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<document ID-CLB-Dataset="110764" ID-DOI="http://dx.doi.org/10.3897/zookeys.1115.83946" ID-GBIF-Dataset="530a7452-695e-4332-babf-897c5fb67f9c" ID-PMC="PMC9848819" ID-Pensoft-Pub="1313-2970-1115-117" ID-Pensoft-UUID="27759136B7BE5C8587DB94CE232B2C99" ID-PubMed="36761074" ID-ZooBank="AB89506184444026B59337331E598259" ModsDocID="1313-2970-1115-117" checkinTime="1659410023474" checkinUser="pensoft" docAuthor="Tsukamoto, Sho, Shimano, Satoshi &amp; Eguchi, Katsuyuki" docDate="2022" docId="1B1A7490A2C854008BEFD6BC3F5DAE32" docLanguage="en" docName="ZooKeys 1115: 117-150" docOrigin="ZooKeys 1115" docPubDate="2022-08-01" docSource="http://dx.doi.org/10.3897/zookeys.1115.83946" docTitle="Nannarrup oyamensis Tsukamoto 2022, sp. nov." docType="treatment" docUuid="1543ADD5-1C03-4471-9B6F-D473E4BB0F22" docUuidSource="ZooBank" docVersion="5" id="27759136B7BE5C8587DB94CE232B2C99" lastPageNumber="117" masterDocId="27759136B7BE5C8587DB94CE232B2C99" masterDocTitle="Two new species of the dwarf centipede genus Nannarrup Foddai, Bonato, Pereira &amp; Minelli, 2003 (Chilopoda, Geophilomorpha, Mecistocephalidae) from Japan" masterLastPageNumber="150" masterPageNumber="117" pageNumber="117" updateTime="1732811224425" updateUser="ExternalLinkService">
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<mods:title id="C2BA612E03B06E7447754959637D9014">Two new species of the dwarf centipede genus Nannarrup Foddai, Bonato, Pereira &amp; Minelli, 2003 (Chilopoda, Geophilomorpha, Mecistocephalidae) from Japan</mods:title>
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<mods:namePart id="3BB7CD3396B16EB498FA447002FD6532">Tsukamoto, Sho</mods:namePart>
<mods:nameIdentifier id="1DF089F4C10012B8AB2BEBBB43983726" type="ORCID">https://orcid.org/0000-0003-3020-8454</mods:nameIdentifier>
<mods:affiliation id="6B3D8BF057D797AB09E615030A434A72">Systematic Zoology Laboratory, Graduate School of Science, Tokyo Metropolitan University, Minami-osawa 1 - 1 Hachioji-shi, Tokyo 192 - 0397, Japan</mods:affiliation>
<mods:nameIdentifier id="F66EB4A657E2C6C10CC62122E0B8DE89" type="email">esutukamoto153@gmail.com</mods:nameIdentifier>
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<mods:name id="AF57247248F9E3783A3F20AE7C3BB3B0" type="personal">
<mods:role id="CC3B8908D829434AB23B64D9454A443C">
<mods:roleTerm id="9C4D96973A18B81EF60AF9EE79D3BB3B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DDD31A4662A38810DD3A988BC0D132B7">Shimano, Satoshi</mods:namePart>
<mods:affiliation id="8E8C9BAC34A1A66E272B560C406E1112">Science Research Center, Hosei University, Fujimi 2 - 17 - 1 Chiyoda-ku, Tokyo 102 - 8160, Japan</mods:affiliation>
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<mods:namePart id="98DCA84403A110E0A7CD2D13F358CA70">Eguchi, Katsuyuki</mods:namePart>
<mods:nameIdentifier id="2D87F0DAB1A5C0ACBD62E6BE16D95EAC" type="ORCID">https://orcid.org/0000-0002-1054-1295</mods:nameIdentifier>
<mods:affiliation id="B18C3621ECE2D293B7477F269A4C0309">Systematic Zoology Laboratory, Graduate School of Science, Tokyo Metropolitan University, Minami-osawa 1 - 1 Hachioji-shi, Tokyo 192 - 0397, Japan &amp; Department of International Health and Medical Anthropology, Institute of Tropical Medicine, Nagasaki University, Sakamoto 1 - 12 - 4, Nagasaki, Nagasaki 852 - 8523, Japan</mods:affiliation>
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<mods:date id="067B7BA0918078EF30922E135C5A52DA">2022</mods:date>
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<mods:number id="23ABD1910CFD2C38A1701B9DBA51F22B">2022-08-01</mods:number>
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<mods:number id="7E448FC6A9090704582E76F04F208693">1115</mods:number>
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<subSection id="5BE503B85DD12B1072C43530BF790EB8" pageId="0" pageNumber="117" type="results">
<treatment id="1B1A7490A2C854008BEFD6BC3F5DAE32" ID-GBIF-Taxon="198401537" LSID="urn:lsid:zoobank.org:act:1543ADD5-1C03-4471-9B6F-D473E4BB0F22" httpUri="http://treatment.plazi.org/id/1B1A7490A2C854008BEFD6BC3F5DAE32" lastPageNumber="117" pageId="0" pageNumber="117" scope_class="Chilopoda" scope_family="Mecistocephalidae" scope_order="Geophilomorpha">
<subSubSection id="915624C3EF1937EA9E2AE583B27EBB59" pageId="0" pageNumber="117" type="nomenclature">
<paragraph id="9F6D1D1D516C76091610AD5A76D9274A" pageId="0" pageNumber="117">
<taxonomicName id="58CCA8DEDF8FA8F6B8554E90B2349B5C" LSID="https://zoobank.org/1543ADD5-1C03-4471-9B6F-D473E4BB0F22" authority="Tsukamoto" authorityName="Tsukamoto" authorityYear="2022" class="Chilopoda" family="Mecistocephalidae" genus="Nannarrup" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nannarrup oyamensis" order="Geophilomorpha" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="oyamensis" status="sp. nov.">Nannarrup oyamensis Tsukamoto</taxonomicName>
