2445." type="journal article" year="2004">Khalaji-Pirbalouty &amp; Sari, 2004</bibRefCitation>
</taxonomicName>
reported and described from the same province as
<taxonomicName id="53B00DFBFFB68A1AFA2823CE9656FC0A" ID-CoL="3F895" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="30" phylum="Arthropoda" rank="species" species="bakhteyaricus">
<emphasis id="A6C4AA6AFFB68A1AFA2823CE9656FC0A" italics="true" pageId="9" pageNumber="30">G. bakhteyaricus</emphasis>
</taxonomicName>
has a higher number and length of setae on A2, shorter setae on both endo- and exopodite of U3 and no plumose setae on exopodite external margin, and shorter setae and spines on telson. Coxal plate of GP1 is much wider at distal than proximal; it has two rows of setae and a row of small spines on the palm face of GP1 and two rows of setae in addition of small accessory spines close to the palmar angle spine in GP2, sharper EP2 and EP3 and a specifically characteristic lobate shape of EP
<quantity id="5348DB9DFFB68A1AFC5C240D934EFBAF" box="[1015,1059,1118,1144]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="9" pageNumber="30" unit="in" value="3.0">3 in</quantity>
male and female; P57 bear no setae on merus and carpus, and P7 basis has a lower length/width ratio.
</paragraph>
<paragraph id="940F7678FFB68A1AFF6E24FF958DFAB7" blockId="9.[151,1437,152,2034]" pageId="9" pageNumber="30">
<taxonomicName id="53B00DFBFFB68A1AFF6E24FF9627FB12" box="[197,330,1196,1221]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="30" phylum="Arthropoda" rank="species" species="balutchi">
<emphasis id="A6C4AA6AFFB68A1AFF6E24FF9627FB12" box="[197,330,1196,1221]" italics="true" pageId="9" pageNumber="30">G. balutchi</emphasis>
</taxonomicName>
(
<bibRefCitation id="F0210B89FFB68A1AFEF224FF960BFB3A" pageId="9" pageNumber="39" refString="Khalaji-Pirbalouty, V. &amp; Sari, A. (2006) A Description of Gammarus balutchi n. sp. (Amphipoda: Gammaridae) from Iran, based on light and electron microscopy. Zoologische Mededelingen (Leiden), 80 1 (6), 91 - 100." type="journal article">Khalaji-Pirbalouty &amp; Sari, 2006 reported and described from Lorestan province (North to Sepidan County)</bibRefCitation>
show long setae on peduncles of A2, much shorter endopodite of U3, and not elevated epimere dorsal; it has two rows of setae and a row of small spines on the palm face of GP1 and two rows of setae in addition of small accessory spines near the palmar angle spine in GP2, P7 basis has a higher length/ width ratio, and no posterior-inferior corner setae.
</paragraph>
<paragraph id="940F7678FFB68A1AFF6E253E9506FA2C" blockId="9.[151,1437,152,2034]" pageId="9" pageNumber="30">
<taxonomicName id="53B00DFBFFB68A1AFF6E253E9570FA50" ID-CoL="6JWX6" authority="Stock et al., 1998" authorityName="Stock et al." authorityYear="1998" box="[197,541,1389,1415]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="30" phylum="Arthropoda" rank="species" species="lobifer">
<emphasis id="A6C4AA6AFFB68A1AFF6E253E9654FA51" box="[197,313,1389,1414]" italics="true" pageId="9" pageNumber="30">G. lobifer</emphasis>
<bibRefCitation id="F0210B89FFB68A1AFEEA253E9570FA50" author="Stock" box="[321,541,1389,1415]" pageId="9" pageNumber="39" refString="Stock, J. H., Mirzajani, A. R., Vonk, R., Naderi, S. &amp; Kiabi, B. H. (1998) Limnic and brackish water Amphipoda (Crustacea) from Iran. Beaufortia, 48 (9), 173 - 234." type="journal article" year="1998">
Stock
<emphasis id="A6C4AA6AFFB68A1AFE25253E96A8FA51" box="[398,453,1389,1414]" italics="true" pageId="9" pageNumber="30">et al</emphasis>
., 1998
</bibRefCitation>
</taxonomicName>
