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<document id="65E2A401F8977679BF1C75B5E7D10998" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D729234FDE8FDD9CCE22E34" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius sougb Telnov 2020, sp. nov." docType="treatment" docUuid="C8545020-E5E7-4EAA-B78C-B68FC1CB885D" docUuidSource="ZooBank" docVersion="10" lastPageNumber="112" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="111" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="D5CDB8006A0C55BEDCE9276AE2A43157">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
</mods:titleInfo>
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<mods:namePart id="6AAAC820FDF08149D4D9E86C4D6D51CF">Telnov, Dmitry</mods:namePart>
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<mods:title id="B7D91BA7FD8795CB998B1C26B61C7423">European Journal of Taxonomy</mods:title>
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<mods:date id="0595DB534C9CD8641D7C798E126A806E">2020</mods:date>
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<mods:number id="94C2A4DC5E4ECEE8DF0269D162DEADF0">2020-10-19</mods:number>
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<mods:identifier id="E8FEC1DE7B1445C207D6E35246E59383" type="DOI">10.5852/ejt.2020.721.1127</mods:identifier>
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<treatment id="1A3787F69D729234FDE8FDD9CCE22E34" ID-DOI="http://doi.org/10.5281/zenodo.4344283" ID-GBIF-Taxon="168648128" ID-Zenodo-Dep="4344283" LSID="urn:lsid:zoobank.org:act:C8545020-E5E7-4EAA-B78C-B68FC1CB885D" httpUri="http://treatment.plazi.org/id/1A3787F69D729234FDE8FDD9CCE22E34" lastPageId="111" lastPageNumber="112" pageId="110" pageNumber="111">
<subSubSection id="DA84656B9D729235FDE8FDD9CBEF2AE0" box="[635,953,615,642]" pageId="110" pageNumber="111" type="nomenclature">
<paragraph id="922136E09D729235FDE8FDD9CBEF2AE0" blockId="110.[360,1227,615,711]" box="[635,953,615,642]" pageId="110" pageNumber="111">
<heading id="C969818C9D729235FDE8FDD9CBEF2AE0" box="[635,953,615,642]" centered="true" fontSize="11" level="2" pageId="110" pageNumber="111" reason="2">
<taxonomicName id="559E4D639D729235FDE8FDD9CB0E2AE0" ID-CoL="7X3XR" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[635,856,615,642]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="species" species="sougb" status="sp. nov.">
<emphasis id="A0EAEAF29D729235FDE8FDD9CB0E2AE0" bold="true" box="[635,856,615,642]" italics="true" pageId="110" pageNumber="111">Maechidius sougb</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D729235FCCCFDD6CBEF2AE0" box="[863,953,616,642]" pageId="110" pageNumber="111" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="DA84656B9D729235FEFBFD34CC452AA5" pageId="110" pageNumber="111" type="description">
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urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
<paragraph id="922136E09D729235FDB2FD12CC452AA5" blockId="110.[360,1227,615,711]" box="[545,1043,684,711]" pageId="110" pageNumber="111">
<figureCitation id="0AA52A659D729235FDB2FD12CA2D2AA5" box="[545,635,684,711]" captionStart="Figs 82–85" captionStartId="106.[189,241,1894,1920]" captionTargetBox="[330,1255,264,1870]" captionTargetId="figure-67@106.[330,1258,252,1883]" captionTargetPageId="106" captionText="Figs 82–85. Papuan and Wallacean Maechidius Macleay, 1819, habitus, dorsal view. 82. M. similis sp. nov., paratype, ♂ (BMNH). 83. M. skalei sp. nov., paratype, ♂ (NME). 84. M. simplex Frey, 1969, holotype, ♂ (photograph courtesy BPBM) (BPBM). 85. M. sougb sp. nov., holotype, ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122165" httpUri="https://zenodo.org/record/4122165/files/figure.png" pageId="110" pageNumber="111">Figs 85</figureCitation>
,