<taxonomicNameLabel id="464971F9D3FE3D2CC3DB59EC181540E2" pageId="0" pageNumber="117">sp. nov.</taxonomicNameLabel>
</paragraph>
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<subSubSection id="84F9B0388826C3E96E0DEB9478D5FFD0" pageId="0" pageNumber="117" type="description">
<paragraph id="F6F4D0129546FA462DFED5AE1E15CF08" pageId="0" pageNumber="117">
<figureCitation id="195CF6FE6E279964CA809053FE93D7D4" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Nannarrup oyamensis sp. nov., holotype (TS 20210217 - 04) A cephalic plate, dorsal B clypeus and clypeal pleurite, ventral. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1115.83946.figure12" httpUri="https://binary.pensoft.net/fig/722238" pageId="0" pageNumber="117">New Japanese name: Amefuri-himejimukade Figs 12</figureCitation>
<figureCitation id="13273D5E115E1E2C7ABDE775E28F29F7" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Nannarrup oyamensis sp. nov., holotype (TS 20210217 - 04) A right antenna, dorsal B right antenna, ventral C antennal article XIV, dorsal D antennal article XIV, ventral. Arrows indicate apical sensillum; arrowheads indicate club-like sensillum. Scale bars: 0.1 mm (A, B); 0.05 mm (C, D)." figureDoi="10.3897/zookeys.1115.83946.figure13" httpUri="https://binary.pensoft.net/fig/722239" pageId="0" pageNumber="117">, 13</figureCitation>
<figureCitation id="17D7F4D9C604053D3D875CB888F87208" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Nannarrup oyamensis sp. nov., holotype (TS 20210217 - 04) A right mandible, ventral B maxillae complex, ventral. Scale bars: 0.05 mm." figureDoi="10.3897/zookeys.1115.83946.figure14" httpUri="https://binary.pensoft.net/fig/722240" pageId="0" pageNumber="117">, 14</figureCitation>
<figureCitation id="F5529B363BFFF7BC59C07E508375695A" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Nannarrup oyamensis sp. nov., holotype (TS 20210217 - 04) A forcipular segment, dorsal B forcipular segment, ventral C denticles on right forcipule, dorsal D left poison calyx, dorsal. Circle indicates poison calyx. Scale bars: 0.1 mm (A, B); 0.05 mm (C, D)." figureDoi="10.3897/zookeys.1115.83946.figure15" httpUri="https://binary.pensoft.net/fig/722241" pageId="0" pageNumber="117">, 15</figureCitation>
<figureCitation id="3A7DAF269FCA5D880B194D7CBE277003" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Nannarrup oyamensis sp. nov., holotype (TS 20210217 - 04) A pretarsus of left leg 40, dorsal B pretarsus of left leg 40, ventral C tergite of leg-bearing segment 40, dorsal D sternite of leg-bearing segment 40, ventral. Scale bars: 0.05 mm (A, B); 0.1 mm (C, D)." figureDoi="10.3897/zookeys.1115.83946.figure16" httpUri="https://binary.pensoft.net/fig/722242" pageId="0" pageNumber="117">, 16</figureCitation>
<figureCitation id="95DC6148CAFA6E767641C833A7429A73" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Nannarrup oyamensis sp. nov. A, B holotype (TS 20210217 - 04) C, D paratype (TS 20210725 - 02). A ultimate leg-bearing segment and male postpedal segment, dorsal B ultimate leg-bearing segment and male postpedal segment, ventral C right ultimate leg, dorsal D right ultimate leg, ventral. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1115.83946.figure17" httpUri="https://binary.pensoft.net/fig/722243" pageId="0" pageNumber="117">, 17</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="FF57571DED23A63303526AC63CAADB90" pageId="0" pageNumber="117" type="type material">
<paragraph id="5FC019F7A536330C3EB0B0862E3EE5AB" pageId="0" pageNumber="117">Type material.</paragraph>
<paragraph id="7DBC5809BA97FE22899EAABDAEB88704" pageId="0" pageNumber="117">
<emphasis id="750DF210847679CBF5309D8E205878AE" bold="true" italics="true" pageId="0" pageNumber="117">Holotype</emphasis>
1 adult male, Hinata, Isehara-shi, Kanagawa prefecture, Japan (
<geoCoordinate id="C7C3E327BD6C81CE42A943E7F9AAF077" degrees="35" direction="north" minutes="26.07" orientation="latitude" precision="9" value="35.4345">35°26.07'N</geoCoordinate>
,
<geoCoordinate id="AD6B27231B57D008F94FD99F2B7DAA7B" degrees="139" direction="east" minutes="14.75" orientation="longitude" precision="9" value="139.24583">139°14.75'E</geoCoordinate>
), 17 February 2021, coll. Sho Tsukamoto (labeled as TS20210217-04), deposited at the Collection of
<taxonomicName id="8129239DDD6DB11F293B1D30C1E9ED3C" authorityName="Foddai, Bonato, Pereira &amp; Minelli" authorityYear="2003" kingdom="Animalia" lsidName="" pageId="0" pageNumber="117" rank="subPhylum" subPhylum="Myriapoda">Myriapoda</taxonomicName>
, Department of Zoology,
<emphasis id="E73BC48DBC061F47FEE5C0EFCC4AD47B" bold="true" pageId="0" pageNumber="117">NSMT</emphasis>
.
