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<mods:title id="0E50B72836D98F0081444BAE6E5B1A82">Notes on the genital morphology and phylogenetics of two Arabian species of the genus Lepidochrysops Hedicke, 1923 (Lepidoptera, Lycaenidae)</mods:title>
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<mods:namePart id="52BE212DE4559C8E0437BFDDC8C04FF2">Verovnik, Rudi</mods:namePart>
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<mods:affiliation id="F1075D407723507E3239D654806C82A8">University of Ljubljana, Biotechnical Faculty, Department of Biology, Jamnikarjeva 101, 1000 Ljubljana, Slovenia. E-mail: rudi. verovnik @ bf. uni-lj. si; https: // orcid. org / 0000 - 0002 - 5841 - 5925</mods:affiliation>
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<mods:namePart id="742205FF73003B820611623BB449A0F3">Seizmair, Michael</mods:namePart>
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<mods:affiliation id="EA8BAC64485D30AAD2BFEC2A41B42BB5">Birkenstrasse 36, D- 82194 Gröbenzell, Germany; https: // orcid. org / 0000 - 0002 - 1385 - 7148 * Corresponding author. E-mail: michael. seizmair @ gmx. net</mods:affiliation>
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<treatment id="1B61EC6EFFFDF229DCE9F902FDA1FACB" LSID="urn:lsid:plazi:treatment:1B61EC6EFFFDF229DCE9F902FDA1FACB" httpUri="http://treatment.plazi.org/id/1B61EC6EFFFDF229DCE9F902FDA1FACB" lastPageId="6" lastPageNumber="206" pageId="4" pageNumber="204">
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<taxonomicName id="54C826FBFFFDF22BDCE9F902FD8EF8D8" ID-CoL="3TB5X" authority="Larsen, 1983" authorityName="Larsen" authorityYear="1983" box="[189,636,1782,1808]" class="Insecta" family="Lycaenidae" genus="Lepidochrysops" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="204" phylum="Arthropoda" rank="species" species="haveni">
<emphasis id="A1BC816AFFFDF22BDCE9F902FD8EF8D8" bold="true" box="[189,636,1782,1808]" pageId="4" pageNumber="204">
<emphasis id="A1BC816AFFFDF22BDCE9F902FE21F8D8" bold="true" box="[189,467,1782,1808]" italics="true" pageId="4" pageNumber="204">Lepidochrysops haveni</emphasis>
<bibRefCitation id="F7592089FFFDF22BDD8FF902FD8EF8D8" author="Larsen, T. B." box="[475,636,1782,1808]" pageId="4" pageNumber="204" pagination="333 - 478" refId="ref5547" refString="Larsen, T. B. (1983) Insects of Saudi Arabia. Lepidoptera; Rhopalocera (A Monograph of the Butterflies of the Arabian Peninsula). Fauna of Saudi Arabia, 5, 333 - 478." type="journal article" year="1983">Larsen, 1983</bibRefCitation>
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<emphasis id="A1BC816AFFFDF22BDCE9F8CCFE5EF89A" bold="true" box="[189,428,1848,1874]" pageId="4" pageNumber="204">Material examined</emphasis>
:
<materialsCitation id="23A05725FFFDF22BDDE9F8CCFA85F8BD" collectingDate="2023-11-10" collectionCode="GP" collectorName="M. Seizmair." country="Saudi Arabia" county="Province" elevation="950" location="Marwah" municipality="Mt. Faifa" pageId="4" pageNumber="204" specimenCode="GP001, GP057, 24, GP004" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Jizan">
