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<document id="FFEA56764ED3544BBFDB9E4B803C19E9" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D659220FDEEFAECCB9F2AF4" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius ursus Telnov 2020, sp. nov." docType="treatment" docUuid="395C3699-19FC-48D0-BAAD-689558BC2BF6" docUuidSource="ZooBank" docVersion="10" lastPageNumber="124" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="122" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="D6AC1015A6DEDB3AB7BBDB017C7BB80A">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
</mods:titleInfo>
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<mods:namePart id="FDFBAB59004504BF650D31B048809AAA">Telnov, Dmitry</mods:namePart>
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<mods:title id="1F6D177F6D64B86BC09B8A5DF94B2ED2">European Journal of Taxonomy</mods:title>
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<mods:date id="0C6DAD12273C0F3F491B1283B20C47FC">2020</mods:date>
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<mods:number id="60087DE8716952F5E4526A2B69524387">2020-10-19</mods:number>
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<mods:classification id="047679D7ADFAA44B6610DCA5CBF62B01">journal article</mods:classification>
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<mods:identifier id="155EC32004756935111B62F634057655" type="DOI">10.5852/ejt.2020.721.1127</mods:identifier>
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<treatment id="1A3787F69D659220FDEEFAECCB9F2AF4" ID-DOI="http://doi.org/10.5281/zenodo.4344309" ID-GBIF-Taxon="168648129" ID-Zenodo-Dep="4344309" LSID="urn:lsid:zoobank.org:act:395C3699-19FC-48D0-BAAD-689558BC2BF6" httpUri="http://treatment.plazi.org/id/1A3787F69D659220FDEEFAECCB9F2AF4" lastPageId="123" lastPageNumber="124" pageId="121" pageNumber="122">
<subSubSection id="DA84656B9D659222FDEEFAECCBE12D0F" box="[637,951,1362,1389]" pageId="121" pageNumber="122" type="nomenclature">
<paragraph id="922136E09D659222FDEEFAECCBE12D0F" blockId="121.[363,1224,1362,1463]" box="[637,951,1362,1389]" pageId="121" pageNumber="122">
<heading id="C969818C9D659222FDEEFAECCBE12D0F" box="[637,951,1362,1389]" centered="true" fontSize="11" level="2" pageId="121" pageNumber="122" reason="2">
<taxonomicName id="559E4D639D659222FDEEFAECCB032D0F" ID-CoL="7X3XX" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[637,853,1362,1389]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="species" species="ursus" status="sp. nov.">
<emphasis id="A0EAEAF29D659222FDEEFAECCB032D0F" bold="true" box="[637,853,1362,1389]" italics="true" pageId="121" pageNumber="122">Maechidius ursus</emphasis>
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<taxonomicNameLabel id="BBD957899D659222FCCFFAEDCBE12D0F" box="[860,951,1363,1389]" pageId="121" pageNumber="122" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="DA84656B9D659222FEF8FAC6CC452DD5" pageId="121" pageNumber="122" type="description">
<paragraph id="922136E09D659222FEF8FAC6CC9E2DF1" blockId="121.[363,1224,1362,1463]" box="[363,1224,1400,1427]" pageId="121" pageNumber="122">
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urn:lsid:zoobank.org:act:
<uuid id="E6380C359D659222FD1AFAC6CC9E2DF1" box="[649,1224,1400,1427]" pageId="121" pageNumber="122">395C3699-19FC-48D0-BAAD-689558BC2BF6</uuid>
</uri>
</paragraph>
<paragraph id="922136E09D659222FDB2FA22CC452DD5" blockId="121.[363,1224,1362,1463]" box="[545,1043,1436,1463]" pageId="121" pageNumber="122">
<figureCitation id="0AA52A659D659222FDB2FA22CA2D2DD5" box="[545,635,1436,1463]" captionStart="Figs 90–93" captionStartId="119.[189,241,1928,1954]" captionTargetBox="[329,1264,264,1883]" captionTargetId="figure-60@119.[321,1264,256,1905]" captionTargetPageId="119" captionText="Figs 90–93. Papuan Maechidius Macleay, 1819, habitus, dorsal view. 90. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 91. M. trivialis sp. nov., holotype, ♂ (BMNH). 92. M. ursus sp. nov., holotype, ♀ (NME). 93. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122169" httpUri="https://zenodo.org/record/4122169/files/figure.png" pageId="121" pageNumber="122">Figs 92</figureCitation>
,
