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<document id="F689009B4A4F0AB1E9CCBEDD40281355" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D7C9239FD9DFC91CB1E2CFC" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius perlatus Telnov 2020, comb. nov." docType="treatment" docVersion="10" lastPageNumber="99" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="97" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="219F6113F8514E8CF3D4EDFB5ED3D418">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
</mods:titleInfo>
<mods:name id="3FB927534B31D0F301D7C4AB27DE4426" type="personal">
<mods:role id="3297CB3978A3D9AA68BAABCD97E5C7CA">
<mods:roleTerm id="93F9890ADB4577951F20831DD708081E">Author</mods:roleTerm>
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<mods:namePart id="5B7A6769D379D3DEECCDC29D47313072">Telnov, Dmitry</mods:namePart>
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<mods:titleInfo id="BB0CA0590ADC0A5254BC1CBB05DB3DBE">
<mods:title id="A885B3E9A4869D75D45362AC0FD6A2ED">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="530B12842333F762F9DDE2775B7BFC6F">
<mods:date id="8B4CB1027D3F9E2A4CFDE1E8C67662FA">2020</mods:date>
<mods:detail id="596071B9EB0F5D7AF6E141E688275D86" type="pubDate">
<mods:number id="A20EA20268F1C772F2C096CDFF115F7B">2020-10-19</mods:number>
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<mods:detail id="4CE044E6F83D2C2BCF065D14614428CB" type="volume">
<mods:number id="0B8A01D95C6EB16A5E488B2EFBF1789B">721</mods:number>
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<mods:start id="414AF8301DDF478CE00EAD85ABE6AA61">1</mods:start>
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<mods:classification id="A58409C838E2F588E9F1AB68CD9A247E">journal article</mods:classification>
<mods:identifier id="94168FFE7E97F614C70CB99F714942F9" type="CLB-Dataset">9693</mods:identifier>
<mods:identifier id="89A7E57F7C84FB5E48BC8EDAE60B39A5" type="DOI">10.5852/ejt.2020.721.1127</mods:identifier>
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<treatment id="1A3787F69D7C9239FD9DFC91CB1E2CFC" ID-DOI="http://doi.org/10.5281/zenodo.4344413" ID-GBIF-Taxon="168648115" ID-Zenodo-Dep="4344413" LSID="urn:lsid:plazi:treatment:1A3787F69D7C9239FD9DFC91CB1E2CFC" httpUri="http://treatment.plazi.org/id/1A3787F69D7C9239FD9DFC91CB1E2CFC" lastPageId="98" lastPageNumber="99" pageId="96" pageNumber="97">
<subSubSection id="DA84656B9D7C923BFD9DFC91CC702B29" box="[526,1062,815,843]" pageId="96" pageNumber="97" type="nomenclature">
<paragraph id="922136E09D7C923BFD9DFC91CC702B29" blockId="96.[427,1160,815,877]" box="[526,1062,815,843]" pageId="96" pageNumber="97">
<heading id="C969818C9D7C923BFD9DFC91CC702B29" box="[526,1062,815,843]" centered="true" fontSize="11" level="2" pageId="96" pageNumber="97" reason="2">
<taxonomicName id="559E4D639D7C923BFD9DFC91CBCB2B29" ID-CoL="7X3XH" authority="(Frey, 1969)" authorityName="Telnov" authorityYear="2020" baseAuthorityName="Frey" baseAuthorityYear="1969" box="[526,925,815,843]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="perlatus" status="comb. nov.">
<emphasis id="A0EAEAF29D7C923BFD9DFC91CB532B28" bold="true" box="[526,773,815,842]" italics="true" pageId="96" pageNumber="97">Maechidius perlatus</emphasis>
(
<bibRefCitation id="F60F4B119D7C923BFC87FC8ECBC22B29" author="Frey G." box="[788,916,816,843]" pageId="96" pageNumber="97" pagination="494 - 509" refId="ref98001" refString="Frey G. 1969. Revision der Maechidius - Arten Neu-Guineas (Col., Melolonth.). Entomologische Arbeiten aus dem Museum G. Frey 20: 494 - 509." type="journal article" year="1969">Frey, 1969</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="BBD957899D7C923BFC37FC8ECC702B29" box="[932,1062,816,843]" pageId="96" pageNumber="97" rank="species">comb. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D7C923BFE38FCECCCDE2B0E" box="[427,1160,850,877]" pageId="96" pageNumber="97" type="description">
<paragraph id="922136E09D7C923BFE38FCECCCDE2B0E" blockId="96.[427,1160,815,877]" box="[427,1160,850,877]" pageId="96" pageNumber="97">
<figureCitation id="0AA52A659D7C923BFE38FCECCA532B0F" box="[427,517,850,877]" captionStart="Figs 74–77" captionStartId="97.[189,241,1928,1954]" captionTargetBox="[328,1255,263,1887]" captionTargetId="figure-56@97.[325,1262,256,1897]" captionTargetPageId="97" captionText="Figs 74–77. Papuan Maechidius Macleay, 1819, habitus, dorsal view. 74. M. perlatus (Frey, 1969) comb. nov., holotype, ♂ (NHMB). 75. Same, ♂ (SMNS). 76. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 77. Same, paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122160" httpUri="https://zenodo.org/record/4122160/files/figure.png" pageId="96" pageNumber="97">Figs 74</figureCitation>
,
<figureCitation id="0AA52A659D7C923BFD82FCECCA2D2B0E" box="[529,635,850,876]" captionStart="Figs 157–168" captionStartId="142.[189,241,1752,1778]" captionTargetBox="[338,1248,270,1715]" captionTargetId="figure-140@142.[338,1249,265,1724]" captionTargetPageId="142" captionText="Figs 157–168. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal (157–159, 161–167) and frontal (160, 168) views. 157. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). 158. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 159. Same, ♂ (paratype of M. arrowi Frey, 1969) (BMNH). 160. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 161. Same, ♀ (SNSD). 162. M. peregrinus Lansberge, 1886, ♂ (DTC). 163. M. pedarioides Arrow, 1941, syntype, ♀ (BMNH). 164. Same, syntype, ♂ (BMNH). 165. M. riedeli sp. nov., holotype, ♂ (SMNS). 166. M. perlatus (Frey, 1969) comb. nov., holotype, ♂ (NHMB). 167. Same, ♀ (NHMB). 168. M. simplex Frey, 1969, holotype, ♂ (photograph courtesy BPBM) (BPBM). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122183" httpUri="https://zenodo.org/record/4122183/files/figure.png" pageId="96" pageNumber="97">166–167</figureCitation>
