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<document id="33A3F028686010DF93C51B9A0DEA1E02" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D7F923EFDEBFAA0CC682D91" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius riedeli Telnov 2020, sp. nov." docType="treatment" docUuid="C67FBA17-03C1-45B1-A5DD-716FBBCB7A7B" docUuidSource="ZooBank" docVersion="10" lastPageNumber="102" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="100" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="5FB7DA3BD10C280E0B47B52DE5CB5417" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="BCA9E05380B5F8F9213CD2B12B7E7FF7">
<mods:title id="02E850F09E1E096F7957F0073A977F0F">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
</mods:titleInfo>
<mods:name id="B55AEA6296AFD9545A332E760B39E964" type="personal">
<mods:role id="8F24B0902D759A788F10C55959D843A5">
<mods:roleTerm id="938504D61978BBA6D6F8797A895CA144">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CF2E99B1F6E5EB8C1F6E4EFB62EED239">Telnov, Dmitry</mods:namePart>
</mods:name>
<mods:typeOfResource id="81FA924B67FF7837A40983E543ED3C66">text</mods:typeOfResource>
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<mods:titleInfo id="486497059A44FB4CE7856F64F826012D">
<mods:title id="EB30AF2625981652DA0EC3F81582E643">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="2762A0957BFD03610A18AB8E526F0C63">
<mods:date id="0EDFBD11F38FB1130DEAFD6D4784DEDB">2020</mods:date>
<mods:detail id="1DC530B2B58C111A285503A4FDF78006" type="pubDate">
<mods:number id="ED977189EE392C2933962B2816C94798">2020-10-19</mods:number>
</mods:detail>
<mods:detail id="4C12BC22604C8200D181DEB000DA451B" type="volume">
<mods:number id="D5B13D2EE40004C9B793AEB0EDAFB4D8">721</mods:number>
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<mods:start id="B87EBE98F4DF8B81C2FA45BDA3414996">1</mods:start>
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<mods:classification id="E5270201D1E8CF4125C99CB5C3B0C42A">journal article</mods:classification>
<mods:identifier id="8FB0D2735EECE32F73C5532BAA04BF73" type="CLB-Dataset">9693</mods:identifier>
<mods:identifier id="733B489B158A92A2B82CBF132AE67583" type="DOI">10.5852/ejt.2020.721.1127</mods:identifier>
<mods:identifier id="1DB3AFEE63C9805283610B97FFB10DE4" type="GBIF-Dataset">0cd61b23-1089-4c1b-bfc0-487887abad84</mods:identifier>
<mods:identifier id="B1F2888691BE6470D7B5A6AB9E21CBF4" type="Zenodo-Dep">4122118</mods:identifier>
<mods:identifier id="F1814A191C3EACC5DD02B5B08F098555" type="ZooBank">89E62EF8-2E45-4C59-94B7-6A5603E8939B</mods:identifier>
</mods:mods>
<treatment id="1A3787F69D7F923EFDEBFAA0CC682D91" ID-DOI="http://doi.org/10.5281/zenodo.4344301" ID-GBIF-Taxon="168648120" ID-Zenodo-Dep="4344301" LSID="urn:lsid:zoobank.org:act:C67FBA17-03C1-45B1-A5DD-716FBBCB7A7B" httpUri="http://treatment.plazi.org/id/1A3787F69D7F923EFDEBFAA0CC682D91" lastPageId="101" lastPageNumber="102" pageId="99" pageNumber="100">
<subSubSection id="DA84656B9D7F9238FDEBFAA0CBEA2D5B" box="[632,956,1310,1337]" pageId="99" pageNumber="100" type="nomenclature">
<paragraph id="922136E09D7F9238FDEBFAA0CBEA2D5B" blockId="99.[353,1235,1310,1407]" box="[632,956,1310,1337]" pageId="99" pageNumber="100">
<heading id="C969818C9D7F9238FDEBFAA0CBEA2D5B" box="[632,956,1310,1337]" centered="true" fontSize="11" level="2" pageId="99" pageNumber="100" reason="2">
<taxonomicName id="559E4D639D7F9238FDEBFAA0CB0C2D5B" ID-CoL="7X3XK" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[632,858,1310,1337]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="99" pageNumber="100" phylum="Arthropoda" rank="species" species="riedeli" status="sp. nov.">
<emphasis id="A0EAEAF29D7F9238FDEBFAA0CB0C2D5B" bold="true" box="[632,858,1310,1337]" italics="true" pageId="99" pageNumber="100">Maechidius riedeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D7F9238FCF2FAA1CBEA2D5B" box="[865,956,1311,1337]" pageId="99" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D7F9238FEF2FAFFCC442D1D" pageId="99" pageNumber="100" type="description">
<paragraph id="922136E09D7F9238FEF2FAFFCC852D3E" blockId="99.[353,1235,1310,1407]" box="[353,1235,1345,1372]" pageId="99" pageNumber="100">
