245 lines
38 KiB
XML
245 lines
38 KiB
XML
<document id="97A35542E7B0705868AD90F798DECEAC" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D4F920FFDD2FD76CBA82DBF" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius nepenthephilus Telnov 2020, sp. nov." docType="treatment" docUuid="C34ACD72-FDCF-42C6-BCFE-0E6E5B06F38C" docUuidSource="ZooBank" docVersion="10" lastPageNumber="85" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="84" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="C9D401DCA50D625EB97A8016ACDD5FB3">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
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<mods:namePart id="C942A69DD224B5C3A342FF0CC11E34DF">Telnov, Dmitry</mods:namePart>
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<mods:typeOfResource id="BB0FC81CC9B924AB6A6B06ACA17AEC02">text</mods:typeOfResource>
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<mods:title id="25A58CEA53A53369BD731BEF51720505">European Journal of Taxonomy</mods:title>
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<mods:part id="2C8175C783E87C87CCFC003803AA2A88">
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<mods:date id="B9EA5BBD841150130B23DAD212DA8926">2020</mods:date>
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<mods:number id="47A25ED2CE82AA4D3241CB6D0A7D85B2">2020-10-19</mods:number>
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<mods:number id="443FDB451154A7B292D3C286BDC14D73">721</mods:number>
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<mods:classification id="CC270A67EF48C3B48D870CDBB3671DD8">journal article</mods:classification>
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<mods:identifier id="A1522434AAB32E99E536FD15F13A4D66" type="DOI">10.5852/ejt.2020.721.1127</mods:identifier>
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<treatment id="1A3787F69D4F920FFDD2FD76CBA82DBF" ID-DOI="http://doi.org/10.5281/zenodo.4344263" ID-GBIF-Taxon="168648163" ID-Zenodo-Dep="4344263" LSID="urn:lsid:zoobank.org:act:C34ACD72-FDCF-42C6-BCFE-0E6E5B06F38C" httpUri="http://treatment.plazi.org/id/1A3787F69D4F920FFDD2FD76CBA82DBF" lastPageId="84" lastPageNumber="85" pageId="83" pageNumber="84">
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<subSubSection id="DA84656B9D4F9208FDD2FD76CBA42A81" box="[577,1010,712,739]" pageId="83" pageNumber="84" type="nomenclature">
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<paragraph id="922136E09D4F9208FDD2FD76CBA42A81" blockId="83.[355,1232,712,810]" box="[577,1010,712,739]" pageId="83" pageNumber="84">
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<heading id="C969818C9D4F9208FDD2FD76CBA42A81" box="[577,1010,712,739]" centered="true" fontSize="11" level="2" pageId="83" pageNumber="84" reason="2">
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<taxonomicName id="559E4D639D4F9208FDD2FD76CBC72A81" ID-CoL="7X3X9" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[577,913,712,739]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="nepenthephilus" status="sp. nov.">
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<emphasis id="A0EAEAF29D4F9208FDD2FD76CBC72A81" bold="true" box="[577,913,712,739]" italics="true" pageId="83" pageNumber="84">Maechidius nepenthephilus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="BBD957899D4F9208FC0BFD77CBA42A81" box="[920,1010,713,739]" pageId="83" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="DA84656B9D4F9208FEF0FD52CC3D2B48" pageId="83" pageNumber="84" type="description">
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<paragraph id="922136E09D4F9208FEF0FD52CC862B65" blockId="83.[355,1232,712,810]" box="[355,1232,748,775]" pageId="83" pageNumber="84">
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urn:lsid:zoobank.org:act:
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<uuid id="E6380C359D4F9208FD13FD52CC862B65" box="[640,1232,748,775]" pageId="83" pageNumber="84">C34ACD72-FDCF-42C6-BCFE-0E6E5B06F38C</uuid>
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</uri>
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</paragraph>