, firstly reported and described from the western adjacent province, has a shorter endopodite of U3, and dorsal of epimere is not elevated; the armature of GP1 and GP2 are similar to
<taxonomicName id="53B00DFBFFB68A1AFA2825C79791FA03" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="30" phylum="Arthropoda" rank="species" species="balutchi">
<emphasis id="A6C4AA6AFFB68A1AFA2825C79791FA03" italics="true" pageId="9" pageNumber="30">G. balutchi</emphasis>
</taxonomicName>
; basis of P7 with lower length/width ratio, and only one short posterior-inferior corner setae in contrast to 14 longer in
<taxonomicName id="53B00DFBFFB68A1AFE1725B29509FA2D" ID-CoL="3F8N7" box="[444,612,1505,1530]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="30" phylum="Arthropoda" rank="species" species="zagrosensis">
<emphasis id="A6C4AA6AFFB68A1AFE1725B29509FA2D" box="[444,612,1505,1530]" italics="true" pageId="9" pageNumber="30">G. zagrosensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="940F7678FFB68A1AFF6E265B94DDF8DD" blockId="9.[151,1437,152,2034]" pageId="9" pageNumber="30">
<taxonomicName id="53B00DFBFFB68A1AFF6E265B9551F9F5" authority="Martynov, 1935" authorityName="Martynov" authorityYear="1935" box="[197,572,1544,1570]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="30" phylum="Arthropoda" rank="species" species="subequalis">
<emphasis id="A6C4AA6AFFB68A1AFF6E265B9609F9F6" box="[197,356,1544,1569]" italics="true" pageId="9" pageNumber="30">G. subequalis</emphasis>
(
<bibRefCitation id="F0210B89FFB68A1AFEDF265B955EF9F5" author="Martynov" box="[372,563,1544,1570]" pageId="9" pageNumber="39" refString="Martynov, A. (1935) K poznaniyu Amphipoda tekuchikh vod Turkestana. Trudy Zoologicheskogo Instituta Akademii Nauk SSSR, 2 (2 - 3), 411 - 508." type="journal article" year="1935">Martynov, 1935</bibRefCitation>
)
</taxonomicName>
(Figs. 30-39) described from Karatau Mountains (Southern
<collectingCountry id="ECA736E8FFB68A1AFAA2265B92F9F9F5" box="[1289,1428,1544,1570]" name="Kazakhstan" pageId="9" pageNumber="30">Kazakhstan</collectingCountry>
) shares with
<taxonomicName id="53B00DFBFFB68A1AFE89267C96A7F99F" ID-CoL="3F8N7" box="[290,458,1583,1608]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="30" phylum="Arthropoda" rank="species" species="zagrosensis">
<emphasis id="A6C4AA6AFFB68A1AFE89267C96A7F99F" box="[290,458,1583,1608]" italics="true" pageId="9" pageNumber="30">G. zagrosensis</emphasis>
</taxonomicName>
similar setation of peduncle and flagellum of A2, armature of merus to propodus of P7 and richly setose U3 with similar ratio of endo-/exopodite. However, in contrast to
<taxonomicName id="53B00DFBFFB68A1AFB7B260692EFF9B9" ID-CoL="3F8N7" box="[1232,1410,1621,1646]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="30" phylum="Arthropoda" rank="species" species="zagrosensis">
<emphasis id="A6C4AA6AFFB68A1AFB7B260692EFF9B9" box="[1232,1410,1621,1646]" italics="true" pageId="9" pageNumber="30">G. zagrosensis</emphasis>
</taxonomicName>
it possesses less setose and longer (ca. 1/2 of total body length) A1, P7 basis with spine accompanied by few equally long setae at postero-interior corner, different shape of epimeral plates armed with spines and short setae only, and much shorter setae around apical spines on telson. Also,
<bibRefCitation id="F0210B89FFB68A1AFC52269A93ABF934" author="Martynov" box="[1017,1222,1737,1763]" pageId="9" pageNumber="39" refString="Martynov, A. (1935) K poznaniyu Amphipoda tekuchikh vod Turkestana. Trudy Zoologicheskogo Instituta Akademii Nauk SSSR, 2 (2 - 3), 411 - 508." type="journal article" year="1935">Martynov (1935)</bibRefCitation>
does not mention presence of setules on dorsal surface of metasomites in this species.