<collectingCountry id="EBDF1DE8FFFDF22BDDE9F8CCFDAEF89B" box="[445,604,1848,1875]" name="Saudi Arabia" pageId="4" pageNumber="204">Saudi Arabia</collectingCountry>
,
<collectingCounty id="7A1625F4FFFDF22BDE3DF8CCFD23F89B" box="[617,721,1848,1875]" pageId="4" pageNumber="204">Province</collectingCounty>
<collectingRegion id="510C939AFFFDF22BDE8EF8CCFCE8F89B" box="[730,794,1848,1875]" country="Saudi Arabia" name="Jizan" pageId="4" pageNumber="204">Jizan</collectingRegion>
,
<collectingMunicipality id="7313C702FFFDF22BDF73F8CDFC6BF89B" box="[807,921,1848,1875]" pageId="4" pageNumber="204">Mt. Faifa</collectingMunicipality>
,
<location id="96170BA3FFFDF22BDFF3F8CCFBFEF89B" LSID="urn:lsid:plazi:treatment:1B61EC6EFFFDF229DCE9F902FDA1FACB:96170BA3FFFDF22BDFF3F8CCFBFEF89B" box="[935,1036,1848,1875]" country="Saudi Arabia" county="Province" municipality="Mt. Faifa" name="Marwah" pageId="4" pageNumber="204" stateProvince="Jizan">Marwah</location>
, 17.281553°, 43.101033°,
<quantity id="5430F09DFFFDF22BD91EF8CCFF24F8BD" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="4" pageNumber="204" unit="m" value="950.0">
<elevation id="18E5BA4BFFFDF22BD91EF8CCFF24F8BD" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="4" pageNumber="204" unit="m" value="950.0">950 m</elevation>
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,
<date id="E7767BB8FFFDF22BDCB5F8AEFE9DF8BD" box="[225,367,1882,1909]" pageId="4" pageNumber="204" value="2023-11-10">
<collectingDate id="F7328250FFFDF22BDCB5F8AEFE9DF8BD" box="[225,367,1882,1909]" pageId="4" pageNumber="204" value="2023-11-10">10-XI-2023</collectingDate>
</date>
,
<specimenCount id="85CE96F1FFFDF22BDD2DF8AEFE6CF8BD" box="[377,414,1882,1909]" count="1" pageId="4" pageNumber="204" type="male">1♂</specimenCount>
, slide no. 24
<specimenCode id="C36EF503FFFDF22BDE67F8AEFD74F8BD" box="[563,646,1882,1909]" collectionCode="GP" country="Brazil" name="Instituto de Geociencias, Universidade de Sao Paulo" pageId="4" pageNumber="204">GP001</specimenCode>
,
<specimenCount id="85CE96F1FFFDF22BDEC5F8AEFD47F8BD" box="[657,693,1882,1909]" count="2" pageId="4" pageNumber="204" type="female">2♀</specimenCount>
, slide no. 23
<specimenCode id="C36EF503FFFDF22BDF1DF8AEFC6CF8BD" box="[841,926,1882,1909]" collectionCode="GP" country="Brazil" name="Instituto de Geociencias, Universidade de Sao Paulo" pageId="4" pageNumber="204">GP057</specimenCode>
,
<specimenCode id="C36EF503FFFDF22BDFFCF8AEFC34F8BD" box="[936,966,1882,1909]" collectionCode="GP" country="Brazil" name="Instituto de Geociencias, Universidade de Sao Paulo" pageId="4" pageNumber="204">24</specimenCode>
<specimenCode id="C36EF503FFFDF22BDF92F8AEFBE8F8BD" box="[966,1050,1882,1909]" collectionCode="GP" country="Brazil" name="Instituto de Geociencias, Universidade de Sao Paulo" pageId="4" pageNumber="204">GP004</specimenCode>
, leg., prep., coll.