<figureCitation id="0AA52A659D729235FD14FD12CAE32AA4" box="[647,693,684,710]" captionStart="Figs 169–180" captionStartId="143.[189,241,1787,1813]" captionTargetBox="[359,1217,271,1750]" captionTargetId="figure-123@143.[359,1228,265,1755]" captionTargetPageId="143" captionText="Figs 169–180. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 169. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 170. Same, paratype, ♂ (BMNH). 171. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 172. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 173. Same, ♂ (ZMHB). 174. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 175. M. similis sp. nov., paratype, ♂ (BMNH). 176. Same, paratype, ♀ (BMNH). 177. M. skalei sp. nov., paratype, ♂ (NME). 178. M. sougb sp. nov., holotype, ♂ (NME). 179. M. speciosus (Frey, 1969) comb. nov., ♂ (NHMB). 180. Same, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122185" httpUri="https://zenodo.org/record/4122185/files/figure.png" pageId="110" pageNumber="111">178</figureCitation>
,
<figureCitation id="0AA52A659D729235FD52FD12CAA62AA5" box="[705,752,684,711]" captionStart="Figs 242–258" captionStartId="148.[189,241,1646,1672]" captionTargetBox="[394,1193,265,1617]" captionTargetId="figure-193@148.[394,1193,265,1626]" captionTargetPageId="148" captionText="Figs 242–258. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 242. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 243. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 244. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 245. M. peregrinus Lansberge, 1886, ♂ (DTC). 246. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 247. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 248. M. riedeli sp. nov., holotype, ♂ (SMNS). 249. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 250. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 251. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 252. M. similis sp. nov., paratype, ♂ (BMNH). 253. M. skalei sp. nov., paratype, ♂ (NME). 254. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 255. M. sougb sp. nov., holotype, ♂ (NME). 256. M. speciosus (Frey, 1969) comb. nov., ♀ (BMNH). 257. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 258. M. suwawa sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122196" httpUri="https://zenodo.org/record/4122196/files/figure.png" pageId="110" pageNumber="111">255</figureCitation>
,
<figureCitation id="0AA52A659D729235FD6FFD12CB7D2AA5" box="[764,811,684,711]" captionStart="Figs 333–348" captionStartId="155.[189,241,1682,1708]" captionTargetBox="[394,1193,265,1641]" captionTargetId="figure-177@155.[394,1193,265,1655]" captionTargetPageId="155" captionText="Figs 333–348. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 333. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 334. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 335. M. peregrinus Lansberge, 1886, ♂ (DTC). 336. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 337. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 338. Same, enlarged (SMNS). 339. M. riedeli sp. nov., holotype, ♂ (SMNS). 340. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 341. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 342. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 343. M. similis sp. nov., paratype, ♂ (BMNH). 344. M. skalei sp. nov., paratype, ♂ (NME). 345. M. sougb sp. nov., holotype, ♂ (NME). 346. M. speciosus (Frey, 1969) comb. nov., ♀(BMNH). 347. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 348. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122212" httpUri="https://zenodo.org/record/4122212/files/figure.png" pageId="110" pageNumber="111">345</figureCitation>
, 417, 518, 715–717
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D729235FF2EFD52CBDF2C21" pageId="110" pageNumber="111" type="diagnosis">
<paragraph id="922136E09D729235FF2EFD52C99F2B64" blockId="110.[189,457,748,774]" box="[189,457,748,774]" pageId="110" pageNumber="111">
<heading id="C969818C9D729235FF2EFD52C99F2B64" bold="true" box="[189,457,748,774]" fontSize="11" level="3" pageId="110" pageNumber="111" reason="3">
<emphasis id="A0EAEAF29D729235FF2EFD52C99F2B64" bold="true" box="[189,457,748,774]" pageId="110" pageNumber="111">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D729235FF2EFCA7CBDF2C21" blockId="110.[189,1399,793,1091]" pageId="110" pageNumber="111">