<figureCitation id="0AA52A659D659222FD14FA23CAE32DD5" box="[647,693,1437,1463]" captionStart="Figs 181–192" captionStartId="144.[189,241,1787,1813]" captionTargetBox="[338,1249,274,1742]" captionTargetId="figure-122@144.[338,1249,265,1749]" captionTargetPageId="144" captionText="Figs 181–192. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 181. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 182. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 183. Same, ♀ (SNSD). 184. M. suwawa sp. nov., holotype, ♂ (BMNH). 185. Same, paratype, ♀ (BMNH). 186. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 187. M. trivialis sp. nov., holotype, ♂ (BMNH). 188. M. ursus sp. nov., holotype, ♀ (NME). 189. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 190. Same, ♂ (SNSD). 191. M. weigeli sp. nov., holotype, ♂ (NME). 192. Same, paratype, ♀ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122187" httpUri="https://zenodo.org/record/4122187/files/figure.png" pageId="121" pageNumber="122">188</figureCitation>
,
<figureCitation id="0AA52A659D659222FD52FA22CAA62DD4" box="[705,752,1436,1462]" captionStart="Figs 259–265" captionStartId="149.[189,241,1372,1398]" captionTargetBox="[189,1379,265,1308]" captionTargetId="figure-88@149.[189,1398,265,1311]" captionTargetPageId="149" captionText="Figs 259–265. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 259. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 260. M. trivialis sp. nov., holotype, ♂ (BMNH). 261. M. ursus sp. nov., holotype, ♀ (NME). 262. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 263. M. weigeli sp. nov., holotype, ♂ (NME). 264. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 265. M. yamdena sp. nov., paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122198" httpUri="https://zenodo.org/record/4122198/files/figure.png" pageId="121" pageNumber="122">261</figureCitation>
,
<figureCitation id="0AA52A659D659222FD6FFA22CB302DD5" box="[764,870,1436,1463]" captionStart="Figs 349–358" captionStartId="156.[189,241,1786,1812]" captionTargetBox="[287,1299,265,1743]" captionTargetId="figure-123@156.[287,1301,265,1760]" captionTargetPageId="156" captionText="Figs 349–358. Papuan and Wallacean Maechidius Macleay, 1819, elytron (349–356) and pronotal setation (357–358), dorsal view. 349. M. suwawa sp. nov., paratype, ♀ (BMNH). 350. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 351. M. trivialis sp. nov., holotype, ♂ (BMNH). 352. M. ursus sp. nov., holotype, ♀ (NME). 353. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 354. M. weigeli sp. nov., holotype, ♂ (NME). 355. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 356. M. yamdena sp. nov., paratype, ♂ (BMNH). 357. M. leucopsar sp. nov., holotype, ♂ (IECA). 358. M. ursus sp. nov., holotype, ♀ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122214" httpUri="https://zenodo.org/record/4122214/files/figure.png" pageId="121" pageNumber="122">352, 358</figureCitation>
,
<figureCitation id="0AA52A659D659222FCE2FA23CBF62DD5" box="[881,928,1437,1463]" captionStart="Figs 424–435" captionStartId="161.[189,241,1394,1420]" captionTargetBox="[238,1349,265,1318]" captionTargetId="figure-136@161.[217,1354,265,1322]" captionTargetPageId="161" captionText="Figs 424–435. Papuan and Wallacean Maechidius Macleay, 1819, protibia, dorsal view (424–428), and abdominal ventrites (429–435), ventral and lateral views. 424. M. ursus sp. nov., holotype, ♀ (NME). 425. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 426. M. weigeli sp. nov., holotype, ♂ (NME). 427. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 428. M. yamdena sp. nov., paratype, ♂ (BMNH). 429. M.dani sp. nov., holotype, ♂ (IECA). 430. Same, different aspect (IECA). 431. M. hamatus sp. nov., holotype, ♂ (IECA). 432. Same, different aspect (IECA). 433. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 434. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 435. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122216" httpUri="https://zenodo.org/record/4122216/files/figure.png" pageId="121" pageNumber="122">424</figureCitation>
,