<emphasis id="A06FB739499F5ED1B5631B4BCBB5DDF4" bold="true" italics="true" pageId="0" pageNumber="117">Paratype</emphasis>
1 subadult male, Hinata, Isehara-shi, Kanagawa prefecture, Japan (
<geoCoordinate id="6B3680D43CE0638BB9BE82EC80BFB757" degrees="35" direction="north" minutes="26.07" orientation="latitude" precision="9" value="35.4345">35°26.07'N</geoCoordinate>
,
<geoCoordinate id="BB7BB9FF255810CE97E234C68D7B54C1" degrees="139" direction="east" minutes="14.75" orientation="longitude" precision="9" value="139.24583">139°14.75'E</geoCoordinate>
), 25 July 2021, coll. Sho Tsukamoto (labeled as TS20210725-02), deposited at
<emphasis id="8C19B51EDF4238337FB6D63FF3522F99" bold="true" pageId="0" pageNumber="117">MNHAH</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="30492DF5B004CDBFC48DCB502A2B4318" pageId="0" pageNumber="117" type="etymology">
<paragraph id="16C7F28BBFB05B66915FB1ACA87B450F" pageId="0" pageNumber="117">Etymology.</paragraph>
<paragraph id="1618F615E433F5404CC84A202136292F" pageId="0" pageNumber="117">
The species name is derived from the name of Japanese mountain, namely Mt. Oyama. The word was further Latinized by adding the Latin masculine adjective suffix -
<emphasis id="85CE40C097996127C3EC2C5E59B46451" italics="true" pageId="0" pageNumber="117">ensis</emphasis>
, to form
<taxonomicName id="6DDF925119C4A31E1B79655D5D2D8B9B" authority="oyamensis" class="Chilopoda" family="Mecistocephalidae" form="oyamensis" genus="Nannarrup" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nannarrup oyamensis" order="Geophilomorpha" pageId="0" pageNumber="117" phylum="Arthropoda" rank="form" species="oyamensis">Nannarrup oyamensis oyamensis</taxonomicName>
. The last
<normalizedToken id="1C3FDDA3EB6D0A266E5329B6AC5C1C41" originalValue="“a”">&quot;a&quot;</normalizedToken>
of Oyama and the first
<normalizedToken id="19D1DE3BF9EF6CDAB9FEC5658121E4D6" originalValue="“e”">&quot;e&quot;</normalizedToken>
of -ensis are merged into
<normalizedToken id="6513CCD9C5E814C1023FF47AF9C3A398" originalValue="“e.”">&quot;e.&quot;</normalizedToken>
Examined specimens were collected from Mt. Oyama, an object of the mountain worship.
</paragraph>
</subSubSection>
<subSubSection id="F296754E1A8721968ECD749C4E4FE16A" pageId="0" pageNumber="117" type="diagnosis">
<paragraph id="4070E841BC20370FDC2B57213D5DFEB8" pageId="0" pageNumber="117">Diagnosis.</paragraph>
<paragraph id="1C07570F5815540E358099DBA94A57F3" pageId="0" pageNumber="117">Clypeus without smooth or weakly areolate areas along the posterior part of the paraclypeal sutures; forcipular trochantroprefemur with a large denticle (longer than wide); tarsungulum with a well-pigmented denticle; metasternite of ultimate leg-bearing segment wider than long.</paragraph>
</subSubSection>
<subSubSection id="ED2435C5EE6E46DF488D9D9F21FB14AD" pageId="0" pageNumber="117" type="description">
<paragraph id="0B8E5F6FEE51A1ADF3FECB93D811B687" pageId="0" pageNumber="117">Description.</paragraph>
<paragraph id="BDE6271FA557CF0D0965923D9C4101E2" pageId="0" pageNumber="117">
<emphasis id="414FAB43990E11E42F048921C55CA11F" bold="true" italics="true" pageId="0" pageNumber="117">General features</emphasis>
: Body 8.6 mm long (holotype), gradually attenuate posterior, almost uniformly very pale yellow, with head and forcipular segment pale ocher.
</paragraph>
<paragraph id="3B6328EBF2C6B250C4ACED7249C82BA8" pageId="0" pageNumber="117">
<emphasis id="26E869556ED63D4C0B86CA0904518194" bold="true" italics="true" pageId="0" pageNumber="117">Cephalic capsule</emphasis>
(Fig.
<figureCitation id="58CE3F7C17F3660BBF021818B1C79C61" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Nannarrup oyamensis sp. nov., holotype (TS 20210217 - 04) A cephalic plate, dorsal B clypeus and clypeal pleurite, ventral. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1115.83946.figure12" httpUri="https://binary.pensoft.net/fig/722238" pageId="0" pageNumber="117">12A, B</figureCitation>
): Cephalic plate ca 1.5
<normalizedToken id="5FBBA17E77F89D83EA8A42258C6634A9" originalValue="×">x</normalizedToken>
as long as wide, lateral margins more distinctly converging anteriorly than posteriorly, posterior margin straight; scutes approximately isometric and up to ca 15
<normalizedToken id="E9DE846E9B17F87FB21CAC13ADC537F4" originalValue="μm">μm</normalizedToken>
wide; transverse suture absent; setae up to ca 50
<normalizedToken id="6C568E868087E67FE800A6643AE0CE12" originalValue="μm">μm</normalizedToken>
long. Clypeus ca 1.5
<normalizedToken id="E2CB8F6C6646564D477C189D69539614" originalValue="×">x</normalizedToken>
as wide as long, with lateral margins complete, almost uniformly areolate, with scutes ca 10
<normalizedToken id="ED313FFCCF3086BD4634AAF21850F0D2" originalValue="μm">μm</normalizedToken>
wide, a pair of clypeal areas absent; 13 setae in holotype, 1+1 postantennal, 1+1 median, 5+4 prelabral; clypeal ratio ca 1: 7; clypeal plagulae without additional smooth area along posterior part of paraclypeal sutures; 17 pore-like organs on entire part of clypeus. Anterior and distolateral parts of pleurites areolate, without setae. Side-pieces of labrum medially in contact, only incompletely divided into anterior and posterior alae by weak chitinous line, without longitudinal stripes on posterior alae; with slightly visible short fringe on posterior margin of side-pieces; mid-piece as long as wide, converging anteriorly and posteriorly.
</paragraph>
<caption id="F1A0EDF670DEC39080453740E84C46BF" doi="10.3897/zookeys.1115.83946.figure12" httpUri="https://binary.pensoft.net/fig/722238" pageId="0" pageNumber="117" start="Figure 12" startId="F12">
<paragraph id="20E152E08D78A25076F9886828886FDC" pageId="0" pageNumber="117">
<emphasis id="2875AB0C7EE2230381D491FB27621802" bold="true" pageId="0" pageNumber="117">Figure 12.</emphasis>
<taxonomicName id="42C0E181FF2EF2E40AE480F3AD11992A" authorityName="Tsukamoto" authorityYear="2022" class="Chilopoda" family="Mecistocephalidae" genus="Nannarrup" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nannarrup oyamensis" order="Geophilomorpha" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="oyamensis">
<emphasis id="7D99AD23A6E77D570BB77B0F988F399C" italics="true" pageId="0" pageNumber="117">Nannarrup oyamensis</emphasis>
</taxonomicName>
sp. nov., holotype (TS20210217-04)
<emphasis id="0D1BB9DB65B8E71428D6E448CA8FFBAE" bold="true" pageId="0" pageNumber="117">A</emphasis>
cephalic plate, dorsal
<emphasis id="4060374CA431332E4CA70E347EB67636" bold="true" pageId="0" pageNumber="117">B</emphasis>
clypeus and clypeal pleurite, ventral. Scale bars: 0.1 mm.
