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<document id="9EF1F5BB808EE29462B507092305B9BA" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D6392DBFDBAFEB6CAF729D4" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius woodlarkianus Heller 1914" docType="treatment" docVersion="10" lastPageNumber="129" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="128" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="4A059C1FADDCE31F3E74BE8A44A558BB">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
</mods:titleInfo>
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<mods:namePart id="2637CE7E940F8A0ABD4A5F8B26214CA6">Telnov, Dmitry</mods:namePart>
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<mods:title id="9DEB52927F8D776351A6192EB77F0B87">European Journal of Taxonomy</mods:title>
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<mods:date id="B75CCED1B346AFA2D8B9AFFEB9277744">2020</mods:date>
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<mods:number id="3FC5BFBA0D9BBDF48CE589ED01C9D80F">2020-10-19</mods:number>
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<treatment id="1A3787F69D6392DBFDBAFEB6CAF729D4" ID-DOI="http://doi.org/10.5281/zenodo.4344471" ID-GBIF-Taxon="168648173" ID-Zenodo-Dep="4344471" LSID="urn:lsid:plazi:treatment:1A3787F69D6392DBFDBAFEB6CAF729D4" httpUri="http://treatment.plazi.org/id/1A3787F69D6392DBFDBAFEB6CAF729D4" lastPageId="128" lastPageNumber="129" pageId="127" pageNumber="128">
<subSubSection id="DA84656B9D639224FDBAFEB6CC5D2941" box="[553,1035,264,291]" pageId="127" pageNumber="128" type="nomenclature">
<paragraph id="922136E09D639224FDBAFEB6CC5D2941" blockId="127.[488,1100,264,327]" box="[553,1035,264,291]" pageId="127" pageNumber="128">
<heading id="C969818C9D639224FDBAFEB6CC5D2941" box="[553,1035,264,291]" centered="true" fontSize="11" level="2" pageId="127" pageNumber="128" reason="2">
<taxonomicName id="559E4D639D639224FDBAFEB6CC5D2941" ID-CoL="3XFLW" authority="Heller, 1914" authorityName="Heller" authorityYear="1914" box="[553,1035,264,291]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="127" pageNumber="128" phylum="Arthropoda" rank="species" species="woodlarkianus">
<emphasis id="A0EAEAF29D639224FDBAFEB6CB272941" bold="true" box="[553,881,264,291]" italics="true" pageId="127" pageNumber="128">Maechidius woodlarkianus</emphasis>
<bibRefCitation id="F60F4B119D639224FCEAFEB6CC5D2941" author="Heller K. M." box="[889,1035,264,291]" pageId="127" pageNumber="128" pagination="615 - 666" refId="ref98213" refString="Heller K. M. 1914. Coleoptera. Nova Guinea. Resultats de l'Expedition scientifique neerlandaise a la Nouvelle-Guinee en 1907 et 1909 sous les auspices du Dr. H. A. Lorentz. Zoologie 9: 615 - 666." type="journal article" year="1914">Heller, 1914</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D639224FE7BFE92CC1A2924" box="[488,1100,300,327]" pageId="127" pageNumber="128" type="description">
<paragraph id="922136E09D639224FE7BFE92CC1A2924" blockId="127.[488,1100,264,327]" box="[488,1100,300,327]" pageId="127" pageNumber="128">
<figureCitation id="0AA52A659D639224FE7BFE92CA682925" box="[488,574,300,327]" captionStart="Figs 94–96" captionStartId="126.[189,241,1928,1954]" captionTargetBox="[346,1191,267,1904]" captionTargetId="figure-55@126.[339,1249,265,1909]" captionTargetPageId="126" captionText="Figs 94–96. Papuan and Wallacean Maechidius Macleay, 1819, habitus, dorsal view. 94. M. weigeli sp. nov., holotype, ♂ (NME). 95. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 96. M. yamdena sp. nov., paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122171" httpUri="https://zenodo.org/record/4122171/files/figure.png" pageId="127" pageNumber="128">Fig. 95</figureCitation>