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<paragraph id="922136E09D4F9208FE5AFCB1CC3D2B48" blockId="83.[355,1232,712,810]" box="[457,1131,783,810]" pageId="83" pageNumber="84">
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<figureCitation id="0AA52A659D4F9208FE5AFCB1CA752B48" box="[457,547,783,810]" captionStart="Figs 62–65" captionStartId="81.[189,241,1926,1952]" captionTargetBox="[260,1331,260,1918]" captionTargetId="figure-59@81.[254,1332,254,1925]" captionTargetPageId="81" captionText="Figs 62–65. Papuan Maechidius Macleay, 1819, habitus, dorsal view. 62. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 63. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 64. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 65. M. nepenthephilus sp. nov., holotype, ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122154" httpUri="https://zenodo.org/record/4122154/files/figure.png" pageId="83" pageNumber="84">Figs 65</figureCitation>
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,
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<figureCitation id="0AA52A659D4F9208FDBCFCB1CACE2B48" box="[559,664,783,810]" captionStart="Figs 145–156" captionStartId="141.[189,241,1787,1813]" captionTargetBox="[328,1254,265,1740]" captionTargetId="figure-124@141.[328,1260,265,1749]" captionTargetPageId="141" captionText="Figs 145–156. Papuan Maechidius Macleay, 1819, head, dorsal view. 145. M. merdeka sp. nov., holotype, ♂ (BMNH). 146. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 147. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 148. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 149. Same, ♂ (SNSD). 150. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 151. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 152. M. nepenthephilus sp. nov., holotype, ♂ (NME). 153. Same, paratype, ♀ (DTC). 154. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 155. Same, paratype, ♀ (BMNH). 156. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122181" httpUri="https://zenodo.org/record/4122181/files/figure.png" pageId="83" pageNumber="84">152–153</figureCitation>
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,
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<figureCitation id="0AA52A659D4F9208FD37FCB1CA852B4B" box="[676,723,783,809]" captionStart="Figs 226–241" captionStartId="147.[189,241,1681,1707]" captionTargetBox="[391,1192,268,1647]" captionTargetId="figure-170@147.[391,1196,265,1656]" captionTargetPageId="147" captionText="Figs 226–241. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 226. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 227. M. lobaticeps Frey, 1969, ♂ (DTC). 228. M. longipes sp. nov., holotype, ♂ (BMNH). 229. M. luniceps Fairmaire, 1883, ♂ (BMNH). 230. M. mailu sp. nov., holotype, ♂ (NHMB). 231. M. maleo sp. nov., holotype, ♂ (IRSN). 232. M. merdeka sp. nov., holotype, ♂ (BMNH). 233. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 234. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 235. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 236. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 237. M. nepenthephilus sp. nov., holotype, ♂ (NME). 238. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 239. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 240. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 241. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122193" httpUri="https://zenodo.org/record/4122193/files/figure.png" pageId="83" pageNumber="84">237</figureCitation>
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,