<collectorName id="3E3D38AEFFFDF22BD88BF8AFFA85F8BD" box="[1247,1399,1882,1909]" pageId="4" pageNumber="204">M. Seizmair.</collectorName>
</materialsCitation>
<materialsCitation id="23A05725FFFDF22BDCE9F888FC53F870" collectingDate="2023-11-08" collectionCode="GP" collectorName="M. Seizmair." country="Saudi Arabia" county="Province" elevation="660" location="Ayban Belghazi" municipality="Mt. Faifa" pageId="4" pageNumber="204" specimenCode="GP055, 23, GP060" specimenCount="2" specimenCount-male="2" stateProvince="Jizan">
<collectingCountry id="EBDF1DE8FFFDF22BDCE9F888FEA8F85F" box="[189,346,1916,1943]" name="Saudi Arabia" pageId="4" pageNumber="204">Saudi Arabia</collectingCountry>
,
<collectingCounty id="7A1625F4FFFDF22BDD30F888FE3EF85F" box="[356,460,1916,1943]" pageId="4" pageNumber="204">Province</collectingCounty>
<collectingRegion id="510C939AFFFDF22BDD87F888FDE0F85F" box="[467,530,1916,1943]" country="Saudi Arabia" name="Jizan" pageId="4" pageNumber="204">Jizan</collectingRegion>
,
<collectingMunicipality id="7313C702FFFDF22BDE49F889FD7EF85F" box="[541,652,1916,1943]" pageId="4" pageNumber="204">Mt. Faifa</collectingMunicipality>
,
<location id="96170BA3FFFDF22BDEC3F888FCA5F85F" LSID="urn:lsid:plazi:treatment:1B61EC6EFFFDF229DCE9F902FDA1FACB:96170BA3FFFDF22BDEC3F888FCA5F85F" box="[663,855,1916,1943]" country="Saudi Arabia" county="Province" municipality="Mt. Faifa" name="Ayban Belghazi" pageId="4" pageNumber="204" stateProvince="Jizan">Ayban Belghazi</location>
, 17.263406°, 43.076147°,
<quantity id="5430F09DFFFDF22BD8DAF888FB2BF85F" box="[1166,1241,1916,1943]" metricMagnitude="2" metricUnit="m" metricValue="6.6" pageId="4" pageNumber="204" unit="m" value="660.0">
<elevation id="18E5BA4BFFFDF22BD8DAF888FB2BF85F" box="[1166,1241,1916,1943]" metricMagnitude="2" metricUnit="m" metricValue="6.6" pageId="4" pageNumber="204" unit="m" value="660.0">660 m</elevation>
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,
<date id="E7767BB8FFFDF22BD8B0F888FA80F85F" box="[1252,1394,1916,1943]" pageId="4" pageNumber="204" value="2023-11-08">
<collectingDate id="F7328250FFFDF22BD8B0F888FA80F85F" box="[1252,1394,1916,1943]" pageId="4" pageNumber="204" value="2023-11-08">08-XI-2023</collectingDate>
</date>
,
<specimenCount id="85CE96F1FFFDF22BDCE9F869FF10F870" box="[189,226,1949,1976]" count="2" pageId="4" pageNumber="204" type="male">2♂</specimenCount>
, slide no. 23
<specimenCode id="C36EF503FFFDF22BDD20F869FE3BF870" box="[372,457,1949,1976]" collectionCode="GP" country="Brazil" name="Instituto de Geociencias, Universidade de Sao Paulo" pageId="4" pageNumber="204">GP055</specimenCode>
,
<specimenCode id="C36EF503FFFDF22BDD87F869FE02F870" box="[467,496,1949,1976]" collectionCode="GP" country="Brazil" name="Instituto de Geociencias, Universidade de Sao Paulo" pageId="4" pageNumber="204">23</specimenCode>
<specimenCode id="C36EF503FFFDF22BDDA4F869FDB7F870" box="[496,581,1949,1976]" collectionCode="GP" country="Brazil" name="Instituto de Geociencias, Universidade de Sao Paulo" pageId="4" pageNumber="204">GP060</specimenCode>
, leg., prep., coll.