,
<figureCitation id="0AA52A659D639224FDD9FE92CAE52924" box="[586,691,300,326]" captionStart="Figs 193–207" captionStartId="145.[189,241,1716,1742]" captionTargetBox="[385,1203,267,1681]" captionTargetId="figure-159@145.[385,1203,265,1694]" captionTargetPageId="145" captionText="Figs 193–207. Papuan and Wallacean Maechidius Macleay, 1819, head (193–196) and pronotum (197– 207), dorsal view. 193. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 194. Same, ♂ (SNSD). 195. M. yamdena sp. nov., paratype, ♂ (BMNH). 196. Same, paratype, ♀ (BMNH). 197. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 198. M. aiyura sp. nov., holotype, ♂ (BMNH). 199. M. alesbezdeki sp. nov., holotype, ♂ (NME). 200. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 201. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 202. M. awu sp. nov., holotype, ♂ (IECA). 203. M. babyrousa sp. nov., holotype, ♂ (BMNH). 204. M. bintang sp. nov., holotype, ♂ (NME). 205. M. boessnecki sp. nov., holotype, ♂ (SNSD). 206. M. brocki sp. nov., holotype, ♂ (BMNH). 207. M. crypticus sp. nov., paratype, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122189" httpUri="https://zenodo.org/record/4122189/files/figure.png" pageId="127" pageNumber="128">193–194</figureCitation>
,
<figureCitation id="0AA52A659D639224FD2CFE92CAB82924" box="[703,750,300,326]" captionStart="Figs 259–265" captionStartId="149.[189,241,1372,1398]" captionTargetBox="[189,1379,265,1308]" captionTargetId="figure-88@149.[189,1398,265,1311]" captionTargetPageId="149" captionText="Figs 259–265. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 259. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 260. M. trivialis sp. nov., holotype, ♂ (BMNH). 261. M. ursus sp. nov., holotype, ♀ (NME). 262. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 263. M. weigeli sp. nov., holotype, ♂ (NME). 264. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 265. M. yamdena sp. nov., paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122198" httpUri="https://zenodo.org/record/4122198/files/figure.png" pageId="127" pageNumber="128">264</figureCitation>
,
<figureCitation id="0AA52A659D639224FD69FE92CB7F2925" box="[762,809,300,327]" captionStart="Figs 278–285" captionStartId="151.[189,241,1697,1723]" captionTargetBox="[211,1375,265,1597]" captionTargetId="figure-104@151.[211,1376,265,1614]" captionTargetPageId="151" captionText="Figs 278–285. Papuan and Wallacean Maechidius Macleay, 1819, pronotal hypomeron, ventro-lateral view. 278. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 279. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 280. M. seriepunctatus Moser, 1920, lectotype, ♂ (SNSD). 281. M. speciosus (Frey, 1969) comb. nov., ♀ (BMNH). 282. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 283. M. trivialis sp. nov., holotype, ♂ (BMNH). 284. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 285. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122202" httpUri="https://zenodo.org/record/4122202/files/figure.png" pageId="127" pageNumber="128">285</figureCitation>
,