</paragraph>
</caption>
<paragraph id="6B11FA3AF75BFD2A96858B613F8A0C67" pageId="0" pageNumber="117">
<emphasis id="307A3AADE9053B842BDB6D8A2C8A51C4" bold="true" italics="true" pageId="0" pageNumber="117">Antenna</emphasis>
(Fig.
<figureCitation id="E780713C2195C4BB4C0A67412B44D8E2" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Nannarrup oyamensis sp. nov., holotype (TS 20210217 - 04) A right antenna, dorsal B right antenna, ventral C antennal article XIV, dorsal D antennal article XIV, ventral. Arrows indicate apical sensillum; arrowheads indicate club-like sensillum. Scale bars: 0.1 mm (A, B); 0.05 mm (C, D)." figureDoi="10.3897/zookeys.1115.83946.figure13" httpUri="https://binary.pensoft.net/fig/722239" pageId="0" pageNumber="117">13A-D</figureCitation>
): Antenna with 14 articles, when stretched, ca 2.3
<normalizedToken id="6DA11B355B6958DBEEAD9378EF4CC0A2" originalValue="×">x</normalizedToken>
as long as head length. Intermediate articles slightly longer than wide. Article XIV ca 2.0
<normalizedToken id="1CDB347DF6D655499B03B30D72E5168D" originalValue="×">x</normalizedToken>
as long as wide, ca 1.9
<normalizedToken id="CCD3C61A62B0861004ED3A0C1B93807A" originalValue="×">x</normalizedToken>
as long as article XIII, and 1.9-2.1
<normalizedToken id="C61EDF16F67A6C63E9613D238522F5DB" originalValue="×">x</normalizedToken>
as long as intermediate articles. Setae on articles VIII-XVI denser than articles I-VII. Setae gradually shorter from article VIII to XIV, up to 50
<normalizedToken id="CA238C01F3E008162F839AD00110B2CA" originalValue="μm">μm</normalizedToken>
long on article I, up to 33
<normalizedToken id="C94F1246F0501CCB1BAB52633141ABBB" originalValue="µm">µm</normalizedToken>
long on article VIII and &lt;18
<normalizedToken id="49D95A00A6529F16989EF596D3AFA617" originalValue="μm">μm</normalizedToken>
long on article XIV. Article XIV with two types of sensilla, apical sensilla (arrows in Fig.
<figureCitation id="DB37CEC9EFCA38400148490390506A86" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Nannarrup oyamensis sp. nov., holotype (TS 20210217 - 04) A right antenna, dorsal B right antenna, ventral C antennal article XIV, dorsal D antennal article XIV, ventral. Arrows indicate apical sensillum; arrowheads indicate club-like sensillum. Scale bars: 0.1 mm (A, B); 0.05 mm (C, D)." figureDoi="10.3897/zookeys.1115.83946.figure13" httpUri="https://binary.pensoft.net/fig/722239" pageId="0" pageNumber="117">13C, D</figureCitation>
) ca 5
<normalizedToken id="D0AB6E84F7F27A220A5C2E0493046C71" originalValue="μm">μm</normalizedToken>
long, with wide flat ring at mid-length; club-like sensilla (arrowheads in Fig.
<figureCitation id="37ACBFA84B532CFC5625AE130A714907" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Nannarrup oyamensis sp. nov., holotype (TS 20210217 - 04) A right antenna, dorsal B right antenna, ventral C antennal article XIV, dorsal D antennal article XIV, ventral. Arrows indicate apical sensillum; arrowheads indicate club-like sensillum. Scale bars: 0.1 mm (A, B); 0.05 mm (C, D)." figureDoi="10.3897/zookeys.1115.83946.figure13" httpUri="https://binary.pensoft.net/fig/722239" pageId="0" pageNumber="117">13C, D</figureCitation>
) ca 10
<normalizedToken id="5B91120E0AAEAC8B6F11681E8348072B" originalValue="μm">μm</normalizedToken>
long, clustered in the distal parts of the internal and external sides of the article. Three longitudinal rows consisted of ca 9 proprioceptive spine-like sensilla at bases of antennal articles II-V, VII-IX, approximately dorsal, ventro-internal and ventro-external; rows reduced to 1-3 spines on antennal articles I and VI, and 0-1 spine on antennal articles X-XIV. A few pointed sensilla, up to 2.5
<normalizedToken id="20CEBD4B2CDA117BCC0E9561ED005CBD" originalValue="μm">μm</normalizedToken>
long, on both dorso-external and ventro-internal position, close to distal margin of articles II, V, IX and XIII.
</paragraph>
<caption id="AEFAC3F9FF7F090588C38173DBF6D9EE" doi="10.3897/zookeys.1115.83946.figure13" httpUri="https://binary.pensoft.net/fig/722239" pageId="0" pageNumber="117" start="Figure 13" startId="F13">
<paragraph id="76588F2D74AA9D712A23020487E2D88A" pageId="0" pageNumber="117">
<emphasis id="DAE48182AEC649605765A5A58B5E5C26" bold="true" pageId="0" pageNumber="117">Figure 13.</emphasis>
<taxonomicName id="3677984487EA85624381B51ADF58ADDC" authorityName="Tsukamoto" authorityYear="2022" class="Chilopoda" family="Mecistocephalidae" genus="Nannarrup" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nannarrup oyamensis" order="Geophilomorpha" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="oyamensis">
<emphasis id="6F86318BC6EA8855E871676ADA14C056" italics="true" pageId="0" pageNumber="117">Nannarrup oyamensis</emphasis>
</taxonomicName>
sp. nov., holotype (TS20210217-04)
<emphasis id="7E982C6FBF48D4F3186D695A6D6E2543" bold="true" pageId="0" pageNumber="117">A</emphasis>
right antenna, dorsal
<emphasis id="880D464EBE585FE81CD7DABCC49F1ADB" bold="true" pageId="0" pageNumber="117">B</emphasis>
right antenna, ventral
<emphasis id="6B42330805A81F69E5BC72FDE023E154" bold="true" pageId="0" pageNumber="117">C</emphasis>
antennal article XIV, dorsal
<emphasis id="C2D7A38E5006E9159ACA9D3BEFCEE249" bold="true" pageId="0" pageNumber="117">D</emphasis>
antennal article XIV, ventral. Arrows indicate apical sensillum; arrowheads indicate club-like sensillum. Scale bars: 0.1 mm (
<emphasis id="69395979819DDF493A771026690A7EB2" bold="true" pageId="0" pageNumber="117">A, B</emphasis>
); 0.05 mm (
<emphasis id="22CC7F7557F48D5BAB76B456401738F3" bold="true" pageId="0" pageNumber="117">C, D</emphasis>
).