<collectorName id="3E3D38AEFFFDF22BDF5EF86AFC53F870" box="[778,929,1949,1976]" pageId="4" pageNumber="204">M. Seizmair.</collectorName>
</materialsCitation>
<materialsCitation id="23A05725FFFDF22BDFF2F869FE29F834" collectionCode="GP" collectorName="M. Seizmair." country="Saudi Arabia" elevation="560" location="Mt. Faifa" municipality="Province" pageId="4" pageNumber="204" specimenCode="GP003, 23, GP061" specimenCount="2" specimenCount-female="2" stateProvince="Jizan">
<collectingCountry id="EBDF1DE8FFFDF22BDFF2F869FBB0F870" box="[934,1090,1949,1976]" name="Saudi Arabia" pageId="4" pageNumber="204">Saudi Arabia</collectingCountry>
,
<collectingMunicipality id="7313C702FFFDF22BD818F869FB46F870" box="[1100,1204,1949,1976]" pageId="4" pageNumber="204">Province</collectingMunicipality>
<collectingRegion id="510C939AFFFDF22BD8EEF869FB0BF870" box="[1210,1273,1949,1976]" country="Saudi Arabia" name="Jizan" pageId="4" pageNumber="204">Jizan</collectingRegion>
,
<location id="96170BA3FFFDF22BD957F86AFA80F870" LSID="urn:lsid:plazi:treatment:1B61EC6EFFFDF229DCE9F902FDA1FACB:96170BA3FFFDF22BD957F86AFA80F870" box="[1283,1394,1949,1976]" country="Saudi Arabia" municipality="Province" name="Mt. Faifa" pageId="4" pageNumber="204" stateProvince="Jizan">Mt. Faifa</location>
,
<locationDeviation id="82C5445AFFFDF22BDCE9F84BFDD2F812" box="[189,544,1983,2010]" location="Ayban Belghazi" pageId="4" pageNumber="204">
<quantity id="5430F09DFFFDF22BDCE9F84BFEFFF812" box="[189,269,1983,2010]" metricMagnitude="3" metricUnit="m" metricValue="8.5" pageId="4" pageNumber="204" unit="km" value="8.5">8.5 km</quantity>
NE of Ayban Belghazi
</locationDeviation>
, 17.302683°, 43.076006°,
<quantity id="5430F09DFFFDF22BDF03F84BFC51F812" box="[855,931,1983,2010]" metricMagnitude="2" metricUnit="m" metricValue="5.6" pageId="4" pageNumber="204" unit="m" value="560.0">
<elevation id="18E5BA4BFFFDF22BDF03F84BFC51F812" box="[855,931,1983,2010]" metricMagnitude="2" metricUnit="m" metricValue="5.6" pageId="4" pageNumber="204" unit="m" value="560.0">560 m</elevation>
</quantity>
,
<specimenCount id="85CE96F1FFFDF22BDFFBF84BFC26F812" box="[943,980,1983,2010]" count="2" pageId="4" pageNumber="204" type="female">2♀</specimenCount>
, slide no. 24
<specimenCode id="C36EF503FFFDF22BD83DF84BFB4FF812" box="[1129,1213,1983,2010]" collectionCode="GP" country="Brazil" name="Instituto de Geociencias, Universidade de Sao Paulo" pageId="4" pageNumber="204">GP003</specimenCode>
,
<specimenCode id="C36EF503FFFDF22BD893F84BFB16F812" box="[1223,1252,1983,2010]" collectionCode="GP" country="Brazil" name="Instituto de Geociencias, Universidade de Sao Paulo" pageId="4" pageNumber="204">23</specimenCode>
<specimenCode id="C36EF503FFFDF22BD8B1F84BFAC8F812" box="[1253,1338,1983,2010]" collectionCode="GP" country="Brazil" name="Instituto de Geociencias, Universidade de Sao Paulo" pageId="4" pageNumber="204">GP061</specimenCode>
, leg., prep., coll.
<collectorName id="3E3D38AEFFFDF22BDD16F816FE29F834" box="[322,475,2017,2044]" pageId="4" pageNumber="204">M. Seizmair.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="93775D78FFFCF22ADD48FF30FB29FDBB" blockId="5.[189,1399,195,964]" pageId="5" pageNumber="205">
<emphasis id="A1BC816AFFFCF22ADD48FF30FB4BFF16" bold="true" box="[284,1209,196,222]" pageId="5" pageNumber="205">
External diagnostic characters (
<figureCitation id="0BF341FDFFFCF22ADEF2FF30FCEBFF16" box="[678,793,196,222]" captionStart="Figure 1" captionStartId="3.[189,264,1123,1147]" captionTargetBox="[196,1388,664,1103]" captionTargetId="figure-666@3.[196,1392,664,1103]" captionTargetPageId="3" captionText="Figure 1. Adults. A D: L. forsskali, A, C: male, upperside and underside, Saudi Arabia, Prov. Jizan, Mt. Faifa, Marwah, 960 m, 10-XI-2023, leg. et coll. M. Seizmair B, D: female, upperside and underside, Saudi Arabia, Prov. Jizan, Mt. Faifa, Faifa Village, 1560 m, 13-XI-2023, leg et coll. M. Seizmair, E H: L. haveni, E, G: male, upperside and underside, Saudi Arabia, Prov. Jizan, Mt. Faifa, Aayban Bengazi, 660 m, 08-XI-2023, leg. et coll. M. Seizmair, F, H: female, upperside and underside, same collection data as B, D. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.13247939" httpUri="https://zenodo.org/record/13247939/files/figure.png" pageId="5" pageNumber="205">Figures 1</figureCitation>