</paragraph>
<paragraph id="940F7678FFB68A1AFF6E274495B9F872" blockId="9.[151,1437,152,2034]" pageId="9" pageNumber="30">
<taxonomicName id="53B00DFBFFB68A1AFF6E27449542F8E7" ID-CoL="3F8LN" authority="Martynov, 1925" authorityName="Martynov" authorityYear="1925" box="[197,559,1814,1840]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="30" phylum="Arthropoda" rank="species" species="suifunensis">
<emphasis id="A6C4AA6AFFB68A1AFF6E27449605F8E7" box="[197,360,1815,1840]" italics="true" pageId="9" pageNumber="30">G. suifunensis</emphasis>
Martynov, 1925
</taxonomicName>
, reported from Ural Mountains,
<collectingCountry id="ECA736E8FFB68A1AFC0427459490F8E7" box="[943,1021,1814,1840]" name="Russia" pageId="9" pageNumber="30">Russia</collectingCountry>
(
<bibRefCitation id="F0210B89FFB68A1AFBA627459226F8E7" author="Karaman" box="[1037,1355,1814,1840]" pageId="9" pageNumber="39" refString="Karaman, G. S. (1991) The survey of described and cited freshwater Gammarus species (Fam. Gammaridae) from Soviet Union with redescription of two taxa. (Contribution to the knowledge of the Amphipoda 205). Poljoprivreda I Sumarstovo, XXXVII, 3 - 4, 37 - 73, Titograd." type="journal article" year="1991">Karaman 1991; Figs. IVI</bibRefCitation>
), has a similar weak setation of P67, very short U3 endopodite, but endopodite is much shorter (one-third of exopodite), gland cone is shorter than the segment, A2, P34, and U3 (exo- and endopodite) are poorly setose, and eyes are smaller compared to
<taxonomicName id="53B00DFBFFB68A1AFD8E27D895A1F873" ID-CoL="3F8N7" box="[549,716,1931,1956]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="30" phylum="Arthropoda" rank="species" species="zagrosensis">
<emphasis id="A6C4AA6AFFB68A1AFD8E27D895A1F873" box="[549,716,1931,1956]" italics="true" pageId="9" pageNumber="30">G. zagrosensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="DCAA25F3FFB68A19FF6E27E2957BFF28" lastPageId="10" lastPageNumber="31" pageId="9" pageNumber="30" type="description">
<paragraph id="940F7678FFB68A19FF6E27E2957BFF28" blockId="9.[151,1437,152,2034]" lastBlockId="10.[151,1437,152,915]" lastPageId="10" lastPageNumber="31" pageId="9" pageNumber="30">
<taxonomicName id="53B00DFBFFB68A1AFF6E27E29684F81C" authority="Ueno, 1940" authorityName="Ueno" authorityYear="1940" box="[197,489,1969,1995]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="30" phylum="Arthropoda" rank="species" species="koreanus">
<emphasis id="A6C4AA6AFFB68A1AFF6E27E2963FF81D" box="[197,338,1969,1994]" italics="true" pageId="9" pageNumber="30">G. koreanus</emphasis>
Ueno, 1940
</taxonomicName>
, originally described from NE
<collectingCountry id="ECA736E8FFB68A1AFCCB27E294C4F81C" box="[864,937,1969,1995]" name="South Korea" pageId="9" pageNumber="30">Korea</collectingCountry>
which is also reported from Khabarovsk, Far East
<collectingCountry id="ECA736E8FFB68A1AFEAA278B9622F825" box="[257,335,2008,2034]" name="Russia" pageId="9" pageNumber="30">Russia</collectingCountry>
(
<bibRefCitation id="F0210B89FFB68A1AFECB278B95D6F825" author="Karaman" box="[352,699,2008,2034]" pageId="9" pageNumber="39" refString="Karaman, G. S. (1991) The survey of described and cited freshwater Gammarus species (Fam. Gammaridae) from Soviet Union with redescription of two taxa. (Contribution to the knowledge of the Amphipoda 205). Poljoprivreda I Sumarstovo, XXXVII, 3 - 4, 37 - 73, Titograd." type="journal article" year="1991">Karaman, 1991, Figs. VIIX</bibRefCitation>
), shares the poor setation of P57, highly setose U3 (exo- and endopodite) and long endopodite, and setose P34 with
<taxonomicName id="53B00DFBFFB58A19FC8320CB94A2FF66" box="[808,975,152,177]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="31" phylum="Arthropoda" rank="species" species="zagrosensis">
<emphasis id="A6C4AA6AFFB58A19FC8320CB94A2FF66" box="[808,975,152,177]" italics="true" pageId="10" pageNumber="31">G. zagrosensis</emphasis>
</taxonomicName>
. On the other hand, G.
