treatments-xml/data/1B/66/69/1B666942FFDDFF92FF59F8E7FEE65081.xml
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<document id="DE20E495E6118C664A811305CB83C88E" ID="10.11646/zootaxa.4247.4.3SLASH10667" ID-CLB-Dataset="33339" ID-DOI="10.11646/zootaxa.4247.4.3" ID-GBIF-Dataset="dfce4512-2062-4cd8-8ce7-bfe59cbbe6aa" ID-ISSN="1175-5326" ID-Zenodo-Dep="438865" ID-ZooBank="B926664F-8D4E-4CBA-8EEA-317B5AB46357" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1490772937598" checkinUser="plazi" docAuthor="Wang, Ying-Yong, Guo, Qiang, Liu, Zu-Yao, Lyu, Zhi-Tong, Wang, Jian, Luo, Lin, Sun, Yan-Jun &amp; Zhang, Yan-Wu" docDate="2017" docId="1B666942FFDDFF92FF59F8E7FEE65081" docLanguage="en" docName="zootaxa.4247.4.3SLASH10667.pdf" docOrigin="Zootaxa 4247 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Opisthotropis shenzhenensis Wang, Guo, Liu, Lyu, Wang, Luo, Sun &amp; Zhang, 2017, sp. nov." docType="treatment" docUuid="64396AA2-30E3-41CB-AAFA-25CB1A998863" docUuidSource="ZooBank" docVersion="9" lastPageNumber="405" masterDocId="E75F113AFFD6FF9CFFCEFFCAFFB95775" masterDocTitle="Revisions of two poorly known species of Opisthotropis Günther, 1872 (Squamata: Colubridae: Natricinae) with description of a new species from China" masterLastPageNumber="412" masterPageNumber="391" pageNumber="402" updateTime="1698438506206" updateUser="ExternalLinkService">
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<mods:title id="BBE8E6E9C464141885549408CC97C952">Revisions of two poorly known species of Opisthotropis Günther, 1872 (Squamata: Colubridae: Natricinae) with description of a new species from China</mods:title>
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<mods:namePart id="E58A9DE01361CAC22A08D334B15D8931">Wang, Ying-Yong</mods:namePart>
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<mods:namePart id="26D9E1BFCD497DA6FC4D46FF7F352B05">Guo, Qiang</mods:namePart>
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<mods:namePart id="FDC532F77FB3A9036B4E6797A71022CB">Liu, Zu-Yao</mods:namePart>
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<mods:namePart id="72DF6BAB46FD016BB57C6C00E425F8C4">Lyu, Zhi-Tong</mods:namePart>
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<mods:namePart id="4407D979454EEB9BCC63F67EE11824F2">Wang, Jian</mods:namePart>
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<mods:namePart id="50210446C295E95389793DFC16192F91">Luo, Lin</mods:namePart>
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<mods:namePart id="C3C5A20475907B33B3F0347760059690">Sun, Yan-Jun</mods:namePart>
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<mods:namePart id="86F1378175C9F793578B2A6CD08BF80B">Zhang, Yan-Wu</mods:namePart>
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<treatment id="1B666942FFDDFF92FF59F8E7FEE65081" ID-DOI="http://doi.org/10.5281/zenodo.5674723" ID-GBIF-Taxon="128009805" ID-Zenodo-Dep="5674723" LSID="urn:lsid:zoobank.org:act:64396AA2-30E3-41CB-AAFA-25CB1A998863" httpUri="http://treatment.plazi.org/id/1B666942FFDDFF92FF59F8E7FEE65081" lastPageId="14" lastPageNumber="405" pageId="11" pageNumber="402">
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<paragraph id="9370D854FFDDFF97FF59F8E7FDED5032" blockId="11.[151,596,1837,1896]" box="[151,596,1837,1863]" pageId="11" pageNumber="402">
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<taxonomicName id="54CFA3D7FFDDFF97FF59F8E7FE565032" box="[151,495,1837,1863]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="402" phylum="Chordata" rank="species" species="shenzhenensis" status="sp. nov.">
<emphasis id="A1BB0446FFDDFF97FF59F8E7FE565032" bold="true" box="[151,495,1837,1863]" italics="true" pageId="11" pageNumber="402">Opisthotropis shenzhenensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BA88B93DFFDDFF97FE38F8E7FDED5032" box="[502,596,1837,1863]" pageId="11" pageNumber="402" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
<paragraph id="9370D854FFDDFF97FF59F885FE8D501D" blockId="11.[151,596,1837,1896]" box="[151,308,1871,1896]" pageId="11" pageNumber="402">
<figureCitation id="0BF4C4D1FFDDFF97FF59F885FF5A501D" box="[151,227,1871,1896]" captionStart="FIGURE 5" captionStartId="13.[151,250,1362,1384]" captionTargetBox="[151,1436,447,1340]" captionTargetId="figure@13.[151,1436,445,1340]" captionTargetPageId="13" captionText="FIGURE 5. General aspect and scalation of head of Opisthotropis shenzhenensis sp. nov.. A: Dorsal view of the holotype SYS r 001018. B: Ventral view of the holotype. C: Dorsolateral view of head and anterior neck of the holotype in life. D: General aspect of SYS r 001145 in life. Photos by JW and JZ." httpUri="https://zenodo.org/record/438870/files/figure.png" pageId="11" pageNumber="402">Figs. 5</figureCitation>
and
<figureCitation id="0BF4C4D1FFDDFF97FED4F885FE9E501D" box="[282,295,1871,1896]" captionStart="FIGURE 6" captionStartId="14.[151,250,1596,1618]" captionTargetBox="[157,1415,864,1558]" captionTargetId="figure@14.[151,1435,844,1574]" captionTargetPageId="14" captionText="FIGURE 6. Comparisions of head scalation of a: Opisthotropis shenzhenensis sp. nov. (holotype SYS r 001018), b: O. maxwelli (SYS r 000841), c and d: O. andersonii (SYS r 001020, SYS r 001082). Line illustration by ZTL." httpUri="https://zenodo.org/record/438871/files/figure.png" pageId="11" pageNumber="402">6</figureCitation>
a
</paragraph>
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<paragraph id="9370D854FFDDFF97FF59F85DFB6450A0" blockId="11.[151,1437,1943,2005]" pageId="11" pageNumber="402">
<materialsCitation id="23A7D209FFDDFF97FF59F85DFB6450A0" ID-GBIF-Occurrence="1451078600" collectingDate="2015-09" collectionCode="SYS" collectorName="Jian Zhao &amp; Run-Lin Li" country="China" elevation="260" latitude="22.58189" location="Mt. Wutong" longLatPrecision="1" longitude="114.200745" municipality="Shenzhen City" pageId="11" pageNumber="402" specimenCount="1" stateProvince="Guangdong Province" typeStatus="holotype">
<emphasis id="A1BB0446FFDDFF97FF59F85DFEB250C5" bold="true" box="[151,267,1943,1968]" pageId="11" pageNumber="402">
<typeStatus id="4C7466F6FFDDFF97FF59F85DFEBF50C5" box="[151,262,1943,1968]" pageId="11" pageNumber="402" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="F5DE4091FFDDFF97FEDBF852FEF150C5" box="[277,328,1944,1968]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14836" name="Zhongshan (Sun Yatsen) University" pageId="11" pageNumber="402">SYS</collectionCode>
r001018, adult male, collected by
<collectorName id="3E3ABD82FFDDFF97FD2CF85DFCEA50C5" box="[738,851,1943,1968]" pageId="11" pageNumber="402">Jian Zhao</collectorName>
and
<collectorName id="3E3ABD82FFDDFF97FC5EF85DFBAB50C5" box="[912,1042,1943,1968]" pageId="11" pageNumber="402">Run-Lin Li</collectorName>
on
<date id="E771FE94FFDDFF97FB8DF85DFB5A50C5" box="[1091,1251,1943,1968]" pageId="11" pageNumber="402" value="2015-09">
<collectingDate id="F735077CFFDDFF97FB8DF85DFB5A50C5" box="[1091,1251,1943,1968]" pageId="11" pageNumber="402" value="2015-09">15 September</collectingDate>
</date>
, 2014 from
<location id="96108E8FFFDDFF97FABAF852FF4950A0" LSID="urn:lsid:plazi:treatment:1B666942FFDDFF92FF59F8E7FEE65081:96108E8FFFDDFF97FABAF852FF4950A0" country="China" latitude="22.58189" longLatPrecision="1" longitude="114.200745" municipality="Shenzhen City" name="Mt. Wutong" pageId="11" pageNumber="402" stateProvince="Guangdong Province">Mt. Wutong</location>
(
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,
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;
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<quantity id="543775B1FFDDFF97FD83F876FD2D50A1" box="[589,660,1980,2005]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="11" pageNumber="402" unit="m" value="260.0">260 m</quantity>
a.s.l.
</elevation>
),
<collectingMunicipality id="7314422EFFDDFF97FD28F876FC3250A0" box="[742,907,1980,2005]" pageId="11" pageNumber="402">Shenzhen City</collectingMunicipality>
,
<collectingRegion id="510B16B6FFDDFF97FC59F876FB3150A0" box="[919,1160,1980,2005]" country="China" name="Guangdong" pageId="11" pageNumber="402">Guangdong Province</collectingRegion>
,
<collectingCountry id="EBD898C4FFDDFF97FB5CF876FB6050A0" box="[1170,1241,1980,2005]" name="China" pageId="11" pageNumber="402">China</collectingCountry>
.
</materialsCitation>
</paragraph>
<paragraph id="9370D854FFDAFF90FF09FF5DFD2C56FD" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="403">
<emphasis id="A1BB0446FFDAFF90FF09FF5DFEFF57C5" bold="true" box="[199,326,151,176]" pageId="12" pageNumber="403">Paratypes.</emphasis>
<materialsCitation id="23A7D209FFDAFF90FE83FF52FEFE57A0" ID-GBIF-Occurrence="1451078601" collectingDate="2011-04" collectionCode="SYS" pageId="12" pageNumber="403" specimenCount="1" typeStatus="holotype">
<collectionCode id="F5DE4091FFDAFF90FE83FF52FE3957C5" box="[333,384,152,176]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14836" name="Zhongshan (Sun Yatsen) University" pageId="12" pageNumber="403">SYS</collectionCode>
r000635, an adult male, collected by JHY and RLL on
<date id="E771FE94FFDAFF90FC3AFF52FBEC57C5" box="[1012,1109,151,176]" pageId="12" pageNumber="403" value="2011-04">
<collectingDate id="F735077CFFDAFF90FC3AFF52FBEC57C5" box="[1012,1109,151,176]" pageId="12" pageNumber="403" value="2011-04">11 April</collectingDate>
</date>
, 2012 from the same locality as the
<typeStatus id="4C7466F6FFDAFF90FF2FFF76FEFE57A0" box="[225,327,188,213]" pageId="12" pageNumber="403" type="holotype">holotype</typeStatus>
</materialsCitation>
.