<figureCitation id="0AA52A659D659222FC3FFA23CB8D2DD5" box="[940,987,1437,1463]" captionStart="Figs 436–450" captionStartId="162.[189,241,1619,1645]" captionTargetBox="[205,1349,265,1554]" captionTargetId="figure-184@162.[205,1377,265,1557]" captionTargetPageId="162" captionText="Figs 436–450. Papuan and Wallacean Maechidius Macleay, 1819, legs, various views. 436. M. hirtipes Arrow, 1941, lectotype, ♂, basal metatarsomere (BMNH). 437. M. tarsalis Arrow, 1941, lectotype, ♂, basal metatarsomere (BMNH). 438. M. babyrousa sp. nov., paratype, ♀, metatibial terminal spurs (BMNH). 439. Same, paratype, ♂ (BMNH). 440. Same, different aspect (BMNH). 441. M. yamdena sp. nov., paratype, ♀, metatibial terminal spurs (BMNH). 442. Same, paratype, ♂ (BMNH). 443. M. longipes sp. nov., holotype, ♂, metatibial terminal spurs (BMNH). 444. M. ursus sp. nov., holotype, ♀, left metatibia and basal metatarsomere (NME). 445. M. dani sp. nov., holotype, ♂, apical portion of metatibia (IECA). 446. M. skalei sp. nov., holotype, ♂, apical portion of metatibia and basal metatarsomere (NME). 447. M. yamdena sp. nov., paratype, ♂, protibial terminal spur (BMNH). 448. Same, paratype, ♀ (BMNH). 449. M. babyrousa sp. nov., paratype, ♀, protibial terminal spur (BMNH). 450. Same, paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122218" httpUri="https://zenodo.org/record/4122218/files/figure.png" pageId="121" pageNumber="122">444</figureCitation>
, 528
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D659222FF2EFA61CC912F33" pageId="121" pageNumber="122" type="diagnosis">
<paragraph id="922136E09D659222FF2EFA61C99F2D9B" blockId="121.[189,457,1503,1529]" box="[189,457,1503,1529]" pageId="121" pageNumber="122">
<heading id="C969818C9D659222FF2EFA61C99F2D9B" bold="true" box="[189,457,1503,1529]" fontSize="11" level="3" pageId="121" pageNumber="122" reason="3">
<emphasis id="A0EAEAF29D659222FF2EFA61C99F2D9B" bold="true" box="[189,457,1503,1529]" pageId="121" pageNumber="122">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D659222FF2EF9AECC912F33" blockId="121.[189,1399,1552,1873]" pageId="121" pageNumber="122">
This is a very distinctive species among all congeners primarily due to the strongly bigibbose frons, the flattened and enlarged lateral margins of the pronotum, the metatibia being smooth and glabrous on the inner surface but dense long setose on the external surface, the single long terminal spur of the metatibia, and the peculiar long fur-like setae. It is closest to
<taxonomicName id="559E4D639D659222FC9AF9C0CCF92EFA" authority="Arrow, 1941" authorityName="Arrow" authorityYear="1941" box="[777,1199,1662,1688]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="species" species="opatroides">
<emphasis id="A0EAEAF29D659222FC9AF9C0CC432EFA" box="[777,1045,1662,1688]" italics="true" pageId="121" pageNumber="122">Maechidius opatroides</emphasis>
<bibRefCitation id="F60F4B119D659222FB88F9C0CCF92EFA" author="Arrow G. E." box="[1051,1199,1662,1688]" pageId="121" pageNumber="122" pagination="448 - 464" refId="ref97558" refString="Arrow G. E. 1941. XXIX. - Some undescribed species of melolonthid Coleoptera from Western New Guinea and the adjacent islands of Waigeu and Japen. Annals and Magazine of Natural History, Series 11 7 (41): 448 - 464. https: // doi. org / 10.1080 / 03745481.1941.9727945" type="journal article" year="1941">Arrow, 1941</bibRefCitation>
</taxonomicName>
comb. rest. from Waigeo Island (consider the new combination above), but in this species (of which only females are known) the clustered brush-like setae are much more slender, appressed and shorter, the lateral margin of the pronotum is evenly emarginate postmedially (not angulate), the sutural carina of the elytron is not indicated, the metatibia is less wide, the anterolateral angles of the labroclypeus are obtuse angulate and not protruding anteriad and the labroclypeus is generally wider than in
<taxonomicName id="559E4D639D659222FC6DF889CC332F32" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1022,1125,1847,1872]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="species" species="ursus" status="sp. nov.">