,
<figureCitation id="0AA52A659D7C923BFD14FCECCAE02B0E" box="[647,694,850,876]" captionStart="Figs 242–258" captionStartId="148.[189,241,1646,1672]" captionTargetBox="[394,1193,265,1617]" captionTargetId="figure-193@148.[394,1193,265,1626]" captionTargetPageId="148" captionText="Figs 242–258. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 242. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 243. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 244. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 245. M. peregrinus Lansberge, 1886, ♂ (DTC). 246. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 247. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 248. M. riedeli sp. nov., holotype, ♂ (SMNS). 249. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 250. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 251. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 252. M. similis sp. nov., paratype, ♂ (BMNH). 253. M. skalei sp. nov., paratype, ♂ (NME). 254. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 255. M. sougb sp. nov., holotype, ♂ (NME). 256. M. speciosus (Frey, 1969) comb. nov., ♀ (BMNH). 257. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 258. M. suwawa sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122196" httpUri="https://zenodo.org/record/4122196/files/figure.png" pageId="96" pageNumber="97">246</figureCitation>
,
<figureCitation id="0AA52A659D7C923BFD52FCEDCAA62B0F" box="[705,752,851,877]" captionStart="Figs 266–277" captionStartId="150.[189,241,1753,1779]" captionTargetBox="[366,1222,265,1687]" captionTargetId="figure-144@150.[366,1222,265,1703]" captionTargetPageId="150" captionText="Figs 266–277. Papuan and Wallacean Maechidius Macleay, 1819, pronotal hypomeron, ventrolateral view. 266. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 267. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 268. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 269. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 270. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 271. M. mailu sp. nov., holotype, ♂ (NHMB). 272. M. lobaticeps Frey, 1969, ♂ (BMNH). 273. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 274. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 275. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 276. M. peregrinus Lansberge, 1886, ♂ (DTC). 277. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122200" httpUri="https://zenodo.org/record/4122200/files/figure.png" pageId="96" pageNumber="97">277</figureCitation>
,
<figureCitation id="0AA52A659D7C923BFD6FFCEDCB302B0F" box="[764,870,851,877]" captionStart="Figs 333–348" captionStartId="155.[189,241,1682,1708]" captionTargetBox="[394,1193,265,1641]" captionTargetId="figure-177@155.[394,1193,265,1655]" captionTargetPageId="155" captionText="Figs 333–348. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 333. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 334. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 335. M. peregrinus Lansberge, 1886, ♂ (DTC). 336. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 337. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 338. Same, enlarged (SMNS). 339. M. riedeli sp. nov., holotype, ♂ (SMNS). 340. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 341. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 342. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 343. M. similis sp. nov., paratype, ♂ (BMNH). 344. M. skalei sp. nov., paratype, ♂ (NME). 345. M. sougb sp. nov., holotype, ♂ (NME). 346. M. speciosus (Frey, 1969) comb. nov., ♀(BMNH). 347. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 348. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122212" httpUri="https://zenodo.org/record/4122212/files/figure.png" pageId="96" pageNumber="97">337–338</figureCitation>
, 409, 510, 557, 694–696
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D7C923BFF2EFC2DCAFA2BCC" box="[189,684,915,942]" pageId="96" pageNumber="97" type="reference_group">
<paragraph id="922136E09D7C923BFF2EFC2DCAFA2BCC" blockId="96.[189,684,915,942]" box="[189,684,915,942]" pageId="96" pageNumber="97">
<treatmentCitationGroup id="B28E11CE9D7C923BFF2EFC2DCAFA2BCC" box="[189,684,915,942]" pageId="96" pageNumber="97">
<treatmentCitation id="133F10F19D7C923BFF2EFC2DCAFF2BCC" author="Frey G." box="[189,681,915,942]" page="507" pageId="96" pageNumber="97" year="1969">
<taxonomicName id="559E4D639D7C923BFF2EFC2DCAFF2BCC" ID-CoL="7XWMN" authority="Frey, 1969: 507" authorityName="Frey" authorityPageNumber="507" authorityYear="1969" box="[189,681,915,942]" class="Insecta" family="Scarabaeidae" genus="Paramaechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="perlatus">
<emphasis id="A0EAEAF29D7C923BFF2EFC2DC9B32BCF" box="[189,485,915,941]" italics="true" pageId="96" pageNumber="97">Paramaechidius perlatus</emphasis>