<figureCitation id="0AA52A659D639224FCA6FE92CB322925" box="[821,868,300,327]" captionStart="Figs 349–358" captionStartId="156.[189,241,1786,1812]" captionTargetBox="[287,1299,265,1743]" captionTargetId="figure-123@156.[287,1301,265,1760]" captionTargetPageId="156" captionText="Figs 349–358. Papuan and Wallacean Maechidius Macleay, 1819, elytron (349–356) and pronotal setation (357–358), dorsal view. 349. M. suwawa sp. nov., paratype, ♀ (BMNH). 350. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 351. M. trivialis sp. nov., holotype, ♂ (BMNH). 352. M. ursus sp. nov., holotype, ♀ (NME). 353. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 354. M. weigeli sp. nov., holotype, ♂ (NME). 355. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 356. M. yamdena sp. nov., paratype, ♂ (BMNH). 357. M. leucopsar sp. nov., holotype, ♂ (IECA). 358. M. ursus sp. nov., holotype, ♀ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122214" httpUri="https://zenodo.org/record/4122214/files/figure.png" pageId="127" pageNumber="128">355</figureCitation>
,
<figureCitation id="0AA52A659D639224FCFCFE92CBC82924" box="[879,926,300,326]" captionStart="Figs 424–435" captionStartId="161.[189,241,1394,1420]" captionTargetBox="[238,1349,265,1318]" captionTargetId="figure-136@161.[217,1354,265,1322]" captionTargetPageId="161" captionText="Figs 424–435. Papuan and Wallacean Maechidius Macleay, 1819, protibia, dorsal view (424–428), and abdominal ventrites (429–435), ventral and lateral views. 424. M. ursus sp. nov., holotype, ♀ (NME). 425. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 426. M. weigeli sp. nov., holotype, ♂ (NME). 427. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 428. M. yamdena sp. nov., paratype, ♂ (BMNH). 429. M.dani sp. nov., holotype, ♂ (IECA). 430. Same, different aspect (IECA). 431. M. hamatus sp. nov., holotype, ♂ (IECA). 432. Same, different aspect (IECA). 433. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 434. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 435. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122216" httpUri="https://zenodo.org/record/4122216/files/figure.png" pageId="127" pageNumber="128">427</figureCitation>
, 533, 742–744
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D639224FF2EFED0CC4A2AF6" pageId="127" pageNumber="128" type="materials_examined">
<paragraph id="922136E09D639224FF2EFED0C93A29EA" blockId="127.[189,364,366,392]" box="[189,364,366,392]" pageId="127" pageNumber="128">
<heading id="C969818C9D639224FF2EFED0C93A29EA" bold="true" box="[189,364,366,392]" fontSize="11" level="3" pageId="127" pageNumber="128" reason="3">
<emphasis id="A0EAEAF29D639224FF2EFED0C93A29EA" bold="true" box="[189,364,366,392]" pageId="127" pageNumber="128">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D639224FF79FE23CA3A29DA" blockId="127.[189,1398,413,512]" box="[234,620,413,440]" pageId="127" pageNumber="128">
<emphasis id="A0EAEAF29D639224FF79FE23C93029D5" bold="true" box="[234,358,413,439]" pageId="127" pageNumber="128">
<typeStatus id="4D2588429D639224FF79FE23C93029D5" box="[234,358,413,439]" pageId="127" pageNumber="128" type="lectotype">Lectotype</typeStatus>
</emphasis>
[herewith designated]
</paragraph>
<paragraph id="922136E09D639224FF2EFE7FCD27299D" blockId="127.[189,1398,413,512]" pageId="127" pageNumber="128">
<materialsCitation id="22F63CBD9D639224FF2EFE7FCD27299D" ID-GBIF-Occurrence="2898878426" collectionCode="SNSD" collectorName="Gehr. W. Muller Vermacht. &amp; A. S. Meek" country="Papua New Guinea" location="PAPUA NEW GUINEA" pageId="127" pageNumber="128" specimenCount="1" specimenCount-female="1" typeStatus="lectotype">
<collectingCountry id="EA8976709D639224FF2EFE7FC98B29B9" box="[189,477,449,476]" name="Papua New Guinea" pageId="127" pageNumber="128">PAPUA NEW GUINEA</collectingCountry>
•
<specimenCount id="8498FD699D639224FE69FE7CCA4729BE" box="[506,529,450,476]" pageId="127" pageNumber="128" type="female">♀</specimenCount>
; “
<collectorName id="3F6B53369D639224FDB8FE7FCB2329B9" box="[555,885,449,476]" pageId="127" pageNumber="128">Gehr. W. Müller Vermächt.</collectorName>
1909 [p, blue label] //Woodlark, III.