<uri id="E60F3AE29D7F9238FEF2FAFFCC852D3E" box="[353,1235,1345,1372]" pageId="99" pageNumber="100">
urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
<paragraph id="922136E09D7F9238FDB2FADACC442D1D" blockId="99.[353,1235,1310,1407]" box="[545,1042,1380,1407]" pageId="99" pageNumber="100">
<figureCitation id="0AA52A659D7F9238FDB2FADACA2D2D1D" box="[545,635,1380,1407]" captionStart="Figs 78–81" captionStartId="100.[189,241,1928,1954]" captionTargetBox="[341,1237,264,1897]" captionTargetId="figure-63@100.[336,1259,251,1916]" captionTargetPageId="100" captionText="Figs 78–81. Papuan Maechidius Macleay, 1819, habitus, dorsal view. 78. M. riedeli sp. nov., holotype, ♂ (SMNS). 79. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 80. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 81. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122163" httpUri="https://zenodo.org/record/4122163/files/figure.png" pageId="99" pageNumber="100">Figs 78</figureCitation>
,
<figureCitation id="0AA52A659D7F9238FD14FADACAE32D1D" box="[647,693,1380,1407]" captionStart="Figs 157–168" captionStartId="142.[189,241,1752,1778]" captionTargetBox="[338,1248,270,1715]" captionTargetId="figure-140@142.[338,1249,265,1724]" captionTargetPageId="142" captionText="Figs 157–168. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal (157–159, 161–167) and frontal (160, 168) views. 157. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). 158. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 159. Same, ♂ (paratype of M. arrowi Frey, 1969) (BMNH). 160. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 161. Same, ♀ (SNSD). 162. M. peregrinus Lansberge, 1886, ♂ (DTC). 163. M. pedarioides Arrow, 1941, syntype, ♀ (BMNH). 164. Same, syntype, ♂ (BMNH). 165. M. riedeli sp. nov., holotype, ♂ (SMNS). 166. M. perlatus (Frey, 1969) comb. nov., holotype, ♂ (NHMB). 167. Same, ♀ (NHMB). 168. M. simplex Frey, 1969, holotype, ♂ (photograph courtesy BPBM) (BPBM). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122183" httpUri="https://zenodo.org/record/4122183/files/figure.png" pageId="99" pageNumber="100">165</figureCitation>
,
<figureCitation id="0AA52A659D7F9238FD51FADBCAA72D1D" box="[706,753,1381,1407]" captionStart="Figs 242–258" captionStartId="148.[189,241,1646,1672]" captionTargetBox="[394,1193,265,1617]" captionTargetId="figure-193@148.[394,1193,265,1626]" captionTargetPageId="148" captionText="Figs 242–258. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 242. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 243. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 244. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 245. M. peregrinus Lansberge, 1886, ♂ (DTC). 246. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 247. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 248. M. riedeli sp. nov., holotype, ♂ (SMNS). 249. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 250. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 251. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 252. M. similis sp. nov., paratype, ♂ (BMNH). 253. M. skalei sp. nov., paratype, ♂ (NME). 254. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 255. M. sougb sp. nov., holotype, ♂ (NME). 256. M. speciosus (Frey, 1969) comb. nov., ♀ (BMNH). 257. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 258. M. suwawa sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122196" httpUri="https://zenodo.org/record/4122196/files/figure.png" pageId="99" pageNumber="100">248</figureCitation>
,
<figureCitation id="0AA52A659D7F9238FD6EFADBCB7A2D1D" box="[765,812,1381,1407]" captionStart="Figs 333–348" captionStartId="155.[189,241,1682,1708]" captionTargetBox="[394,1193,265,1641]" captionTargetId="figure-177@155.[394,1193,265,1655]" captionTargetPageId="155" captionText="Figs 333–348. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 333. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 334. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 335. M. peregrinus Lansberge, 1886, ♂ (DTC). 336. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 337. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 338. Same, enlarged (SMNS). 339. M. riedeli sp. nov., holotype, ♂ (SMNS). 340. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 341. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 342. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 343. M. similis sp. nov., paratype, ♂ (BMNH). 344. M. skalei sp. nov., paratype, ♂ (NME). 345. M. sougb sp. nov., holotype, ♂ (NME). 346. M. speciosus (Frey, 1969) comb. nov., ♀(BMNH). 347. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 348. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122212" httpUri="https://zenodo.org/record/4122212/files/figure.png" pageId="99" pageNumber="100">339</figureCitation>