<bibRefCitation id="F60F4B119D7C923BFE7FFC2ACAFF2BCC" author="Frey G." box="[492,681,915,942]" pageId="96" pageNumber="97" pagination="494 - 509" refId="ref98001" refString="Frey G. 1969. Revision der Maechidius - Arten Neu-Guineas (Col., Melolonth.). Entomologische Arbeiten aus dem Museum G. Frey 20: 494 - 509." type="journal article" year="1969">Frey, 1969: 507</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D7C923BFF2EFC6ACCB02D71" pageId="96" pageNumber="97" type="materials_examined">
<paragraph id="922136E09D7C923BFF2EFC6AC93A2B8F" blockId="96.[189,364,979,1006]" box="[189,364,979,1006]" pageId="96" pageNumber="97">
<heading id="C969818C9D7C923BFF2EFC6AC93A2B8F" bold="true" box="[189,364,979,1006]" fontSize="11" level="3" pageId="96" pageNumber="97" reason="3">
<emphasis id="A0EAEAF29D7C923BFF2EFC6AC93A2B8F" bold="true" box="[189,364,979,1006]" pageId="96" pageNumber="97">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D7C923BFF79FBBFC90A2C79" blockId="96.[189,1398,1025,1121]" box="[234,348,1025,1051]" pageId="96" pageNumber="97">
<emphasis id="A0EAEAF29D7C923BFF79FBBFC90A2C79" bold="true" box="[234,348,1025,1051]" pageId="96" pageNumber="97">
<typeStatus id="4D2588429D7C923BFF79FBBFC90A2C79" box="[234,348,1025,1051]" pageId="96" pageNumber="97" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="922136E09D7C923BFF2EFB9ACB4B2C03" blockId="96.[189,1398,1025,1121]" pageId="96" pageNumber="97">
<materialsCitation id="22F63CBD9D7C923BFF2EFB9ACB4B2C03" ID-GBIF-Occurrence="2898878430" collectionCode="TYPE, NHMB" country="Papua New Guinea" determinerName="G. Frey" location="PAPUA NEW GUINEA" pageId="96" pageNumber="97" specimenCount="2" typeStatus="holotype">
<collectingCountry id="EA8976709D7C923BFF2EFB9AC9832C5C" box="[189,469,1060,1086]" name="Papua New Guinea" pageId="96" pageNumber="97">PAPUA NEW GUINEA</collectingCountry>
•
<specimenCount id="8498FD699D7C923BFE78FB9ACA552C5C" box="[491,515,1060,1086]" pageId="96" pageNumber="97">♂</specimenCount>
; “
<typeStatus id="4D2588429D7C923BFD8BFB9ACA322C5C" box="[536,612,1060,1086]" pageId="96" pageNumber="97">
<collectionCode id="F48FAE259D7C923BFD8BFB9ACA322C5C" box="[536,612,1060,1086]" pageId="96" pageNumber="97">TYPE</collectionCode>
</typeStatus>
[p, red label]// Bismarck Geb. v. Guinea [h] //
<typeStatus id="4D2588429D7C923BFBE5FB9ACCE62C5C" box="[1142,1200,1060,1086]" pageId="96" pageNumber="97">Type</typeStatus>
<taxonomicName id="559E4D639D7C923BFB25FB9AC94B2C02" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" baseAuthorityName="Frey" baseAuthorityYear="1969" class="Insecta" family="Scarabaeidae" genus="Paramaechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="perlatus" status="sp. nov.">
<emphasis id="A0EAEAF29D7C923BFB25FB9AC94B2C02" italics="true" pageId="96" pageNumber="97">Paramaechidius perlatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D7C923BFEB7FBF9C9352C03" box="[292,355,1095,1121]" pageId="96" pageNumber="97" rank="species">n. sp.</taxonomicNameLabel>
[h] det.
<determinerName id="67FDDA249D7C923BFE2DFBF8CA472C02" box="[446,529,1094,1120]" pageId="96" pageNumber="97">G.Frey</determinerName>
,1967/68 [p]”;
<collectionCode id="F48FAE259D7C923BFD2EFBF9CB4F2C03" box="[701,793,1095,1121]" collectionName="Switzerland, Basel, Naturhistorisches Museum" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="96" pageNumber="97">NHMB</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="922136E09D7C923BFF2EFB39C9302CC2" blockId="96.[189,358,1158,1185]" box="[189,358,1158,1185]" pageId="96" pageNumber="97">
<heading id="C969818C9D7C923BFF2EFB39C9302CC2" bold="true" box="[189,358,1158,1185]" fontSize="11" level="3" pageId="96" pageNumber="97" reason="3">
<emphasis id="A0EAEAF29D7C923BFF2EFB39C9302CC2" bold="true" box="[189,358,1158,1185]" pageId="96" pageNumber="97">New material</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D7C923BFF2EFB0BCCB02D71" blockId="96.[189,1398,1204,1300]" pageId="96" pageNumber="97">
<materialsCitation id="22F63CBD9D7C923BFF2EFB0BCAD12C93" ID-GBIF-Occurrence="2898878466" collectingDate="2000-12-23" collectingDateMax="2000-12-25" collectingDateMin="2000-12-23" collectionCode="SMNS" country="Indonesia" county="Japen" elevation="650" location="Ambaidru" municipality="Kontiunai" pageId="96" pageNumber="97" specimenCount="1" specimenCount-female="1" stateProvince="Papua Barat">
<collectingCountry id="EA8976709D7C923BFF2EFB0BC90D2CAD" box="[189,347,1205,1231]" name="Indonesia" pageId="96" pageNumber="97">INDONESIA</collectingCountry>
•
<specimenCount id="8498FD699D7C923BFEE0FB0BC9F62CAD" box="[371,416,1205,1231]" pageId="96" pageNumber="97" type="male">1 ♂</specimenCount>
,
<specimenCount id="8498FD699D7C923BFE3EFB0BC98C2CAD" box="[429,474,1205,1231]" pageId="96" pageNumber="97" type="female">1 ♀</specimenCount>
; “
<collectingRegion id="505AF8029D7C923BFE61FB0BCAC32CAD" box="[498,661,1205,1231]" country="Indonesia" name="Papua Barat" pageId="96" pageNumber="97">IRIAN JAYA</collectingRegion>
,
<collectingCounty id="7B404E6C9D7C923BFD33FB0BCAB52CAD" box="[672,739,1205,1231]" pageId="96" pageNumber="97">Japen</collectingCounty>
[sic
<specimenCount id="8498FD699D7C923BFC88FB0ACB492CAD" box="[795,799,1204,1231]" pageId="96" pageNumber="97">♂</specimenCount>
] Isl.,
<collectingMunicipality id="7245AC9A9D7C923BFCF1FB0ACB8C2CAC" box="[866,986,1204,1230]" pageId="96" pageNumber="97">Kontiunai</collectingMunicipality>
, road to