</paragraph>
</caption>
<paragraph id="71D183D010360BADAA2B8B4B414F7A41" pageId="0" pageNumber="117">
<emphasis id="C6E02A6F160A712AF6762D8C3757CEFF" bold="true" italics="true" pageId="0" pageNumber="117">Mandible</emphasis>
(Fig.
<figureCitation id="5C4250F36A2EA0CE63F133B5C03AC8BA" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Nannarrup oyamensis sp. nov., holotype (TS 20210217 - 04) A right mandible, ventral B maxillae complex, ventral. Scale bars: 0.05 mm." figureDoi="10.3897/zookeys.1115.83946.figure14" httpUri="https://binary.pensoft.net/fig/722240" pageId="0" pageNumber="117">14A</figureCitation>
). At least four pectinate lamellae, with elongated teeth. Each tooth ca 2
<normalizedToken id="559A07A7A356094605DE9FEB2C831EE4" originalValue="×">x</normalizedToken>
as long as wide.
</paragraph>
<caption id="64EFC87D7832B8CBA8778BBA40C55A60" doi="10.3897/zookeys.1115.83946.figure14" httpUri="https://binary.pensoft.net/fig/722240" pageId="0" pageNumber="117" start="Figure 14" startId="F14">
<paragraph id="74B117B56F7C19053FFEDA4ED6D12CF5" pageId="0" pageNumber="117">
<emphasis id="A5E754F68501AF5A379A477C2380051C" bold="true" pageId="0" pageNumber="117">Figure 14.</emphasis>
<taxonomicName id="382532B20C5FAE2DAECB868DC39F760F" authorityName="Tsukamoto" authorityYear="2022" class="Chilopoda" family="Mecistocephalidae" genus="Nannarrup" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nannarrup oyamensis" order="Geophilomorpha" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="oyamensis">
<emphasis id="8656C50563F49456C63AFE1C9449284C" italics="true" pageId="0" pageNumber="117">Nannarrup oyamensis</emphasis>
</taxonomicName>
sp. nov., holotype (TS20210217-04)
<emphasis id="12BB99E05EB84645FEBE605003F922B2" bold="true" pageId="0" pageNumber="117">A</emphasis>
right mandible, ventral
<emphasis id="52E6749C4191DFDA512D4FC004901BD7" bold="true" pageId="0" pageNumber="117">B</emphasis>
maxillae complex, ventral. Scale bars: 0.05 mm.
</paragraph>
</caption>
<paragraph id="F2C33F1589CF98E31B53CF61B7513B7D" pageId="0" pageNumber="117">
<emphasis id="89D8C56C091130E9938D869881D71D62" bold="true" italics="true" pageId="0" pageNumber="117">First maxillae</emphasis>
(Fig.
<figureCitation id="6BDE3D5170E37D3AF423C424D04940E3" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Nannarrup oyamensis sp. nov., holotype (TS 20210217 - 04) A right mandible, ventral B maxillae complex, ventral. Scale bars: 0.05 mm." figureDoi="10.3897/zookeys.1115.83946.figure14" httpUri="https://binary.pensoft.net/fig/722240" pageId="0" pageNumber="117">14B</figureCitation>
): Coxosternite medially divided but slightly, without setae, faintly areolate. Coxal projections well developed and hyaline distally, provided with 1+1 setae and 3+4 small sensilla. Telopodite uni-articulated and hyaline distally, with one (two) seta(e). No lobes on either coxosternite or telopodites.
</paragraph>
<paragraph id="87E398126E0D04D7D2F20B1CB875903C" pageId="0" pageNumber="117">
<emphasis id="422FE784AEDDE2BFA468A89945A17EA0" bold="true" italics="true" pageId="0" pageNumber="117">Second maxillae</emphasis>
(Fig.
<figureCitation id="DCE211F10585857DAB6786CD34296FB9" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Nannarrup oyamensis sp. nov., holotype (TS 20210217 - 04) A right mandible, ventral B maxillae complex, ventral. Scale bars: 0.05 mm." figureDoi="10.3897/zookeys.1115.83946.figure14" httpUri="https://binary.pensoft.net/fig/722240" pageId="0" pageNumber="117">14B</figureCitation>
): Coxosternite medially undivided, without suture, with 2+3 setae along the anterior margin, with 4+4 setae located behind anterior margin, with anterior margin slightly concave, with metameric pores on posterior part. Telopodites tri-articulate overreaching medial projections and telopodites of first maxillae. Claw of telopodite virtually absent, represented by short spine only.
</paragraph>
<paragraph id="FCB51B02CE1215D904A48D58E6E1A473" pageId="0" pageNumber="117">
<emphasis id="6C8CB1721B1ED5FBEB4D6A5787FB171E" bold="true" italics="true" pageId="0" pageNumber="117">Forcipular segment</emphasis>
(Fig.