<materialsCitation id="23A7D209FFDAFF90FE9DFF76FDBE578D" ID-GBIF-Occurrence="1451078602" collectingDate="2011-04" collectionCode="SYS" pageId="12" pageNumber="403" specimenCount="1" typeStatus="holotype">
Three adult female specimens:
<collectionCode id="F5DE4091FFDAFF90FD78FF76FD5357A1" box="[694,746,188,212]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14836" name="Zhongshan (Sun Yatsen) University" pageId="12" pageNumber="403">SYS</collectionCode>
r000636 collected by JHY and RLL on
<date id="E771FE94FFDAFF90FB76FF76FAA057A0" box="[1208,1305,188,213]" pageId="12" pageNumber="403" value="2011-04">
<collectingDate id="F735077CFFDAFF90FB76FF76FAA057A0" box="[1208,1305,188,213]" pageId="12" pageNumber="403" value="2011-04">11 April</collectingDate>
</date>
, 2012 from the same locality as the
<typeStatus id="4C7466F6FFDAFF90FE6CFF15FDBE578D" box="[418,519,223,248]" pageId="12" pageNumber="403" type="holotype">holotype</typeStatus>
</materialsCitation>
;
<materialsCitation id="23A7D209FFDAFF90FDDCFF2AFCC85668" ID-GBIF-Occurrence="1451078603" collectingDate="1917-09" collectionCode="SYS" elevation="314" latitude="22.647821" location="Sanzhoutian" longLatPrecision="1" longitude="114.27558" municipality="Shenzhen City" pageId="12" pageNumber="403" specimenCount="1" typeStatus="holotype">
<collectionCode id="F5DE4091FFDAFF90FDDCFF2AFDFF578D" box="[530,582,224,248]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14836" name="Zhongshan (Sun Yatsen) University" pageId="12" pageNumber="403">SYS</collectionCode>
r001021 collected by JZ and RLL on
<date id="E771FE94FFDAFF90FC21FF2AFB32578D" box="[1007,1163,223,248]" pageId="12" pageNumber="403" value="1917-09">
<collectingDate id="F735077CFFDAFF90FC21FF2AFB32578D" box="[1007,1163,223,248]" pageId="12" pageNumber="403" value="1917-09">17 September</collectingDate>
</date>
, 2014 from
<location id="96108E8FFFDAFF90FADCFF15FA25578D" LSID="urn:lsid:plazi:treatment:1B666942FFDDFF92FF59F8E7FEE65081:96108E8FFFDAFF90FADCFF15FA25578D" box="[1298,1436,223,248]" latitude="22.647821" longLatPrecision="1" longitude="114.27558" municipality="Shenzhen City" name="Sanzhoutian" pageId="12" pageNumber="403">Sanzhoutian</location>
(
<geoCoordinate id="F6FBBE93FFDAFF90FF51FECEFEF25668" box="[159,331,260,285]" direction="north" orientation="latitude" pageId="12" pageNumber="403" precision="1" value="22.647821">22°3852.16″N</geoCoordinate>
,
<geoCoordinate id="F6FBBE93FFDAFF90FE95FECEFDA95668" box="[347,528,260,285]" direction="east" orientation="longitude" pageId="12" pageNumber="403" precision="1" value="114.27558">114°1632.08″E</geoCoordinate>
;
<elevation id="18E23F67FFDAFF90FDD1FECEFD135668" box="[543,682,260,285]" metricMagnitude="2" metricUnit="m" metricValue="3.1399999999999997" pageId="12" pageNumber="403" unit="m" value="314.0">
<quantity id="543775B1FFDAFF90FDD1FECEFDD25669" box="[543,619,260,284]" metricMagnitude="2" metricUnit="m" metricValue="3.1399999999999997" pageId="12" pageNumber="403" unit="m" value="314.0">314 m</quantity>
a.s.l.
</elevation>
),
<collectingMunicipality id="7314422EFFDAFF90FD0BFECEFCC85668" box="[709,881,260,285]" pageId="12" pageNumber="403">Shenzhen City</collectingMunicipality>
</materialsCitation>
;
<materialsCitation id="23A7D209FFDAFF90FC4EFECEFB4C5635" ID-GBIF-Occurrence="1451078604" collectingDate="1919-09" collectionCode="SYS" elevation="327" latitude="22.683819" location="Mt. Tiantou" longLatPrecision="1" longitude="114.404884" municipality="Shenzhen City" pageId="12" pageNumber="403" specimenCount="1" typeStatus="holotype">
<collectionCode id="F5DE4091FFDAFF90FC4EFECEFC0A5669" box="[896,947,260,284]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14836" name="Zhongshan (Sun Yatsen) University" pageId="12" pageNumber="403">SYS</collectionCode>
r001032 collected by JZ and RLL on
<date id="E771FE94FFDAFF90FA4EFECEFEA85635" pageId="12" pageNumber="403" value="1919-09">
<collectingDate id="F735077CFFDAFF90FA4EFECEFEA85635" pageId="12" pageNumber="403" value="1919-09">19 September</collectingDate>
</date>
, 2014 from
<location id="96108E8FFFDAFF90FE6EFEE2FD9E5635" LSID="urn:lsid:plazi:treatment:1B666942FFDDFF92FF59F8E7FEE65081:96108E8FFFDAFF90FE6EFEE2FD9E5635" box="[416,551,295,320]" latitude="22.683819" longLatPrecision="1" longitude="114.404884" municipality="Shenzhen City" name="Mt. Tiantou" pageId="12" pageNumber="403">Mt. Tiantou</location>
(
<geoCoordinate id="F6FBBE93FFDAFF90FDF7FEEDFD6E5635" box="[569,727,295,320]" direction="north" orientation="latitude" pageId="12" pageNumber="403" precision="1" value="22.683819">22°411.75″N</geoCoordinate>
,
<geoCoordinate id="F6FBBE93FFDAFF90FD28FEEDFC225635" box="[742,923,295,320]" direction="east" orientation="longitude" pageId="12" pageNumber="403" precision="1" value="114.404884">114°2417.58″E</geoCoordinate>
;
<elevation id="18E23F67FFDAFF90FC67FEE2FB885635" box="[937,1073,295,320]" metricMagnitude="2" metricUnit="m" metricValue="3.27" pageId="12" pageNumber="403" unit="m" value="327.0">
<quantity id="543775B1FFDAFF90FC67FEE2FC4A564A" box="[937,1011,296,320]" metricMagnitude="2" metricUnit="m" metricValue="3.27" pageId="12" pageNumber="403" unit="m" value="327.0">327 m</quantity>
a.s.l.
</elevation>
),
<collectingMunicipality id="7314422EFFDAFF90FB84FEEDFB4C5635" box="[1098,1269,295,320]" pageId="12" pageNumber="403">Shenzhen City</collectingMunicipality>
</materialsCitation>
;
<materialsCitation id="23A7D209FFDAFF90FACDFEE2FD2C56FD" ID-GBIF-Occurrence="1451078605" collectingDate="2013-04" collectionCode="SYS" country="China" elevation="155" latitude="22.877928" location="Mt. Yingping" longLatPrecision="1" longitude="114.15341" municipality="Dongguan City" pageId="12" pageNumber="403" specimenCount="1" stateProvince="Guangdong Province" typeStatus="holotype">
<collectionCode id="F5DE4091FFDAFF90FACDFEE2FA8F5635" box="[1283,1334,296,320]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14836" name="Zhongshan (Sun Yatsen) University" pageId="12" pageNumber="403">SYS</collectionCode>
r001145 collected by JW and ZTL on
<date id="E771FE94FFDAFF90FE3CFE86FDEF5610" box="[498,598,332,357]" pageId="12" pageNumber="403" value="2013-04">
<collectingDate id="F735077CFFDAFF90FE3CFE86FDEF5610" box="[498,598,332,357]" pageId="12" pageNumber="403" value="2013-04">13 April</collectingDate>
</date>
, 2015 from
<location id="96108E8FFFDAFF90FD29FE87FC395610" LSID="urn:lsid:plazi:treatment:1B666942FFDDFF92FF59F8E7FEE65081:96108E8FFFDAFF90FD29FE87FC395610" box="[743,896,332,357]" country="China" latitude="22.877928" longLatPrecision="1" longitude="114.15341" municipality="Dongguan City" name="Mt. Yingping" pageId="12" pageNumber="403" stateProvince="Guangdong Province">Mt. Yingping</location>
(
<geoCoordinate id="F6FBBE93FFDAFF90FC5DFE86FB865610" box="[915,1087,332,357]" direction="north" orientation="latitude" pageId="12" pageNumber="403" precision="1" value="22.877928">22°5240.54″N</geoCoordinate>
,
<geoCoordinate id="F6FBBE93FFDAFF90FB80FE86FB4C5610" box="[1102,1269,332,357]" direction="east" orientation="longitude" pageId="12" pageNumber="403" precision="1" value="114.15341">114°912.27″E</geoCoordinate>
;
<elevation id="18E23F67FFDAFF90FACAFE86FA355610" box="[1284,1420,332,357]" metricMagnitude="2" metricUnit="m" metricValue="1.55" pageId="12" pageNumber="403" unit="m" value="155.0">
<quantity id="543775B1FFDAFF90FACAFE86FAF75611" box="[1284,1358,332,357]" metricMagnitude="2" metricUnit="m" metricValue="1.55" pageId="12" pageNumber="403" unit="m" value="155.0">155 m</quantity>
a.s.l.