<location id="9741603B9D7C923BFBAFFB0ACCE02CAC" LSID="urn:lsid:plazi:treatment:1A3787F69D7C9239FD9DFC91CB1E2CFC:9741603B9D7C923BFBAFFB0ACCE02CAC" box="[1084,1206,1204,1230]" country="Indonesia" county="Japen" municipality="Kontiunai" name="Ambaidru" pageId="96" pageNumber="97" stateProvince="Papua Barat">Ambaidru</location>
,
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,
<date id="E62010209D7C923BFADCFB0BC9192C90" pageId="96" pageNumber="97" value="2000-12-23" valueMax="2000-12-25" valueMin="2000-12-23">
<collectingDate id="F664E9C89D7C923BFADCFB0BC9192C90" pageId="96" pageNumber="97" value="2000-12-23" valueMax="2000-12-25" valueMin="2000-12-23">23- 25.XII.2000</collectingDate>
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, leg. A. RIEDEL”;
<collectionCode id="F48FAE259D7C923BFDA4FB69CAD12C93" box="[567,647,1239,1265]" collectionName="Germany, Stuttgart, Staatliches Museum fuer Naturkunde" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="96" pageNumber="97">SMNS</collectionCode>
</materialsCitation>
•
<materialsCitation id="22F63CBD9D7C923BFD32FB69CC3B2C93" ID-GBIF-Occurrence="2898878423" box="[673,1133,1239,1265]" pageId="96" pageNumber="97">
<specimenCount id="8498FD699D7C923BFD32FB69CA862C93" box="[673,720,1239,1265]" pageId="96" pageNumber="97" type="male">1 ♂</specimenCount>
; same label as for preceding;
<collectionCode id="F48FAE259D7C923BFBA1FB69CC3B2C93" box="[1074,1133,1239,1265]" collectionName="Dmitrijs Telnovs" pageId="96" pageNumber="97">DTC</collectionCode>
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•
<materialsCitation id="22F63CBD9D7C923BFB14FB69CCB52D71" ID-GBIF-Occurrence="2898878348" collectingDate="2001-01-05" collectionCode="SMNS" country="Indonesia" county="Wandammen Bay" elevation="575" location="Mts. Wasior" municipality="Wondiwoi Mts." pageId="96" pageNumber="97" specimenCount="1" stateProvince="Papua Barat">
<specimenCount id="8498FD699D7C923BFB14FB69CCE32C93" box="[1159,1205,1239,1265]" pageId="96" pageNumber="97" type="male">1 ♂</specimenCount>
; “
<collectingRegion id="505AF8029D7C923BFB5DFB69CD242C93" box="[1230,1394,1239,1265]" country="Indonesia" name="Papua Barat" pageId="96" pageNumber="97">IRIAN JAYA</collectingRegion>
,
<collectingCounty id="7B404E6C9D7C923BFF2EFB47C9D82D76" box="[189,398,1273,1300]" pageId="96" pageNumber="97">Wandammen Bay</collectingCounty>
,
<collectingMunicipality id="7245AC9A9D7C923BFE09FB47CA062D76" box="[410,592,1273,1300]" pageId="96" pageNumber="97">
Wondiwoi
<location id="9741603B9D7C923BFD88FB44CA062D76" LSID="urn:lsid:plazi:treatment:1A3787F69D7C9239FD9DFC91CB1E2CFC:9741603B9D7C923BFD88FB44CA062D76" box="[539,592,1274,1300]" country="Indonesia" county="Wandammen Bay" municipality="Wondiwoi Mts." name="Mts. Wasior" pageId="96" pageNumber="97" stateProvince="Papua Barat">Mts.</location>
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Wasior,
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,
<date id="E62010209D7C923BFCD6FB47CBFF2D76" box="[837,937,1273,1300]" pageId="96" pageNumber="97" value="2001-01-05">
<collectingDate id="F664E9C89D7C923BFCD6FB47CBFF2D76" box="[837,937,1273,1300]" pageId="96" pageNumber="97" value="2001-01-05">5.I.2001</collectingDate>
</date>
, leg. A. RIEDEL”;
<collectionCode id="F48FAE259D7C923BFB1CFB47CCB52D71" box="[1167,1251,1273,1299]" collectionName="Germany, Stuttgart, Staatliches Museum fuer Naturkunde" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="96" pageNumber="97">SMNS</collectionCode>
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.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D7C9239FF2EFA84CB1E2CFC" lastPageId="98" lastPageNumber="99" pageId="96" pageNumber="97" type="description">
<paragraph id="922136E09D7C923BFF2EFA84C9182D36" blockId="96.[189,334,1338,1364]" box="[189,334,1338,1364]" pageId="96" pageNumber="97">
<heading id="C969818C9D7C923BFF2EFA84C9182D36" bold="true" box="[189,334,1338,1364]" fontSize="11" level="3" pageId="96" pageNumber="97" reason="3">
<emphasis id="A0EAEAF29D7C923BFF2EFA84C9182D36" bold="true" box="[189,334,1338,1364]" pageId="96" pageNumber="97">Description</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D7C923BFF2EFAD6CB552DE0" blockId="96.[189,771,1383,1410]" box="[189,771,1383,1410]" pageId="96" pageNumber="97">MEASUREMENTS. Total body length 7.50–9.00 mm.</paragraph>
<paragraph id="922136E09D7C9239FF2EFA17CD3A2C6C" blockId="96.[189,1399,1448,2025]" lastBlockId="98.[189,1399,265,1039]" lastPageId="98" lastPageNumber="99" pageId="96" pageNumber="97">
Dorsum uniformly black to black-brown with castaneous brown labroclypeus, mouthparts and antennae. Entire dorsum covered with dirty yellowish dense microscopical fur-like velvety pubescence (
<figureCitation id="0AA52A659D7C923BFA90FA74CD3F2D87" box="[1283,1385,1482,1509]" captionStart="Figs 333–348" captionStartId="155.[189,241,1682,1708]" captionTargetBox="[394,1193,265,1641]" captionTargetId="figure-177@155.[394,1193,265,1655]" captionTargetPageId="155" captionText="Figs 333–348. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 333. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 334. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 335. M. peregrinus Lansberge, 1886, ♂ (DTC). 336. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 337. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 338. Same, enlarged (SMNS). 339. M. riedeli sp. nov., holotype, ♂ (SMNS). 340. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 341. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 342. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 343. M. similis sp. nov., paratype, ♂ (BMNH). 344. M. skalei sp. nov., paratype, ♂ (NME). 345. M. sougb sp. nov., holotype, ♂ (NME). 346. M. speciosus (Frey, 1969) comb. nov., ♀(BMNH). 347. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 348. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122212" httpUri="https://zenodo.org/record/4122212/files/figure.png" pageId="96" pageNumber="97">Fig. 338</figureCitation>