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<figureCitation id="0AA52A659D4F9208FD4CFCB1CB582B4B" box="[735,782,783,809]" captionStart="Figs 317–332" captionStartId="154.[189,241,1681,1707]" captionTargetBox="[401,1187,265,1655]" captionTargetId="figure-172@154.[401,1187,265,1669]" captionTargetPageId="154" captionText="Figs 317–332. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 317. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 318. M. lobaticeps Frey, 1969, ♂ (DTC). 319. M. longipes sp. nov., holotype, ♂ (BMNH). 320. M. luniceps Fairmaire, 1883, ♂ (BMNH). 321. M. mailu sp. nov., holotype, ♂ (NHMB). 322. M. maleo sp. nov., holotype, ♂ (IRSN). 323. M. merdeka sp. nov., holotype, ♂ (BMNH). 324. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 325. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 326. M. muticusArrow, 1941, syntype, ♂ (BMNH). 327. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 328. M. nepenthephilus sp. nov., holotype, ♂ (NME). 329. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 330. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 331. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 332. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122210" httpUri="https://zenodo.org/record/4122210/files/figure.png" pageId="83" pageNumber="84">328</figureCitation>
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, 401, 497, 679–681, 751–753
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="DA84656B9D4F9208FF2EFCEECCBD2BDE" pageId="83" pageNumber="84" type="diagnosis">
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<paragraph id="922136E09D4F9208FF2EFCEEC99F2B08" blockId="83.[189,457,848,874]" box="[189,457,848,874]" pageId="83" pageNumber="84">
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<heading id="C969818C9D4F9208FF2EFCEEC99F2B08" bold="true" box="[189,457,848,874]" fontSize="11" level="3" pageId="83" pageNumber="84" reason="3">
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||
<emphasis id="A0EAEAF29D4F9208FF2EFCEEC99F2B08" bold="true" box="[189,457,848,874]" pageId="83" pageNumber="84">Differential diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="922136E09D4F9208FF2EFCC1CCBD2BDE" blockId="83.[189,1399,895,957]" pageId="83" pageNumber="84">This species is primarily different among congeners due to the shape of the aedeagus in combination with the sculpture (irregularly paired rows of linear punctures) and the setae of the dorsum.</paragraph>
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</subSubSection>
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<subSubSection id="DA84656B9D4F9208FF2EFC5ACAEE2C11" pageId="83" pageNumber="84" type="etymology">
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<paragraph id="922136E09D4F9208FF2EFC5AC9122B9C" blockId="83.[189,324,996,1022]" box="[189,324,996,1022]" pageId="83" pageNumber="84">
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<heading id="C969818C9D4F9208FF2EFC5AC9122B9C" bold="true" box="[189,324,996,1022]" fontSize="11" level="3" pageId="83" pageNumber="84" reason="3">
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<emphasis id="A0EAEAF29D4F9208FF2EFC5AC9122B9C" bold="true" box="[189,324,996,1022]" pageId="83" pageNumber="84">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph id="922136E09D4F9208FF2EFBADCAEE2C11" blockId="83.[189,1398,1043,1139]" pageId="83" pageNumber="84">
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The name is derived from a combination of
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<taxonomicName id="559E4D639D4F9208FD40FBADCC582C4C" authority="Linnaeus, 1753" authorityName="Linnaeus" authorityYear="1753" box="[723,1038,1043,1070]" class="Magnoliopsida" family="Nepenthaceae" genus="Nepenthes" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="83" pageNumber="84" phylum="Tracheophyta" rank="genus">
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<emphasis id="A0EAEAF29D4F9208FD40FBADCB1B2C4F" box="[723,845,1043,1069]" italics="true" pageId="83" pageNumber="84">Nepenthes</emphasis>
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Linnaeus, 1753
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</taxonomicName>
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(genus of carnivorous pitcher plants) and the Greek ‘
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<emphasis id="A0EAEAF29D4F9208FE59FB88CA582C32" box="[458,526,1078,1104]" italics="true" pageId="83" pageNumber="84">philia</emphasis>
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’ (φιλία, Ancient Greek for love and friendship), indicating the association of the imago with flowers of pitcher plants.