<emphasis id="A0EAEAF29D659222FC6DF889CC332F32" box="[1022,1125,1847,1872]" italics="true" pageId="121" pageNumber="122">M. ursus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D659222FBFFF889CC912F33" box="[1132,1223,1847,1873]" pageId="121" pageNumber="122" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D659222FF2EF8C7CB5D2F8A" pageId="121" pageNumber="122" type="etymology">
<paragraph id="922136E09D659222FF2EF8C7C9122FF1" blockId="121.[189,324,1913,1939]" box="[189,324,1913,1939]" pageId="121" pageNumber="122">
<heading id="C969818C9D659222FF2EF8C7C9122FF1" bold="true" box="[189,324,1913,1939]" fontSize="11" level="3" pageId="121" pageNumber="122" reason="3">
<emphasis id="A0EAEAF29D659222FF2EF8C7C9122FF1" bold="true" box="[189,324,1913,1939]" pageId="121" pageNumber="122">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D659222FF2EF817CB5D2F8A" blockId="121.[189,1398,1961,2025]" pageId="121" pageNumber="122">
The specific name is derived from the Latin
<emphasis id="A0EAEAF29D659222FD28F814CB572FA1" box="[699,769,1962,1987]" italics="true" pageId="121" pageNumber="122">Ursus</emphasis>
(bear), indicating the robust body and peculiar, dense, mammal-fur-like pubescence. Noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D669221FF2EFEB7CA7629A3" pageId="122" pageNumber="123" type="materials_examined">
<paragraph id="922136E09D669221FF2EFEB7C93A2940" blockId="122.[189,364,264,291]" box="[189,364,264,291]" pageId="122" pageNumber="123">
<heading id="C969818C9D669221FF2EFEB7C93A2940" bold="true" box="[189,364,264,291]" fontSize="11" level="3" pageId="122" pageNumber="123" reason="3">
<emphasis id="A0EAEAF29D669221FF2EFEB7C93A2940" bold="true" box="[189,364,264,291]" pageId="122" pageNumber="123">Type material</emphasis>
</heading>
</paragraph>
<materialsCitation id="22F63CBD9D669221FF79FE87CA7629A3" ID-GBIF-Occurrence="2898878508" collectingDate="2009-03-28" collectionCode="NME" country="Indonesia" latitude="-1.9780556" location="River Ifeylo" longLatPrecision="21" longitude="129.92168" pageId="122" pageNumber="123" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<paragraph id="922136E09D669221FF79FE87C90A2931" blockId="122.[189,1399,313,449]" box="[234,348,313,339]" pageId="122" pageNumber="123">
<emphasis id="A0EAEAF29D669221FF79FE87C90A2931" bold="true" box="[234,348,313,339]" pageId="122" pageNumber="123">
<typeStatus id="4D2588429D669221FF79FE87C90A2931" box="[234,348,313,339]" pageId="122" pageNumber="123" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="922136E09D669221FF2EFEE3CA7629A3" blockId="122.[189,1399,313,449]" pageId="122" pageNumber="123">
<collectingCountry id="EA8976709D669221FF2EFEE3C90D2915" box="[189,347,349,375]" name="Indonesia" pageId="122" pageNumber="123">INDONESIA</collectingCountry>
•
<specimenCount id="8498FD699D669221FEE1FEE0C9DF291A" box="[370,393,350,376]" pageId="122" pageNumber="123" type="female">♀</specimenCount>
; “
<collectingCountry id="EA8976709D669221FE33FEE3CA172915" box="[416,577,349,375]" name="Indonesia" pageId="122" pageNumber="123">INDONESIA</collectingCountry>
E, Prov. Raja Ampat, Misool SW, distr. Misool Utara, Aduwey (Adua) vill. ~
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NNE, valley of
<location id="9741603B9D669221FE60FE3CCAD429FE" LSID="urn:lsid:plazi:treatment:1A3787F69D659220FDEEFAECCB9F2AF4:9741603B9D669221FE60FE3CCAD429FE" box="[499,642,386,412]" country="Indonesia" latitude="-1.9780556" longLatPrecision="21" longitude="129.92168" name="River Ifeylo" pageId="122" pageNumber="123">River Ifeylo</location>
,
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,
<geoCoordinate id="F7AA50279D669221FCB0FE3CCB9729FF" box="[803,961,386,413]" degrees="129" direction="east" minutes="55" orientation="longitude" pageId="122" pageNumber="123" precision="15" seconds="18" value="129.92168">129°55’18”E</geoCoordinate>
,
<date id="E62010209D669221FC58FE3CCC0429FE" box="[971,1106,386,412]" pageId="122" pageNumber="123" value="2009-03-28">
<collectingDate id="F664E9C89D669221FC58FE3CCC0429FE" box="[971,1106,386,412]" pageId="122" pageNumber="123" value="2009-03-28">28.III.2009</collectingDate>
</date>
, edge of primary lowland rainforest, white light”;
<collectionCode id="F48FAE259D669221FE44FE19CA4A29A3" box="[471,540,423,449]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="122" pageNumber="123">NME</collectionCode>
.