This is one of the largest
<taxonomicName id="559E4D639D729235FE7FFCA7CA232B51" authorityName="Macleay" authorityYear="1819" box="[492,629,793,819]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="genus">
<emphasis id="A0EAEAF29D729235FE7FFCA7CA232B51" box="[492,629,793,819]" italics="true" pageId="110" pageNumber="111">Maechidius</emphasis>
</taxonomicName>
in the Papuan Region. It is distinguishable among all its Papuan congeners with an (almost) glabrous dorsum and bidentate protibiae primarily in the shape of the male genital organs, the 8-segmented antennae, the strongly protruding anterolateral angles of the male labroclypeus (which, in addition, is deeply emarginate anteriorly) in combination with rather delicate punctures on the pronotum and elytra. The most similar species,
<taxonomicName id="559E4D639D729235FC5EFC1CC9572BBF" authority="(Narakusumo &amp; Balke, 2019)" authorityName="Telnov" authorityYear="2020" baseAuthorityName="Narakusumo &amp; Balke" baseAuthorityYear="2019" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="species" species="obiensis" status="comb. nov.">
<emphasis id="A0EAEAF29D729235FC5EFC1CCC002BD9" box="[973,1110,929,955]" italics="true" pageId="110" pageNumber="111">M. obiensis</emphasis>
(
<bibRefCitation id="F60F4B119D729235FBFBFC1FC8AF2BBF" author="Narakusumo R. P. &amp; Balke M." pageId="110" pageNumber="111" pagination="35 - 50" refId="ref98467" refString="Narakusumo R. P. &amp; Balke M. 2019. Four new species of Epholcis Waterhouse, 1875 (Coleoptera: Scarabaeidae: Melolonthinae: Maechidiini) from the Moluccas, Indonesia. Treubia 46: 35 - 50. https: // doi. org / 10.14203 / treubia. v 46 i 0.3791" type="journal article" year="2019">Narakusumo &amp; Balke, 2019</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="BBD957899D729235FE9CFC7DC9CF2BBC" box="[271,409,963,990]" pageId="110" pageNumber="111" rank="species">comb. nov.</taxonomicNameLabel>
, is generally smaller (body length under
<quantity id="55669B059D729235FC30FC7ACBA72BBC" box="[931,1009,964,990]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="110" pageNumber="111" unit="mm" value="7.0">7 mm</quantity>
), male metatibia thickened and somewhat curved on inner margin and with differently shaped aedeagus
<taxonomicName id="559E4D639D729235FB95FC5BC8AE2C43" authority="Arrow, 1941" authorityName=", Arrow" authorityYear="1941" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="species" species="pedarioides">
<emphasis id="A0EAEAF29D729235FB95FC5BCD762B9D" box="[1030,1312,997,1023]" italics="true" pageId="110" pageNumber="111">Maechidius pedarioides</emphasis>
<bibRefCitation id="F60F4B119D729235FAB7FC58C8AE2C43" author="Arrow G. E." pageId="110" pageNumber="111" pagination="448 - 464" refId="ref97558" refString="Arrow G. E. 1941. XXIX. - Some undescribed species of melolonthid Coleoptera from Western New Guinea and the adjacent islands of Waigeu and Japen. Annals and Magazine of Natural History, Series 11 7 (41): 448 - 464. https: // doi. org / 10.1080 / 03745481.1941.9727945" type="journal article" year="1941">Arrow, 1941</bibRefCitation>
</taxonomicName>
(Raja Ampat Islands: Waigeo), has tridentate protibiae and stronger dorsal punctures, as well as less protruding anterolateral angles of the male labroclypeus.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D729235FF2EFBD4CAA82C97" pageId="110" pageNumber="111" type="etymology">
<paragraph id="922136E09D729235FF2EFBD4C9122CE6" blockId="110.[189,324,1130,1156]" box="[189,324,1130,1156]" pageId="110" pageNumber="111">
<heading id="C969818C9D729235FF2EFBD4C9122CE6" bold="true" box="[189,324,1130,1156]" fontSize="11" level="3" pageId="110" pageNumber="111" reason="3">
<emphasis id="A0EAEAF29D729235FF2EFBD4C9122CE6" bold="true" box="[189,324,1130,1156]" pageId="110" pageNumber="111">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D729235FF2EFB29CAA82C97" blockId="110.[189,1398,1175,1269]" pageId="110" pageNumber="111">
Named after Sougb, the local name of one of the main tribes and languages in the Anggi Lakes area of the Arfak Mountains. Sougb has the alternative names of Soug and Mantion, but is listed as Manikion by