<date id="E62010209D639224FAB5FE7CCD2529B9" box="[1318,1395,449,476]" pageId="127" pageNumber="128" value="1997-04">IV. 97</date>
. (
<collectorName id="3F6B53369D639224FF55FE5BC9042A62" box="[198,338,485,512]" pageId="127" pageNumber="128">A. S. Meek</collectorName>
). [p] //
<typeStatus id="4D2588429D639224FE0DFE5BC9BA299D" box="[414,492,485,511]" pageId="127" pageNumber="128">Typus</typeStatus>
! [p]
<taxonomicName id="559E4D639D639224FDB3FE5BCA98299D" authorityName="Heller" authorityYear="1914" box="[544,718,485,511]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="127" pageNumber="128" phylum="Arthropoda" rank="species" species="woodlarkianus">
<emphasis id="A0EAEAF29D639224FDB3FE5BCA98299D" box="[544,718,485,511]" italics="true" pageId="127" pageNumber="128">woodlarkianus</emphasis>
</taxonomicName>
[h] // Staatl. Museum für Tierkunde Dresden [p]”;
<collectionCode id="F48FAE259D639224FA8CFE5BCD38299D" box="[1311,1390,485,511]" collectionName="SNSD" pageId="127" pageNumber="128">SNSD</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="922136E09D639224FF2EFD99C9302A23" blockId="127.[189,358,551,577]" box="[189,358,551,577]" pageId="127" pageNumber="128">
<heading id="C969818C9D639224FF2EFD99C9302A23" bold="true" box="[189,358,551,577]" fontSize="11" level="3" pageId="127" pageNumber="128" reason="3">
<emphasis id="A0EAEAF29D639224FF2EFD99C9302A23" bold="true" box="[189,358,551,577]" pageId="127" pageNumber="128">New material</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D639224FF2EFDE8CC4A2AF6" blockId="127.[189,1399,597,660]" pageId="127" pageNumber="128">
<materialsCitation id="22F63CBD9D639224FF2EFDE8CC4A2AF6" ID-GBIF-Occurrence="2898878395" pageId="127" pageNumber="128">
<collectingCountry id="EA8976709D639224FF2EFDE8C98D2A12" box="[189,475,598,624]" name="Papua New Guinea" pageId="127" pageNumber="128">PAPUA NEW GUINEA</collectingCountry>
•
<specimenCount id="8498FD699D639224FE64FDE8CA6A2A12" box="[503,572,598,624]" pageId="127" pageNumber="128" type="male">4 ♂♂</specimenCount>
; “Woodlark, III.