, 411, 513, 700–702
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D7F9238FF2EFA1BCC772D8C" pageId="99" pageNumber="100" type="diagnosis">
<paragraph id="922136E09D7F9238FF2EFA1BC99F2DDD" blockId="99.[189,457,1445,1471]" box="[189,457,1445,1471]" pageId="99" pageNumber="100">
<heading id="C969818C9D7F9238FF2EFA1BC99F2DDD" bold="true" box="[189,457,1445,1471]" fontSize="11" level="3" pageId="99" pageNumber="100" reason="3">
<emphasis id="A0EAEAF29D7F9238FF2EFA1BC99F2DDD" bold="true" box="[189,457,1445,1471]" pageId="99" pageNumber="100">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D7F9238FF2EFA6ACC772D8C" blockId="99.[189,1057,1492,1518]" box="[189,1057,1492,1518]" pageId="99" pageNumber="100">This new species is primarily peculiar in the shape of the male parameres.</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D7F9238FF2EF9ABCD742EE2" pageId="99" pageNumber="100" type="etymology">
<paragraph id="922136E09D7F9238FF2EF9ABC9122E4D" blockId="99.[189,324,1557,1583]" box="[189,324,1557,1583]" pageId="99" pageNumber="100">
<heading id="C969818C9D7F9238FF2EF9ABC9122E4D" bold="true" box="[189,324,1557,1583]" fontSize="11" level="3" pageId="99" pageNumber="100" reason="3">
<emphasis id="A0EAEAF29D7F9238FF2EF9ABC9122E4D" bold="true" box="[189,324,1557,1583]" pageId="99" pageNumber="100">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D7F9238FF2EF9FDCD742EE2" blockId="99.[189,1398,1603,1665]" pageId="99" pageNumber="100">
Patronymic. The species is named after its collector, Alexander Riedel (Staatliches Museum für Naturkunde Karlsruhe,
<collectingCountry id="EA8976709D7F9238FE42F9D9CA152EE3" box="[465,579,1639,1665]" name="Germany" pageId="99" pageNumber="100">Germany</collectingCountry>
), a well-known coleopterist and explorer of Eastern
<collectingCountry id="EA8976709D7F9238FB3AF9D8CD492EE2" box="[1193,1311,1638,1664]" name="Indonesia" pageId="99" pageNumber="100">Indonesia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D7F9238FF2EF916CA3E2F54" pageId="99" pageNumber="100" type="materials_examined">
<paragraph id="922136E09D7F9238FF2EF916C93A2EA3" blockId="99.[189,364,1703,1730]" box="[189,364,1703,1730]" pageId="99" pageNumber="100">
<heading id="C969818C9D7F9238FF2EF916C93A2EA3" bold="true" box="[189,364,1703,1730]" fontSize="11" level="3" pageId="99" pageNumber="100" reason="3">
<emphasis id="A0EAEAF29D7F9238FF2EF916C93A2EA3" bold="true" box="[189,364,1703,1730]" pageId="99" pageNumber="100">Type material</emphasis>
</heading>
</paragraph>
<materialsCitation id="22F63CBD9D7F9238FF79F968CA3E2F54" ID-GBIF-Occurrence="2898878499" collectingDate="2000-12-23" collectingDateMax="2000-12-25" collectingDateMin="2000-12-23" collectionCode="SMNS" country="Indonesia" county="Japen" elevation="650" location="Ambaidru" municipality="Kontiunai" pageId="99" pageNumber="100" specimenCount="1" stateProvince="Papua Barat" typeStatus="holotype">
<paragraph id="922136E09D7F9238FF79F968C90A2E92" blockId="99.[189,1399,1750,1847]" box="[234,348,1750,1776]" pageId="99" pageNumber="100">
<emphasis id="A0EAEAF29D7F9238FF79F968C90A2E92" bold="true" box="[234,348,1750,1776]" pageId="99" pageNumber="100">
<typeStatus id="4D2588429D7F9238FF79F968C90A2E92" box="[234,348,1750,1776]" pageId="99" pageNumber="100" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="922136E09D7F9238FF2EF947CA3E2F54" blockId="99.[189,1399,1750,1847]" pageId="99" pageNumber="100">
<collectingCountry id="EA8976709D7F9238FF2EF947C90D2F71" box="[189,347,1785,1811]" name="Indonesia" pageId="99" pageNumber="100">INDONESIA</collectingCountry>
•