<bibRefCitation id="F60F4B119D729235FF71FB65CA5C2C97" author="Simons G. F. &amp; Fennig C. D." box="[226,522,1243,1269]" pageId="110" pageNumber="111" refId="ref98633" refString="Simons G. F. &amp; Fennig C. D. (eds) 2018. Ethnologue: Languages of the World. 21 st Edition. SIL International, Dallas, Texas. Available from http: // www. ethnologue. com [accessed 6 Jan. 2019]." type="url" year="2018">Simons &amp; Fennig (2018)</bibRefCitation>
. Noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D729235FF2EFAA5CB7C2E49" pageId="110" pageNumber="111" type="materials_examined">
<paragraph id="922136E09D729235FF2EFAA5C93A2D57" blockId="110.[189,364,1307,1333]" box="[189,364,1307,1333]" pageId="110" pageNumber="111">
<heading id="C969818C9D729235FF2EFAA5C93A2D57" bold="true" box="[189,364,1307,1333]" fontSize="11" level="3" pageId="110" pageNumber="111" reason="3">
<emphasis id="A0EAEAF29D729235FF2EFAA5C93A2D57" bold="true" box="[189,364,1307,1333]" pageId="110" pageNumber="111">Type material</emphasis>
</heading>
</paragraph>
<materialsCitation id="22F63CBD9D729235FF79FAF7CC772DAB" ID-GBIF-Occurrence="2898878452" collectingDate="2015-09-10" collectingDateMax="2015-09-11" collectingDateMin="2015-09-10" collectionCode="NME" country="Indonesia" elevation="2200" latitude="-1.3013889" location="Arfak mts" longLatPrecision="21" longitude="133.90666" municipality="Anggi Gigi Lake" pageId="110" pageNumber="111" specimenCount="1" typeStatus="holotype">
<paragraph id="922136E09D729235FF79FAF7C90A2D01" blockId="110.[189,1398,1353,1481]" box="[234,348,1353,1379]" pageId="110" pageNumber="111">
<emphasis id="A0EAEAF29D729235FF79FAF7C90A2D01" bold="true" box="[234,348,1353,1379]" pageId="110" pageNumber="111">
<typeStatus id="4D2588429D729235FF79FAF7C90A2D01" box="[234,348,1353,1379]" pageId="110" pageNumber="111" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="922136E09D729235FF2EFAD5CC772DAB" blockId="110.[189,1398,1353,1481]" pageId="110" pageNumber="111">
<collectingCountry id="EA8976709D729235FF2EFAD5C90D2DE7" box="[189,347,1387,1413]" name="Indonesia" pageId="110" pageNumber="111">INDONESIA</collectingCountry>
• ♂; “
<collectingCountry id="EA8976709D729235FE30FAD5CA122DE7" box="[419,580,1387,1413]" name="Indonesia" pageId="110" pageNumber="111">INDONESIA</collectingCountry>
E, W New Guinea, Doberai Peninsula,
<location id="9741603B9D729235FB86FAD4CCD82DE7" LSID="urn:lsid:plazi:treatment:1A3787F69D729234FDE8FDD9CCE22E34:9741603B9D729235FB86FAD4CCD82DE7" box="[1045,1166,1386,1413]" country="Indonesia" latitude="-1.3013889" longLatPrecision="21" longitude="133.90666" municipality="Anggi Gigi Lake" name="Arfak mts" pageId="110" pageNumber="111">Arfak mts</location>
,
<collectingMunicipality id="7245AC9A9D729235FB0BFAD4CD092DE6" box="[1176,1375,1386,1412]" pageId="110" pageNumber="111">Anggi Gigi Lake</collectingMunicipality>
S env., Uper vill.,
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,
<geoCoordinate id="F7AA50279D729235FD93FA32CACB2DC5" box="[512,669,1420,1447]" degrees="133" direction="east" minutes="54" orientation="longitude" pageId="110" pageNumber="111" precision="15" seconds="24" value="133.90666">133°54’24”E</geoCoordinate>
, 9-10 &amp;
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<collectingDate id="F664E9C89D729235FC92FA32CBF92DC5" box="[769,943,1420,1447]" pageId="110" pageNumber="111" value="2015-09-10" valueMax="2015-09-11" valueMin="2015-09-10">10-11.IX.2015</collectingDate>
</date>
,
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<elevation id="19B3D1D39D729235FC29FA33CC422DC5" box="[954,1044,1421,1447]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="110" pageNumber="111" unit="m" value="2200.0">2200 m</elevation>
</quantity>
, edge of primary mid montane rainforest, white light”;
<collectionCode id="F48FAE259D729235FE44FA11CA4E2DAB" box="[471,536,1455,1481]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="110" pageNumber="111">NME</collectionCode>
[tarsomeres 4–5 of right protarsus missing].