</elevation>
),
<collectingMunicipality id="7314422EFFDAFF90FF59FEBAFEFD56FD" box="[151,324,367,392]" pageId="12" pageNumber="403">Dongguan City</collectingMunicipality>
,
<collectingRegion id="510B16B6FFDAFF90FE81FEA5FD8656FD" box="[335,575,367,392]" country="China" name="Guangdong" pageId="12" pageNumber="403">Guangdong Province</collectingRegion>
,
<collectingCountry id="EBD898C4FFDAFF90FD84FEA5FD2856FD" box="[586,657,367,392]" name="China" pageId="12" pageNumber="403">China</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="DBD58BDFFFDAFF90FF09FE5EFDC35680" pageId="12" pageNumber="403" type="etymology">
<paragraph id="9370D854FFDAFF90FF09FE5EFDC35680" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="403">
<emphasis id="A1BB0446FFDAFF90FF09FE5EFEF756D8" bold="true" box="[199,334,404,429]" pageId="12" pageNumber="403">Etymology.</emphasis>
The specific name “
<taxonomicName id="54CFA3D7FFDAFF90FDFAFE5EFD6A56D9" box="[564,723,404,428]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="403" phylum="Chordata" rank="species" species="shenzhenensis">
<emphasis id="A1BB0446FFDAFF90FDFAFE5EFD6A56D9" box="[564,723,404,428]" italics="true" pageId="12" pageNumber="403">shenzhenensis</emphasis>
</taxonomicName>
” is derived from the
<typeStatus id="4C7466F6FFDAFF90FC0FFE5FFC4856D8" box="[961,1009,405,429]" pageId="12" pageNumber="403">type</typeStatus>
locality of the new species, Shenzhen City,
<collectingRegion id="510B16B6FFDAFF90FF1AFE7DFE7D56A5" box="[212,452,439,464]" country="China" name="Guangdong" pageId="12" pageNumber="403">Guangdong Province</collectingRegion>
,
<collectingCountry id="EBD898C4FFDAFF90FE1EFE7DFDAE56A5" box="[464,535,439,464]" name="China" pageId="12" pageNumber="403">China</collectingCountry>
. We propose the common name in English “Shenzhen Mountain Stream Snake” and “Shenzhen Hou Leng She” in Chinese.
</paragraph>
</subSubSection>
<subSubSection id="DBD58BDFFFDAFF90FF09FE35FDE7544D" pageId="12" pageNumber="403" type="diagnosis">
<paragraph id="9370D854FFDAFF90FF09FE35FDE7544D" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="403">
<emphasis id="A1BB0446FFDAFF90FF09FE35FEF8556D" bold="true" box="[199,321,511,536]" pageId="12" pageNumber="403">Diagnosis.</emphasis>
<taxonomicName id="54CFA3D7FFDAFF90FE86FDCAFD31556D" box="[328,648,512,536]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="403" phylum="Chordata" rank="species" species="shenzhenensis" status="sp. nov.">
<emphasis id="A1BB0446FFDAFF90FE86FDCAFD31556D" box="[328,648,512,536]" italics="true" pageId="12" pageNumber="403">Opisthotropis shenzhenensis</emphasis>
</taxonomicName>
<emphasis id="A1BB0446FFDAFF90FD41FE35FD50556D" bold="true" box="[655,745,511,536]" pageId="12" pageNumber="403">
<taxonomicNameLabel id="BA88B93DFFDAFF90FD41FE35FD50556D" box="[655,745,511,536]" pageId="12" pageNumber="403" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from other species of
<taxonomicName id="54CFA3D7FFDAFF90FBF6FDCAFB68556D" box="[1080,1233,512,536]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="403" phylum="Chordata" rank="genus">
<emphasis id="A1BB0446FFDAFF90FBF6FDCAFB68556D" box="[1080,1233,512,536]" italics="true" pageId="12" pageNumber="403">Opisthotropis</emphasis>
</taxonomicName>
by a combination of the following characters: (1) TL
<quantity id="543775B1FFDAFF90FDEAFDEEFD045549" box="[548,701,548,572]" metricMagnitude="-1" metricUnit="m" metricValue="3.7" metricValueMax="4.12" metricValueMin="3.28" pageId="12" pageNumber="403" unit="mm" value="370.0" valueMax="412.0" valueMin="328.0">328412 mm</quantity>
; (2) tail short, TaL 1619% of TL; (3) rostral, narrowed, RW 25 30% of HW; (4) nasal in contact with first, second and third supralabials; (5) nasal cleft invariably pointing to the middle of upper edge of second supralabial; (6) loreal 1.31.6 times as long as deep; (7) loreal not entering the orbit, not in contact with second supralabials in all known specimens; (8) supralabials 910, the last one smaller than adjacent preceding one; (9) infralabials 810; (10) dorsal scale rows 19:19:19; (11) dorsal scales keeled throughout, weakly keeled on neck, posteriorly strongly keeled; (12) ventrals 162179; subcaudals 5360; (13) olive-green above in life, each scale with black edge, forming fine mesh pattern; yellow beneath, ventral surface of head and tail mottled with blackish grey.
</paragraph>
</subSubSection>
<subSubSection id="DBD58BDFFFDAFF92FF09FC8EFEA156D8" lastPageId="14" lastPageNumber="405" pageId="12" pageNumber="403" type="description">
<paragraph id="9370D854FFDAFF90FF09FC8EFBE052B5" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="403">
<emphasis id="A1BB0446FFDAFF90FF09FC8EFE5E5428" bold="true" box="[199,487,836,861]" pageId="12" pageNumber="403">
Description of
<typeStatus id="4C7466F6FFDAFF90FEB7FC8EFE5B5428" box="[377,482,836,861]" pageId="12" pageNumber="403" type="holotype">holotype</typeStatus>
.
</emphasis>
Body cylindrical; TL
<quantity id="543775B1FFDAFF90FD29FC8EFCFF5429" box="[743,838,836,860]" metricMagnitude="-1" metricUnit="m" metricValue="4.07" pageId="12" pageNumber="403" unit="mm" value="407.0">407 mm</quantity>
(SVL
<quantity id="543775B1FFDAFF90FC5AFC8EFC4D5429" box="[916,1012,836,860]" metricMagnitude="-1" metricUnit="m" metricValue="3.37" pageId="12" pageNumber="403" unit="mm" value="337.0">337 mm</quantity>
, TaL
<quantity id="543775B1FFDAFF90FBF9FC8EFB325429" box="[1079,1163,836,860]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="12" pageNumber="403" unit="mm" value="70.0">70 mm</quantity>
); tail short, TaL 17% of TL; head small, indistinct from neck; rostral nearly flatted, small, slightly less than twice as broad as deep, RW 25 30% of HW, barely visible from above; two internasals, crescentic-shaped, in contact with each other medially behind the rostral, not in contact with loreal, posteriorly in contact with prefrontal; a single prefrontal, in contact with loreal, preocular and supraocular laterally, with frontal posteriorly; a single frontal, pentagonal, in contact with supraocular laterally, with two parietals posteriorly; parietals large, in contact with each other medially; nasal directed dorsolaterally, polygonal, in contact with first, second and third supralabials ventrally, with loreal and prefrontal posteriorly, with internasal dorsally, with rostral anteriorly; nostril horizontally oval, in the upper part of nasal; a short vertical cleft below the nostril and dividing nasal into anterior and posterior parts, pointing to the middle of upper edge of second supralabial; a single loreal, trapezoid, 1.4 times as long as deep, not entering the orbit, in contact with third and fourth supralabials, but not with second infralabial; a single supraocular; a single preocular; two postoculars; a single anterior temporal, significantly elongate; two posterior temporals; supralabials 9/9, the last one shorter than the adjacent preceding supralabial; fifth and sixth supralabials entering orbit; infralabials 9/9, the first one in contact with its fellow behind the mental; two pairs of chin shields; anterior chinshields large, in contact with each other medially, and in contact with the first five infralabials on both sides; posterior chin shields small and separated from each other by small scales; dorsal scale rows 19:19:19; dorsal scales weakly keeled on neck, posteriorly gradually strongly keeled; the outermost a dorsal scale row smooth at midbody, those following keeled; V 172; cloacal plate divided; subcaudals 53, paired.
</paragraph>
<paragraph id="9370D854FFDAFF90FF09FA06FC5D5158" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="403">
<emphasis id="A1BB0446FFDAFF90FF09FA06FE185290" bold="true" box="[199,417,1484,1509]" pageId="12" pageNumber="403">Coloration in life.</emphasis>
Olive-green above, each scale with black edge, forming a fine mesh pattern; the outer second and third scale rows with yellow spots, upper part of the most outer scale row dark green, lower part yellow; yellow beneath, ventral surface of head and tail mottled with blackish-grey.
</paragraph>
<paragraph id="9370D854FFDAFF90FF09F9FDFA245125" blockId="12.[151,1437,151,2013]" box="[199,1437,1591,1616]" pageId="12" pageNumber="403">
<emphasis id="A1BB0446FFDAFF90FF09F9FDFE745125" bold="true" box="[199,461,1591,1616]" pageId="12" pageNumber="403">Coloration in ethanol.</emphasis>
Greenish-black above, barely perceptible black edge on each scale; yellowish beneath.
</paragraph>
<paragraph id="9370D854FFDAFF90FF09F996FB765195" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="403">
<emphasis id="A1BB0446FFDAFF90FF09F996FE865100" bold="true" box="[199,319,1628,1653]" pageId="12" pageNumber="403">Variation.</emphasis>
Measurements, scalation and body proportions of
<typeStatus id="4C7466F6FFDAFF90FC68F997FC6F5100" box="[934,982,1629,1653]" pageId="12" pageNumber="403">type</typeStatus>
series of
<taxonomicName id="54CFA3D7FFDAFF90FB98F996FA255101" box="[1110,1436,1628,1652]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="403" phylum="Chordata" rank="species" species="shenzhenensis" status="sp. nov.">
<emphasis id="A1BB0446FFDAFF90FB98F996FA255101" box="[1110,1436,1628,1652]" italics="true" pageId="12" pageNumber="403">Opisthotropis shenzhenensis</emphasis>
</taxonomicName>
<emphasis id="A1BB0446FFDAFF90FF59F9B5FF5051ED" bold="true" box="[151,233,1663,1688]" pageId="12" pageNumber="403">
<taxonomicNameLabel id="BA88B93DFFDAFF90FF59F9B5FF5051ED" box="[151,233,1663,1688]" pageId="12" pageNumber="403" rank="species">sp.nov.</taxonomicNameLabel>
</emphasis>
are given in Table 5. In specimen SYS r001032, anterior chin-shields in contact with first six infralabials in both sides. In the SYS r001021, only fifth supralabial entering the orbit in both sides. In the SYS r001032 and 1145, fifth and six supralabials entering the orbit in left side, fifth entering the orbit in right side.