<specimenCount id="8498FD699D7F9238FEEAF947C9C72F71" box="[377,401,1785,1811]" pageId="99" pageNumber="100">♂</specimenCount>
; “
<collectingRegion id="505AF8029D7F9238FE38F947CA042F71" box="[427,594,1785,1811]" country="Indonesia" name="Papua Barat" pageId="99" pageNumber="100">IRIAN JAYA</collectingRegion>
,
<collectingCounty id="7B404E6C9D7F9238FDF3F947CAF52F71" box="[608,675,1785,1811]" pageId="99" pageNumber="100">Japen</collectingCounty>
[sic
<specimenCount id="8498FD699D7F9238FD4DF946CAB42F71" box="[734,738,1784,1811]" pageId="99" pageNumber="100">♂</specimenCount>
] Isl.,
<collectingMunicipality id="7245AC9A9D7F9238FCB8F947CBF52F71" box="[811,931,1785,1811]" pageId="99" pageNumber="100">Kontiunai</collectingMunicipality>
, road to
<location id="9741603B9D7F9238FB9CF947CCDF2F71" LSID="urn:lsid:plazi:treatment:1A3787F69D7F923EFDEBFAA0CC682D91:9741603B9D7F9238FB9CF947CCDF2F71" box="[1039,1161,1785,1811]" country="Indonesia" county="Japen" municipality="Kontiunai" name="Ambaidru" pageId="99" pageNumber="100" stateProvince="Papua Barat">Ambaidru</location>
,
<quantity id="55669B059D7F9238FB04F947CD4A2F71" box="[1175,1308,1785,1811]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="99" pageNumber="100" unit="m" value="650.0" valueMax="700.0" valueMin="600.0">
<elevation id="19B3D1D39D7F9238FB04F947CD4A2F71" box="[1175,1308,1785,1811]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="99" pageNumber="100" unit="m" value="650.0" valueMax="700.0" valueMin="600.0">600-700 m</elevation>
</quantity>
,
<date id="E62010209D7F9238FAB8F947C97D2F54" pageId="99" pageNumber="100" value="2000-12-23" valueMax="2000-12-25" valueMin="2000-12-23">
<collectingDate id="F664E9C89D7F9238FAB8F947C97D2F54" pageId="99" pageNumber="100" value="2000-12-23" valueMax="2000-12-25" valueMin="2000-12-23">23-25. XII.2000</collectingDate>
</date>
, leg. A. RIEDEL”;
<collectionCode id="F48FAE259D7F9238FD83F8A2CA322F54" box="[528,612,1820,1846]" collectionName="Germany, Stuttgart, Staatliches Museum fuer Naturkunde" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="99" pageNumber="100">SMNS</collectionCode>
.
</paragraph>
</materialsCitation>
</subSubSection>
<subSubSection id="DA84656B9D7F923EFF2EF8E0C9FD2D78" lastPageId="101" lastPageNumber="102" pageId="99" pageNumber="100" type="description">
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<emphasis id="A0EAEAF29D7F9238FF2EF8E0C9182F1A" bold="true" box="[189,334,1886,1912]" pageId="99" pageNumber="100">Description</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D7F9238FF2EF832C97B2F8B" blockId="99.[189,1399,1931,2025]" pageId="99" pageNumber="100">
MEASUREMENTS.
<typeStatus id="4D2588429D7F9238FEEFF835C9BA2FC7" box="[380,492,1931,1957]" pageId="99" pageNumber="100" type="holotype">Holotype</typeStatus>
, total body length
<quantity id="55669B059D7F9238FD54F835CB652FC4" box="[711,819,1931,1958]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="99" pageNumber="100" unit="mm" value="6.6">6.60 mm</quantity>
. Head
<quantity id="55669B059D7F9238FC16F835CBB82FC4" box="[901,1006,1931,1958]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="99" pageNumber="100" unit="mm" value="1.1">1.10 mm</quantity>
long, across eyes
<quantity id="55669B059D7F9238FB55F835CD792FC4" box="[1222,1327,1931,1958]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="99" pageNumber="100" unit="mm" value="1.3">1.30 mm</quantity>
wide. Pronotum
<quantity id="55669B059D7F9238FEA4F813C9C92FA5" box="[311,415,1965,1991]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="99" pageNumber="100" unit="mm" value="1.6">1.60 mm</quantity>
long, maximum width
<quantity id="55669B059D7F9238FD3EF813CB412FA5" box="[685,791,1965,1991]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="99" pageNumber="100" unit="mm" value="2.2">2.20 mm</quantity>
. Elytral length
<quantity id="55669B059D7F9238FC54F813CC642FA5" box="[967,1074,1965,1991]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="99" pageNumber="100" unit="mm" value="3.9">3.90 mm</quantity>
, maximum combined width
<quantity id="55669B059D7F9238FF2EF870C97F2F8B" box="[189,297,1998,2025]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="99" pageNumber="100" unit="mm" value="2.9">2.90 mm</quantity>
.