<date id="E62010209D639224FC9BFDE8CB022A12" box="[776,852,598,624]" pageId="127" pageNumber="128" value="1997-04">IV. 97</date>
. (A. S. Meek). [p] // Coll. C. Felsche Kauf 20, 1918 [p, blue label]// Staatl. Museum für Tierkunde Dresden [p]”;
<collectionCode id="F48FAE259D639224FC5AFDC4CC4E2AF6" box="[969,1048,634,660]" collectionName="SNSD" pageId="127" pageNumber="128">SNSD</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D639224FF2EFD02CD252BBE" pageId="127" pageNumber="128" type="discussion">
<paragraph id="922136E09D639224FF2EFD02C9792AB4" blockId="127.[189,303,700,726]" box="[189,303,700,726]" pageId="127" pageNumber="128">
<heading id="C969818C9D639224FF2EFD02C9792AB4" bold="true" box="[189,303,700,726]" fontSize="11" level="3" pageId="127" pageNumber="128" reason="3">
<emphasis id="A0EAEAF29D639224FF2EFD02C9792AB4" bold="true" box="[189,303,700,726]" pageId="127" pageNumber="128">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D639224FF2EFD55CD252BBE" blockId="127.[189,1399,747,988]" pageId="127" pageNumber="128">
<bibRefCitation id="F60F4B119D639224FF2EFD55C9CA2B64" author="Heller K. M." box="[189,412,747,774]" pageId="127" pageNumber="128" pagination="615 - 666" refId="ref98213" refString="Heller K. M. 1914. Coleoptera. Nova Guinea. Resultats de l'Expedition scientifique neerlandaise a la Nouvelle-Guinee en 1907 et 1909 sous les auspices du Dr. H. A. Lorentz. Zoologie 9: 615 - 666." type="journal article" year="1914">Heller (1914: 628)</bibRefCitation>
likely based his description of
<taxonomicName id="559E4D639D639224FC8DFD52CBA32B67" authorityName="Heller" authorityYear="1914" box="[798,1013,747,773]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="127" pageNumber="128" phylum="Arthropoda" rank="species" species="woodlarkianus">
<emphasis id="A0EAEAF29D639224FC8DFD52CBA32B67" box="[798,1013,747,773]" italics="true" pageId="127" pageNumber="128">M. woodlarkianus</emphasis>
</taxonomicName>
on a single specimen, although not explicitly stated. The single SNSD specimen labelled “Typus♂” is herewith designated as
<typeStatus id="4D2588429D639224FA9FFCB0CD202B4A" box="[1292,1398,782,808]" pageId="127" pageNumber="128" type="lectotype">lectotype</typeStatus>
in order to enhance the stability of nomenclature and fix the specimen I based my redescription on, in case other (non-conspecific)
<typeStatus id="4D2588429D639224FDA6FCE9CACC2B13" box="[565,666,855,881]" pageId="127" pageNumber="128" type="syntype">syntypes</typeStatus>
are discovered in the future. Four additional specimens with identical collecting data labels are allocated in SNSD (see Checklist). However, these specimens are not considered
<typeStatus id="4D2588429D639224FED2FC20C9FF2BDA" box="[321,425,926,952]" pageId="127" pageNumber="128" type="syntype">syntypes</typeStatus>
, because the original description of
<taxonomicName id="559E4D639D639224FCD0FC20CC422BD5" authorityName="Heller" authorityYear="1914" box="[835,1044,925,951]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="127" pageNumber="128" phylum="Arthropoda" rank="species" species="woodlarkianus">
<emphasis id="A0EAEAF29D639224FCD0FC20CC422BD5" box="[835,1044,925,951]" italics="true" pageId="127" pageNumber="128">M. woodlarkianus</emphasis>
</taxonomicName>
was published in 1914, but the additional
<specimenCount id="8498FD699D639224FEABFC7FC9912BB9" box="[312,455,961,987]" pageId="127" pageNumber="128" type="generic">4 specimens</specimenCount>
in question were acquired by SNSD in 1918, as their labels state (see Checklist).
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D6392DBFF2EFBBDCAF729D4" lastPageId="128" lastPageNumber="129" pageId="127" pageNumber="128" type="description">
<paragraph id="922136E09D639224FF2EFBBDC9182C7F" blockId="127.[189,334,1027,1053]" box="[189,334,1027,1053]" pageId="127" pageNumber="128">
<heading id="C969818C9D639224FF2EFBBDC9182C7F" bold="true" box="[189,334,1027,1053]" fontSize="11" level="3" pageId="127" pageNumber="128" reason="3">
<emphasis id="A0EAEAF29D639224FF2EFBBDC9182C7F" bold="true" box="[189,334,1027,1053]" pageId="127" pageNumber="128">Description</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D639224FF2EFB8DCBB42C2F" blockId="127.[189,994,1074,1101]" box="[189,994,1074,1101]" pageId="127" pageNumber="128">
MEASUREMENTS. Total body length 7.10–7.20 (
<typeStatus id="4D2588429D639224FD46FB8CCB122C2F" box="[725,836,1074,1101]" pageId="127" pageNumber="128" type="lectotype">lectotype</typeStatus>
) to
<quantity id="55669B059D639224FCE1FB8CCB8B2C2F" box="[882,989,1074,1101]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="127" pageNumber="128" unit="mm" value="7.6">7.60 mm</quantity>
.