), surface structure therefore almost completely hidden. Head transverse, opaque dorsally and ventrally, convex between eyes. Compound eye large, occupying about half side of head. Male labroclypeus (
<figureCitation id="0AA52A659D7C923BFF55F98CC97E2E2F" box="[198,296,1586,1613]" captionStart="Figs 157–168" captionStartId="142.[189,241,1752,1778]" captionTargetBox="[338,1248,270,1715]" captionTargetId="figure-140@142.[338,1249,265,1724]" captionTargetPageId="142" captionText="Figs 157–168. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal (157–159, 161–167) and frontal (160, 168) views. 157. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). 158. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 159. Same, ♂ (paratype of M. arrowi Frey, 1969) (BMNH). 160. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 161. Same, ♀ (SNSD). 162. M. peregrinus Lansberge, 1886, ♂ (DTC). 163. M. pedarioides Arrow, 1941, syntype, ♀ (BMNH). 164. Same, syntype, ♂ (BMNH). 165. M. riedeli sp. nov., holotype, ♂ (SMNS). 166. M. perlatus (Frey, 1969) comb. nov., holotype, ♂ (NHMB). 167. Same, ♀ (NHMB). 168. M. simplex Frey, 1969, holotype, ♂ (photograph courtesy BPBM) (BPBM). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122183" httpUri="https://zenodo.org/record/4122183/files/figure.png" pageId="96" pageNumber="97">Fig. 166</figureCitation>
) deeply and broadly emarginate anteriorly, its lateral margins sinuous in both dorsal and lateral views, male anterolateral angles strongly protruding, acute in dorsal view, bent up at ~90° to frons in lateral view. Female labroclypeus (
<figureCitation id="0AA52A659D7C923BFDCEF9C8CA952EF3" box="[605,707,1654,1681]" captionStart="Figs 157–168" captionStartId="142.[189,241,1752,1778]" captionTargetBox="[338,1248,270,1715]" captionTargetId="figure-140@142.[338,1249,265,1724]" captionTargetPageId="142" captionText="Figs 157–168. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal (157–159, 161–167) and frontal (160, 168) views. 157. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). 158. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 159. Same, ♂ (paratype of M. arrowi Frey, 1969) (BMNH). 160. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 161. Same, ♀ (SNSD). 162. M. peregrinus Lansberge, 1886, ♂ (DTC). 163. M. pedarioides Arrow, 1941, syntype, ♀ (BMNH). 164. Same, syntype, ♂ (BMNH). 165. M. riedeli sp. nov., holotype, ♂ (SMNS). 166. M. perlatus (Frey, 1969) comb. nov., holotype, ♂ (NHMB). 167. Same, ♀ (NHMB). 168. M. simplex Frey, 1969, holotype, ♂ (photograph courtesy BPBM) (BPBM). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122183" httpUri="https://zenodo.org/record/4122183/files/figure.png" pageId="96" pageNumber="97">Fig. 167</figureCitation>
) with broad anterior emargination, its anterolateral angles broadly rounded and not protruding, sinuous in both dorsal and lateral views. Upper- and underside of labroclypeus with sparse long setae along anterior and lateral margins. Canthus nearly straight in dorsal view. Anterior and lateral margins of labroclypeus smooth. Dorsal punctures of irregular ovoid shape, very deep. Intervening spaces generally distinctly larger than punctures. Dirty-yellow suberect seta rises from anterior margin of each puncture; setae vary in size and shape from longer than to not surpassing length of corresponding punctures. Antenna in both sexes 9-segmented, club 3-lamellate. Scape large, with bulbous predistal projection on dorsal side, provided with two very long setae near distal margin. Antennomere 2 transverse. Pronotum anterior margin broadly emarginate. Basal margin of pronotum obtusely angulate medially. Lateral margin of pronotum widened towards obtuse postmedian angulation and afterwards broadly emarginate before base (
<figureCitation id="0AA52A659D7C923BFC81F870CB292F8B" box="[786,895,1998,2025]" captionStart="Figs 242–258" captionStartId="148.[189,241,1646,1672]" captionTargetBox="[394,1193,265,1617]" captionTargetId="figure-193@148.[394,1193,265,1626]" captionTargetPageId="148" captionText="Figs 242–258. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 242. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 243. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 244. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 245. M. peregrinus Lansberge, 1886, ♂ (DTC). 246. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 247. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 248. M. riedeli sp. nov., holotype, ♂ (SMNS). 249. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 250. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 251. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 252. M. similis sp. nov., paratype, ♂ (BMNH). 253. M. skalei sp. nov., paratype, ♂ (NME). 254. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 255. M. sougb sp. nov., holotype, ♂ (NME). 256. M. speciosus (Frey, 1969) comb. nov., ♀ (BMNH). 257. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 258. M. suwawa sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122196" httpUri="https://zenodo.org/record/4122196/files/figure.png" pageId="96" pageNumber="97">Fig. 246</figureCitation>