</paragraph>
<caption id="C6E166689D78923FFF2EF836CD5D2F8A" ID-DOI="http://doi.org/10.5281/zenodo.4122163" ID-Zenodo-Dep="4122163" httpUri="https://zenodo.org/record/4122163/files/figure.png" pageId="100" pageNumber="101" startId="100.[189,241,1928,1954]" targetBox="[341,1237,264,1897]" targetPageId="100">
<paragraph id="922136E09D78923FFF2EF836CD5D2F8A" blockId="100.[189,1399,1928,2025]" pageId="100" pageNumber="101">
<emphasis id="A0EAEAF29D78923FFF2EF836C9112FC0" bold="true" box="[189,327,1928,1954]" pageId="100" pageNumber="101">Figs 78–81.</emphasis>
Papuan
<taxonomicName id="559E4D639D78923FFE39F836CAB02FC0" authority="Macleay, 1819" authorityName="Macleay" authorityYear="1819" box="[426,742,1928,1954]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="100" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="A0EAEAF29D78923FFE39F836CA652FC0" box="[426,563,1928,1954]" italics="true" pageId="100" pageNumber="101">Maechidius</emphasis>
<bibRefCitation id="F60F4B119D78923FFDABF836CAB02FC0" author="Macleay W. S." box="[568,742,1928,1954]" pageId="100" pageNumber="101" refId="ref98319" refString="Macleay W. S. 1819. Horae Entomologicae: or Essays on the Annulose Animals. Volume I, Part I. Containing General Observations on the Geography, Manners, and the Affinities of the Insects which compose the Genus Scarabaeus of Linnaeus; to which are added a few incidental Remarks on the Genera Lucanus and Hister of the same Author. With Appendix and Plates. R. and A. Taylor, London." type="book" year="1819">Macleay, 1819</bibRefCitation>
</taxonomicName>
, habitus, dorsal view.
<emphasis id="A0EAEAF29D78923FFC77F836CC572FC0" bold="true" box="[996,1025,1928,1954]" pageId="100" pageNumber="101">78</emphasis>
.
<taxonomicName id="559E4D639D78923FFB9CF837CCD72FC0" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1039,1153,1928,1954]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="100" pageNumber="101" phylum="Arthropoda" rank="species" species="riedeli" status="sp. nov.">
<emphasis id="A0EAEAF29D78923FFB9CF837CCD72FC0" box="[1039,1153,1928,1954]" italics="true" pageId="100" pageNumber="101">M. riedeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D78923FFB14F836CCB42FC0" box="[1159,1250,1928,1954]" pageId="100" pageNumber="101" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (SMNS).
<emphasis id="A0EAEAF29D78923FFEA0F815C9062FA7" bold="true" box="[307,336,1963,1989]" pageId="100" pageNumber="101">79</emphasis>
.
<taxonomicName id="559E4D639D78923FFEF0F812CAF22FA7" authority="Moser, 1920" authorityName=", Moser Determined" authorityYear="1920" box="[355,676,1963,1990]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="100" pageNumber="101" phylum="Arthropoda" rank="species" species="rugicollis">
<emphasis id="A0EAEAF29D78923FFEF0F812C9A82FA7" box="[355,510,1963,1989]" italics="true" pageId="100" pageNumber="101">M. rugicollis</emphasis>
<bibRefCitation id="F60F4B119D78923FFD99F812CAF22FA7" author="Moser J." box="[522,676,1963,1990]" pageId="100" pageNumber="101" pagination="3 - 28" refId="ref98398" refString="Moser J. 1920. Beitrag zur Kenntnis der Melolonthiden XI. Stettiner Entomologische Zeitung 81: 3 - 28." type="journal article" year="1920">Moser, 1920</bibRefCitation>
</taxonomicName>
, lectotype, ♀ (ZMHB).