</paragraph>
<paragraph id="922136E09D6392DBFF2EFBCBCC942924" blockId="127.[189,1399,1140,2025]" lastBlockId="128.[189,1398,265,327]" lastPageId="128" lastPageNumber="129" pageId="127" pageNumber="128">
Dorsum and venter uniformly brown, appendages and abdomen paler brown. Head flattened dorsally between compound eyes, glossy dorsally and ventrally. Male labroclypeus (
<figureCitation id="0AA52A659D639224FBAAFB26CCC82CD0" box="[1081,1182,1176,1202]" captionStart="Figs 193–207" captionStartId="145.[189,241,1716,1742]" captionTargetBox="[385,1203,267,1681]" captionTargetId="figure-159@145.[385,1203,265,1694]" captionTargetPageId="145" captionText="Figs 193–207. Papuan and Wallacean Maechidius Macleay, 1819, head (193–196) and pronotum (197– 207), dorsal view. 193. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 194. Same, ♂ (SNSD). 195. M. yamdena sp. nov., paratype, ♂ (BMNH). 196. Same, paratype, ♀ (BMNH). 197. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 198. M. aiyura sp. nov., holotype, ♂ (BMNH). 199. M. alesbezdeki sp. nov., holotype, ♂ (NME). 200. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 201. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 202. M. awu sp. nov., holotype, ♂ (IECA). 203. M. babyrousa sp. nov., holotype, ♂ (BMNH). 204. M. bintang sp. nov., holotype, ♂ (NME). 205. M. boessnecki sp. nov., holotype, ♂ (SNSD). 206. M. brocki sp. nov., holotype, ♂ (BMNH). 207. M. crypticus sp. nov., paratype, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122189" httpUri="https://zenodo.org/record/4122189/files/figure.png" pageId="127" pageNumber="128">Fig. 194</figureCitation>
) very broadly and shallowly emarginate anteriorly, its lateral margins nearly straight to slightly sinuous in dorsal, sinuous in lateral view, anterolateral angles of labroclypeus not projecting anteriad, raised up at angle of nearly 70–80° to frons in lateral view. Female labroclypeus (
<figureCitation id="0AA52A659D639224FCDBFABDCBF92D7F" box="[840,943,1283,1309]" captionStart="Figs 193–207" captionStartId="145.[189,241,1716,1742]" captionTargetBox="[385,1203,267,1681]" captionTargetId="figure-159@145.[385,1203,265,1694]" captionTargetPageId="145" captionText="Figs 193–207. Papuan and Wallacean Maechidius Macleay, 1819, head (193–196) and pronotum (197– 207), dorsal view. 193. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 194. Same, ♂ (SNSD). 195. M. yamdena sp. nov., paratype, ♂ (BMNH). 196. Same, paratype, ♀ (BMNH). 197. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 198. M. aiyura sp. nov., holotype, ♂ (BMNH). 199. M. alesbezdeki sp. nov., holotype, ♂ (NME). 200. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 201. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 202. M. awu sp. nov., holotype, ♂ (IECA). 203. M. babyrousa sp. nov., holotype, ♂ (BMNH). 204. M. bintang sp. nov., holotype, ♂ (NME). 205. M. boessnecki sp. nov., holotype, ♂ (SNSD). 206. M. brocki sp. nov., holotype, ♂ (BMNH). 207. M. crypticus sp. nov., paratype, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122189" httpUri="https://zenodo.org/record/4122189/files/figure.png" pageId="127" pageNumber="128">Fig. 193</figureCitation>
) similar to that of male. Anterior and lateral margins of labroclypeus smooth. Upper- and underside of labroclypeus with sparse inconspicuous setae along anterior and lateral margins. Canthus very broadly rounded in dorsal view. Punctures of frons ovoid, moderately deep and large. Intervening spaces glossy, variably large, in part wrinkled. Inconspicuous appressed seta rises from anterior margin of each puncture, not or hardly surpassing length of corresponding puncture. Antenna 9-segmented, club 3-lamellate. Pronotum transverse, flattened dorsally, glossy dorsally and laterally. Anterior margin of pronotum broadly emarginate with anterolateral angles slightly protruding anteriad. Basal margin of pronotum broadly rounded. Lateral margin or pronotum broadly rounded, crenulate all along (