<taxonomicName id="53B00DFBFFB58A19FB4A20CB9226FF66" box="[1249,1355,152,177]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="31" phylum="Arthropoda" rank="species" species="koreanus">
<emphasis id="A6C4AA6AFFB58A19FB4A20CB9226FF66" box="[1249,1355,152,177]" italics="true" pageId="10" pageNumber="31">koreanus</emphasis>
</taxonomicName>
shows poorly setose A1 and A2 peduncles and flagellum, shorter gland cone, longer U3 endopodite, and seate on posterodorsal corner of P7 basis.
</paragraph>
</subSubSection>
<subSubSection id="DCAA25F3FFB58A19FF6E215F9632FD55" pageId="10" pageNumber="31" type="reference_group">
<paragraph id="940F7678FFB58A19FF6E215F935CFE4D" blockId="10.[151,1437,152,915]" pageId="10" pageNumber="31">
<taxonomicName id="53B00DFBFFB58A19FF6E215F952DFEF1" ID-CoL="6K995" authority="Hou &amp; Li, 2002" authorityName="Hou &amp; Li" authorityYear="2002" box="[197,576,268,294]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="31" phylum="Arthropoda" rank="species" species="paucispinus">
<emphasis id="A6C4AA6AFFB58A19FF6E215F9619FEF2" box="[197,372,268,293]" italics="true" pageId="10" pageNumber="31">G. paucispinus</emphasis>
<bibRefCitation id="F0210B89FFB58A19FED7215F952DFEF1" author="Hou" box="[380,576,268,294]" pageId="10" pageNumber="39" refString="Hou, Z. &amp; Li, S. (2002) Descriptions of two new species of the genus Gammarus (Crustacea: Amphipoda: Gammaridae) from Yunnan, China. The Raffles Bulletin Of Zoology. 50 (1), 37 - 52." type="journal article" year="2002">Hou &amp; Li, 2002</bibRefCitation>
</taxonomicName>
, distributed in Yunnan Province, South
<collectingCountry id="ECA736E8FFB58A19FBB7215F930BFEF1" box="[1052,1126,268,294]" name="China" pageId="10" pageNumber="31">China</collectingCountry>
, show similarities with
<taxonomicName id="53B00DFBFFB58A19FA28215F964FFE9C" ID-CoL="3F8N7" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="31" phylum="Arthropoda" rank="species" species="zagrosensis">
<emphasis id="A6C4AA6AFFB58A19FA28215F964FFE9C" italics="true" pageId="10" pageNumber="31">G. zagrosensis</emphasis>
</taxonomicName>
in its non-setose P57, having long setae on telson, non-pointed EP with small spines, and U3 endopodite 1/2 the exopodite. However, it has a poorly setose A1, less setose A2, shorter antennal gland cone, less setose P34, and poorely spinous urosomites (
<bibRefCitation id="F0210B89FFB58A19FD4C21D3934EFE4D" author="Hou" box="[743,1059,384,410]" pageId="10" pageNumber="39" refString="Hou, Z. &amp; Li, S. (2002) Descriptions of two new species of the genus Gammarus (Crustacea: Amphipoda: Gammaridae) from Yunnan, China. The Raffles Bulletin Of Zoology. 50 (1), 37 - 52." type="journal article" year="2002">
Hou &amp; Li, 2002;
<figureCitation id="0C8B6AFDFFB58A19FC1821D3934EFE4D" box="[947,1059,384,410]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="1.[151,254,1376,1400]" captionStartId-1="3.[151,254,1942,1966]" captionStartId-2="5.[151,254,2006,2030]" captionStartId-3="7.[151,257,1972,1996]" captionTargetBox-0="[173,1426,313,1341]" captionTargetBox-1="[214,1370,193,1913]" captionTargetBox-2="[191,1404,191,1980]" captionTargetBox-3="[212,1368,193,1938]" captionTargetId-0="figure@1.[165,1450,313,1351]" captionTargetId-1="figure@3.[194,1399,165,1937]" captionTargetId-2="figure@5.[185,1409,182,1982]" captionTargetId-3="figure@7.