<figureCitation id="3A6C8308635B864EDAC626F56E2E1F72" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Nannarrup oyamensis sp. nov., holotype (TS 20210217 - 04) A forcipular segment, dorsal B forcipular segment, ventral C denticles on right forcipule, dorsal D left poison calyx, dorsal. Circle indicates poison calyx. Scale bars: 0.1 mm (A, B); 0.05 mm (C, D)." figureDoi="10.3897/zookeys.1115.83946.figure15" httpUri="https://binary.pensoft.net/fig/722241" pageId="0" pageNumber="117">15A-D</figureCitation>
): Tergite trapezoidal, ca 1.9
<normalizedToken id="B24261A196069B094BCF99C4B6517480" originalValue="×">x</normalizedToken>
as wide as long, with lateral margins converging anteriorly, approximately as wide as cephalic plate and ca 0.7
<normalizedToken id="C7E93E801085970D3096EA57C4AD61E2" originalValue="×">x</normalizedToken>
as wide as following tergite, 1+1 setae of similar length arranged in an anterior row, and 3+3 setae of similar length arranged in a posterior row, one pair of longitudinal rows of three tiny setae located between middle and distal setae in posterior row. Mid-longitudinal sulcus of tergite not visible. Exposed part of coxosternite ca 1.3
<normalizedToken id="0116473073D3B7F362D0BD0BCDD38BC1" originalValue="×">x</normalizedToken>
as wide as long; anterior margin with shallow medial concavity and with one pair of denticles; coxopleural sutures complete in entirely ventrum, sinuous and diverging anteriorly; chitin-lines absent. Basal distance between forcipules ca 0.1
<normalizedToken id="539FF436C526BEE4B994BBF4A5829328" originalValue="×">x</normalizedToken>
of maximum width of coxosternite. Trochanteroprefemur ca 1.3
<normalizedToken id="3F93F8D8FD080D4EF25FB20117BCF314" originalValue="×">x</normalizedToken>
as long as wide; with a well-developed and strong pigmented denticle at distal internal margin, ca 1.3
<normalizedToken id="B42C3E0CD95E4CF20EFC082BDDE8FA8C" originalValue="×">x</normalizedToken>
as long as wide. Intermediate articles distinct, tubercle on tibia not visible. Tarsungulum with basal denticle well-pigmented; both external and internal margins uniformly curved, except for moderate mesal basal bulge; ungulum not distinctly flattened. Elongated poison calyx (circle in Fig.
<figureCitation id="44D570155D06BC782B5AA9AF2BE07975" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Nannarrup oyamensis sp. nov., holotype (TS 20210217 - 04) A forcipular segment, dorsal B forcipular segment, ventral C denticles on right forcipule, dorsal D left poison calyx, dorsal. Circle indicates poison calyx. Scale bars: 0.1 mm (A, B); 0.05 mm (C, D)." figureDoi="10.3897/zookeys.1115.83946.figure15" httpUri="https://binary.pensoft.net/fig/722241" pageId="0" pageNumber="117">15D</figureCitation>
), ca 6
<normalizedToken id="D902C7C7BC1D4D36BB3D31820ABB3869" originalValue="×">x</normalizedToken>
as long as wide, lodged inside intermediate forcipular articles.
</paragraph>
<caption id="BDB804989F356BF56DCA35DD199D6AD8" doi="10.3897/zookeys.1115.83946.figure15" httpUri="https://binary.pensoft.net/fig/722241" pageId="0" pageNumber="117" start="Figure 15" startId="F15">
<paragraph id="0C30331EB0D91ECB8F294A316AFE8ECA" pageId="0" pageNumber="117">
<emphasis id="405013C7433D635CF481085183E33158" bold="true" pageId="0" pageNumber="117">Figure 15.</emphasis>
<taxonomicName id="EFF72B0BE710FA7E50637D8662A23BF4" authorityName="Tsukamoto" authorityYear="2022" class="Chilopoda" family="Mecistocephalidae" genus="Nannarrup" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nannarrup oyamensis" order="Geophilomorpha" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="oyamensis">
<emphasis id="21345310D621508C4F5373EEF28B881A" italics="true" pageId="0" pageNumber="117">Nannarrup oyamensis</emphasis>
</taxonomicName>
sp. nov., holotype (TS20210217-04)
<emphasis id="AF150FC265390FFA4AB551AC4EF0299F" bold="true" pageId="0" pageNumber="117">A</emphasis>
forcipular segment, dorsal
<emphasis id="6692DF09788CA48806BA628D572A0675" bold="true" pageId="0" pageNumber="117">B</emphasis>
forcipular segment, ventral
<emphasis id="67B7D9948F1B15BBFB7D9048FE707C98" bold="true" pageId="0" pageNumber="117">C</emphasis>
denticles on right forcipule, dorsal
<emphasis id="48BDD04C3A257AFB614B2CC772BFD4A6" bold="true" pageId="0" pageNumber="117">D</emphasis>
left poison calyx, dorsal. Circle indicates poison calyx. Scale bars: 0.1 mm (
<emphasis id="FBFF1F731C4D26746328BCDB0244198A" bold="true" pageId="0" pageNumber="117">A, B</emphasis>
); 0.05 mm (
<emphasis id="97C135E740A8D484141E4322448E83CF" bold="true" pageId="0" pageNumber="117">C, D</emphasis>
).
</paragraph>
</caption>
<paragraph id="24614E81BD8FB125907D9B68CBAF6149" pageId="0" pageNumber="117">
<emphasis id="6055EE5DA052EC827830768FE0F1445A" bold="true" italics="true" pageId="0" pageNumber="117">Leg-bearing segments</emphasis>
(Fig.
<figureCitation id="028F9224451A65354C87E89CD6E4CC4E" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Nannarrup oyamensis sp. nov., holotype (TS 20210217 - 04) A pretarsus of left leg 40, dorsal B pretarsus of left leg 40, ventral C tergite of leg-bearing segment 40, dorsal D sternite of leg-bearing segment 40, ventral. Scale bars: 0.05 mm (A, B); 0.1 mm (C, D)." figureDoi="10.3897/zookeys.1115.83946.figure16" httpUri="https://binary.pensoft.net/fig/722242" pageId="0" pageNumber="117">16A-D</figureCitation>
): Forty-one pairs of legs present. Metatergite 1 slightly wider than subsequent one, with two paramedian sulci visible on tergites of anterior half of body, without pretergite. No paratergites. Walking legs shorter than width of trunk; legs of first pair much smaller than following ones; claws simple, uniformly bent, with two accessory spines; anterior spine reaching at most 10% of length of claw; posterior spine shorter than anterior spine. Metasternites slightly longer than wide. Sternal sulcus visible on a few anterior sternites, represented by very shallow mid-longitudinal thickening, anterior not furcate. No ventral glandular pores on each metasternite.