</paragraph>
<paragraph id="9370D854FFDAFF91FF09F926FAD456FD" blockId="12.[151,1437,151,2013]" lastBlockId="13.[151,1436,151,392]" lastPageId="13" lastPageNumber="404" pageId="12" pageNumber="403">
<emphasis id="A1BB0446FFDAFF90FF09F926FED45070" bold="true" box="[199,365,1772,1797]" pageId="12" pageNumber="403">Comparisons.</emphasis>
<taxonomicName id="54CFA3D7FFDAFF90FEB9F926FD015071" box="[375,696,1772,1796]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="403" phylum="Chordata" rank="species" species="shenzhenensis" status="sp. nov.">
<emphasis id="A1BB0446FFDAFF90FEB9F926FD015071" box="[375,696,1772,1796]" italics="true" pageId="12" pageNumber="403">Opisthotropis shenzhenensis</emphasis>
</taxonomicName>
<emphasis id="A1BB0446FFDAFF90FD0DF926FC995070" bold="true" box="[707,800,1772,1797]" pageId="12" pageNumber="403">
<taxonomicNameLabel id="BA88B93DFFDAFF90FD0DF926FC995070" box="[707,800,1772,1797]" pageId="12" pageNumber="403" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is mostly similar to
<taxonomicName id="54CFA3D7FFDAFF90FBD8F926FB0D5071" box="[1046,1204,1772,1796]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="403" phylum="Chordata" rank="species" species="andersonii">
<emphasis id="A1BB0446FFDAFF90FBD8F926FB0D5071" box="[1046,1204,1772,1796]" italics="true" pageId="12" pageNumber="403">O. andersonii</emphasis>
</taxonomicName>
and
<taxonomicName id="54CFA3D7FFDAFF90FB3FF926FAC55071" box="[1265,1404,1772,1796]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="403" phylum="Chordata" rank="species" species="maxwelli">
<emphasis id="A1BB0446FFDAFF90FB3FF926FAC55071" box="[1265,1404,1772,1796]" italics="true" pageId="12" pageNumber="403">O. maxwelli</emphasis>
</taxonomicName>
in habitus, color and pattern. In our phylogenetic tree, the new species is the sister taxon of
<taxonomicName id="54CFA3D7FFDAFF90FBBAF8DAFAB4505D" box="[1140,1293,1808,1832]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="403" phylum="Chordata" rank="species" species="andersonii">
<emphasis id="A1BB0446FFDAFF90FBBAF8DAFAB4505D" box="[1140,1293,1808,1832]" italics="true" pageId="12" pageNumber="403">O. andersonii</emphasis>
</taxonomicName>
, from which it differs in having (1) DSR 19:19:19 vs. 17:17:17; (2) dorsal scales keeled throughout, weakly keeled on neck, posteriorly strongly keeled vs. smooth on the anterior neck, posteriorly feebly keeled, rather strongly keeled on posterior part of body; (3) rostral small, RW 2530% of HW vs. rostral slightly large, RW 3137% of HW; (4) nasal in contact with first, second and third supralabials vs. only with first and second, not in contact with third; (5) loreal 1.31.6 times as long as deep vs. 2.12.9 times as long as deep; (6) loreal not in contact with second supralabial vs. loreal in contact with second supralabial.
<taxonomicName id="54CFA3D7FFDBFF91FCF9FF52FBC257C5" box="[823,1147,152,176]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="404" phylum="Chordata" rank="species" species="shenzhenensis" status="sp. nov.">
<emphasis id="A1BB0446FFDBFF91FCF9FF52FBC257C5" box="[823,1147,152,176]" italics="true" pageId="13" pageNumber="404">Opisthotropis shenzhenensis</emphasis>
</taxonomicName>
<emphasis id="A1BB0446FFDBFF91FB48FF5DFB6657C5" bold="true" box="[1158,1247,151,176]" pageId="13" pageNumber="404">
<taxonomicNameLabel id="BA88B93DFFDBFF91FB48FF5DFB6657C5" box="[1158,1247,151,176]" pageId="13" pageNumber="404" rank="species">sp. nov</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="54CFA3D7FFDBFF91FA4FFF52FF4557A1" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="404" phylum="Chordata" rank="species" species="maxwelli">
<emphasis id="A1BB0446FFDBFF91FA4FFF52FF4557A1" italics="true" pageId="13" pageNumber="404">O. maxwelli</emphasis>
</taxonomicName>
in having (1) DSR 19:19:19 vs. 17:17:17; (2) tail short, TaL 1619% of TL vs. TaL 2023% of TL; (3) the last supralabial smaller than adjacent preceding one vs. the last supralabial is the longest one; (4) dorsal scales keeled throughout, weakly keeled on neck, posteriorly strongly keeled vs. smooth on the anterior neck, feebly keeled posteriorly, rather strongly keeled on posterior part of body; (5) nasal cleft invariably pointing to the middle of upper edge of second supralabial vs. invariable pointing to the apex angle of first supralabial; (6) nasal in contact with first, second and third supralabials vs. in contact only with first and second, but not with third. See
<figureCitation id="0BF4C4D1FFDBFF91FAEBFEA5FAD056FD" box="[1317,1385,367,392]" captionStart="FIGURE 6" captionStartId="14.[151,250,1596,1618]" captionTargetBox="[157,1415,864,1558]" captionTargetId="figure@14.[151,1435,844,1574]" captionTargetPageId="14" captionText="FIGURE 6. Comparisions of head scalation of a: Opisthotropis shenzhenensis sp. nov. (holotype SYS r 001018), b: O. maxwelli (SYS r 000841), c and d: O. andersonii (SYS r 001020, SYS r 001082). Line illustration by ZTL." httpUri="https://zenodo.org/record/438871/files/figure.png" pageId="13" pageNumber="404">Fig. 6</figureCitation>
.
</paragraph>
<caption id="C7B088DCFFDBFF91FF59FA98FD1452D0" httpUri="https://zenodo.org/record/438870/files/figure.png" pageId="13" pageNumber="404" targetBox="[151,1436,447,1340]" targetPageId="13">
<paragraph id="9370D854FFDBFF91FF59FA98FD1452D0" blockId="13.[151,1436,1362,1445]" pageId="13" pageNumber="404">
<emphasis id="A1BB0446FFDBFF91FF59FA98FEAB521D" bold="true" box="[151,274,1362,1384]" pageId="13" pageNumber="404">FIGURE 5.</emphasis>
General aspect and scalation of head of
<taxonomicName id="54CFA3D7FFDBFF91FD64FA98FC73521D" box="[682,970,1362,1384]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="404" phylum="Chordata" rank="species" species="shenzhenensis" status="sp. nov.">
<emphasis id="A1BB0446FFDBFF91FD64FA98FC73521D" box="[682,970,1362,1384]" italics="true" pageId="13" pageNumber="404">Opisthotropis shenzhenensis</emphasis>
</taxonomicName>
<emphasis id="A1BB0446FFDBFF91FC1EFA98FB99521D" bold="true" box="[976,1056,1362,1384]" pageId="13" pageNumber="404">
<taxonomicNameLabel id="BA88B93DFFDBFF91FC1EFA98FB99521D" box="[976,1056,1362,1384]" pageId="13" pageNumber="404" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
.
<emphasis id="A1BB0446FFDBFF91FBE3FA98FBFE521D" bold="true" box="[1069,1095,1362,1384]" pageId="13" pageNumber="404">A:</emphasis>
Dorsal view of the holotype SYS r001018.
<emphasis id="A1BB0446FFDBFF91FF37FABBFEAA52F2" bold="true" box="[249,275,1393,1415]" pageId="13" pageNumber="404">B:</emphasis>
Ventral view of the holotype.
<emphasis id="A1BB0446FFDBFF91FD83FABBFDD052F2" bold="true" box="[589,617,1393,1415]" pageId="13" pageNumber="404">C:</emphasis>
Dorsolateral view of head and anterior neck of the holotype in life.
<emphasis id="A1BB0446FFDBFF91FAE4FABBFAFD52F2" bold="true" box="[1322,1348,1393,1415]" pageId="13" pageNumber="404">D:</emphasis>
General aspect of SYS r001145 in life. Photos by JW and JZ.
</paragraph>
</caption>
<paragraph id="9370D854FFDBFF91FF09FA07FD465194" blockId="13.[151,1436,1484,2014]" pageId="13" pageNumber="404">
<taxonomicName id="54CFA3D7FFDBFF91FF09FA07FDB05290" box="[199,521,1485,1509]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="404" phylum="Chordata" rank="species" species="shenzhenensis" status="sp. nov.">
<emphasis id="A1BB0446FFDBFF91FF09FA07FDB05290" box="[199,521,1485,1509]" italics="true" pageId="13" pageNumber="404">Opisthotropis shenzhenensis</emphasis>
</taxonomicName>
<emphasis id="A1BB0446FFDBFF91FDDDFA06FDC95290" bold="true" box="[531,624,1484,1509]" pageId="13" pageNumber="404">
<taxonomicNameLabel id="BA88B93DFFDBFF91FDDDFA06FDC95290" box="[531,624,1484,1509]" pageId="13" pageNumber="404" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="54CFA3D7FFDBFF91FD4CFA07FCA65290" box="[642,799,1485,1509]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="404" phylum="Chordata" rank="species" species="andersonii">
<emphasis id="A1BB0446FFDBFF91FD4CFA07FCA65290" box="[642,799,1485,1509]" italics="true" pageId="13" pageNumber="404">O. andersonii</emphasis>
</taxonomicName>
,
<taxonomicName id="54CFA3D7FFDBFF91FCFFFA07FC0A5290" box="[817,947,1485,1509]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="404" phylum="Chordata" rank="species" species="lateralis">
<emphasis id="A1BB0446FFDBFF91FCFFFA07FC0A5290" box="[817,947,1485,1509]" italics="true" pageId="13" pageNumber="404">O. lateralis</emphasis>
</taxonomicName>
and
<taxonomicName id="54CFA3D7FFDBFF91FC3FFA07FB225290" box="[1009,1179,1485,1509]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="404" phylum="Chordata" rank="species" species="kuatunensis">
<emphasis id="A1BB0446FFDBFF91FC3FFA07FB225290" box="[1009,1179,1485,1509]" italics="true" pageId="13" pageNumber="404">O. kuatunensis</emphasis>
</taxonomicName>
are sympatric species from Mt. Wutong, Shenzhen.