</paragraph>
</materialsCitation>
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<typeStatus id="4D2588429D729235FF79FA51C90A2E6B" box="[234,348,1519,1545]" pageId="110" pageNumber="111" type="paratype">Paratype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="922136E09D729235FF2EF9AFCB7C2E49" blockId="110.[189,810,1519,1579]" box="[189,810,1553,1579]" pageId="110" pageNumber="111">
<materialsCitation id="22F63CBD9D729235FF2EF9AFCB702E49" ID-GBIF-Occurrence="2898878588" box="[189,806,1553,1579]" collectionCode="DTC" country="Indonesia" location="INDONESIA" pageId="110" pageNumber="111" specimenCount="1" typeStatus="holotype">
<collectingCountry id="EA8976709D729235FF2EF9AFC90D2E49" box="[189,347,1553,1579]" name="Indonesia" pageId="110" pageNumber="111">INDONESIA</collectingCountry>
•
<specimenCount id="8498FD699D729235FEE7F9AFC9F42E49" box="[372,418,1553,1579]" pageId="110" pageNumber="111" type="male">1 ♂</specimenCount>
; same label as in
<typeStatus id="4D2588429D729235FDE2F9AFCA8D2E49" box="[625,731,1553,1579]" pageId="110" pageNumber="111" type="holotype">holotype</typeStatus>
;
<collectionCode id="F48FAE259D729235FD7BF9AFCB702E49" box="[744,806,1553,1579]" collectionName="Dmitrijs Telnovs" pageId="110" pageNumber="111">DTC</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D729234FF2EF9EFC9FD2D1B" lastPageId="111" lastPageNumber="112" pageId="110" pageNumber="111" type="description">
<paragraph id="922136E09D729235FF2EF9EFC9182E09" blockId="110.[189,334,1617,1643]" box="[189,334,1617,1643]" pageId="110" pageNumber="111">
<heading id="C969818C9D729235FF2EF9EFC9182E09" bold="true" box="[189,334,1617,1643]" fontSize="11" level="3" pageId="110" pageNumber="111" reason="3">
<emphasis id="A0EAEAF29D729235FF2EF9EFC9182E09" bold="true" box="[189,334,1617,1643]" pageId="110" pageNumber="111">Description</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D729235FF2EF9C1CA1B2EBE" blockId="110.[189,1399,1662,1757]" pageId="110" pageNumber="111">
MEASUREMENTS.
<typeStatus id="4D2588429D729235FEE8F9C0C9BD2EFA" box="[379,491,1662,1688]" pageId="110" pageNumber="111" type="holotype">Holotype</typeStatus>
, total body length
<quantity id="55669B059D729235FD51F9C0CB6A2EFB" box="[706,828,1662,1689]" metricMagnitude="-2" metricUnit="m" metricValue="1.005" pageId="110" pageNumber="111" unit="mm" value="10.05">10.05 mm</quantity>
. Head
<quantity id="55669B059D729235FC1FF9C0CBA22EFB" box="[908,1012,1662,1689]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" pageId="110" pageNumber="111" unit="mm" value="1.75">1.75 mm</quantity>
long, across eyes
<quantity id="55669B059D729235FB5BF9C0CD662EFB" box="[1224,1328,1662,1689]" metricMagnitude="-3" metricUnit="m" metricValue="2.07" pageId="110" pageNumber="111" unit="mm" value="2.07">2.07 mm</quantity>
wide. Pronotum
<quantity id="55669B059D729235FEA4F91EC9C92ED9" box="[311,415,1696,1723]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="110" pageNumber="111" unit="mm" value="2.2">2.20 mm</quantity>
long, maximum width
<quantity id="55669B059D729235FD3EF91ECB412ED9" box="[685,791,1696,1723]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="110" pageNumber="111" unit="mm" value="3.1">3.10 mm</quantity>
. Elytral length
<quantity id="55669B059D729235FC54F91ECC642ED9" box="[967,1074,1696,1723]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="110" pageNumber="111" unit="mm" value="6.1">6.10 mm</quantity>
, maximum combined width
<quantity id="55669B059D729235FF2EF97CC97F2EBF" box="[189,297,1730,1757]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="110" pageNumber="111" unit="mm" value="4.5">4.50 mm</quantity>
.