<figureCitation id="0AA52A659D639224FCCCF99FCB902E5E" box="[863,966,1569,1596]" captionStart="Figs 259–265" captionStartId="149.[189,241,1372,1398]" captionTargetBox="[189,1379,265,1308]" captionTargetId="figure-88@149.[189,1398,265,1311]" captionTargetPageId="149" captionText="Figs 259–265. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 259. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 260. M. trivialis sp. nov., holotype, ♂ (BMNH). 261. M. ursus sp. nov., holotype, ♀ (NME). 262. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 263. M. weigeli sp. nov., holotype, ♂ (NME). 264. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 265. M. yamdena sp. nov., paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122198" httpUri="https://zenodo.org/record/4122198/files/figure.png" pageId="127" pageNumber="128">Fig. 264</figureCitation>
). Short suberect seta present between every two crenulae. Lateral margin of pronotum slightly arched in lateral view. Hypomeron separated from prosternum by low nearly straight carina, with long setae on anterolateral margin opposite to compound eye (
<figureCitation id="0AA52A659D639224FEE9F932C9B72EC5" box="[378,481,1676,1703]" captionStart="Figs 278–285" captionStartId="151.[189,241,1697,1723]" captionTargetBox="[211,1375,265,1597]" captionTargetId="figure-104@151.[211,1376,265,1614]" captionTargetPageId="151" captionText="Figs 278–285. Papuan and Wallacean Maechidius Macleay, 1819, pronotal hypomeron, ventro-lateral view. 278. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 279. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 280. M. seriepunctatus Moser, 1920, lectotype, ♂ (SNSD). 281. M. speciosus (Frey, 1969) comb. nov., ♀ (BMNH). 282. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 283. M. trivialis sp. nov., holotype, ♂ (BMNH). 284. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 285. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122202" httpUri="https://zenodo.org/record/4122202/files/figure.png" pageId="127" pageNumber="128">Fig. 285</figureCitation>
). Antennal pocket shallow. Pronotal punctures ovoid to irregularly hexagonal, variably large and moderately deep. Intervening spaces glossy, generally larger than punctures. Setae of pronotal disc as those on head. Scutellar shield rounded apically. Elytron glossy, with tracks of four flat glabrous longitudinal carinae, including sutural one. Punctures linear (elongate and narrow), incisionshaped. Intervening spaces glossy, larger than punctures (
<figureCitation id="0AA52A659D639224FCF5F8A5CB982F54" box="[870,974,1819,1846]" captionStart="Figs 349–358" captionStartId="156.[189,241,1786,1812]" captionTargetBox="[287,1299,265,1743]" captionTargetId="figure-123@156.[287,1301,265,1760]" captionTargetPageId="156" captionText="Figs 349–358. Papuan and Wallacean Maechidius Macleay, 1819, elytron (349–356) and pronotal setation (357–358), dorsal view. 349. M. suwawa sp. nov., paratype, ♀ (BMNH). 350. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 351. M. trivialis sp. nov., holotype, ♂ (BMNH). 352. M. ursus sp. nov., holotype, ♀ (NME). 353. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 354. M. weigeli sp. nov., holotype, ♂ (NME). 355. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 356. M. yamdena sp. nov., paratype, ♂ (BMNH). 357. M. leucopsar sp. nov., holotype, ♂ (IECA). 358. M. ursus sp. nov., holotype, ♀ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122214" httpUri="https://zenodo.org/record/4122214/files/figure.png" pageId="127" pageNumber="128">Fig. 355</figureCitation>