</paragraph>
</materialsCitation>
</subSubSection>
<subSubSection id="DA84656B9D669220FF2EFE57C92029D4" lastPageId="123" lastPageNumber="124" pageId="122" pageNumber="123" type="description">
<paragraph id="922136E09D669221FF2EFE57C9182A61" blockId="122.[189,334,489,515]" box="[189,334,489,515]" pageId="122" pageNumber="123">
<heading id="C969818C9D669221FF2EFE57C9182A61" bold="true" box="[189,334,489,515]" fontSize="11" level="3" pageId="122" pageNumber="123" reason="3">
<emphasis id="A0EAEAF29D669221FF2EFE57C9182A61" bold="true" box="[189,334,489,515]" pageId="122" pageNumber="123">Description</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D669221FF2EFDA4C97B2A1F" blockId="122.[189,1399,537,637]" pageId="122" pageNumber="123">
MEASUREMENTS.
<typeStatus id="4D2588429D669221FEEFFDA7C9BA2A51" box="[380,492,537,563]" pageId="122" pageNumber="123" type="holotype">Holotype</typeStatus>
, total body length
<quantity id="55669B059D669221FD54FDA7CB652A56" box="[711,819,537,564]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="122" pageNumber="123" unit="mm" value="8.6">8.60 mm</quantity>
. Head
<quantity id="55669B059D669221FC16FDA4CBB82A56" box="[901,1006,538,564]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="122" pageNumber="123" unit="mm" value="1.7">1.70 mm</quantity>
long, across eyes
<quantity id="55669B059D669221FB55FDA7CD792A56" box="[1222,1327,537,564]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" pageId="122" pageNumber="123" unit="mm" value="1.75">1.75 mm</quantity>
wide. Pronotum
<quantity id="55669B059D669221FEA4FD80C9C92A3B" box="[311,415,574,601]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="122" pageNumber="123" unit="mm" value="1.8">1.80 mm</quantity>
long, maximum width
<quantity id="55669B059D669221FD3EFD80CB412A3B" box="[685,791,574,601]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="122" pageNumber="123" unit="mm" value="3.2">3.20 mm</quantity>
. Elytral length
<quantity id="55669B059D669221FC54FD80CC642A3B" box="[967,1074,574,601]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="122" pageNumber="123" unit="mm" value="5.1">5.10 mm</quantity>
, maximum combined width
<quantity id="55669B059D669221FF2EFDDDC97F2A1F" box="[189,297,611,637]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="122" pageNumber="123" unit="mm" value="3.7">3.70 mm</quantity>
.
</paragraph>
<paragraph id="922136E09D669220FF2EFD18CD212925" blockId="122.[189,1399,678,2025]" lastBlockId="123.[189,1399,265,327]" lastPageId="123" lastPageNumber="124" pageId="122" pageNumber="123">
Dorsum and venter covered with microscopical velvety pubescence, uniformly dark brown with reddish brown mouthparts, antennae and legs. Head slightly transverse, opaque dorsally and ventrally except for glossy labroclypeus, vaguely impressed on labroclypeus anterior to each compound eye and with paired strong obtuse hump between eyes. Compound eye large, occupying more than half side of head. Female labroclypeus (
<figureCitation id="0AA52A659D669221FEFBFC86C9842B31" box="[360,466,824,851]" captionStart="Figs 181–192" captionStartId="144.[189,241,1787,1813]" captionTargetBox="[338,1249,274,1742]" captionTargetId="figure-122@144.[338,1249,265,1749]" captionTargetPageId="144" captionText="Figs 181–192. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 181. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 182. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 183. Same, ♀ (SNSD). 184. M. suwawa sp. nov., holotype, ♂ (BMNH). 185. Same, paratype, ♀ (BMNH). 186. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 187. M. trivialis sp. nov., holotype, ♂ (BMNH). 188. M. ursus sp. nov., holotype, ♀ (NME). 189. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 190. Same, ♂ (SNSD). 191. M. weigeli sp. nov., holotype, ♂ (NME). 192. Same, paratype, ♀ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122187" httpUri="https://zenodo.org/record/4122187/files/figure.png" pageId="122" pageNumber="123">Fig. 188</figureCitation>
) broadly and moderately deep emarginate anteriorly, lateral margins nearly straight in dorsal, slightly sinuous in lateral view, anterolateral angles moderately protruding, acute in dorsal view, bent up at ~90° to frons in lateral view. Canthus straight in dorsal view. Anterior and lateral margins of labroclypeus smooth. Head punctures circular, moderately deep, variably large. Intervening spaces microreticulate and glabrous, variably large. Conspicuous rather long appressed to suberect brush of clustered setae rises from each puncture (