<typeStatus id="4D2588429D729235FEA7F97DC9CD2EBF" box="[308,411,1731,1757]" pageId="110" pageNumber="111" type="paratype">Paratype</typeStatus>
<quantity id="55669B059D729235FE31F97CCA5D2EBF" box="[418,523,1730,1757]" metricMagnitude="-3" metricUnit="m" metricValue="9.7" pageId="110" pageNumber="111" unit="mm" value="9.7">9.70 mm</quantity>
long.
</paragraph>
<paragraph id="922136E09D729234FF2EF8BDCB712D69" blockId="110.[189,1399,1794,2025]" lastBlockId="111.[189,1399,265,1291]" lastPageId="111" lastPageNumber="112" pageId="110" pageNumber="111">
Dorsum and venter uniformly dark castaneous with labroclypeus, mouthparts and legs somewhat paler reddish brown. Head transverse, glossy dorsally and ventrally, with large slightly prominent compound eyes occupying about half side of head. Male labroclypeus (
<figureCitation id="0AA52A659D729235FC03F8F8CBA02F02" box="[912,1014,1862,1888]" captionStart="Figs 169–180" captionStartId="143.[189,241,1787,1813]" captionTargetBox="[359,1217,271,1750]" captionTargetId="figure-123@143.[359,1228,265,1755]" captionTargetPageId="143" captionText="Figs 169–180. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 169. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 170. Same, paratype, ♂ (BMNH). 171. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 172. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 173. Same, ♂ (ZMHB). 174. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 175. M. similis sp. nov., paratype, ♂ (BMNH). 176. Same, paratype, ♀ (BMNH). 177. M. skalei sp. nov., paratype, ♂ (NME). 178. M. sougb sp. nov., holotype, ♂ (NME). 179. M. speciosus (Frey, 1969) comb. nov., ♂ (NHMB). 180. Same, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122185" httpUri="https://zenodo.org/record/4122185/files/figure.png" pageId="110" pageNumber="111">Fig. 178</figureCitation>
) with deep and broad U-shaped emargination, its lateral margins sinuous in both dorsal and lateral views. Anterolateral angles large, acute angulate and raised up at angle of nearly 60–70° to axis of frons in lateral view. Anterior margin of labroclypeus smooth. Punctures of head irregularly circular, shallow, in part annular, larger in posterior part. Intervening spaces glossy, smaller than to as large as punctures. Pubescence yellowish, sparse, generally appressed and directed posteriad; seta rises from anterior margin of each puncture, surpassing length of corresponding puncture. Setae between compound eyes distinctly longer than those on forehead. Male antenna 8-segmented, antennal club 3-segmented. Scape large, with bulbous denticle-like predistal projection on upper side, provided with few long erect setae on its posterior and two very long on distal margin. Antennomere 2 strongly widened distally but not transverse. Pronotum transverse, glossy dorsally and laterally. Anterior margin of pronotum slightly sinuous with anterolateral angles (stronger) and mesal portion (slightly) protruding anteriad. Basal margin of pronotum very broadly rounded. Lateral margin of pronotum almost straight in anterior half, gradually widened towards median part, rounded in basal half and shortly emarginate just in front of posterolateral angles (
<figureCitation id="0AA52A659D739234FB1FFDA3CCA22A55" box="[1164,1268,540,567]" captionStart="Figs 242–258" captionStartId="148.[189,241,1646,1672]" captionTargetBox="[394,1193,265,1617]" captionTargetId="figure-193@148.[394,1193,265,1626]" captionTargetPageId="148" captionText="Figs 242–258. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 242. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 243. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 244. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 245. M. peregrinus Lansberge, 1886, ♂ (DTC). 246. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 247. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 248. M. riedeli sp. nov., holotype, ♂ (SMNS). 249. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 250. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 251. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 252. M. similis sp. nov., paratype, ♂ (BMNH). 253. M. skalei sp. nov., paratype, ♂ (NME). 254. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 255. M. sougb sp. nov., holotype, ♂ (NME). 256. M. speciosus (Frey, 1969) comb. nov., ♀ (BMNH). 257. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 258. M. suwawa sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122196" httpUri="https://zenodo.org/record/4122196/files/figure.png" pageId="111" pageNumber="112">Fig. 255</figureCitation>