). Lateral margin somewhat flattened and expanded laterally, delicately crenulate all along – a scale-like appressed seta present between every two crenulae. Lateral margin of pronotum nearly straight in lateral view. Punctures deep, of irregular ovoid shape, on disc generally larger than those on head, near lateral margins small. Intervening spaces variably large, generally larger than punctures. Setae scale-like, longer on anterior margin facing frons and at posterolateral angles, but medially generally short and not surpassing length of corresponding punctures. Conspicuous rather short appressed brush of clustered setae rises from each puncture along lateral margins of pronotum. Hypomeron emarginate and very long setose on anterior margin which is flange-like (
<figureCitation id="0AA52A659D7E9239FEDAFE49C9F82A73" box="[329,430,503,529]" captionStart="Figs 266–277" captionStartId="150.[189,241,1753,1779]" captionTargetBox="[366,1222,265,1687]" captionTargetId="figure-144@150.[366,1222,265,1703]" captionTargetPageId="150" captionText="Figs 266–277. Papuan and Wallacean Maechidius Macleay, 1819, pronotal hypomeron, ventrolateral view. 266. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 267. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 268. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 269. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 270. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 271. M. mailu sp. nov., holotype, ♂ (NHMB). 272. M. lobaticeps Frey, 1969, ♂ (BMNH). 273. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 274. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 275. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 276. M. peregrinus Lansberge, 1886, ♂ (DTC). 277. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122200" httpUri="https://zenodo.org/record/4122200/files/figure.png" pageId="98" pageNumber="99">Fig. 277</figureCitation>
), covered by fur-like pubescence.Antennal pocket deep. Median anterior process of prosternum long brushy setose, short, slightly raised. Scutellar shield narrowly rounded apically. Elytra with maximum width across midlength, with distinct humeri. Tracks of four convex longitudinal carinae on each elytron (including sutural one) formed by irregular large glossy glabrous humps. Remnants of sutural carinae indicated by more delicate humps. Punctures of disc of irregular oblong shape, deep, generally smaller than those on forebody (
<figureCitation id="0AA52A659D7E9239FD2DFD1FCB7E2AD9" box="[702,808,673,699]" captionStart="Figs 333–348" captionStartId="155.[189,241,1682,1708]" captionTargetBox="[394,1193,265,1641]" captionTargetId="figure-177@155.[394,1193,265,1655]" captionTargetPageId="155" captionText="Figs 333–348. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 333. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 334. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 335. M. peregrinus Lansberge, 1886, ♂ (DTC). 336. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 337. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 338. Same, enlarged (SMNS). 339. M. riedeli sp. nov., holotype, ♂ (SMNS). 340. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 341. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 342. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 343. M. similis sp. nov., paratype, ♂ (BMNH). 344. M. skalei sp. nov., paratype, ♂ (NME). 345. M. sougb sp. nov., holotype, ♂ (NME). 346. M. speciosus (Frey, 1969) comb. nov., ♀(BMNH). 347. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 348. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122212" httpUri="https://zenodo.org/record/4122212/files/figure.png" pageId="98" pageNumber="99">Fig. 337</figureCitation>
). Entire elytral surface covered with velvety furlike setae (
<figureCitation id="0AA52A659D7E9239FEAEFD7DC9F32ABF" box="[317,421,707,733]" captionStart="Figs 333–348" captionStartId="155.[189,241,1682,1708]" captionTargetBox="[394,1193,265,1641]" captionTargetId="figure-177@155.[394,1193,265,1655]" captionTargetPageId="155" captionText="Figs 333–348. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 333. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 334. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 335. M. peregrinus Lansberge, 1886, ♂ (DTC). 336. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 337. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 338. Same, enlarged (SMNS). 339. M. riedeli sp. nov., holotype, ♂ (SMNS). 340. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 341. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 342. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 343. M. similis sp. nov., paratype, ♂ (BMNH). 344. M. skalei sp. nov., paratype, ♂ (NME). 345. M. sougb sp. nov., holotype, ♂ (NME). 346. M. speciosus (Frey, 1969) comb. nov., ♀(BMNH). 347. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 348. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122212" httpUri="https://zenodo.org/record/4122212/files/figure.png" pageId="98" pageNumber="99">Fig. 338</figureCitation>
). Ordinary setae scale-like, moderately long, suberect, arranged in irregular paired longitudinal rows (