<figureCitation id="0AA52A659D669221FD5CFC4ECB6E2C69" box="[719,824,1008,1035]" captionStart="Figs 349–358" captionStartId="156.[189,241,1786,1812]" captionTargetBox="[287,1299,265,1743]" captionTargetId="figure-123@156.[287,1301,265,1760]" captionTargetPageId="156" captionText="Figs 349–358. Papuan and Wallacean Maechidius Macleay, 1819, elytron (349–356) and pronotal setation (357–358), dorsal view. 349. M. suwawa sp. nov., paratype, ♀ (BMNH). 350. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 351. M. trivialis sp. nov., holotype, ♂ (BMNH). 352. M. ursus sp. nov., holotype, ♀ (NME). 353. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 354. M. weigeli sp. nov., holotype, ♂ (NME). 355. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 356. M. yamdena sp. nov., paratype, ♂ (BMNH). 357. M. leucopsar sp. nov., holotype, ♂ (IECA). 358. M. ursus sp. nov., holotype, ♀ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122214" httpUri="https://zenodo.org/record/4122214/files/figure.png" pageId="122" pageNumber="123">Fig. 358</figureCitation>
). Antenna 9-segmented, club 3-lamellate. Scape large, with bulbous predistal projection on dorsal side, provided with two very long setae near distal margin. Antennomere 2 bulbous, subspherical, slightly longer than wide. Pronotum opaque dorsally and laterally. Anterior margin of pronotum slightly sinuous with anterolateral angles (stronger) and mesal portion (inconspicuously) protruding anteriad. Basal margin of pronotum sinuous. Circular moderately deep impression on either side of pronotum opposite posterolateral angle. Lateral margin of pronotum strongly flattened and expanded laterally, bent up with regard to axis of pronotal disc, evenly widened towards midlength, strongly emarginate postmedially towards nearly posterolateral angles (
<figureCitation id="0AA52A659D669221FA93FB4FCD3F2D6E" box="[1280,1385,1265,1292]" captionStart="Figs 259–265" captionStartId="149.[189,241,1372,1398]" captionTargetBox="[189,1379,265,1308]" captionTargetId="figure-88@149.[189,1398,265,1311]" captionTargetPageId="149" captionText="Figs 259–265. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 259. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 260. M. trivialis sp. nov., holotype, ♂ (BMNH). 261. M. ursus sp. nov., holotype, ♀ (NME). 262. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 263. M. weigeli sp. nov., holotype, ♂ (NME). 264. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 265. M. yamdena sp. nov., paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122198" httpUri="https://zenodo.org/record/4122198/files/figure.png" pageId="122" pageNumber="123">Fig. 261</figureCitation>
). Crenulae of lateral margin large, long erect scale-like seta present between every two crenulae. Lateral margin of pronotum straight in lateral view. Pronotal punctures circular, moderately large and deep. Intervening spaces microreticulate and glabrous, generally larger than punctures, in part covered with microscopical velvety pubescence. Setae as conspicuous as on head (
<figureCitation id="0AA52A659D669221FC60FA3DCC0D2DFC" box="[1011,1115,1411,1438]" captionStart="Figs 349–358" captionStartId="156.[189,241,1786,1812]" captionTargetBox="[287,1299,265,1743]" captionTargetId="figure-123@156.[287,1301,265,1760]" captionTargetPageId="156" captionText="Figs 349–358. Papuan and Wallacean Maechidius Macleay, 1819, elytron (349–356) and pronotal setation (357–358), dorsal view. 349. M. suwawa sp. nov., paratype, ♀ (BMNH). 350. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 351. M. trivialis sp. nov., holotype, ♂ (BMNH). 352. M. ursus sp. nov., holotype, ♀ (NME). 353. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 354. M. weigeli sp. nov., holotype, ♂ (NME). 355. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 356. M. yamdena sp. nov., paratype, ♂ (BMNH). 357. M. leucopsar sp. nov., holotype, ♂ (IECA). 358. M. ursus sp. nov., holotype, ♀ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122214" httpUri="https://zenodo.org/record/4122214/files/figure.png" pageId="122" pageNumber="123">Fig. 358</figureCitation>