<taxonomicName id="54CFA3D7FFDBFF91FE25FA3BFD14517C" box="[491,685,1521,1545]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="404" phylum="Chordata" rank="species" species="shenzhenensis">
<emphasis id="A1BB0446FFDBFF91FE25FA3BFD14517C" box="[491,685,1521,1545]" italics="true" pageId="13" pageNumber="404">O. shenzhenensis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="54CFA3D7FFDBFF91FC8BFA3BFC7F517C" box="[837,966,1521,1545]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="404" phylum="Chordata" rank="species" species="lateralis">
<emphasis id="A1BB0446FFDBFF91FC8BFA3BFC7F517C" box="[837,966,1521,1545]" italics="true" pageId="13" pageNumber="404">O. lateralis</emphasis>
</taxonomicName>
by having DSR 19:19:19 vs. 17:17:17; a single preocular vs. two preoculars; uniformly olive-green above with black mesh pattern on dorsal surface of body and tail vs. brown above, with a distinct longitudinal black stripe on both sides of body.
<taxonomicName id="54CFA3D7FFDBFF91FB47F9F3FAF25124" box="[1161,1355,1593,1617]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="404" phylum="Chordata" rank="species" species="shenzhenensis">
<emphasis id="A1BB0446FFDBFF91FB47F9F3FAF25124" box="[1161,1355,1593,1617]" italics="true" pageId="13" pageNumber="404">O. shenzhenensis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="54CFA3D7FFDBFF91FF1AF997FEC05100" box="[212,377,1629,1653]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="404" phylum="Chordata" rank="species" species="kuatunensis">
<emphasis id="A1BB0446FFDBFF91FF1AF997FEC05100" box="[212,377,1629,1653]" italics="true" pageId="13" pageNumber="404">O. kuatunensis</emphasis>
</taxonomicName>
by having 910 supralabials, all of which entire vs. 1316 supralabials, first six to eight entire, posterior ones divided horizontally; dorsal scales weakly keeled on neck, strongly keeled on body and tail vs. extremely strongly keeled throughout; olive-green above with black mesh pattern on dorsal body and tail vs. brown above, with black longitudinal stripes on body and tail.
</paragraph>
<paragraph id="9370D854FFDBFF92FF09F927FEA156D8" blockId="13.[151,1436,1484,2014]" lastBlockId="14.[151,1436,151,824]" lastPageId="14" lastPageNumber="405" pageId="13" pageNumber="404">
<taxonomicName id="54CFA3D7FFDBFF91FF09F927FDBE5070" box="[199,519,1773,1797]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="404" phylum="Chordata" rank="species" species="shenzhenensis" status="sp. nov.">
<emphasis id="A1BB0446FFDBFF91FF09F927FDBE5070" box="[199,519,1773,1797]" italics="true" pageId="13" pageNumber="404">Opisthotropis shenzhenensis</emphasis>
</taxonomicName>
<emphasis id="A1BB0446FFDBFF91FDC1F927FDD35073" bold="true" box="[527,618,1773,1798]" pageId="13" pageNumber="404">
<taxonomicNameLabel id="BA88B93DFFDBFF91FDC1F927FDD35073" box="[527,618,1773,1798]" pageId="13" pageNumber="404" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be easily distinguished from other Chinese congeners as follows: It differs from
<taxonomicName id="54CFA3D7FFDBFF91FEE8F8DBFE15505C" box="[294,428,1809,1833]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="404" phylum="Chordata" rank="species" species="latouchii">
<emphasis id="A1BB0446FFDBFF91FEE8F8DBFE15505C" box="[294,428,1809,1833]" italics="true" pageId="13" pageNumber="404">O. latouchii</emphasis>
</taxonomicName>
in having DSR 19:19:19 vs. 17:17:17; loreal not entering the orbit vs. entering the orbit; olive-green above with black mesh pattern vs. yellowish green above, with alternate longitudinal black and yellow stripes. It differs from
<taxonomicName id="54CFA3D7FFDBFF91FE5BF893FE4C5004" box="[405,501,1881,1905]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="404" phylum="Chordata" rank="species" species="cheni">
<emphasis id="A1BB0446FFDBFF91FE5BF893FE4C5004" box="[405,501,1881,1905]" italics="true" pageId="13" pageNumber="404">O. cheni</emphasis>
</taxonomicName>
in having DSR 19:19:19 vs. 17:17:17; loreal not entering the orbit vs. entering the orbit; olive-green above with black mesh pattern vs. dark olive brown above, with yellow crossbars or marks. It differs from
<taxonomicName id="54CFA3D7FFDBFF91FEEDF86BFEC950CC" box="[291,368,1953,1977]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="404" phylum="Chordata" rank="species" species="laui">
<emphasis id="A1BB0446FFDBFF91FEEDF86BFEC950CC" box="[291,368,1953,1977]" italics="true" pageId="13" pageNumber="404">O. laui</emphasis>
</taxonomicName>
in having DSR 19:19:19 vs. 25:23:23; olive-green above with black mesh pattern vs. dark olive above, with pale yellow crossbars. It differs from
<taxonomicName id="54CFA3D7FFDBFF91FD71F80FFC8450A8" box="[703,829,1989,2013]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="404" phylum="Chordata" rank="species" species="balteata">
<emphasis id="A1BB0446FFDBFF91FD71F80FFC8450A8" box="[703,829,1989,2013]" italics="true" pageId="13" pageNumber="404">O. balteata</emphasis>
</taxonomicName>
in having DSR 19:19:19 vs. 19:19:17; V 162179, SC 5360 vs.V 190205, SC 5686; dorsal scales keeled throughout, weakly keeled on neck, posteriorly strongly keeled vs. smooth anteriorly, keeled posteriorly; olive-green above with black mesh pattern on dorsal body and tail vs. yellow or orange-yellow above, with at least 40 pairs of black bands. It differs from
<taxonomicName id="54CFA3D7FFD8FF92FBB8FF2AFA935782" box="[1142,1322,224,247]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="guangxiensis">
<emphasis id="A1BB0446FFD8FF92FBB8FF2AFA935782" box="[1142,1322,224,247]" italics="true" pageId="14" pageNumber="405">O. guangxiensis</emphasis>
</taxonomicName>
in having DSR 19:19:19 vs.17:15:15; dorsal scales keeled vs. smooth; olive-green above with black mesh pattern vs. olivebrown above, with narrow light yellow crossbars. It differs from
<taxonomicName id="54CFA3D7FFD8FF92FCA6FEE2FC4D5635" box="[872,1012,296,320]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="maculosa">
<emphasis id="A1BB0446FFD8FF92FCA6FEE2FC4D5635" box="[872,1012,296,320]" italics="true" pageId="14" pageNumber="405">O. maculosa</emphasis>
</taxonomicName>
in having DSR 19:19:19 vs.15:15:15; dorsal scales keeled vs. smooth; olive-green above, each scale with black edge vs. olive-brown above, each scale with single yellow spot. Lastly, it differs from
<taxonomicName id="54CFA3D7FFD8FF92FD6FFEBAFCB356FD" box="[673,778,368,392]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="jacobi">
<emphasis id="A1BB0446FFD8FF92FD6FFEBAFCB356FD" box="[673,778,368,392]" italics="true" pageId="14" pageNumber="405">O. jacobi</emphasis>
</taxonomicName>
in having DSR 19:19:19 vs.15:15:15; dorsal scales keeled vs. smooth.
</paragraph>
</subSubSection>
<subSubSection id="DBD58BDFFFD8FF92FF09FE72FAA55442" pageId="14" pageNumber="405" type="reference_group">
<paragraph id="9370D854FFD8FF92FF09FE72FAA55442" blockId="14.[151,1436,151,824]" pageId="14" pageNumber="405">
<taxonomicName id="54CFA3D7FFD8FF92FF09FE72FDB256A5" box="[199,523,440,464]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="shenzhenensis">
<emphasis id="A1BB0446FFD8FF92FF09FE72FDB256A5" box="[199,523,440,464]" italics="true" pageId="14" pageNumber="405">Opisthotropis shenzhenensis</emphasis>
</taxonomicName>
differs from the other 10 congeners not occurring in
<collectingCountry id="EBD898C4FFD8FF92FB54FE7DFB6456A5" box="[1178,1245,439,464]" name="China" pageId="14" pageNumber="405">China</collectingCountry>
in having DSR 19:19:19 vs. DSR 17:17:
<quantity id="543775B1FFD8FF92FE7BFE16FE4D5680" box="[437,500,476,501]" metricMagnitude="-1" metricUnit="m" metricValue="4.318" pageId="14" pageNumber="405" unit="in" value="17.0">17 in</quantity>
<taxonomicName id="54CFA3D7FFD8FF92FE30FE16FB855681" authority="Orlov, Darevsky &amp; Murphy, 1998" authorityName="Orlov, Darevsky &amp; Murphy" authorityYear="1998" box="[510,1084,476,501]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="daovantieni">
<emphasis id="A1BB0446FFD8FF92FE30FE16FD115681" box="[510,680,476,500]" italics="true" pageId="14" pageNumber="405">O. daovantieni</emphasis>
<bibRefCitation id="F75EA5A5FFD8FF92FD7DFE16FB855681" author="Orlov" box="[691,1084,476,501]" pageId="14" pageNumber="405" refString="Orlov, N., Darevsky, I. S. &amp; Murphy, R. W. (1998) A new species of mountain stream snake, genus Opisthotrophis Gunther, 1872 (Serpentes: Colubridae: Natricinae) from the tropical rain forests of southern Vietnam. Russian Journal of Herpetology, 5 (1), 61 - 64." type="journal article" year="1998">Orlov, Darevsky &amp; Murphy, 1998</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="54CFA3D7FFD8FF92FBB7FE16FA215681" authority="Smith, 1918" authorityName="Smith" authorityYear="1918" box="[1145,1432,476,501]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="spenceri">
<emphasis id="A1BB0446FFD8FF92FBB7FE16FB475681" box="[1145,1278,476,500]" italics="true" pageId="14" pageNumber="405">O. spenceri</emphasis>