[192,1397,185,1957]" captionTargetPageId-0="1" captionTargetPageId-1="3" captionTargetPageId-2="5" captionTargetPageId-3="7" captionText-0="FIGURE 1. Map of sampling area of the western part of Zohre River catchments area, Sepidan County, Fars Province. Sampling sites (all springs): Cheshme Owsip (1); Vezge-Morad (2); Khani Varg (3); Baraghan (4); Shesh-Peer (5)." captionText-1="FIGURE 2. Gammarus zagrosensis n. sp., 3, 17.4 mm. A (HC), B (A 1), and C (A 2) on scale i; D (Mn) and E (MnPl) on scale ii, F (exopodite of Maxilipede), G (left palp of maxilla 1), and H (right palp of maxilla 1) on scale iii, and I (T) on scale iv." captionText-2="FIGURE 3. Gammarus zagrosensis n. sp., 3, 17.4 mm. A (G 1) and B (G 2) on scale i, C (P 3) and D (P 4) on scale ii." captionText-3="FIGURE 4. Gammarus zagrosensis n. sp., 3, 17.4 mm. A (P 5), B (P 6), and C (P 7), Ci and Cii (bases of P 7 in two paratypes) on scale i; D (US) and E (EP) on scale ii; F (U 3) on scale iii." httpUri-0="https://zenodo.org/record/188494/files/figure.png" httpUri-1="https://zenodo.org/record/188495/files/figure.png" httpUri-2="https://zenodo.org/record/188496/files/figure.png" httpUri-3="https://zenodo.org/record/188497/files/figure.png" pageId="10" pageNumber="31">Figs. 14</figureCitation>
</bibRefCitation>
).
</paragraph>
<paragraph id="940F7678FFB58A19FF6D21F59632FD55" blockId="10.[151,1437,152,915]" pageId="10" pageNumber="31">
Additionally, some shared features can be distinguished between
<taxonomicName id="53B00DFBFFB58A19FC7F21F49313FE17" ID-CoL="3F8N7" box="[980,1150,423,448]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="31" phylum="Arthropoda" rank="species" species="zagrosensis">
<emphasis id="A6C4AA6AFFB58A19FC7F21F49313FE17" box="[980,1150,423,448]" italics="true" pageId="10" pageNumber="31">G. zagrosensis</emphasis>
</taxonomicName>
and
<emphasis id="A6C4AA6AFFB58A19FB1021F49251FE17" box="[1211,1340,423,448]" italics="true" pageId="10" pageNumber="31">G. m a d i d u s</emphasis>
<bibRefCitation id="F0210B89FFB58A19FAE221F59796FE30" author="Hou" pageId="10" pageNumber="39" refString="Hou, Z. &amp; Li, S. (2005) Gammarus species from River Jumahe, China (Crustacea: Amphipoda: Gammaridae). Revue Suisse De Zoologie. 112 (2), 313 - 327." type="journal article" year="2005">Hou &amp; Li, 2005</bibRefCitation>
(from around Beijing,
<collectingCountry id="ECA736E8FFB58A19FDA0219E953BFE30" box="[523,598,461,487]" name="China" pageId="10" pageNumber="31">China</collectingCountry>
), including long setae on P4, bared P57, and short U3 endopodite, but
<taxonomicName id="53B00DFBFFB58A19FF3C21A79672FDDA" ID-CoL="3F8FL" box="[151,287,500,525]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="31" phylum="Arthropoda" rank="species" species="madidus">
<emphasis id="A6C4AA6AFFB58A19FF3C21A79672FDDA" box="[151,287,500,525]" italics="true" pageId="10" pageNumber="31">G. madidus</emphasis>
</taxonomicName>
can be easily recognized from the new species by its small eye, poorly setose A1, moderately setose A2, shorter gland cone, short setae on P3, postero-distal corner setae on P7 basis, non-pointed EP, U3 endopodite still shorter than 1/2 the length of exdopodite, and less seate on the U3 endopodite (
<bibRefCitation id="F0210B89FFB58A19FAB62212963CFD55" author="Hou" pageId="10" pageNumber="39" refString="Hou, Z. &amp; Li, S. (2005) Gammarus species from River Jumahe, China (Crustacea: Amphipoda: Gammaridae). Revue Suisse De Zoologie. 112 (2), 313 - 327." type="journal article" year="2005">
Hou &amp; Li, 2005;