E-H,
<bibRefCitation id="F7592089FFFCF22ADF09FF30FC0DFF16" author="Larsen, T. B." box="[861,1023,196,222]" pageId="5" pageNumber="205" pagination="333 - 478" refId="ref5547" refString="Larsen, T. B. (1983) Insects of Saudi Arabia. Lepidoptera; Rhopalocera (A Monograph of the Butterflies of the Arabian Peninsula). Fauna of Saudi Arabia, 5, 333 - 478." type="journal article" year="1983">Larsen, 1983</bibRefCitation>
,
<bibRefCitation id="F7592089FFFCF22AD85FFF30FB58FF16" author="Turlin, B." box="[1035,1194,196,222]" pageId="5" pageNumber="205" pagination="13 - 19" refId="ref5681" refString="Turlin, B. (2000) Notes complementaires sur le Yemen. Entomologia Africana, 5, 13 - 19." type="journal article" year="2000">Turlin, 2000</bibRefCitation>
):
</emphasis>
Wingspan: 23.0 mm 27.0 mm. Thorax and abdomen grey scaled. FW and HW ground of the male silverish blue. Basal and sub-basal areas in the FW and HW UPS in both sexes of a shining metallic blue. End-cellular spot prominent, claviform in the FW UPS of the male and in the FW and HW
<collectionCode id="F5D9C5BDFFFCF22AD87DFEDCFB96FE8B" box="[1065,1124,296,323]" country="Viet Nam" httpUri="http://grbio.org/cool/0nv9-p2k7" name="University of Science, Ho Chi Minh City, Vietnam" pageId="5" pageNumber="205">UNS</collectionCode>
of both sexes. HW E
<superScript id="64BDF030FFFCF22AD930FED1FA9FFEFE" attach="right" box="[1380,1389,293,310]" fontSize="7" pageId="5" pageNumber="205">2</superScript>
- cells in the male UPS with weakly indicated extensive blackish spots - except for cell E
<superScript id="64BDF030FFFCF22AD88FFEB3FB01FEAD" attach="right" box="[1243,1267,327,357]" fontSize="7" pageId="5" pageNumber="205">22</superScript>
, where the black spot is considerably more distinct and surrounded by greyish-white scales, orange scales weakly indicated in the proximal half of the spot in E
<superScript id="64BDF030FFFCF22ADEB4FE7FFD09FE61" attach="right" box="[736,763,395,425]" fontSize="7" pageId="5" pageNumber="205">22</superScript>
. Spots in the cells E
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E
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in
</quantity>
the HW
<collectionCode id="F5D9C5BDFFFCF22AD88AFE7AFAEBFE61" box="[1246,1305,398,425]" country="Viet Nam" httpUri="http://grbio.org/cool/0nv9-p2k7" name="University of Science, Ho Chi Minh City, Vietnam" pageId="5" pageNumber="205">UNS</collectionCode>
of both sexes brownish, slightly sagittate, spot in cell E
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metallic blue in its distal part, yellowish in its proximal part. FW and HW ground of the UPS in the female brownish. Female FW and HW UPS margin with weakly indicated interneural strokes, HW cell E
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with a distinct blackish spot, with pale orange scales above its proximal margin, cell E
<superScript id="64BDF030FFFCF22ADE18FDE6FD96FDF8" attach="left" box="[588,612,530,560]" fontSize="7" pageId="5" pageNumber="205">21</superScript>
bare from spots, brownish spots in cells E
<superScript id="64BDF030FFFCF22AD836FDE6FB88FDF8" attach="right" box="[1122,1146,530,560]" fontSize="7" pageId="5" pageNumber="205">23</superScript>
E
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surrounded by greyish-white scales. M
<superScript id="64BDF030FFFCF22ADD86FDC7FE29FD8C" attach="left" box="[466,475,563,580]" fontSize="7" pageId="5" pageNumber="205">1</superScript>
spot row in the female FW
<collectionCode id="F5D9C5BDFFFCF22ADF43FDC2FCA0FD99" box="[791,850,566,593]" country="Viet Nam" httpUri="http://grbio.org/cool/0nv9-p2k7" name="University of Science, Ho Chi Minh City, Vietnam" pageId="5" pageNumber="205">UNS</collectionCode>
strongly reduced in size and nearly evanescent. The
<collectionCode id="F5D9C5BDFFFCF22ADCA6FDACFEDFFDBB" box="[242,301,600,627]" country="Viet Nam" httpUri="http://grbio.org/cool/0nv9-p2k7" name="University of Science, Ho Chi Minh City, Vietnam" pageId="5" pageNumber="205">UNS</collectionCode>
maculation is visible through the UPS in both wings. The hind wing is untailed.