<bibRefCitation id="F75EA5A5FFD8FF92FAC6FE16FA215681" author="Smith" box="[1288,1432,476,501]" pageId="14" pageNumber="405" refString="Smith, M. A. (1918) Description of a new snake (Opisthotropis spenceri) from Siam. The Journal of the Natural History Society of Siam, 3 (1), 13." type="journal article" year="1918">Smith, 1918</bibRefCitation>
</taxonomicName>
, 19:17:
<quantity id="543775B1FFD8FF92FF12FDCAFEA0556D" box="[220,281,511,536]" metricMagnitude="-1" metricUnit="m" metricValue="4.318" pageId="14" pageNumber="405" unit="in" value="17.0">17 in</quantity>
<taxonomicName id="54CFA3D7FFD8FF92FEEFFDCAFCB6556D" authority="Ziegler, David &amp; Vu, 2008" authorityName="Ziegler, David &amp; Vu" authorityYear="2008" box="[289,783,511,536]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="tamdaoensis">
<emphasis id="A1BB0446FFD8FF92FEEFFDCAFE68556D" box="[289,465,512,536]" italics="true" pageId="14" pageNumber="405">O. tamdaoensis</emphasis>
Ziegler,
<bibRefCitation id="F75EA5A5FFD8FF92FDF7FE35FCB6556D" author="David" box="[569,783,511,536]" pageId="14" pageNumber="405" refString="Ziegler, T., David, P. &amp; Thana, V. N. (2008) A new natricine snake of the genus Opisthotropis from Tam Dao, Vinh Phuc Province, northern Vietnam (Squamata, Colubridae). Zoosystematics and Evolution, 84, 197 - 203. https: // doi. org / 10.1002 / zoos. 200800004" type="journal article" year="2008">David &amp; Vu, 2008</bibRefCitation>
</taxonomicName>
, 19:17:
<quantity id="543775B1FFD8FF92FCAFFE35FC27556D" box="[865,926,511,536]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" pageId="14" pageNumber="405" unit="in" value="15.0">15 in</quantity>
<taxonomicName id="54CFA3D7FFD8FF92FC69FDCAFA9F556D" authority="Lidth de Jeude, 1890" authorityName="Lidth de Jeude" authorityYear="1890" box="[935,1318,511,536]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="rugosa">
<emphasis id="A1BB0446FFD8FF92FC69FDCAFBA0556D" box="[935,1049,512,536]" italics="true" pageId="14" pageNumber="405">O. rugosa</emphasis>
(Lidth de
<bibRefCitation id="F75EA5A5FFD8FF92FB5DFE35FAA7556D" author="Jeude" box="[1171,1310,511,536]" pageId="14" pageNumber="405" refString="Lithe de Jeude, T. W. van (1890) Reptilia from the Malay Archipelago. II. Ophidia. In: Weber, M. (Ed.), Zoologische Ergebnisse einer Reise in Niederlandisch Ost-Indien, 1, 178 - 192, pls. 15 - 16." type="journal article" year="1890">Jeude, 1890</bibRefCitation>
)
</taxonomicName>
, 17 dorsal scale rows at midbody in
<taxonomicName id="54CFA3D7FFD8FF92FE77FDEEFD0E5549" authority="Gunther, 1872" authorityName="Gunther" authorityYear="1872" box="[441,695,548,573]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="atra">
<emphasis id="A1BB0446FFD8FF92FE77FDEEFDB25549" box="[441,523,548,572]" italics="true" pageId="14" pageNumber="405">O. atra</emphasis>
<bibRefCitation id="F75EA5A5FFD8FF92FDDDFDEEFD0E5549" author="Gunther" box="[531,695,548,573]" pageId="14" pageNumber="405" refString="Gunther, A. C. L. G. (1872) Seventh account of new species of snakes in the collection of the British Museum. Annals and Magazine of Natural History, 9 (4), 13 - 37." type="journal article" year="1872">Günther, 1872</bibRefCitation>
</taxonomicName>
, 19:21:
<quantity id="543775B1FFD8FF92FCC6FDEEFCFD5548" box="[776,836,548,573]" metricMagnitude="-1" metricUnit="m" metricValue="4.318" pageId="14" pageNumber="405" unit="in" value="17.0">17 in</quantity>
<taxonomicName id="54CFA3D7FFD8FF92FC82FDEEFF6B5515" authority="Teynie, Lottier, David, Nguyen &amp; Vogel, 2013" authorityName="Teynie, Lottier, David, Nguyen &amp; Vogel" authorityYear="2013" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="durandi">
<emphasis id="A1BB0446FFD8FF92FC82FDEEFC7E5549" box="[844,967,548,572]" italics="true" pageId="14" pageNumber="405">O. durandi</emphasis>
<bibRefCitation id="F75EA5A5FFD8FF92FC01FDEEFF6B5515" author="Teynie" pageId="14" pageNumber="405" refString="Teynie, A., Lottier, A., David, P., Nguyen, T. Q. &amp; Vogel, G. (2013) A new species of the genus Opisthotropis Gunther, 1872 from northern Laos (Squamata: Natricidae). Zootaxa, 3774 (2), 165 - 182." type="journal article" year="2013">Teynié, Lottier, David, Nguyen &amp; Vogel, 2013</bibRefCitation>
</taxonomicName>
, 15 dorsal scale rows at midbody in
<taxonomicName id="54CFA3D7FFD8FF92FDA0FD82FBB85515" authority="Okada &amp; Takara, 1958" authorityName="Okada &amp; Takara" authorityYear="1958" box="[622,1025,583,608]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="kikuzatoi">
<emphasis id="A1BB0446FFD8FF92FDA0FD82FD4F5515" box="[622,758,584,608]" italics="true" pageId="14" pageNumber="405">O. kikuzatoi</emphasis>
<bibRefCitation id="F75EA5A5FFD8FF92FD30FD8DFBB85515" author="Okada" box="[766,1025,583,608]" pageId="14" pageNumber="405" refString="Okada, Y. &amp; Takara, T. (1958) A new species of Liopeltis (Ophidia, Colubridae) from Ryukyu Islands. Bulletin of the Biogeographical Society of Japan, 20 (3), 1 - 3." type="journal article" year="1958">Okada &amp; Takara, 1958</bibRefCitation>
</taxonomicName>
; loreal not in contact with internasal in both sides of head vs. in contact in
<taxonomicName id="54CFA3D7FFD8FF92FDAEFDA6FC4055F0" authority="Brown &amp; Leviton, 1961" authorityName="Brown &amp; Leviton" authorityYear="1961" box="[608,1017,620,645]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="alcalai">
<emphasis id="A1BB0446FFD8FF92FDAEFDA6FD6A55F1" box="[608,723,620,644]" italics="true" pageId="14" pageNumber="405">O. alcalai</emphasis>
<bibRefCitation id="F75EA5A5FFD8FF92FD10FDA7FC4055F0" author="Brown" box="[734,1017,620,645]" pageId="14" pageNumber="405" refString="Brown, W. C. &amp; Leviton, A. E. (1961) Discovery of the snake genus Opisthotropis in the Philippine Islands, with description of a new species. Occasional Papers of the Natural History Museum of Stanford University, 8, 1 - 5." type="journal article" year="1961">Brown &amp; Leviton, 1961</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="54CFA3D7FFD8FF92FBC7FDA6FE9755DD" authority="David, Pham, Nguyen &amp; Ziegler, 2011" authorityName="David, Pham, Nguyen &amp; Ziegler" authorityYear="2011" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="cucae">
<emphasis id="A1BB0446FFD8FF92FBC7FDA6FBC955F1" box="[1033,1136,620,644]" italics="true" pageId="14" pageNumber="405">O. cucae</emphasis>
<bibRefCitation id="F75EA5A5FFD8FF92FBB5FDA6FE9755DD" author="David" pageId="14" pageNumber="405" refString="David, P., Pham, T. C., Nguyen, Q. T. &amp; Ziegler, T. (2011) A new species of the genus Opisthotropis Gunther, 1872 (Squamata: Natricidae) from the highlands of Kon Tum Province, Vietnam. Zootaxa, 2758, 43 - 56." type="journal article" year="2011">David, Pham, Nguyen &amp; Ziegler, 2011</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="54CFA3D7FFD8FF92FEF6FD5AFDCB55DD" authority="Mocquard, 1890" authorityName="Mocquard" authorityYear="1890" box="[312,626,655,680]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="typica">
<emphasis id="A1BB0446FFD8FF92FEF6FD5AFE2655D2" box="[312,415,656,679]" italics="true" pageId="14" pageNumber="405">O. typica</emphasis>
(
<bibRefCitation id="F75EA5A5FFD8FF92FE62FD45FDD355DD" author="Mocquard" box="[428,618,655,680]" pageId="14" pageNumber="405" refString="Mocquard, F. (1890) Recherches sur la faune herpetologique des iles de Borneo et de Palawan. Nouvelles Archives du Museum d'Histoire Naturelle de Paris, 3 (2), 115 - 168." type="journal article" year="1890">Mocquard, 1890</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="54CFA3D7FFD8FF92FDB3FD5AFD7455DD" box="[637,717,656,680]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="atra">
<emphasis id="A1BB0446FFD8FF92FDB3FD5AFD7455DD" box="[637,717,656,680]" italics="true" pageId="14" pageNumber="405">O. atra</emphasis>
</taxonomicName>
,
<taxonomicName id="54CFA3D7FFD8FF92FD15FD5AFC3955DD" box="[731,896,656,680]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="daovantieni">
<emphasis id="A1BB0446FFD8FF92FD15FD5AFC3955DD" box="[731,896,656,680]" italics="true" pageId="14" pageNumber="405">O. daovantieni</emphasis>
</taxonomicName>
,
<taxonomicName id="54CFA3D7FFD8FF92FC40FD5AFBB155DD" box="[910,1032,656,680]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="durandi">
<emphasis id="A1BB0446FFD8FF92FC40FD5AFBB155DD" box="[910,1032,656,680]" italics="true" pageId="14" pageNumber="405">O. durandi</emphasis>
</taxonomicName>
,
<taxonomicName id="54CFA3D7FFD8FF92FBD8FD5AFB3F55DD" box="[1046,1158,656,680]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="rugosa">
<emphasis id="A1BB0446FFD8FF92FBD8FD5AFB3F55DD" box="[1046,1158,656,680]" italics="true" pageId="14" pageNumber="405">O. rugosa</emphasis>
</taxonomicName>
and
<taxonomicName id="54CFA3D7FFD8FF92FB75FD5AFA8255D2" box="[1211,1339,656,679]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="spenceri">
<emphasis id="A1BB0446FFD8FF92FB75FD5AFA8255D2" box="[1211,1339,656,679]" italics="true" pageId="14" pageNumber="405">O. spenceri</emphasis>
</taxonomicName>
; SPL 9 10 vs.