<figureCitation id="0C8B6AFDFFB58A19FF4A223B963CFD55" box="[225,337,616,642]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="1.[151,254,1376,1400]" captionStartId-1="3.[151,254,1942,1966]" captionStartId-2="5.[151,254,2006,2030]" captionStartId-3="7.[151,257,1972,1996]" captionStartId-4="8.[151,254,1109,1133]" captionTargetBox-0="[173,1426,313,1341]" captionTargetBox-1="[214,1370,193,1913]" captionTargetBox-2="[191,1404,191,1980]" captionTargetBox-3="[212,1368,193,1938]" captionTargetBox-4="[172,1410,191,1075]" captionTargetId-0="figure@1.[165,1450,313,1351]" captionTargetId-1="figure@3.[194,1399,165,1937]" captionTargetId-2="figure@5.[185,1409,182,1982]" captionTargetId-3="figure@7.[192,1397,185,1957]" captionTargetId-4="figure@8.[151,1436,178,1085]" captionTargetPageId-0="1" captionTargetPageId-1="3" captionTargetPageId-2="5" captionTargetPageId-3="7" captionTargetPageId-4="8" captionText-0="FIGURE 1. Map of sampling area of the western part of Zohre River catchments area, Sepidan County, Fars Province. Sampling sites (all springs): Cheshme Owsip (1); Vezge-Morad (2); Khani Varg (3); Baraghan (4); Shesh-Peer (5)." captionText-1="FIGURE 2. Gammarus zagrosensis n. sp., 3, 17.4 mm. A (HC), B (A 1), and C (A 2) on scale i; D (Mn) and E (MnPl) on scale ii, F (exopodite of Maxilipede), G (left palp of maxilla 1), and H (right palp of maxilla 1) on scale iii, and I (T) on scale iv." captionText-2="FIGURE 3. Gammarus zagrosensis n. sp., 3, 17.4 mm. A (G 1) and B (G 2) on scale i, C (P 3) and D (P 4) on scale ii." captionText-3="FIGURE 4. Gammarus zagrosensis n. sp., 3, 17.4 mm. A (P 5), B (P 6), and C (P 7), Ci and Cii (bases of P 7 in two paratypes) on scale i; D (US) and E (EP) on scale ii; F (U 3) on scale iii." captionText-4="FIGURE 5. Gammarus zagrosensis n. sp., Ƥ, 12 mm. A (basis of P 5), B (basis of P 6), C (basis of P 7) and E (oostegite 1) on scale i; D (EP) on scale ii." httpUri-0="https://zenodo.org/record/188494/files/figure.png" httpUri-1="https://zenodo.org/record/188495/files/figure.png" httpUri-2="https://zenodo.org/record/188496/files/figure.png" httpUri-3="https://zenodo.org/record/188497/files/figure.png" httpUri-4="https://zenodo.org/record/188498/files/figure.png" pageId="10" pageNumber="31">Figs. 15</figureCitation>
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="DCAA25F3FFB58A19FF6E22DD9553FC44" pageId="10" pageNumber="31" type="description">
<paragraph id="940F7678FFB58A19FF6E22DD9553FC44" blockId="10.[151,1437,152,915]" pageId="10" pageNumber="31">
Comparison of cuticular ultra-structures shows strict differences between
<taxonomicName id="53B00DFBFFB58A19FBE322DC939BFD7F" box="[1096,1270,655,680]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="31" phylum="Arthropoda" rank="species" species="zagrosensis">
<emphasis id="A6C4AA6AFFB58A19FBE322DC939BFD7F" box="[1096,1270,655,680]" italics="true" pageId="10" pageNumber="31">G. zagrosensis</emphasis>
</taxonomicName>
and the three studied species. Previous studies have exclusively reported L-type pore arrangement for
<taxonomicName id="53B00DFBFFB58A19FB0622E697A0FD21" authority="Stype" authorityName="Stype" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="31" phylum="Arthropoda" rank="species" species="bakhteyaricus">
<emphasis id="A6C4AA6AFFB58A19FB0622E6921FFD19" box="[1197,1394,693,718]" italics="true" pageId="10" pageNumber="31">G. bakhteyaricus</emphasis>
, Stype
</taxonomicName>
for
<taxonomicName id="53B00DFBFFB58A19FEAF228F94E2FD21" authority="Khalaji-Pirbalouty &amp; Sari, 2004" authorityName="Khalaji-Pirbalouty &amp; Sari" authorityYear="2004" box="[260,911,732,758]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="31" phylum="Arthropoda" rank="species" species="lordeganensis">