), shorter and sparser on disc, much longer and denser on anterior part and along lateral sides. Hypomeron slightly emarginate and very long setose on anterior margin, which is well-projected and flange-like.Antennal pocket moderately deep. Median anterior process of prosternum long, narrow, flange-like raised. Scutellar shield obtusely pointed apically. Elytra slightly widened in median part, maximum width across median third, opaque dorsally, with distinct humeri.Distinct obtuse triangular hump at apex of each elytron.Two flat longitudinal carinae on each elytron. Sutural carinae well developed, raised, nearly complete. Elytral disc with rather large deep cylindrical punctures which are irregular in apical half, becoming arranged in striae in apical third (
<figureCitation id="0AA52A659D669221FE94F917C9242EA6" box="[263,370,1705,1732]" captionStart="Figs 349–358" captionStartId="156.[189,241,1786,1812]" captionTargetBox="[287,1299,265,1743]" captionTargetId="figure-123@156.[287,1301,265,1760]" captionTargetPageId="156" captionText="Figs 349–358. Papuan and Wallacean Maechidius Macleay, 1819, elytron (349–356) and pronotal setation (357–358), dorsal view. 349. M. suwawa sp. nov., paratype, ♀ (BMNH). 350. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 351. M. trivialis sp. nov., holotype, ♂ (BMNH). 352. M. ursus sp. nov., holotype, ♀ (NME). 353. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 354. M. weigeli sp. nov., holotype, ♂ (NME). 355. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 356. M. yamdena sp. nov., paratype, ♂ (BMNH). 357. M. leucopsar sp. nov., holotype, ♂ (IECA). 358. M. ursus sp. nov., holotype, ♀ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122214" httpUri="https://zenodo.org/record/4122214/files/figure.png" pageId="122" pageNumber="123">Fig. 352</figureCitation>
). Intervening spaces microreticulate, covered with microscopical velvety pubescence, variably large but generally about as large as punctures. Setae inconspicuous, short and sparse, suberect seta rises from anterior part of each puncture. Epipleuron all along with conspicuous short brush-like setae. At apical margin with transverse brush of dense posteriad-directed brush-like setae, elytral apices therefore look flattened and slightly prolonged (
<figureCitation id="0AA52A659D669221FD74F885CB6B2F34" box="[743,829,1851,1878]" captionStart="Figs 90–93" captionStartId="119.[189,241,1928,1954]" captionTargetBox="[329,1264,264,1883]" captionTargetId="figure-60@119.[321,1264,256,1905]" captionTargetPageId="119" captionText="Figs 90–93. Papuan Maechidius Macleay, 1819, habitus, dorsal view. 90. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 91. M. trivialis sp. nov., holotype, ♂ (BMNH). 92. M. ursus sp. nov., holotype, ♀ (NME). 93. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122169" httpUri="https://zenodo.org/record/4122169/files/figure.png" pageId="122" pageNumber="123">Fig. 92</figureCitation>
). Female pygidium dorsally impressed, regularly circular punctured, antero-medially with a group of dense brush-like setae (Fig. 528). Venter covered with sparse shallow circular punctures, each provided with appressed conspicuous brush-like seta. Legs long and slender. Protibia nearly straight (crenulate) on external margin, with delicate incomplete dorsal carina. Female protibia with obtuse distal tooth and rudimentary predistal one (
<figureCitation id="0AA52A659D669221FB1EF870CCAC2F8A" box="[1165,1274,1998,2024]" captionStart="Figs 424–435" captionStartId="161.[189,241,1394,1420]" captionTargetBox="[238,1349,265,1318]" captionTargetId="figure-136@161.[217,1354,265,1322]" captionTargetPageId="161" captionText="Figs 424–435. Papuan and Wallacean Maechidius Macleay, 1819, protibia, dorsal view (424–428), and abdominal ventrites (429–435), ventral and lateral views. 424. M. ursus sp. nov., holotype, ♀ (NME). 425. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 426. M. weigeli sp. nov., holotype, ♂ (NME). 427. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 428. M. yamdena sp. nov., paratype, ♂ (BMNH). 429. M.dani sp. nov., holotype, ♂ (IECA). 430. Same, different aspect (IECA). 431. M. hamatus sp. nov., holotype, ♂ (IECA). 432. Same, different aspect (IECA). 433. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 434. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 435. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122216" httpUri="https://zenodo.org/record/4122216/files/figure.png" pageId="122" pageNumber="123">Fig. 424</figureCitation>