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</paragraph>
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</subSubSection>
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<subSubSection id="DA84656B9D4F9208FF2EFB25CB632DD0" pageId="83" pageNumber="84" type="materials_examined">
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<paragraph id="922136E09D4F9208FF2EFB25C93A2CD7" blockId="83.[189,364,1179,1205]" box="[189,364,1179,1205]" pageId="83" pageNumber="84">
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<heading id="C969818C9D4F9208FF2EFB25C93A2CD7" bold="true" box="[189,364,1179,1205]" fontSize="11" level="3" pageId="83" pageNumber="84" reason="3">
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<emphasis id="A0EAEAF29D4F9208FF2EFB25C93A2CD7" bold="true" box="[189,364,1179,1205]" pageId="83" pageNumber="84">Type material</emphasis>
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||
</heading>
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||
</paragraph>
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<materialsCitation id="22F63CBD9D4F9208FF79FB74CA932D2C" ID-GBIF-Occurrence="2898878569" collectingDate="2018-03-31" collectionCode="NME" country="Indonesia" county="Doyo Lama" elevation="158" latitude="-2.5638888" location="New Guinea" longLatPrecision="21" longitude="140.45555" municipality="Nimbokrang" pageId="83" pageNumber="84" specimenCount="1" typeStatus="holotype">
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<paragraph id="922136E09D4F9208FF79FB74C90A2C86" blockId="83.[189,1398,1226,1358]" box="[234,348,1226,1252]" pageId="83" pageNumber="84">
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<emphasis id="A0EAEAF29D4F9208FF79FB74C90A2C86" bold="true" box="[234,348,1226,1252]" pageId="83" pageNumber="84">
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<typeStatus id="4D2588429D4F9208FF79FB74C90A2C86" box="[234,348,1226,1252]" pageId="83" pageNumber="84" type="holotype">Holotype</typeStatus>
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</emphasis>
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</paragraph>
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<paragraph id="922136E09D4F9208FF2EFB53CA932D2C" blockId="83.[189,1398,1226,1358]" pageId="83" pageNumber="84">
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<collectingCountry id="EA8976709D4F9208FF2EFB53C90D2D65" box="[189,347,1261,1287]" name="Indonesia" pageId="83" pageNumber="84">INDONESIA</collectingCountry>
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•
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<specimenCount id="8498FD699D4F9208FEE0FB53C9DD2D65" box="[371,395,1261,1287]" pageId="83" pageNumber="84" type="male">♂</specimenCount>
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; “
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<collectingCountry id="EA8976709D4F9208FE31FB53CA152D65" box="[418,579,1261,1287]" name="Indonesia" pageId="83" pageNumber="84">INDONESIA</collectingCountry>
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E,
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<location id="9741603B9D4F9208FDF9FB53CB562D65" LSID="urn:lsid:plazi:treatment:1A3787F69D4F920FFDD2FD76CBA82DBF:9741603B9D4F9208FDF9FB53CB562D65" box="[618,768,1261,1287]" country="Indonesia" county="Doyo Lama" latitude="-2.5638888" longLatPrecision="21" longitude="140.45555" municipality="Nimbokrang" name="New Guinea" pageId="83" pageNumber="84">New Guinea</location>
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||
,
|
||