<emphasis id="A0EAEAF29D78923FFC5DF815CBBD2FA7" bold="true" box="[974,1003,1963,1989]" pageId="100" pageNumber="101">80</emphasis>
.
<taxonomicName id="559E4D639D78923FFC6CF812CD272FA7" authority="Heller, 1914" authorityName="Heller" authorityYear="1914" box="[1023,1393,1963,1989]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="100" pageNumber="101" phylum="Arthropoda" rank="species" species="seriegranosus">
<emphasis id="A0EAEAF29D78923FFC6CF812CC992FA7" box="[1023,1231,1964,1989]" italics="true" pageId="100" pageNumber="101">M. seriegranosus</emphasis>
<bibRefCitation id="F60F4B119D78923FFB49F815CD272FA7" author="Heller K. M." box="[1242,1393,1963,1989]" pageId="100" pageNumber="101" pagination="615 - 666" refId="ref98213" refString="Heller K. M. 1914. Coleoptera. Nova Guinea. Resultats de l'Expedition scientifique neerlandaise a la Nouvelle-Guinee en 1907 et 1909 sous les auspices du Dr. H. A. Lorentz. Zoologie 9: 615 - 666." type="journal article" year="1914">Heller, 1914</bibRefCitation>
</taxonomicName>
, lectotype, ♂ (SNSD).
<emphasis id="A0EAEAF29D78923FFE51F870C9892F8A" bold="true" box="[450,479,1998,2024]" pageId="100" pageNumber="101">81</emphasis>
.
<taxonomicName id="559E4D639D78923FFE7DF871CB0B2F8A" authority="Moser, 1920" authorityName="Moser" authorityYear="1920" box="[494,861,1998,2025]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="100" pageNumber="101" phylum="Arthropoda" rank="species" species="seriepunctatus">
<emphasis id="A0EAEAF29D78923FFE7DF871CA962F8A" box="[494,704,1999,2024]" italics="true" pageId="100" pageNumber="101">M. seriepunctatus</emphasis>
<bibRefCitation id="F60F4B119D78923FFD54F871CB0B2F8A" author="Moser J." box="[711,861,1998,2025]" pageId="100" pageNumber="101" pagination="3 - 28" refId="ref98398" refString="Moser J. 1920. Beitrag zur Kenntnis der Melolonthiden XI. Stettiner Entomologische Zeitung 81: 3 - 28." type="journal article" year="1920">Moser, 1920</bibRefCitation>
</taxonomicName>
, lectotype, ♂ (ZMHB). Not to scale.
</paragraph>
</caption>
<paragraph id="922136E09D79923EFF2EFEB7CC1C2CCF" blockId="101.[189,1399,265,1197]" pageId="101" pageNumber="102">
Body uniformly pale brown. Head glossy dorsally and ventrally, frons flattened dorsally. Male labroclypeus (
<figureCitation id="0AA52A659D79923EFEF0FE94C9912927" box="[355,455,298,325]" captionStart="Figs 157–168" captionStartId="142.[189,241,1752,1778]" captionTargetBox="[338,1248,270,1715]" captionTargetId="figure-140@142.[338,1249,265,1724]" captionTargetPageId="142" captionText="Figs 157–168. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal (157–159, 161–167) and frontal (160, 168) views. 157. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). 158. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 159. Same, ♂ (paratype of M. arrowi Frey, 1969) (BMNH). 160. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 161. Same, ♀ (SNSD). 162. M. peregrinus Lansberge, 1886, ♂ (DTC). 163. M. pedarioides Arrow, 1941, syntype, ♀ (BMNH). 164. Same, syntype, ♂ (BMNH). 165. M. riedeli sp. nov., holotype, ♂ (SMNS). 166. M. perlatus (Frey, 1969) comb. nov., holotype, ♂ (NHMB). 167. Same, ♀ (NHMB). 168. M. simplex Frey, 1969, holotype, ♂ (photograph courtesy BPBM) (BPBM). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122183" httpUri="https://zenodo.org/record/4122183/files/figure.png" pageId="101" pageNumber="102">Fig. 165</figureCitation>
) broadly deeply emarginate on anterior margin, its anterolateral angles rounded, strongly protruding. Lateral margins of labroclypeus sinuous in both dorsal and lateral views, raised up at angle of nearly 80° to frons in lateral view. Anterior and lateral margins of labroclypeus smooth. Upper- and underside of labroclypeus with sparse long setae along anterior and lateral margins. Canthus broadly rounded in dorsal view. Head dorsal punctures circular, in part annular. Intervening spaces glossy, variably large. Head setae scale-like, elongate, appressed; each seta rises from anterior margin of puncture, generally longer to much longer than length of corresponding puncture. Three extraordinarily long erect setae along inner margin of either compound eye and additional one on either canthus. Antenna 9-segmented, club 3-lamellate. Pronotum glossy dorsally and laterally. Anterior margin of pronotum sinuous, anterolateral angles protruding anteriad, basal margin broadly rounded. Lateral margin of pronotum in dorsal view broadly rounded, delicately crenulate all along (