<figureCitation id="0AA52A659D7E9239FE0FFD5BC9A02A9D" box="[412,502,741,767]" captionStart="Figs 74–77" captionStartId="97.[189,241,1928,1954]" captionTargetBox="[328,1255,263,1887]" captionTargetId="figure-56@97.[325,1262,256,1897]" captionTargetPageId="97" captionText="Figs 74–77. Papuan Maechidius Macleay, 1819, habitus, dorsal view. 74. M. perlatus (Frey, 1969) comb. nov., holotype, ♂ (NHMB). 75. Same, ♂ (SMNS). 76. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 77. Same, paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122160" httpUri="https://zenodo.org/record/4122160/files/figure.png" pageId="98" pageNumber="99">Fig. 74</figureCitation>
). Each seta rises from anterior margin of corresponding puncture. Pygidium in both sexes slightly convex, deeply irregularly oblong-shaped punctate, intervening spaces covered with fur-like pubescence (
<figureCitation id="0AA52A659D7E9239FE61FC97CA0D2B21" box="[498,603,808,835]" captionStart="Figs 504–521" captionStartId="166.[189,241,1681,1707]" captionTargetBox="[433,1155,265,1647]" captionTargetId="figure-177@166.[433,1155,265,1658]" captionTargetPageId="166" captionText="Figs 504–521. Papuan and Wallacean Maechidius Macleay, 1819, pygidium, dorsal view. 504. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 505. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 506. Same, ♀ (BMNH). 507. M. pedarioides Arrow, 1941, syntype, ♀ (BMNH). 508. Same, syntype, ♂ (BMNH). 509. M. peregrinus Lansberge, 1886, ♂ (DTC). 510. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 511. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 512. Same, paratype, ♂ (BMNH). 513. M. riedeli sp. nov., holotype, ♂ (SMNS). 514. M. rugicollis Moser, 1920, ♂ (BMNH). 515. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 516. M. similis sp. nov., paratype, ♂ (BMNH). 517. M. skalei sp. nov., paratype, ♂ (NME). 518. M. sougb sp. nov., holotype, ♂ (NME). 519. M. speciosus (Frey, 1969) comb. nov., ♂ (BMNH). 520. Same, different ♂ (BMNH). 521. Same, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122228" httpUri="https://zenodo.org/record/4122228/files/figure.png" pageId="98" pageNumber="99">Fig. 510</figureCitation>
). Setae elongate clavate, rather long, suberect. Venter covered with sparse small shallow punctures, each provided with a very short seta, in part covered with microscopical velvety pubescence. Legs long and slender. Male protibia nearly straight on external margin, with complete dorsal carina. Two small obtuse distal teeth on protibia in both sexes (Fig. 409). Male protibial terminal spur large, curved, female one shorter and straight. Male metatibial terminal spurs paired, almost equally long, subacute, lower one curved; female ones shorter, nearly straight, obtuse. Tarsal claws with pulvilli in both sexes. Spiculum gastrale as in
<figureCitation id="0AA52A659D7E9239FCCCFC4ACB932C6D" box="[863,965,1012,1039]" captionStart="Figs 545–569" captionStartId="169.[189,241,1541,1567]" captionTargetBox="[308,1266,258,1476]" captionTargetId="figure-249@169.[299,1280,255,1481]" captionTargetPageId="169" captionText="Figs 545–569. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, spiculum gastrale. 545. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 546. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 547. M. yamdena sp. nov., paratype, ♂ (BMNH). 548. M. kazantsevi sp. nov., holotype, ♂ tBMNH). 549. M. legalovi sp. nov., holotype, ♂ (BMNH). 550. M. leucopsar sp. nov., holotype, ♂ (IECA). 551. M. longipes sp. nov., holotype, ♂ (BMNH). 552. M. luniceps Fairmaire, 1883, ♂ (BMNH). 553. M. maleo sp. nov., holotype, ♂ (IRSN). 554. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 555. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 556. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 557. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 558. M. popei (Frey, 1969) comb. nov., paratype, ♂ (BMNH). 559. M. rugicollis Moser, 1920, ♂ (BMNH). 560. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 561. M. esau Heller, 1914, ♂ (BMNH). 562. M. speciosus (Frey, 1969) comb. nov., ♂. 563. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 564. M. suwawa sp. nov., paratype, ♂ (BMNH). 565. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 566. M. trivialis sp. nov., holotype, ♂ (BMNH). 567. M. alesbezdeki sp. nov., holotype, ♂ (NME). 568. M. bintang sp. nov., holotype, ♂ (NME). 569. M. lobaticeps Frey, 1969, ♂ (DTC). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122230" httpUri="https://zenodo.org/record/4122230/files/figure.png" pageId="98" pageNumber="99">Fig. 557</figureCitation>
. Male aedeagus as in
<figureCitation id="0AA52A659D7E9239FB57FC4ACD3E2C6C" box="[1220,1384,1012,1038]" captionStart="Figs 682–696" captionStartId="177.[189,241,1892,1918]" captionTargetBox="[436,1163,269,1860]" captionTargetId="figure-72@177.[423,1174,265,1866]" captionTargetPageId="177" captionText="Figs 682–696. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 682– 684. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 685–687. M. pauxillus Heller, 1910 comb. rest., ♂ (BMNH). 688–690. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 691–693. M. peregrinus Lansberge, 1886, ♂ (DTC). 694-696. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122248" httpUri="https://zenodo.org/record/4122248/files/figure.png" pageId="98" pageNumber="99">Figs 694–696</figureCitation>
.