). Protibial terminal spur long, nearly straight, obtuse. Female metatibia flattened and glabrous on inner margin (
<figureCitation id="0AA52A659D679220FF55FE92C97B2924" box="[198,301,300,326]" captionStart="Figs 436–450" captionStartId="162.[189,241,1619,1645]" captionTargetBox="[205,1349,265,1554]" captionTargetId="figure-184@162.[205,1377,265,1557]" captionTargetPageId="162" captionText="Figs 436–450. Papuan and Wallacean Maechidius Macleay, 1819, legs, various views. 436. M. hirtipes Arrow, 1941, lectotype, ♂, basal metatarsomere (BMNH). 437. M. tarsalis Arrow, 1941, lectotype, ♂, basal metatarsomere (BMNH). 438. M. babyrousa sp. nov., paratype, ♀, metatibial terminal spurs (BMNH). 439. Same, paratype, ♂ (BMNH). 440. Same, different aspect (BMNH). 441. M. yamdena sp. nov., paratype, ♀, metatibial terminal spurs (BMNH). 442. Same, paratype, ♂ (BMNH). 443. M. longipes sp. nov., holotype, ♂, metatibial terminal spurs (BMNH). 444. M. ursus sp. nov., holotype, ♀, left metatibia and basal metatarsomere (NME). 445. M. dani sp. nov., holotype, ♂, apical portion of metatibia (IECA). 446. M. skalei sp. nov., holotype, ♂, apical portion of metatibia and basal metatarsomere (NME). 447. M. yamdena sp. nov., paratype, ♂, protibial terminal spur (BMNH). 448. Same, paratype, ♀ (BMNH). 449. M. babyrousa sp. nov., paratype, ♀, protibial terminal spur (BMNH). 450. Same, paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122218" httpUri="https://zenodo.org/record/4122218/files/figure.png" pageId="123" pageNumber="124">Fig. 444</figureCitation>
). Metatibial terminal spur single, long, obtuse. Tarsal claws without pulvilli (in female only?).
</paragraph>
<paragraph id="922136E09D679220FF2EFED3C9E629E5" blockId="123.[189,432,365,391]" box="[189,432,365,391]" pageId="123" pageNumber="124">
<heading id="C969818C9D679220FF2EFED3C9E629E5" bold="true" box="[189,432,365,391]" fontSize="11" level="3" pageId="123" pageNumber="124" reason="3">
<emphasis id="A0EAEAF29D679220FF2EFED3C9E629E5" bold="true" box="[189,432,365,391]" pageId="123" pageNumber="124">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D679220FF2EFE22C92029D4" blockId="123.[189,374,412,438]" box="[189,374,412,438]" pageId="123" pageNumber="124">Male unknown.</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D679220FF2EFE60CC222A44" box="[189,1140,478,551]" pageId="123" pageNumber="124" type="biology_ecology">
<paragraph id="922136E09D679220FF2EFE60C976299A" blockId="123.[189,288,478,504]" box="[189,288,478,504]" pageId="123" pageNumber="124">
<heading id="C969818C9D679220FF2EFE60C976299A" bold="true" box="[189,288,478,504]" fontSize="11" level="3" pageId="123" pageNumber="124" reason="3">
<emphasis id="A0EAEAF29D679220FF2EFE60C976299A" bold="true" box="[189,288,478,504]" pageId="123" pageNumber="124">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D679220FF2EFDB2CC222A44" blockId="123.[189,1140,524,551]" box="[189,1140,524,551]" pageId="123" pageNumber="124">Attracted to white light in river valley surrounded by primary lowland rainforest.</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D679220FF2EFDF0CB9F2AF4" box="[189,969,590,663]" pageId="123" pageNumber="124" type="distribution">
<paragraph id="922136E09D679220FF2EFDF0C9002A0A" blockId="123.[189,342,590,616]" box="[189,342,590,616]" pageId="123" pageNumber="124">
<heading id="C969818C9D679220FF2EFDF0C9002A0A" bold="true" box="[189,342,590,616]" fontSize="11" level="3" pageId="123" pageNumber="124" reason="3">
<emphasis id="A0EAEAF29D679220FF2EFDF0C9002A0A" bold="true" box="[189,342,590,616]" pageId="123" pageNumber="124">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D679220FF2EFDC2CB9F2AF4" blockId="123.[189,969,636,663]" box="[189,969,636,663]" pageId="123" pageNumber="124">
Hitherto only known from Misool, Raja Ampat Islands,
<collectingCountry id="EA8976709D679220FCDCFDC2CB932AF4" box="[847,965,636,662]" name="Indonesia" pageId="123" pageNumber="124">Indonesia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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