<quantity id="543775B1FFD8FF92FF2EFD7EFEFA55B8" box="[224,323,692,717]" metricMagnitude="-1" metricUnit="m" metricValue="3.175" metricValueMax="3.302" metricValueMin="3.048" pageId="14" pageNumber="405" unit="in" value="12.5" valueMax="13.0" valueMin="12.0">1213 in</quantity>
<taxonomicName id="54CFA3D7FFD8FF92FE87FD7EFE0155B9" box="[329,440,692,716]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="alcalai">
<emphasis id="A1BB0446FFD8FF92FE87FD7EFE0155B9" box="[329,440,692,716]" italics="true" pageId="14" pageNumber="405">O. alcalai</emphasis>
</taxonomicName>
,
<quantity id="543775B1FFD8FF92FE08FD7FFE4855B8" box="[454,497,692,717]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="14" pageNumber="405" unit="in" value="7.0">7 in</quantity>
<taxonomicName id="54CFA3D7FFD8FF92FE36FD7EFDF055B9" box="[504,585,692,716]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="atra">
<emphasis id="A1BB0446FFD8FF92FE36FD7EFDF055B9" box="[504,585,692,716]" italics="true" pageId="14" pageNumber="405">O. atra</emphasis>
</taxonomicName>
and
<taxonomicName id="54CFA3D7FFD8FF92FDB0FD7EFD5855B9" box="[638,737,692,716]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="cucae">
<emphasis id="A1BB0446FFD8FF92FDB0FD7EFD5855B9" box="[638,737,692,716]" italics="true" pageId="14" pageNumber="405">O. cucae</emphasis>
</taxonomicName>
,
<quantity id="543775B1FFD8FF92FD21FD7EFC8C55B8" box="[751,821,692,717]" metricMagnitude="-1" metricUnit="m" metricValue="1.651" metricValueMax="1.778" metricValueMin="1.524" pageId="14" pageNumber="405" unit="in" value="6.5" valueMax="7.0" valueMin="6.0">67 in</quantity>
<taxonomicName id="54CFA3D7FFD8FF92FCF2FD7EFC0F55B9" box="[828,950,692,716]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="durandi">
<emphasis id="A1BB0446FFD8FF92FCF2FD7EFC0F55B9" box="[828,950,692,716]" italics="true" pageId="14" pageNumber="405">O. durandi</emphasis>
</taxonomicName>
,
<quantity id="543775B1FFD8FF92FC0AFD7EFB9E55B8" box="[964,1063,692,717]" metricMagnitude="-1" metricUnit="m" metricValue="3.175" metricValueMax="3.302" metricValueMin="3.048" pageId="14" pageNumber="405" unit="in" value="12.5" valueMax="13.0" valueMin="12.0">1213 in</quantity>
<taxonomicName id="54CFA3D7FFD8FF92FBE3FD7EFB2455B9" box="[1069,1181,692,716]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="rugosa">
<emphasis id="A1BB0446FFD8FF92FBE3FD7EFB2455B9" box="[1069,1181,692,716]" italics="true" pageId="14" pageNumber="405">O. rugosa</emphasis>
</taxonomicName>
,
<quantity id="543775B1FFD8FF92FB65FD7EFAB555B8" box="[1195,1292,692,717]" metricMagnitude="-1" metricUnit="m" metricValue="2.921" metricValueMax="3.048" metricValueMin="2.794" pageId="14" pageNumber="405" unit="in" value="11.5" valueMax="12.0" valueMin="11.0">1112 in</quantity>
<taxonomicName id="54CFA3D7FFD8FF92FADDFD7EFAC055B9" box="[1299,1401,692,716]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="typica">
<emphasis id="A1BB0446FFD8FF92FADDFD7EFAC055B9" box="[1299,1401,692,716]" italics="true" pageId="14" pageNumber="405">O. typica</emphasis>
</taxonomicName>
; V 162179 and SC 5360 vs. V
<quantity id="543775B1FFD8FF92FE3AFD1DFD845585" box="[500,573,727,752]" metricMagnitude="0" metricUnit="m" metricValue="4.953" pageId="14" pageNumber="405" unit="in" value="195.0">195 in</quantity>
<taxonomicName id="54CFA3D7FFD8FF92FD8BFD12FD015585" box="[581,696,728,752]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="alcalai">
<emphasis id="A1BB0446FFD8FF92FD8BFD12FD015585" box="[581,696,728,752]" italics="true" pageId="14" pageNumber="405">O. alcalai</emphasis>
</taxonomicName>
, V 191 and SC
<quantity id="543775B1FFD8FF92FCBBFD12FC095585" box="[885,944,727,752]" metricMagnitude="0" metricUnit="m" metricValue="1.1176" pageId="14" pageNumber="405" unit="in" value="44.0">44 in</quantity>
<taxonomicName id="54CFA3D7FFD8FF92FC76FD12FBA45585" box="[952,1053,728,752]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="cucae">
<emphasis id="A1BB0446FFD8FF92FC76FD12FBA45585" box="[952,1053,728,752]" italics="true" pageId="14" pageNumber="405">O. cucae</emphasis>
</taxonomicName>
, V 189194 and SC
<quantity id="543775B1FFD8FF92FADDFD12FAC15585" box="[1299,1400,727,752]" metricMagnitude="0" metricUnit="m" metricValue="1.0922" metricValueMax="1.1938" metricValueMin="0.9906" pageId="14" pageNumber="405" unit="in" value="43.0" valueMax="47.0" valueMin="39.0">3947 in</quantity>
<taxonomicName id="54CFA3D7FFD8FF92FA4FFD12FEA25461" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="daovantieni">
<emphasis id="A1BB0446FFD8FF92FA4FFD12FEA25461" italics="true" pageId="14" pageNumber="405">O. daovantieni</emphasis>
</taxonomicName>
, SC
<quantity id="543775B1FFD8FF92FE97FD36FE065460" box="[345,447,764,789]" metricMagnitude="0" metricUnit="m" metricValue="2.2606" metricValueMax="2.286" metricValueMin="2.2352" pageId="14" pageNumber="405" unit="in" value="89.0" valueMax="90.0" valueMin="88.0">8890 in</quantity>
<taxonomicName id="54CFA3D7FFD8FF92FE07FD36FDFF5461" box="[457,582,764,788]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="durandi">
<emphasis id="A1BB0446FFD8FF92FE07FD36FDFF5461" box="[457,582,764,788]" italics="true" pageId="14" pageNumber="405">O. durandi</emphasis>
</taxonomicName>
, V
<quantity id="543775B1FFD8FF92FDB9FD36FD405460" box="[631,761,764,789]" metricMagnitude="0" metricUnit="m" metricValue="4.8006" metricValueMax="5.0292" metricValueMin="4.572" pageId="14" pageNumber="405" unit="in" value="189.0" valueMax="198.0" valueMin="180.0">180198 in</quantity>
<taxonomicName id="54CFA3D7FFD8FF92FCCDFD36FC345461" box="[771,909,764,788]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="kikuzatoi">
<emphasis id="A1BB0446FFD8FF92FCCDFD36FC345461" box="[771,909,764,788]" italics="true" pageId="14" pageNumber="405">O. kikuzatoi</emphasis>
</taxonomicName>
, SC
<quantity id="543775B1FFD8FF92FC05FD36FB885460" box="[971,1073,764,789]" metricMagnitude="0" metricUnit="m" metricValue="2.2479" metricValueMax="2.413" metricValueMin="2.0827999999999998" pageId="14" pageNumber="405" unit="in" value="88.5" valueMax="95.0" valueMin="82.0">8295 in</quantity>
<taxonomicName id="54CFA3D7FFD8FF92FBF5FD36FB1C5461" box="[1083,1189,764,788]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="typica">
<emphasis id="A1BB0446FFD8FF92FBF5FD36FB1C5461" box="[1083,1189,764,788]" italics="true" pageId="14" pageNumber="405">O. typica</emphasis>
</taxonomicName>
; dorsal scales keeled throughout vs. smooth throughout in
<taxonomicName id="54CFA3D7FFD8FF92FDF6FCEAFD11544D" box="[568,680,800,824]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="alcalai">
<emphasis id="A1BB0446FFD8FF92FDF6FCEAFD11544D" box="[568,680,800,824]" italics="true" pageId="14" pageNumber="405">O. alcalai</emphasis>
</taxonomicName>
,
<taxonomicName id="54CFA3D7FFD8FF92FD7BFCEAFCA0544D" box="[693,793,800,824]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="cucae">
<emphasis id="A1BB0446FFD8FF92FD7BFCEAFCA0544D" box="[693,793,800,824]" italics="true" pageId="14" pageNumber="405">O. cucae</emphasis>
</taxonomicName>
, O.
<taxonomicName id="54CFA3D7FFD8FF92FC87FCEAFC74544D" box="[841,973,800,824]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="daovantieni">
<emphasis id="A1BB0446FFD8FF92FC87FCEAFC74544D" box="[841,973,800,824]" italics="true" pageId="14" pageNumber="405">daovantieni</emphasis>
</taxonomicName>
,
<taxonomicName id="54CFA3D7FFD8FF92FC12FCEAFBEE544D" box="[988,1111,800,824]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="durandi">
<emphasis id="A1BB0446FFD8FF92FC12FCEAFBEE544D" box="[988,1111,800,824]" italics="true" pageId="14" pageNumber="405">O. durandi</emphasis>
</taxonomicName>
, and
<taxonomicName id="54CFA3D7FFD8FF92FB5AFCEAFAAC5442" box="[1172,1301,800,823]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="spenceri">
<emphasis id="A1BB0446FFD8FF92FB5AFCEAFAAC5442" box="[1172,1301,800,823]" italics="true" pageId="14" pageNumber="405">O. spenceri</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="C7B088DCFFD8FF92FF59F9F6FB7D5104" httpUri="https://zenodo.org/record/438871/files/figure.png" pageId="14" pageNumber="405" targetBox="[157,1415,864,1558]" targetPageId="14">
<paragraph id="9370D854FFD8FF92FF59F9F6FB7D5104" blockId="14.[151,1436,1596,1649]" pageId="14" pageNumber="405">
<emphasis id="A1BB0446FFD8FF92FF59F9F6FEAE5127" bold="true" box="[151,279,1596,1618]" pageId="14" pageNumber="405">FIGURE 6.</emphasis>
Comparisions of head scalation of
<emphasis id="A1BB0446FFD8FF92FD52F9F6FD085127" bold="true" box="[668,689,1596,1618]" pageId="14" pageNumber="405">a:</emphasis>
<taxonomicName id="54CFA3D7FFD8FF92FD72F9F6FC585127" box="[700,993,1596,1618]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="shenzhenensis" status="sp. nov.">
<emphasis id="A1BB0446FFD8FF92FD72F9F6FC585127" box="[700,993,1596,1618]" italics="true" pageId="14" pageNumber="405">Opisthotropis shenzhenensis</emphasis>
</taxonomicName>
<emphasis id="A1BB0446FFD8FF92FC22F9F6FBF85127" bold="true" box="[1004,1089,1596,1618]" pageId="14" pageNumber="405">
<taxonomicNameLabel id="BA88B93DFFD8FF92FC22F9F6FBF85127" box="[1004,1089,1596,1618]" pageId="14" pageNumber="405" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype SYS r001018),
<emphasis id="A1BB0446FFD8FF92FAACF9F6FAC15127" bold="true" box="[1378,1400,1596,1618]" pageId="14" pageNumber="405">b:</emphasis>
<taxonomicName id="54CFA3D7FFD8FF92FA4DF9F7FF4B5104" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="maxwelli">
<emphasis id="A1BB0446FFD8FF92FA4DF9F7FF4B5104" italics="true" pageId="14" pageNumber="405">O. maxwelli</emphasis>
</taxonomicName>
(SYS r000841),
<emphasis id="A1BB0446FFD8FF92FE51F991FE135104" bold="true" box="[415,426,1627,1649]" pageId="14" pageNumber="405">c</emphasis>
and
<emphasis id="A1BB0446FFD8FF92FE12F991FE4B5104" bold="true" box="[476,498,1627,1649]" pageId="14" pageNumber="405">d:</emphasis>
<taxonomicName id="54CFA3D7FFD8FF92FE36F991FD3A5104" box="[504,643,1627,1649]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="andersonii">
<emphasis id="A1BB0446FFD8FF92FE36F991FD3A5104" box="[504,643,1627,1649]" italics="true" pageId="14" pageNumber="405">O. andersonii</emphasis>
</taxonomicName>
(SYS r001020, SYS r001082). Line illustration by ZTL.