</paragraph>
<paragraph id="93775D78FFFCF22ADD48FD8EFAB7FC0C" blockId="5.[189,1399,195,964]" pageId="5" pageNumber="205">
<emphasis id="A1BC816AFFFCF22ADD48FD8EFCD2FD5C" bold="true" box="[284,800,634,660]" pageId="5" pageNumber="205">
Male genitalia (
<figureCitation id="0BF341FDFFFCF22ADDB1FD8EFD42FD5C" box="[485,688,634,660]" captionStart="Figure 2" captionStartId="4.[189,264,1621,1645]" captionTargetBox="[191,1398,189,1602]" captionTargetId="figure-165@4.[189,1398,189,1602]" captionTargetPageId="4" captionText="Figure 2. Male genitalia. A C: L. forsskali, slide no. 23GP056, A: genitalia capsule, phallus omitted, B: phallus, lateral view, C: ventral view. D F: L. haveni, slide no. 24GP001, D: genitalia capsule, phallus omitted, E: lateral view, F: ventro-lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13247941" httpUri="https://zenodo.org/record/13247941/files/figure.png" pageId="5" pageNumber="205">Figures 2E G</figureCitation>
,
<figureCitation id="0BF341FDFFFCF22ADE95FD8EFD3DFD5C" box="[705,719,634,660]" captionStart="Figure 3" captionStartId="5.[189,264,1881,1905]" captionTargetBox="[189,1398,1002,1863]" captionTargetId="figure-500@5.[189,1398,1002,1863]" captionTargetPageId="5" captionText="Figure 3. Male genitalia, close-ups. A C: L. forsskali, slide no. 22GP056, A. Basal valva, sacculus, B: postbasal valva, apex, C: juxta. D F: L. haveni, slide no. 24GP001, D: basal valva, sacculus, E: postbasal valva, apex, F: juxta. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13247943" httpUri="https://zenodo.org/record/13247943/files/figure.png" pageId="5" pageNumber="205">3</figureCitation>
D- F):
</emphasis>
Uncus lobes quasi digitiform, medially rounded, dorso-distally with tufts of long chaetae. Subunci dilated and strongly sclerotized in the basis, curved and tapered after the anterior fourth, tip weakly hooked, obtusely angled. Vinculum of constant width, rounded in the anterior border. Juxta lobes weakly sclerotized with thin, thorn-shaped sclerites at the posterior ends, bare from chaetae. Basal valva chaetose in the ventral area, costal border convexly projected at the anterior end. Sacculus of triangular shape and strongly prominent from the ventral border of the valva, medially with a concave projection. Post-basal valva constant in width, ventral and costal borders parallelled, with several tufts of chaetae in the costal area, sub-apical area straight, apex acuminate and right-angled. Lateral parts of the distal portion of the aedeagus slightly concave in the anterior half, convex in the posterior half. Lateral flanges of the proximal portion broadly rounded.