</paragraph>
<caption id="E6DCE60CDA70163470CF568A2DBA9D15" doi="10.3897/zookeys.1115.83946.figure16" httpUri="https://binary.pensoft.net/fig/722242" pageId="0" pageNumber="117" start="Figure 16" startId="F16">
<paragraph id="DD6D2FBEF21FE7A9364F45C2DBC27A15" pageId="0" pageNumber="117">
<emphasis id="555E9800E55B99EC080F6F7FF827785D" bold="true" pageId="0" pageNumber="117">Figure 16.</emphasis>
<taxonomicName id="D8503CA824B52F7B84327BF70C393782" authorityName="Tsukamoto" authorityYear="2022" class="Chilopoda" family="Mecistocephalidae" genus="Nannarrup" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nannarrup oyamensis" order="Geophilomorpha" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="oyamensis">
<emphasis id="A08AEDA26CBDA487D2D6809A83CC8929" italics="true" pageId="0" pageNumber="117">Nannarrup oyamensis</emphasis>
</taxonomicName>
sp. nov., holotype (TS20210217-04)
<emphasis id="2DFAB658635CCC946AA752DBA0D8BBA8" bold="true" pageId="0" pageNumber="117">A</emphasis>
pretarsus of left leg 40, dorsal
<emphasis id="301265042D2776A98BE421E4DABA2FD6" bold="true" pageId="0" pageNumber="117">B</emphasis>
pretarsus of left leg 40, ventral
<emphasis id="F7C5922CFE59A775E78F6C171D0C41CC" bold="true" pageId="0" pageNumber="117">C</emphasis>
tergite of leg-bearing segment 40, dorsal
<emphasis id="916A927461BC7A6A8954E32265B39CF2" bold="true" pageId="0" pageNumber="117">D</emphasis>
sternite of leg-bearing segment 40, ventral. Scale bars: 0.05 mm (
<emphasis id="1724EC042A7C25A5BF30C80AF6F5BD43" bold="true" pageId="0" pageNumber="117">A, B</emphasis>
); 0.1 mm (
<emphasis id="6FCA2AF2E33CC35261B961B03BA84179" bold="true" pageId="0" pageNumber="117">C, D</emphasis>
).
</paragraph>
</caption>
<paragraph id="853A6BC6F0A0D0557112F6F72B8BED01" pageId="0" pageNumber="117">
<emphasis id="2AE2DF3DEFA0E3CED0EA50F1B2CE535B" bold="true" italics="true" pageId="0" pageNumber="117">Ultimate leg-bearing segment</emphasis>
(Fig.
<figureCitation id="F217E5E30BBB1CF3B9B4EBDF0DD18E54" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Nannarrup oyamensis sp. nov. A, B holotype (TS 20210217 - 04) C, D paratype (TS 20210725 - 02). A ultimate leg-bearing segment and male postpedal segment, dorsal B ultimate leg-bearing segment and male postpedal segment, ventral C right ultimate leg, dorsal D right ultimate leg, ventral. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1115.83946.figure17" httpUri="https://binary.pensoft.net/fig/722243" pageId="0" pageNumber="117">17A-D</figureCitation>
): Pretergite not accompanied by pleurites but incomplete traces of sutures present at both sides. Metatergite subtrapezoidal, almost as wide as long, lateral margins convex and converging posteriorly; posterior margin slightly curved. Coxopleuron ca 1.2
<normalizedToken id="86F5C3A3CFC05B1483A102216CA2701F" originalValue="×">x</normalizedToken>
as long as metasternite; coxal organs of each coxopleuron opening through five or six independent pores, placed ventrally. Metasternite subtriangular, ca 1.6 as wide as long, anteriorly ca 3.5
<normalizedToken id="71862C1901AFFF10EF439D61F17DF93B" originalValue="×">x</normalizedToken>
as wide as posteriorly; lateral margins slightly convex and converging backward; setae almost arranged symmetrically, dense on posterior margin. Telopodite ca 11
<normalizedToken id="9E9FDFB662473B0CC0AE1FDDC9F53D69" originalValue="×">x</normalizedToken>
as long as wide, ca 1.9
<normalizedToken id="D007C7FB212A17EDBA606A13E5B2E329" originalValue="×">x</normalizedToken>
as long and ca 1.3
<normalizedToken id="EEE6AFE5ABF5915A453278BA65411597" originalValue="×">x</normalizedToken>
as wide as penultimate telopodite, with 6 articles; tarsus 2 ca 2.7
<normalizedToken id="5F37B64B6CA7B4153FE3881EC97A43E9" originalValue="×">x</normalizedToken>
as long as wide and ca 1.5
<normalizedToken id="5E03DC78442D11B31073D812FB92A57E" originalValue="×">x</normalizedToken>
as long as tarsus 1; setae arranged uniformly, &lt;50
<normalizedToken id="32A90DC8250082AEC921464BC453C9AA" originalValue="μm">μm</normalizedToken>
long. Pretarsus represented by spines, up to 5
<normalizedToken id="08B4212A20BD8E1FD41F1D36EC187C92" originalValue="µm">µm</normalizedToken>
.
</paragraph>
<caption id="5F46C0AD0ED24CC318E38135C10784A7" doi="10.3897/zookeys.1115.83946.figure17" httpUri="https://binary.pensoft.net/fig/722243" pageId="0" pageNumber="117" start="Figure 17" startId="F17">
<paragraph id="4C3646B1E362EA050AC6924AB166D1F1" pageId="0" pageNumber="117">
<emphasis id="2FD9EF40FCAD3DE62D564282926EDC69" bold="true" pageId="0" pageNumber="117">Figure 17.</emphasis>
<taxonomicName id="A095F69886371D1B47A8657A1033A031" authorityName="Tsukamoto" authorityYear="2022" class="Chilopoda" family="Mecistocephalidae" genus="Nannarrup" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nannarrup oyamensis" order="Geophilomorpha" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="oyamensis">
<emphasis id="859A668392723EA019A8A866AB17D038" italics="true" pageId="0" pageNumber="117">Nannarrup oyamensis</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="D66F7BB29142924D204858BC1616863A" bold="true" pageId="0" pageNumber="117">A, B</emphasis>
holotype (TS20210217-04)
<emphasis id="EFD280BAE39342411BA9269F7138D8DF" bold="true" pageId="0" pageNumber="117">C, D</emphasis>
paratype (TS20210725-02).