</paragraph>
</caption>
<subSubSection id="DBD58BDFFFD8FF92FF09F952FD225010" pageId="14" pageNumber="405" type="distribution">
<paragraph id="9370D854FFD8FF92FF09F952FD225010" blockId="14.[151,1436,1688,2036]" pageId="14" pageNumber="405">
<emphasis id="A1BB0446FFD8FF92FF09F952FE4D51C4" bold="true" box="[199,500,1688,1713]" pageId="14" pageNumber="405">Distribution and habitat.</emphasis>
<taxonomicName id="54CFA3D7FFD8FF92FE32F952FC8451C5" box="[508,829,1688,1712]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="shenzhenensis" status="sp. nov.">
<emphasis id="A1BB0446FFD8FF92FE32F952FC8451C5" box="[508,829,1688,1712]" italics="true" pageId="14" pageNumber="405">Opisthotropis shenzhenensis</emphasis>
</taxonomicName>
<emphasis id="A1BB0446FFD8FF92FC8BF952FC1851C4" bold="true" box="[837,929,1688,1713]" pageId="14" pageNumber="405">
<taxonomicNameLabel id="BA88B93DFFD8FF92FC8BF952FC1851C4" box="[837,929,1688,1713]" pageId="14" pageNumber="405" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
occurs from the low mountain ranges in the eastern Shenzhen and southern Dongguan,
<collectingRegion id="510B16B6FFD8FF92FD4DF976FCCC51A0" box="[643,885,1724,1749]" country="China" name="Guangdong" pageId="14" pageNumber="405">Guangdong Province</collectingRegion>
,
<collectingCountry id="EBD898C4FFD8FF92FC4CF976FC7151A0" box="[898,968,1724,1749]" name="China" pageId="14" pageNumber="405">China</collectingCountry>
, including Mt. Wutong, Sanzhoutian and Mt. Tiantou, and Mt. Yinping (four red dots in
<figureCitation id="0BF4C4D1FFD8FF92FD7BF92AFD4E518D" box="[693,759,1760,1785]" captionStart="FIGURE 1" captionStartId="3.[151,250,1179,1201]" captionTargetBox="[151,1436,195,1155]" captionTargetId="figure@3.[151,1436,195,1155]" captionTargetPageId="3" captionText="FIGURE 1. Sampling localities of Opisthotropis in China. 1: Hong Kong; 2: Mt. Wutong; 3: Sanzhoutian; 4: Mt. Tiantou; 5: Mt. Yinping; 6: Mt. Maofeng; 7: Nanao Island; 8: Huboliao Nature Reserve; 9: Wulong Village; 10: Guadun Village; 11: Qixiling Nature Reserve; 12: Shimentai Nature Reserve; 13: Heishiding Nature Reserve; 14: Shangchuan Island. The red dot represent localities of Opisthotropis shenzhenensis sp. nov.. See Table 1 for locality codes and detailed locality information." httpUri="https://zenodo.org/record/438866/files/figure.png" pageId="14" pageNumber="405">Fig. 1</figureCitation>
). All specimens were collected at night from slow-flowing streams covered with bare rocks at elevations between 155 and
<quantity id="543775B1FFD8FF92FCADF8CFFC125069" box="[867,939,1797,1821]" metricMagnitude="2" metricUnit="m" metricValue="3.27" pageId="14" pageNumber="405" unit="m" value="327.0">327 m</quantity>
a.s.l., where we observed a large number of freshwater snails, aquatic insects, shrimps, crabs, and fishes, and amphibians such as
<taxonomicName id="54CFA3D7FFD8FF92FB7CF8E2FADC5035" box="[1202,1381,1832,1856]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="laui">
<emphasis id="A1BB0446FFD8FF92FB7CF8E2FADC5035" box="[1202,1381,1832,1856]" italics="true" pageId="14" pageNumber="405">Leptolalax laui</emphasis>
</taxonomicName>
and
<taxonomicName id="54CFA3D7FFD8FF92FF59F887FE4A5011" box="[151,499,1868,1892]" class="Amphibia" family="Salamandridae" genus="Paramesotriton" kingdom="Animalia" order="Caudata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="hongkongensis">
<emphasis id="A1BB0446FFD8FF92FF59F887FE4A5011" box="[151,499,1868,1892]" italics="true" pageId="14" pageNumber="405">Paramesotriton hongkongensis</emphasis>
</taxonomicName>
occur (
<figureCitation id="0BF4C4D1FFD8FF92FD86F886FD375010" box="[584,654,1868,1893]" captionStart="FIGURE 7" captionStartId="15.[151,250,1074,1096]" captionTargetBox="[152,1435,195,1052]" captionTargetId="figure@15.[151,1435,195,1052]" captionTargetPageId="15" captionText="FIGURE 7. Habitat of Opisthotropis shenzhenensis sp. nov. at the type locality, Mt. Wutong, Shenzhen City, Guangdong Province, China, which is inhabited also by the other three sympatric species O. andersonii, O. kuatunensis and O. lateralis." httpUri="https://zenodo.org/record/438872/files/figure.png" pageId="14" pageNumber="405">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="DBD58BDFFFD8FF92FF09F8BAFEE65081" pageId="14" pageNumber="405" type="discussion">
<paragraph id="9370D854FFD8FF92FF09F8BAFEE65081" blockId="14.[151,1436,1688,2036]" pageId="14" pageNumber="405">
<emphasis id="A1BB0446FFD8FF92FF09F8BAFE8250FC" bold="true" box="[199,315,1904,1929]" pageId="14" pageNumber="405">Remarks.</emphasis>
<taxonomicName id="54CFA3D7FFD8FF92FE8CF8BAFD3850FD" box="[322,641,1904,1928]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="shenzhenensis">
<emphasis id="A1BB0446FFD8FF92FE8CF8BAFD3850FD" box="[322,641,1904,1928]" italics="true" pageId="14" pageNumber="405">Opisthotropis shenzhenensis</emphasis>
</taxonomicName>
,
<taxonomicName id="54CFA3D7FFD8FF92FD41F8BAFC9350FD" box="[655,810,1904,1928]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="andersonii">
<emphasis id="A1BB0446FFD8FF92FD41F8BAFC9350FD" box="[655,810,1904,1928]" italics="true" pageId="14" pageNumber="405">O. andersonii</emphasis>
</taxonomicName>
,
<taxonomicName id="54CFA3D7FFD8FF92FCF6F8BAFC6750FD" box="[824,990,1904,1928]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="kuatunensis">
<emphasis id="A1BB0446FFD8FF92FCF6F8BAFC6750FD" box="[824,990,1904,1928]" italics="true" pageId="14" pageNumber="405">O. kuatunensis</emphasis>
</taxonomicName>
and
<taxonomicName id="54CFA3D7FFD8FF92FBDBF8BAFB2F50FD" box="[1045,1174,1904,1928]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="lateralis">
<emphasis id="A1BB0446FFD8FF92FBDBF8BAFB2F50FD" box="[1045,1174,1904,1928]" italics="true" pageId="14" pageNumber="405">O. lateralis</emphasis>
</taxonomicName>
are sympatric and they were found frequently in two streams on Mt. Wutong, where a total of
<specimenCount id="85C913DDFFD8FF92FC78F85FFBF750D8" box="[950,1102,1940,1965]" pageId="14" pageNumber="405" type="generic">17 specimens</specimenCount>
were collected between
<date id="E771FE94FFD8FF92FAAAF85FFEBB50A4" pageId="14" pageNumber="405" value="2012" valueMax="2015">2012 and 2015</date>
, including five of
<taxonomicName id="54CFA3D7FFD8FF92FE1FF872FDC150A5" box="[465,632,1976,2000]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="kuatunensis">
<emphasis id="A1BB0446FFD8FF92FE1FF872FDC150A5" box="[465,632,1976,2000]" italics="true" pageId="14" pageNumber="405">O. kuatunensis</emphasis>
</taxonomicName>
, five of
<taxonomicName id="54CFA3D7FFD8FF92FD14F872FCE050A5" box="[730,857,1976,2000]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="lateralis">
<emphasis id="A1BB0446FFD8FF92FD14F872FCE050A5" box="[730,857,1976,2000]" italics="true" pageId="14" pageNumber="405">O. lateralis</emphasis>
</taxonomicName>
(see Appendix), four of
<taxonomicName id="54CFA3D7FFD8FF92FBBCF872FAB450A5" box="[1138,1293,1976,2000]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="andersonii">
<emphasis id="A1BB0446FFD8FF92FBBCF872FAB450A5" box="[1138,1293,1976,2000]" italics="true" pageId="14" pageNumber="405">O. andersonii</emphasis>
</taxonomicName>
and three of
<taxonomicName id="54CFA3D7FFD8FF92FF59F817FEE15081" box="[151,344,2012,2036]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="405" phylum="Chordata" rank="species" species="shenzhenensis">
<emphasis id="A1BB0446FFD8FF92FF59F817FEE15081" box="[151,344,2012,2036]" italics="true" pageId="14" pageNumber="405">O. shenzhenensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>