<figureCitation id="0AA52A659D79923EFB24FDC4CD742AF6" box="[1207,1314,634,660]" captionStart="Figs 242–258" captionStartId="148.[189,241,1646,1672]" captionTargetBox="[394,1193,265,1617]" captionTargetId="figure-193@148.[394,1193,265,1626]" captionTargetPageId="148" captionText="Figs 242–258. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 242. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 243. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 244. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 245. M. peregrinus Lansberge, 1886, ♂ (DTC). 246. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 247. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 248. M. riedeli sp. nov., holotype, ♂ (SMNS). 249. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 250. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 251. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 252. M. similis sp. nov., paratype, ♂ (BMNH). 253. M. skalei sp. nov., paratype, ♂ (NME). 254. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 255. M. sougb sp. nov., holotype, ♂ (NME). 256. M. speciosus (Frey, 1969) comb. nov., ♀ (BMNH). 257. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 258. M. suwawa sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122196" httpUri="https://zenodo.org/record/4122196/files/figure.png" pageId="101" pageNumber="102">Fig. 248</figureCitation>
). Long erect, more or less strongly curved seta present between every two crenulae. Lateral margin of pronotum nearly straight in lateral view. Pronotal punctures circular to ovoid, variably large, dense and shallow; inner margin of punctures with delicate membrane. Intervening spaces glossy, in part microreticulate, generally as large as punctures except near lateral margins where punctures are denser. Setae similar to those on frons. Lateral margin, antero- and posterolateral angles partly covered with dense microscopical velvety pubescence. Hypomeron separated from prosternum by low nearly straight carina, with long setae on anterolateral margin opposite of compound eye. Antennal pocket shallow. Median anterior process of prosternum long brushy setose, short, strongly raised. Scutellar shield pointed apically. Elytron subopaque, with vague tracks of two glabrous longitudinal carinae. Sutural striae or carinae not present. Punctures of elytral disc linear (long and narrow), incision-shaped (
<figureCitation id="0AA52A659D79923EFBB5FC74CCDA2B86" box="[1062,1164,970,996]" captionStart="Figs 333–348" captionStartId="155.[189,241,1682,1708]" captionTargetBox="[394,1193,265,1641]" captionTargetId="figure-177@155.[394,1193,265,1655]" captionTargetPageId="155" captionText="Figs 333–348. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 333. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 334. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 335. M. peregrinus Lansberge, 1886, ♂ (DTC). 336. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 337. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 338. Same, enlarged (SMNS). 339. M. riedeli sp. nov., holotype, ♂ (SMNS). 340. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 341. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 342. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 343. M. similis sp. nov., paratype, ♂ (BMNH). 344. M. skalei sp. nov., paratype, ♂ (NME). 345. M. sougb sp. nov., holotype, ♂ (NME). 346. M. speciosus (Frey, 1969) comb. nov., ♀(BMNH). 347. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 348. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122212" httpUri="https://zenodo.org/record/4122212/files/figure.png" pageId="101" pageNumber="102">Fig. 339</figureCitation>
). Intervening spaces opaque to subopaque, microscopically wrinkled, much larger than incisions. Elytral setae appressed, not surpassing length of corresponding incisions. Male pygidium slightly convex dorsally, with large and shallow annular punctures which are denser in median part (