</paragraph>
<caption id="C6E166689D7D923AFF2EF836CBB32F8A" ID-DOI="http://doi.org/10.5281/zenodo.4122160" ID-Zenodo-Dep="4122160" httpUri="https://zenodo.org/record/4122160/files/figure.png" pageId="97" pageNumber="98" startId="97.[189,241,1928,1954]" targetBox="[328,1255,263,1887]" targetPageId="97">
<paragraph id="922136E09D7D923AFF2EF836CBB32F8A" blockId="97.[189,1399,1928,2025]" pageId="97" pageNumber="98">
<emphasis id="A0EAEAF29D7D923AFF2EF836C91B2FC0" bold="true" box="[189,333,1928,1954]" pageId="97" pageNumber="98">Figs 74–77.</emphasis>
Papuan
<taxonomicName id="559E4D639D7D923AFE28F836CB552FC0" authority="Macleay, 1819" authorityName="Macleay" authorityYear="1819" box="[443,771,1928,1954]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="97" pageNumber="98" phylum="Arthropoda" rank="genus">
<emphasis id="A0EAEAF29D7D923AFE28F836CA122FC0" box="[443,580,1928,1954]" italics="true" pageId="97" pageNumber="98">Maechidius</emphasis>
<bibRefCitation id="F60F4B119D7D923AFDC3F836CB552FC0" author="Macleay W. S." box="[592,771,1928,1954]" pageId="97" pageNumber="98" refId="ref98319" refString="Macleay W. S. 1819. Horae Entomologicae: or Essays on the Annulose Animals. Volume I, Part I. Containing General Observations on the Geography, Manners, and the Affinities of the Insects which compose the Genus Scarabaeus of Linnaeus; to which are added a few incidental Remarks on the Genera Lucanus and Hister of the same Author. With Appendix and Plates. R. and A. Taylor, London." type="book" year="1819">Macleay, 1819</bibRefCitation>
</taxonomicName>
, habitus, dorsal view.
<emphasis id="A0EAEAF29D7D923AFB8AF836CC602FC0" bold="true" box="[1049,1078,1928,1954]" pageId="97" pageNumber="98">74</emphasis>
.
<taxonomicName id="559E4D639D7D923AFBD9F837CD202FC1" authority="(Frey, 1969)" authorityName="Telnov" authorityYear="2020" baseAuthorityName="Frey" baseAuthorityYear="1969" box="[1098,1398,1928,1955]" class="Insecta" family="Scarabaeidae" genus="Paramaechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="perlatus" status="comb. nov.">
<emphasis id="A0EAEAF29D7D923AFBD9F837CC832FC0" box="[1098,1237,1928,1954]" italics="true" pageId="97" pageNumber="98">M. perlatus</emphasis>
(
<bibRefCitation id="F60F4B119D7D923AFB7AF836CD382FC1" author="Frey G." box="[1257,1390,1928,1955]" pageId="97" pageNumber="98" pagination="494 - 509" refId="ref98001" refString="Frey G. 1969. Revision der Maechidius - Arten Neu-Guineas (Col., Melolonth.). Entomologische Arbeiten aus dem Museum G. Frey 20: 494 - 509." type="journal article" year="1969">Frey, 1969</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="BBD957899D7D923AFF2EF815C9112FA4" box="[189,327,1963,1990]" pageId="97" pageNumber="98" rank="species">comb. nov.</taxonomicNameLabel>
, holotype, ♂ (NHMB).
<emphasis id="A0EAEAF29D7D923AFDE5F815CAC52FA7" bold="true" box="[630,659,1963,1989]" pageId="97" pageNumber="98">75</emphasis>
. Same, ♂ (SMNS).
<emphasis id="A0EAEAF29D7D923AFC0BF815CBE32FA7" bold="true" box="[920,949,1963,1989]" pageId="97" pageNumber="98">76</emphasis>
.
<taxonomicName id="559E4D639D7D923AFC59F812CC8D2FA4" authority="(Frey, 1969)" authorityName="Telnov" authorityYear="2020" baseAuthorityName="Frey" baseAuthorityYear="1969" box="[970,1243,1963,1990]" class="Insecta" family="Scarabaeidae" genus="Paramaechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="popei" status="comb. nov.">
<emphasis id="A0EAEAF29D7D923AFC59F812CC6E2FA7" box="[970,1080,1964,1989]" italics="true" pageId="97" pageNumber="98">M. popei</emphasis>
(
<bibRefCitation id="F60F4B119D7D923AFBDEF815CC852FA4" author="Frey G." box="[1101,1235,1963,1990]" pageId="97" pageNumber="98" pagination="494 - 509" refId="ref98001" refString="Frey G. 1969. Revision der Maechidius - Arten Neu-Guineas (Col., Melolonth.). Entomologische Arbeiten aus dem Museum G. Frey 20: 494 - 509." type="journal article" year="1969">Frey, 1969</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="BBD957899D7D923AFB7BF815CD242FA4" box="[1256,1394,1963,1990]" pageId="97" pageNumber="98" rank="species">comb. nov.</taxonomicNameLabel>
, holotype, ♀ (BMNH).
<emphasis id="A0EAEAF29D7D923AFE59F870C9B12F8A" bold="true" box="[458,487,1998,2024]" pageId="97" pageNumber="98">77</emphasis>
. Same, paratype, ♂ (BMNH). Not to scale.
</paragraph>
</caption>
<paragraph id="922136E09D7E9239FF2EFB8AC9E62C2D" blockId="98.[189,432,1076,1103]" box="[189,432,1076,1103]" pageId="98" pageNumber="99">
<heading id="C969818C9D7E9239FF2EFB8AC9E62C2D" bold="true" box="[189,432,1076,1103]" fontSize="11" level="3" pageId="98" pageNumber="99" reason="3">
<emphasis id="A0EAEAF29D7E9239FF2EFB8AC9E62C2D" bold="true" box="[189,432,1076,1103]" pageId="98" pageNumber="99">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D7E9239FF2EFBDCCB1E2CFC" blockId="98.[189,1398,1122,1183]" pageId="98" pageNumber="99">Female body generally larger, labroclypeus shallower emarginate on anterior margin; lamellae of antennal club shorter; terminal spur of protibia straight.</paragraph>
</subSubSection>
</treatment>
</document>
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