). Appressed seta rises from anterior margin of each puncture, not surpassing length of corresponding incision. Male and female pygidium slightly convex dorsally, with large and shallow annular punctures (Fig. 533). Intervening spaces glossy, generally larger than punctures. Setae of pygidium inconspicuous, appressed; seta rises from anterior margin of each corresponding puncture, not surpassing its length. Longer suberect setae on apical margin of pygidium. Protibia hardly widened distally, in male with three external teeth, basal one of which rudimentary (
<figureCitation id="0AA52A659D9C92DBFECFFEB7C9922941" box="[348,452,265,291]" captionStart="Figs 424–435" captionStartId="161.[189,241,1394,1420]" captionTargetBox="[238,1349,265,1318]" captionTargetId="figure-136@161.[217,1354,265,1322]" captionTargetPageId="161" captionText="Figs 424–435. Papuan and Wallacean Maechidius Macleay, 1819, protibia, dorsal view (424–428), and abdominal ventrites (429–435), ventral and lateral views. 424. M. ursus sp. nov., holotype, ♀ (NME). 425. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 426. M. weigeli sp. nov., holotype, ♂ (NME). 427. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 428. M. yamdena sp. nov., paratype, ♂ (BMNH). 429. M.dani sp. nov., holotype, ♂ (IECA). 430. Same, different aspect (IECA). 431. M. hamatus sp. nov., holotype, ♂ (IECA). 432. Same, different aspect (IECA). 433. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 434. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 435. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122216" httpUri="https://zenodo.org/record/4122216/files/figure.png" pageId="128" pageNumber="129">Fig. 427</figureCitation>
). Protibial terminal spur absent in both sexes. Male metatibial terminal spurs not equal, lower spur curved. Tarsal claws with pulvilli. Male aedeagus as in
<figureCitation id="0AA52A659D9C92DBFB8AFE92CCEB2924" box="[1049,1213,300,326]" captionStart="Figs 739–750" captionStartId="181.[189,241,1892,1918]" captionTargetBox="[360,1207,271,1867]" captionTargetId="figure-66@181.[347,1240,265,1871]" captionTargetPageId="181" captionText="Figs 739–750. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 739– 741. M. weigeli sp. nov., paratype, ♂ (NME). 742–744. M. woodlarkianus Heller, 1914, ♂ (SNSD). 745–747. M. yamdena sp. nov., paratype, ♂ (BMNH). 748–750. M. paupianus Heller, 1910 (paratype ♂ of M. arrowi Frey, 1969) (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122256" httpUri="https://zenodo.org/record/4122256/files/figure.png" pageId="128" pageNumber="129">Figs 742–744</figureCitation>
.
</paragraph>
<paragraph id="922136E09D9C92DBFF2EFED3C9E629E5" blockId="128.[189,432,365,391]" box="[189,432,365,391]" pageId="128" pageNumber="129">
<heading id="C969818C9D9C92DBFF2EFED3C9E629E5" bold="true" box="[189,432,365,391]" fontSize="11" level="3" pageId="128" pageNumber="129" reason="3">
<emphasis id="A0EAEAF29D9C92DBFF2EFED3C9E629E5" bold="true" box="[189,432,365,391]" pageId="128" pageNumber="129">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D9C92DBFF2EFE22CAF729D4" blockId="128.[189,673,412,438]" box="[189,673,412,438]" pageId="128" pageNumber="129">Female lamellae of antennal club shorter.</paragraph>
</subSubSection>
</treatment>
</document>
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