</paragraph>
<caption id="C7B70DF0FFFCF22ADCE9F8ADFE3FF866" ID-DOI="http://doi.org/10.5281/zenodo.13247943" ID-Zenodo-Dep="13247943" httpUri="https://zenodo.org/record/13247943/files/figure.png" pageId="5" pageNumber="205" startId="5.[189,264,1881,1905]" targetBox="[189,1398,1002,1863]" targetPageId="5" targetType="figure">
<paragraph id="93775D78FFFCF22ADCE9F8ADFE3FF866" blockId="5.[189,1398,1881,1966]" pageId="5" pageNumber="205">
<emphasis id="A1BC816AFFFCF22ADCE9F8ADFEEEF8B9" bold="true" box="[189,284,1881,1905]" pageId="5" pageNumber="205">Figure 3</emphasis>
. Male genitalia, close-ups. A C:
<taxonomicName id="54C826FBFFFCF22ADED9F8AEFCF7F8B9" authorityName="Larsen" authorityYear="1982" box="[653,773,1881,1905]" class="Insecta" family="Lycaenidae" genus="Lepidochrysops" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="205" phylum="Arthropoda" rank="species" species="forsskali">
<emphasis id="A1BC816AFFFCF22ADED9F8AEFCF7F8B9" box="[653,773,1881,1905]" italics="true" pageId="5" pageNumber="205">L. forsskali</emphasis>
</taxonomicName>
, slide no. 22GP056, A. Basal valva, sacculus, B: postbasal valva, apex, C: juxta. D F:
<taxonomicName id="54C826FBFFFCF22ADDA2F88DFDA8F858" authorityName="Larsen" authorityYear="1983" box="[502,602,1912,1936]" class="Insecta" family="Lycaenidae" genus="Lepidochrysops" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="205" phylum="Arthropoda" rank="species" species="haveni">
<emphasis id="A1BC816AFFFCF22ADDA2F88DFDA8F858" box="[502,602,1912,1936]" italics="true" pageId="5" pageNumber="205">L. haveni</emphasis>
</taxonomicName>
, slide no. 24GP001, D: basal valva, sacculus, E: postbasal valva, apex, F: juxta. Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="93775D78FFFFF229DD48FF30FE56FEAD" blockId="6.[189,1399,195,357]" pageId="6" pageNumber="206">
<emphasis id="A1BC816AFFFFF229DD48FF30FD4BFF16" bold="true" box="[284,697,196,222]" pageId="6" pageNumber="206">
Female genitalia (
<figureCitation id="0BF341FDFFFFF229DDA8FF30FD86FF16" box="[508,628,196,222]" captionStart="Figure 4" captionStartId="6.[189,264,992,1016]" captionTargetBox="[193,1397,382,985]" captionTargetId="figure-513@6.[189,1398,382,985]" captionTargetPageId="6" captionText="Figure 4. Female genitalia. A C: L. forsskali, slide no. 24GP002, A. genitalia capsule, B: close-up, genitalia plate (lateral view), colliculum, C: close-up, anterior apophyses. D F: L. haveni, slide no. 24GP04, D: genitalia capsule, E: close-up, genitalia plate (lateral view), colliculum, F: close-up, anterior apophyses. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13247945" httpUri="https://zenodo.org/record/13247945/files/figure.png" pageId="6" pageNumber="206">Figures 4</figureCitation>
D-F):
</emphasis>
Ovipositor lobes ovate, chaetose, strongly narrowed at the ventral end. Posterior apophyses slender, basally angled, oblong, distal end slightly dilated. Anterior apophyses very short, of triangular shape. The genitalia plate is with several elongate rod-shaped sclerites. Colliculum with numerous short stroke-shaped sclerites. Ductus bursae membranous, bare from sclerotization.