<location id="9741603B9D4F9208FC98FB53CBCC2D65" LSID="urn:lsid:plazi:treatment:1A3787F69D4F920FFDD2FD76CBA82DBF:9741603B9D4F9208FC98FB53CBCC2D65" box="[779,922,1261,1287]" country="Indonesia" county="Doyo Lama" latitude="-2.5638888" longLatPrecision="21" longitude="140.45555" municipality="Nimbokrang" name="Papua Prov." pageId="83" pageNumber="84">Papua Prov.</location>
|
||
,
|
||
<location id="9741603B9D4F9208FC36FB53CBAB2D65" LSID="urn:lsid:plazi:treatment:1A3787F69D4F920FFDD2FD76CBA82DBF:9741603B9D4F9208FC36FB53CBAB2D65" box="[933,1021,1261,1287]" country="Indonesia" county="Doyo Lama" latitude="-2.5638888" longLatPrecision="21" longitude="140.45555" municipality="Nimbokrang" name="Sentani" pageId="83" pageNumber="84">Sentani</location>
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<locationDeviation id="83932FC29D4F9208FB97FB53CC022D65" box="[1028,1108,1261,1288]" metricMagnitude="3" metricUnit="m" metricValue="5.5" pageId="83" pageNumber="84" unit="km" value="5.5">5.5 km</locationDeviation>
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||
</quantity>
|
||
W, road to
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||
<collectingMunicipality id="7245AC9A9D4F9208FB48FB53CD242D65" box="[1243,1394,1261,1287]" pageId="83" pageNumber="84">Nimbokrang</collectingMunicipality>
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,
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<collectingCounty id="7B404E6C9D4F9208FF2EFAAFC91A2D49" box="[189,332,1297,1323]" pageId="83" pageNumber="84">Doyo Lama</collectingCounty>
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vill.,
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,
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,
|
||
<quantity id="55669B059D4F9208FD70FAAECB3F2D49" box="[739,873,1296,1323]" metricMagnitude="2" metricUnit="m" metricValue="1.575" metricValueMax="1.65" metricValueMin="1.5" pageId="83" pageNumber="84" unit="m" value="157.5" valueMax="165.0" valueMin="150.0">
|
||
<elevation id="19B3D1D39D4F9208FD70FAAECB3F2D49" box="[739,873,1296,1323]" metricMagnitude="2" metricUnit="m" metricValue="1.575" metricValueMax="1.65" metricValueMin="1.5" pageId="83" pageNumber="84" unit="m" value="157.5" valueMax="165.0" valueMin="150.0">150-165 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="E62010209D4F9208FCEAFAAECC562D48" box="[889,1024,1296,1322]" pageId="83" pageNumber="84" value="2018-03-31">
|
||
<collectingDate id="F664E9C89D4F9208FCEAFAAECC562D48" box="[889,1024,1296,1322]" pageId="83" pageNumber="84" value="2018-03-31">31.III.2018</collectingDate>
|
||
</date>
|
||
, semidry eucalypt forest, from flowers & pitchers of
|
||
<taxonomicName id="559E4D639D4F9208FE2EFA8DCA612D2F" authorityName="Linnaeus" authorityYear="1753" box="[445,567,1331,1357]" class="Magnoliopsida" family="Nepenthaceae" genus="Nepenthes" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="83" pageNumber="84" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="A0EAEAF29D4F9208FE2EFA8DCA612D2F" box="[445,567,1331,1357]" italics="true" pageId="83" pageNumber="84">Nepenthes</emphasis>
|
||
</taxonomicName>
|
||
sp.”;
|
||
<collectionCode id="F48FAE259D4F9208FDEFFA8ACA972D2C" box="[636,705,1332,1358]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="83" pageNumber="84">NME</collectionCode>
|
||
.
|
||
</paragraph>
|
||
</materialsCitation>
|
||
<paragraph id="922136E09D4F9208FF79FACBC90A2DED" blockId="83.[189,821,1397,1459]" box="[234,348,1397,1423]" pageId="83" pageNumber="84">
|
||
<emphasis id="A0EAEAF29D4F9208FF79FACBC90A2DED" bold="true" box="[234,348,1397,1423]" pageId="83" pageNumber="84">
|
||
<typeStatus id="4D2588429D4F9208FF79FACBC90A2DED" box="[234,348,1397,1423]" pageId="83" pageNumber="84" type="paratype">Paratype</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="922136E09D4F9208FF2EFA26CB632DD0" blockId="83.[189,821,1397,1459]" box="[189,821,1432,1459]" pageId="83" pageNumber="84">