<emphasis id="A8660EEF582C736C728CC792DA289E1E" bold="true" pageId="0" pageNumber="117">A</emphasis>
ultimate leg-bearing segment and male postpedal segment, dorsal
<emphasis id="DDA6C038B2ECEE745E3EBA3F45A4A231" bold="true" pageId="0" pageNumber="117">B</emphasis>
ultimate leg-bearing segment and male postpedal segment, ventral
<emphasis id="1CA7D871A427F50A6148AAE844D47B1B" bold="true" pageId="0" pageNumber="117">C</emphasis>
right ultimate leg, dorsal
<emphasis id="04C70E1C6C122CB86107BDCEFB55FF44" bold="true" pageId="0" pageNumber="117">D</emphasis>
right ultimate leg, ventral. Scale bars: 0.1 mm.
</paragraph>
</caption>
<paragraph id="B02581811890CC93EDEAC8C1C105F1A1" pageId="0" pageNumber="117">
<emphasis id="0BF41DA518EBE3B209FBBC5419C1134B" bold="true" italics="true" pageId="0" pageNumber="117">Male postpedal segments</emphasis>
(Fig.
<figureCitation id="B061DEC16EBE5DDB12A7F97245ECC271" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Nannarrup oyamensis sp. nov. A, B holotype (TS 20210217 - 04) C, D paratype (TS 20210725 - 02). A ultimate leg-bearing segment and male postpedal segment, dorsal B ultimate leg-bearing segment and male postpedal segment, ventral C right ultimate leg, dorsal D right ultimate leg, ventral. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1115.83946.figure17" httpUri="https://binary.pensoft.net/fig/722243" pageId="0" pageNumber="117">17A, B</figureCitation>
): Two gonopods, very widely separated from one another, conical in outline, uni-articulated without any sutures, covered with setae. Anal pore present.
</paragraph>
<paragraph id="BFC9422178CC96B6BF3D8AB8903D757B" pageId="0" pageNumber="117">Female postpedal segments unknown (female unknown).</paragraph>
</subSubSection>
<subSubSection id="A98058B832ED83E18AE6E4F11852BDBD" pageId="0" pageNumber="117" type="distribution">
<paragraph id="1F36B40613E7B425E5D4153911A35C11" pageId="0" pageNumber="117">Distribution.</paragraph>
<paragraph id="5761BDD57E5EA1A1BB70D39788CE9FE7" pageId="0" pageNumber="117">Only known from Mt. Oyama, located in Isehara-shi, Kanagawa prefecture.</paragraph>
</subSubSection>
<subSubSection id="7470B3BCC21DC364DA78CEEFF24E0891" pageId="0" pageNumber="117" type="remarks">
<paragraph id="08A70116F49FA6C70539D64255681C0A" pageId="0" pageNumber="117">Remarks.</paragraph>
<paragraph id="EFC871135DA1EE91FCAD01F592BCEF65" pageId="0" pageNumber="117">
<taxonomicName id="D13AD6109C5F4D12B8487812C6D2A0A5" authorityName="Tsukamoto" authorityYear="2022" class="Chilopoda" family="Mecistocephalidae" genus="Nannarrup" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nannarrup oyamensis" order="Geophilomorpha" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="oyamensis">
<emphasis id="E51076E043BA141C9AA5C84109257F37" italics="true" pageId="0" pageNumber="117">Nannarrup oyamensis</emphasis>
</taxonomicName>
sp. nov. is distinguishable from the two congeners by the absence of smooth or weakly areolate areas along the posterior part of the paraclypeal sutures. Specifically,
<taxonomicName id="94411022AC9AB3DDC0B04990DFA5913C" lsidName="N. oyamensis" pageId="0" pageNumber="117" rank="species" species="oyamensis">
<emphasis id="A2FCCBFF3CED1001FD3FBCBA435AC6A3" italics="true" pageId="0" pageNumber="117">N. oyamensis</emphasis>
</taxonomicName>
sp. nov. can be clearly distinguished from
<taxonomicName id="90931BAB4085034D0A9FD8319B13EB82" lsidName="N. hoffmani" pageId="0" pageNumber="117" rank="species" species="hoffmani">
<emphasis id="75B16D0BE9A686234C46F7E8520368C0" italics="true" pageId="0" pageNumber="117">N. hoffmani</emphasis>
</taxonomicName>
by the presence of a well-developed denticle on the trochanteroprefemur (width: length = 1:1.3) and the absence of smooth or weakly areolate areas along the posterior part of the paraclypeal sutures. Furthermore,
<taxonomicName id="8D30918F8716E114F08D2BA2DB007726" lsidName="N. oyamensis" pageId="0" pageNumber="117" rank="species" species="oyamensis">
<emphasis id="FE746FD5E47A3DFB502BAB9B11CD6B18" italics="true" pageId="0" pageNumber="117">N. oyamensis</emphasis>
</taxonomicName>
sp. nov. can be distinguished from
<taxonomicName id="28FD9748D60A24D4FA15910ED912C43F" lsidName="N. innuptus" pageId="0" pageNumber="117" rank="species" species="innuptus">
<emphasis id="21FDC95169287F2629228EE4826365D0" italics="true" pageId="0" pageNumber="117">N. innuptus</emphasis>
</taxonomicName>
sp. nov. by the absence of a pair of smooth or weakly areolate areas along the posterior part of the paraclypeal sutures (see Table
<tableCitation id="742E47CEC7E7A3E87C800962E4597C44" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Morphological comparison among three species of the genus Nannarrup." httpUri="http://table.plazi.org/id/D55B129FFB76FCFABDC8265582807BEC" pageId="0" pageNumber="117" tableUuid="D55B129FFB76FCFABDC8265582807BEC">3</tableCitation>
for a comparison of characteristics).
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>