<figureCitation id="0AA52A659D79923EFC32FB90CC5A2C2B" box="[929,1036,1070,1097]" captionStart="Figs 504–521" captionStartId="166.[189,241,1681,1707]" captionTargetBox="[433,1155,265,1647]" captionTargetId="figure-177@166.[433,1155,265,1658]" captionTargetPageId="166" captionText="Figs 504–521. Papuan and Wallacean Maechidius Macleay, 1819, pygidium, dorsal view. 504. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 505. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 506. Same, ♀ (BMNH). 507. M. pedarioides Arrow, 1941, syntype, ♀ (BMNH). 508. Same, syntype, ♂ (BMNH). 509. M. peregrinus Lansberge, 1886, ♂ (DTC). 510. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 511. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 512. Same, paratype, ♂ (BMNH). 513. M. riedeli sp. nov., holotype, ♂ (SMNS). 514. M. rugicollis Moser, 1920, ♂ (BMNH). 515. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 516. M. similis sp. nov., paratype, ♂ (BMNH). 517. M. skalei sp. nov., paratype, ♂ (NME). 518. M. sougb sp. nov., holotype, ♂ (NME). 519. M. speciosus (Frey, 1969) comb. nov., ♂ (BMNH). 520. Same, different ♂ (BMNH). 521. Same, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122228" httpUri="https://zenodo.org/record/4122228/files/figure.png" pageId="101" pageNumber="102">Fig. 513</figureCitation>
). Setae of pygidium suberect, moderately long in anterior, longer in apical half. Male protibia modified on inner margin, concave predistally. Male protibia with three distal teeth on external margin, median and distal ones rather acute (Fig. 411). Male terminal protibial spur absent. Aedeagus as in
<figureCitation id="0AA52A659D79923EFC31FB2DCC102CCF" box="[930,1094,1171,1197]" captionStart="Figs 697–711" captionStartId="178.[189,241,1892,1918]" captionTargetBox="[435,1146,268,1853]" captionTargetId="figure-75@178.[425,1163,265,1860]" captionTargetPageId="178" captionText="Figs 697–711. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 697– 699. M. popei (Frey, 1969) comb. nov., paratype, ♂ (BMNH?). 700–702. M. riedeli sp. nov., holotype, ♂ (SMNS). 703–705. M. rugicollis Moser, 1920, ♂ (NHMB). 706–708. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 709–711. M. similis sp. nov., paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122250" httpUri="https://zenodo.org/record/4122250/files/figure.png" pageId="101" pageNumber="102">Figs 700–702</figureCitation>
.
</paragraph>
<paragraph id="922136E09D79923EFF2EFB6CC9E62C8E" blockId="101.[189,432,1234,1260]" box="[189,432,1234,1260]" pageId="101" pageNumber="102">
<heading id="C969818C9D79923EFF2EFB6CC9E62C8E" bold="true" box="[189,432,1234,1260]" fontSize="11" level="3" pageId="101" pageNumber="102" reason="3">
<emphasis id="A0EAEAF29D79923EFF2EFB6CC9E62C8E" bold="true" box="[189,432,1234,1260]" pageId="101" pageNumber="102">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D79923EFF2EFABEC9FD2D78" blockId="101.[189,427,1280,1306]" box="[189,427,1280,1306]" pageId="101" pageNumber="102">Female is unknown.</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D79923EFF2EFAFECA492DE4" box="[189,543,1344,1414]" pageId="101" pageNumber="102" type="biology_ecology">
<paragraph id="922136E09D79923EFF2EFAFEC9762D38" blockId="101.[189,543,1344,1414]" box="[189,288,1344,1370]" pageId="101" pageNumber="102">
<heading id="C969818C9D79923EFF2EFAFEC9762D38" bold="true" box="[189,288,1344,1370]" fontSize="11" level="3" pageId="101" pageNumber="102" reason="3">
<emphasis id="A0EAEAF29D79923EFF2EFAFEC9762D38" bold="true" box="[189,288,1344,1370]" pageId="101" pageNumber="102">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D79923EFF2EFAD2CA492DE4" blockId="101.[189,543,1344,1414]" box="[189,543,1388,1414]" pageId="101" pageNumber="102">Occurs in lowland rainforests.</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D79923EFF2EFA12CC682D91" box="[189,1086,1452,1524]" pageId="101" pageNumber="102" type="distribution">
<paragraph id="922136E09D79923EFF2EFA12C9002DA4" blockId="101.[189,342,1452,1478]" box="[189,342,1452,1478]" pageId="101" pageNumber="102">
<heading id="C969818C9D79923EFF2EFA12C9002DA4" bold="true" box="[189,342,1452,1478]" fontSize="11" level="3" pageId="101" pageNumber="102" reason="3">
<emphasis id="A0EAEAF29D79923EFF2EFA12C9002DA4" bold="true" box="[189,342,1452,1478]" pageId="101" pageNumber="102">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D79923EFF2EFA67CC682D91" blockId="101.[189,1086,1497,1524]" box="[189,1086,1497,1524]" pageId="101" pageNumber="102">
Hitherto only known from Yapen Island, Cenderawasih Bay of New
<collectingCountry id="EA8976709D79923EFC71FA67CC6D2D91" box="[994,1083,1497,1523]" name="Guinea-Bissau" pageId="101" pageNumber="102">Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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