|
||
<materialsCitation id="22F63CBD9D4F9208FF2EFA26CB672DD0" ID-GBIF-Occurrence="2898878576" box="[189,817,1432,1459]" collectingDate="2018-03-31" collectionCode="DTC" country="Indonesia" county="Doyo Lama" elevation="158" latitude="-2.5638888" location="INDONESIA" longLatPrecision="21" longitude="140.45555" municipality="Nimbokrang" pageId="83" pageNumber="84" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||
<collectingCountry id="EA8976709D4F9208FF2EFA26C90D2DD0" box="[189,347,1432,1458]" name="Indonesia" pageId="83" pageNumber="84">INDONESIA</collectingCountry>
|
||
•
|
||
<specimenCount id="8498FD699D4F9208FEE7FA26C9F72DD1" box="[372,417,1432,1459]" pageId="83" pageNumber="84" type="female">1 ♀</specimenCount>
|
||
; same label as for holotype;
|
||
<collectionCode id="F48FAE259D4F9208FD60FA26CB672DD0" box="[755,817,1432,1458]" collectionName="Dmitrijs Telnovs" pageId="83" pageNumber="84">DTC</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DA84656B9D4F920FFF2EFA64CB432CB7" lastPageId="84" lastPageNumber="85" pageId="83" pageNumber="84" type="description">
|
||
<paragraph id="922136E09D4F9208FF2EFA64C9182D96" blockId="83.[189,334,1498,1524]" box="[189,334,1498,1524]" pageId="83" pageNumber="84">
|
||
<heading id="C969818C9D4F9208FF2EFA64C9182D96" bold="true" box="[189,334,1498,1524]" fontSize="11" level="3" pageId="83" pageNumber="84" reason="3">
|
||
<emphasis id="A0EAEAF29D4F9208FF2EFA64C9182D96" bold="true" box="[189,334,1498,1524]" pageId="83" pageNumber="84">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="922136E09D4F9208FF2EF9B7CAE92E0B" blockId="83.[189,1399,1544,1642]" pageId="83" pageNumber="84">
|
||
MEASUREMENTS.
|
||
<typeStatus id="4D2588429D4F9208FEEFF9B6C9BA2E40" box="[380,492,1544,1570]" pageId="83" pageNumber="84" type="holotype">Holotype</typeStatus>
|
||
, total body length
|
||
<quantity id="55669B059D4F9208FD54F9B7CB652E41" box="[711,819,1545,1571]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="83" pageNumber="84" unit="mm" value="7.7">7.70 mm</quantity>
|
||
. Head
|
||
<quantity id="55669B059D4F9208FC16F9B7CBB82E41" box="[901,1006,1545,1571]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="83" pageNumber="84" unit="mm" value="1.3">1.30 mm</quantity>
|
||
long, across eyes
|
||
<quantity id="55669B059D4F9208FB55F9B6CD792E41" box="[1222,1327,1544,1571]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="83" pageNumber="84" unit="mm" value="1.6">1.60 mm</quantity>
|
||
wide. Pronotum
|
||
<quantity id="55669B059D4F9208FEA4F992C9C92E24" box="[311,415,1580,1606]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="83" pageNumber="84" unit="mm" value="1.9">1.90 mm</quantity>
|
||
long, maximum width
|
||
<quantity id="55669B059D4F9208FD3EF992CB412E24" box="[685,791,1580,1606]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="83" pageNumber="84" unit="mm" value="2.8">2.80 mm</quantity>
|
||
. Elytral length
|
||
<quantity id="55669B059D4F9208FC54F992CC642E24" box="[967,1074,1580,1607]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="83" pageNumber="84" unit="mm" value="4.5">4.50 mm</quantity>
|
||
, maximum combined width
|
||
<quantity id="55669B059D4F9208FF2EF9F1C97F2E08" box="[189,297,1615,1642]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="83" pageNumber="84" unit="mm" value="3.4">3.40 mm</quantity>
|
||
.
|
||
<typeStatus id="4D2588429D4F9208FEA7F9EEC9CD2E08" box="[308,411,1616,1642]" pageId="83" pageNumber="84" type="paratype">Paratype</typeStatus>
|
||
female is
|
||
<quantity id="55669B059D4F9208FD87F9F1CA2B2E08" box="[532,637,1615,1642]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="83" pageNumber="84" unit="mm" value="6.7">6.70 mm</quantity>
|
||
long.