</paragraph>
<caption id="C7B70DF0FFFFF229DCE9FC14FA85FBFD" ID-DOI="http://doi.org/10.5281/zenodo.13247945" ID-Zenodo-Dep="13247945" httpUri="https://zenodo.org/record/13247945/files/figure.png" pageId="6" pageNumber="206" startId="6.[189,264,992,1016]" targetBox="[193,1397,382,985]" targetPageId="6" targetType="figure">
<paragraph id="93775D78FFFFF229DCE9FC14FA85FBFD" blockId="6.[189,1399,992,1077]" pageId="6" pageNumber="206">
<emphasis id="A1BC816AFFFFF229DCE9FC14FEEDFC30" bold="true" box="[189,287,992,1016]" pageId="6" pageNumber="206">Figure 4</emphasis>
. Female genitalia. A C:
<taxonomicName id="54C826FBFFFFF229DE12FC15FD4DFC30" authorityName="Larsen" authorityYear="1982" box="[582,703,992,1016]" class="Insecta" family="Lycaenidae" genus="Lepidochrysops" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="206" phylum="Arthropoda" rank="species" species="forsskali">
<emphasis id="A1BC816AFFFFF229DE12FC15FD4DFC30" box="[582,703,992,1016]" italics="true" pageId="6" pageNumber="206">L. forsskali</emphasis>
</taxonomicName>
, slide no. 24GP002, A. genitalia capsule, B: close-up, genitalia plate (lateral view), colliculum, C: close-up, anterior apophyses. D F:
<taxonomicName id="54C826FBFFFFF229DF94FBF4FBD6FBDF" authorityName="Larsen" authorityYear="1983" box="[960,1060,1023,1047]" class="Insecta" family="Lycaenidae" genus="Lepidochrysops" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="206" phylum="Arthropoda" rank="species" species="haveni">
<emphasis id="A1BC816AFFFFF229DF94FBF4FBD6FBDF" box="[960,1060,1023,1047]" italics="true" pageId="6" pageNumber="206">L. haveni</emphasis>
</taxonomicName>
, slide no. 24GP04, D: genitalia capsule, E: close-up, genitalia plate (lateral view), colliculum, F: close-up, anterior apophyses. Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="93775D78FFFFF229DCE9FB76FB09FB57" blockId="6.[189,1275,1154,1183]" box="[189,1275,1154,1183]" pageId="6" pageNumber="206">
<emphasis id="A1BC816AFFFFF229DCE9FB76FB09FB57" bold="true" box="[189,1275,1154,1183]" pageId="6" pageNumber="206">
Main genital morphological characters differentiating
<taxonomicName id="54C826FBFFFFF229DFF1FB76FBC9FB57" authorityName="Larsen" authorityYear="1982" box="[933,1083,1154,1183]" class="Insecta" family="Lycaenidae" genus="Lepidochrysops" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="206" phylum="Arthropoda" rank="species" species="forsskali">
<emphasis id="A1BC816AFFFFF229DFF1FB76FBC9FB57" bold="true" box="[933,1083,1154,1183]" italics="true" pageId="6" pageNumber="206">L. forsskali</emphasis>
</taxonomicName>
and
<taxonomicName id="54C826FBFFFFF229D82AFB76FB09FB57" authorityName="Larsen" authorityYear="1983" box="[1150,1275,1154,1183]" class="Insecta" family="Lycaenidae" genus="Lepidochrysops" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="206" phylum="Arthropoda" rank="species" species="haveni">
<emphasis id="A1BC816AFFFFF229D82AFB76FB09FB57" bold="true" box="[1150,1275,1154,1183]" italics="true" pageId="6" pageNumber="206">L. haveni</emphasis>
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="93775D78FFFFF229DCE9FB33FDA1FACB" blockId="6.[189,1399,1223,1283]" pageId="6" pageNumber="206">
The main diagnostic characters in the male and female genitalia for differentiating
<taxonomicName id="54C826FBFFFFF229D8C9FB3CFAD1FB2A" authorityName="Larsen" authorityYear="1982" box="[1181,1315,1223,1250]" class="Insecta" family="Lycaenidae" genus="Lepidochrysops" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="206" phylum="Arthropoda" rank="species" species="forsskali">
<emphasis id="A1BC816AFFFFF229D8C9FB3CFAD1FB2A" box="[1181,1315,1223,1250]" italics="true" pageId="6" pageNumber="206">L. forsskali</emphasis>
</taxonomicName>
and
<taxonomicName id="54C826FBFFFFF229D934FB3CFEF9FACB" authorityName="Larsen" authorityYear="1983" class="Insecta" family="Lycaenidae" genus="Lepidochrysops" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="206" phylum="Arthropoda" rank="species" species="haveni">
<emphasis id="A1BC816AFFFFF229D934FB3CFEF9FACB" italics="true" pageId="6" pageNumber="206">L. haveni</emphasis>
</taxonomicName>
are summarized in
<tableCitation id="DE4A68C3FFFFF229DDA7FB1CFDBFFACB" box="[499,589,1256,1283]" captionStart="Table 1" captionStartId="6.[189,254,1339,1363]" captionText="Table 1. Diagnostic characters in the male and female genitalia of L. forsskali and L. haveni." pageId="6" pageNumber="206">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>