). All along with well-defined crenulae which becomes protruding and denticle-like near prebasal emargination. Long erect seta present between each pair of lateral crenulae. Lateral margin of pronotum straight in lateral view. Pronotal punctures irregularly circular, rather small and shallow. Intervening spaces glossy, generally as large as to twice as large as punctures. Pronotal setae inconspicuous and short; seta rises from anterior margin of each puncture, slightly surpassing its length (pronotum looks glabrous). Hypomeron nearly straight, separated from prosternum by low slightly arched carina, with long setae on its anterolateral margin. Antennal pocket shallow. Scutellar shield narrowly triangular, rounded apically. Elytra slightly rounded laterally, maximum width across median third, glossy and flattened dorsally, with rounded humeri. Inconspicuous obtuse hump on each elytron near apex. No tracks of longitudinal carinae and no sutural carinae present. Each elytron between suture and humerus with 5 paired rows of delicate circular shallow punctures (some rows partly confused). Lateral sides of elytra more confusedly punctate (
<figureCitation id="0AA52A659D739234FEA3FC04C9C12BB7" box="[304,407,954,981]" captionStart="Figs 333–348" captionStartId="155.[189,241,1682,1708]" captionTargetBox="[394,1193,265,1641]" captionTargetId="figure-177@155.[394,1193,265,1655]" captionTargetPageId="155" captionText="Figs 333–348. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 333. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 334. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 335. M. peregrinus Lansberge, 1886, ♂ (DTC). 336. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 337. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 338. Same, enlarged (SMNS). 339. M. riedeli sp. nov., holotype, ♂ (SMNS). 340. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 341. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 342. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 343. M. similis sp. nov., paratype, ♂ (BMNH). 344. M. skalei sp. nov., paratype, ♂ (NME). 345. M. sougb sp. nov., holotype, ♂ (NME). 346. M. speciosus (Frey, 1969) comb. nov., ♀(BMNH). 347. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 348. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122212" httpUri="https://zenodo.org/record/4122212/files/figure.png" pageId="111" pageNumber="112">Fig. 345</figureCitation>
). Intervening spaces between punctures in rows larger than punctures, between rows about 3–4× as large as punctures. Elytral setae minute; seta rises from anterior margin of each puncture, reaches posterior margin or shortly surpassing its length. Lateral margin of elytra with moderately long suberect setae. Male pygidium (
<figureCitation id="0AA52A659D739234FDAAFB9CCAC92C5E" box="[569,671,1057,1084]" captionStart="Figs 504–521" captionStartId="166.[189,241,1681,1707]" captionTargetBox="[433,1155,265,1647]" captionTargetId="figure-177@166.[433,1155,265,1658]" captionTargetPageId="166" captionText="Figs 504–521. Papuan and Wallacean Maechidius Macleay, 1819, pygidium, dorsal view. 504. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 505. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 506. Same, ♀ (BMNH). 507. M. pedarioides Arrow, 1941, syntype, ♀ (BMNH). 508. Same, syntype, ♂ (BMNH). 509. M. peregrinus Lansberge, 1886, ♂ (DTC). 510. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 511. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 512. Same, paratype, ♂ (BMNH). 513. M. riedeli sp. nov., holotype, ♂ (SMNS). 514. M. rugicollis Moser, 1920, ♂ (BMNH). 515. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 516. M. similis sp. nov., paratype, ♂ (BMNH). 517. M. skalei sp. nov., paratype, ♂ (NME). 518. M. sougb sp. nov., holotype, ♂ (NME). 519. M. speciosus (Frey, 1969) comb. nov., ♂ (BMNH). 520. Same, different ♂ (BMNH). 521. Same, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122228" httpUri="https://zenodo.org/record/4122228/files/figure.png" pageId="111" pageNumber="112">Fig. 518</figureCitation>
) and abdominal ventrites densely covered with large shallow annular punctures, each provided with suberect moderately long seta. Abdominal ventrites with no track of lateral longitudinal ridge. Legs long and slender, femora and tibiae covered with long erect setae arranged in 5 lines. Protibiae hardly sinuous on external margin. Protibiae with delicate carina on dorsal surface. Two large obtuse teeth on external margin of male protibia: distal one narrower, basal one broader and more obtuse (Fig. 417). Male protibial terminal spur absent. Tarsal claws with large pulvilli (in males only?). Male aedeagus as in
<figureCitation id="0AA52A659D739234FDECFB4ECB752D69" box="[639,803,1264,1291]" captionStart="Figs 712–725" captionStartId="179.[189,241,1892,1918]" captionTargetBox="[433,1144,277,1858]" captionTargetId="figure-72@179.[427,1160,265,1870]" captionTargetPageId="179" captionText="Figs 712–725. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 712– 714. M. skalei sp. nov., holotype, ♂ (NME). 715–717. M. sougb sp. nov., holotype, ♂ (NME). 718–720. M. speciosus (Frey, 1969) comb. nov., ♂ (BMNH). 721–723. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 724–725. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122252" httpUri="https://zenodo.org/record/4122252/files/figure.png" pageId="111" pageNumber="112">Figs 715–717</figureCitation>
.