|
||
</paragraph>
|
||
<paragraph id="922136E09D4F920FFF2EF92FCBD22C5D" blockId="83.[189,1399,1681,2025]" lastBlockId="84.[189,1399,265,1088]" lastPageId="84" lastPageNumber="85" pageId="83" pageNumber="84">
|
||
Dorsum uniformly black-brown with brown labroclypeus, antennae, legs and venter. Head transverse, subopaque dorsally and ventrally, flattened dorsally. Compound eye moderately large, occupying less than half side of head. Male labroclypeus (
|
||
<figureCitation id="0AA52A659D4F9208FC9BF969CB262E90" box="[776,880,1751,1778]" captionStart="Figs 145–156" captionStartId="141.[189,241,1787,1813]" captionTargetBox="[328,1254,265,1740]" captionTargetId="figure-124@141.[328,1260,265,1749]" captionTargetPageId="141" captionText="Figs 145–156. Papuan Maechidius Macleay, 1819, head, dorsal view. 145. M. merdeka sp. nov., holotype, ♂ (BMNH). 146. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 147. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 148. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 149. Same, ♂ (SNSD). 150. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 151. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 152. M. nepenthephilus sp. nov., holotype, ♂ (NME). 153. Same, paratype, ♀ (DTC). 154. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 155. Same, paratype, ♀ (BMNH). 156. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122181" httpUri="https://zenodo.org/record/4122181/files/figure.png" pageId="83" pageNumber="84">Fig. 152</figureCitation>
|
||
) broadly emarginate, in female (
|
||
<figureCitation id="0AA52A659D4F9208FA95F969CD382E90" box="[1286,1390,1751,1778]" captionStart="Figs 145–156" captionStartId="141.[189,241,1787,1813]" captionTargetBox="[328,1254,265,1740]" captionTargetId="figure-124@141.[328,1260,265,1749]" captionTargetPageId="141" captionText="Figs 145–156. Papuan Maechidius Macleay, 1819, head, dorsal view. 145. M. merdeka sp. nov., holotype, ♂ (BMNH). 146. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 147. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 148. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 149. Same, ♂ (SNSD). 150. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 151. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 152. M. nepenthephilus sp. nov., holotype, ♂ (NME). 153. Same, paratype, ♀ (DTC). 154. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 155. Same, paratype, ♀ (BMNH). 156. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122181" httpUri="https://zenodo.org/record/4122181/files/figure.png" pageId="83" pageNumber="84">Fig. 153</figureCitation>
|
||
) shallower emarginate anteriorly. Lateral margins of labroclypeus in both sexes sinuous in both dorsal and lateral views, anterolateral angles weakly protruding, obtuse in dorsal view. Upper- and underside of labroclypeus with sparse long setae along anterior and lateral margins. Canthus nearly straight in dorsal view. Anterior and lateral margins of labroclypeus smooth. Head dorsal punctures circular, moderately deep and dense, filled or not with microscopical velvety pubescence. Intervening spaces glossy to delicately microreticulate, variably large. Suberect long seta rises from anterior margin of each puncture; four extraordinarily long erect setae at inner margin of either compound eye and one single on either canthus. Antenna 9-segmented, club 3-lamellate. Scape large, with bulbous predistal projection on dorsal side, provided with two very long setae near distal margin. Antennomere 2 trapezoid, transverse. Pronotum strongly transverse, subopaque dorsally and laterally. Anterior margin of pronotum broadly emarginate, anterolateral angles protruding anteriad. Basal and lateral margins broadly rounded in dorsal view. Crenulae of lateral margin moderately strong; a long erect curved seta present between every two crenulae (
|
||