<emphasis id="A6C4AA6AFFB58A19FEAF228F96BBFD22" box="[260,470,732,757]" italics="true" pageId="10" pageNumber="31">G. lordeganensis</emphasis>
(
<bibRefCitation id="F0210B89FFB58A19FE43228F94EBFD21" author="Khalaji-Pirbalouty" box="[488,902,732,758]" pageId="10" pageNumber="39" refString="Khalaji-Pirbalouty, V. &amp; Sari, A. (2004) Biogeography of amphipods (Crustacea: Amphipoda: Gammaridae) from the central Zagros Mountains, Iran, with descriptions of two new species. Journal of Natural History, 38, 2425 - 2445." type="journal article" year="2004">Khalaji-Pirbalouty &amp; Sari, 2004</bibRefCitation>
)
</taxonomicName>
, and also S-type for
<taxonomicName id="53B00DFBFFB58A19FB3C228F924FFD22" box="[1175,1314,732,757]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="31" phylum="Arthropoda" rank="species" species="balutchi">
<emphasis id="A6C4AA6AFFB58A19FB3C228F924FFD22" box="[1175,1314,732,757]" italics="true" pageId="10" pageNumber="31">G. balutchi</emphasis>
</taxonomicName>
(Khalaji-
<bibRefCitation id="F0210B89FFB58A19FF3C235196DBFCCB" author="Pirbalouty" box="[151,438,770,796]" pageId="10" pageNumber="39" refString="Khalaji-Pirbalouty, V. &amp; Sari, A. (2006) A Description of Gammarus balutchi n. sp. (Amphipoda: Gammaridae) from Iran, based on light and electron microscopy. Zoologische Mededelingen (Leiden), 80 1 (6), 91 - 100." type="journal article" year="2006">Pirbalouty &amp; Sari, 2006</bibRefCitation>
), while the new species shows both
<typeStatus id="4B0BC8DAFFB58A19FCC8235194CCFCCB" box="[867,929,770,796]" pageId="10" pageNumber="31">types</typeStatus>
of pore arrangements on head capsule. On the other hand, the densities of pore distribution in
<taxonomicName id="53B00DFBFFB58A19FD59237A94DBFC95" box="[754,950,809,834]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="31" phylum="Arthropoda" rank="species" species="lordeganensis">
<emphasis id="A6C4AA6AFFB58A19FD59237A94DBFC95" box="[754,950,809,834]" italics="true" pageId="10" pageNumber="31">G. lordeganensis</emphasis>
</taxonomicName>
and
<taxonomicName id="53B00DFBFFB58A19FC44237A93D9FC95" box="[1007,1204,809,834]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="31" phylum="Arthropoda" rank="species" species="bakhteyaricus">
<emphasis id="A6C4AA6AFFB58A19FC44237A93D9FC95" box="[1007,1204,809,834]" italics="true" pageId="10" pageNumber="31">G. bakhteyaricus</emphasis>
</taxonomicName>
have been reported to be 1820 and 4550 per 25 µm2, while they are found in much higher densities (
<quantity id="5348DB9DFFB58A19FB3423019263FCBB" box="[1183,1294,850,876]" metricMagnitude="0" metricUnit="m" metricValue="1.8034" metricValueMax="2.032" metricValueMin="1.5748" pageId="10" pageNumber="31" unit="in" value="71.0" valueMax="80.0" valueMin="62.0">6280 in</quantity>
L-type and
<quantity id="5348DB9DFFB58A19FF3C232A9660FC44" box="[151,269,889,915]" metricMagnitude="0" metricUnit="m" metricValue="4.0512999999999995" metricValueMax="6.1975999999999996" metricValueMin="1.905" pageId="10" pageNumber="31" unit="in" value="159.5" valueMax="244.0" valueMin="75.0">75244 in</quantity>
S-type) in
<taxonomicName id="53B00DFBFFB58A19FE3B232A955AFC45" box="[400,567,889,914]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="31" phylum="Arthropoda" rank="species" species="zagrosensis">
<emphasis id="A6C4AA6AFFB58A19FE3B232A955AFC45" box="[400,567,889,914]" italics="true" pageId="10" pageNumber="31">G. zagrosensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>