</paragraph>
<paragraph id="922136E09D739234FF2EFA8FC9E62D29" blockId="111.[189,432,1329,1355]" box="[189,432,1329,1355]" pageId="111" pageNumber="112">
<heading id="C969818C9D739234FF2EFA8FC9E62D29" bold="true" box="[189,432,1329,1355]" fontSize="11" level="3" pageId="111" pageNumber="112" reason="3">
<emphasis id="A0EAEAF29D739234FF2EFA8FC9E62D29" bold="true" box="[189,432,1329,1355]" pageId="111" pageNumber="112">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D739234FF2EFAE1C9FD2D1B" blockId="111.[189,427,1375,1401]" box="[189,427,1375,1401]" pageId="111" pageNumber="112">Female is unknown.</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D739234FF2EFA1ECCC32D85" box="[189,1173,1440,1512]" pageId="111" pageNumber="112" type="biology_ecology">
<paragraph id="922136E09D739234FF2EFA1EC9762DD8" blockId="111.[189,288,1440,1466]" box="[189,288,1440,1466]" pageId="111" pageNumber="112">
<heading id="C969818C9D739234FF2EFA1EC9762DD8" bold="true" box="[189,288,1440,1466]" fontSize="11" level="3" pageId="111" pageNumber="112" reason="3">
<emphasis id="A0EAEAF29D739234FF2EFA1EC9762DD8" bold="true" box="[189,288,1440,1466]" pageId="111" pageNumber="112">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D739234FF2EFA70CCC32D85" blockId="111.[189,1173,1485,1512]" box="[189,1173,1485,1512]" pageId="111" pageNumber="112">
Occurs in primary mid-montane rainforests at ~
<quantity id="55669B059D739234FD7FFA70CB102D8A" box="[748,838,1486,1512]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="111" pageNumber="112" unit="m" value="2200.0">2200 m</quantity>
altitude. Possibly nocturnal.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D739234FF2EF9B0CCE22E34" box="[189,1204,1550,1623]" pageId="111" pageNumber="112" type="distribution">
<paragraph id="922136E09D739234FF2EF9B0C9002E4A" blockId="111.[189,342,1550,1576]" box="[189,342,1550,1576]" pageId="111" pageNumber="112">
<heading id="C969818C9D739234FF2EF9B0C9002E4A" bold="true" box="[189,342,1550,1576]" fontSize="11" level="3" pageId="111" pageNumber="112" reason="3">
<emphasis id="A0EAEAF29D739234FF2EF9B0C9002E4A" bold="true" box="[189,342,1550,1576]" pageId="111" pageNumber="112">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D739234FF2EF982CCE22E34" blockId="111.[189,1204,1596,1623]" box="[189,1204,1596,1623]" pageId="111" pageNumber="112">
Hitherto only known from Arfak Mountains, Doberai Peninsula, western New
<collectingCountry id="EA8976709D739234FBC4F982CCE62E34" box="[1111,1200,1596,1622]" name="Guinea-Bissau" pageId="111" pageNumber="112">Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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