<figureCitation id="0AA52A659D48920FFEA6FE00C9F729BA" box="[309,417,446,472]" captionStart="Figs 226–241" captionStartId="147.[189,241,1681,1707]" captionTargetBox="[391,1192,268,1647]" captionTargetId="figure-170@147.[391,1196,265,1656]" captionTargetPageId="147" captionText="Figs 226–241. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 226. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 227. M. lobaticeps Frey, 1969, ♂ (DTC). 228. M. longipes sp. nov., holotype, ♂ (BMNH). 229. M. luniceps Fairmaire, 1883, ♂ (BMNH). 230. M. mailu sp. nov., holotype, ♂ (NHMB). 231. M. maleo sp. nov., holotype, ♂ (IRSN). 232. M. merdeka sp. nov., holotype, ♂ (BMNH). 233. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 234. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 235. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 236. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 237. M. nepenthephilus sp. nov., holotype, ♂ (NME). 238. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 239. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 240. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 241. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122193" httpUri="https://zenodo.org/record/4122193/files/figure.png" pageId="84" pageNumber="85">Fig. 237</figureCitation>
|
||
). Lateral margin of pronotum slightly arched in lateral view. Pronotal punctures ovoid to oblong-ovoid, annular, deep and dense, variably large (generally larger along lateral margins and on base), filled or not with microscopical velvety pubescence. Intervening spaces in part glossy and glabrous, in part microreticulate, generally smaller than punctures except in some areas. Setae similar to those on head, somewhat stronger curved. Lateral margins partly (narrowly) covered with microscopical velvety pubescence. Hypomeron separated from prosternum by low nearly straight carina which is long setose opposite to compound eye. Antennal pocket shallow. Median anterior process of prosternum long brushy setose, short, strongly raised. Scutellar shield narrowly rounded apically. Elytra cylindrical, maximum width across midlength, opaque dorsally, with distinct humeri. Elytral disc irregularly, mainly transversely wrinkled. Punctures of elytral disc linear (elongate and narrow), incision-like, arranged in irregular paired longitudinal rows (
|
||
<figureCitation id="0AA52A659D48920FFD6DFC96CB3C2B20" box="[766,874,808,834]" captionStart="Figs 317–332" captionStartId="154.[189,241,1681,1707]" captionTargetBox="[401,1187,265,1655]" captionTargetId="figure-172@154.[401,1187,265,1669]" captionTargetPageId="154" captionText="Figs 317–332. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 317. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 318. M. lobaticeps Frey, 1969, ♂ (DTC). 319. M. longipes sp. nov., holotype, ♂ (BMNH). 320. M. luniceps Fairmaire, 1883, ♂ (BMNH). 321. M. mailu sp. nov., holotype, ♂ (NHMB). 322. M. maleo sp. nov., holotype, ♂ (IRSN). 323. M. merdeka sp. nov., holotype, ♂ (BMNH). 324. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 325. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 326. M. muticusArrow, 1941, syntype, ♂ (BMNH). 327. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 328. M. nepenthephilus sp. nov., holotype, ♂ (NME). 329. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 330. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 331. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 332. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122210" httpUri="https://zenodo.org/record/4122210/files/figure.png" pageId="84" pageNumber="85">Fig. 328</figureCitation>
|
||
). Moderately long suberect seta rises from anterior margin of each incision, surpassing its length (elytral setae shorter than those on forebody). Lateral margin of elytra all along with moderately long erect setae. Male and female pygidium flattened dorsally, shallowly impressed in apical part. Punctures of pygidium large and dense, circular, annular, very shallow. Intervening spaces much smaller than punctures, glossy to subopaque (
|
||
<figureCitation id="0AA52A659D48920FFB2BFC07CD772BB1" box="[1208,1313,953,979]" captionStart="Figs487–503" captionStartId="165.[189,241,1681,1707]" captionTargetBox="[422,1160,265,1655]" captionTargetId="figure-171@165.[421,1167,265,1658]" captionTargetPageId="165" captionText="Figs487–503.Papuan andWallaceanMaechidius Macleay, 1819,pygidium, dorsalview.487.M.lobaticeps Frey, 1969, ♂ (DTC). 488. M. longipes sp. nov., holotype, ♂ (BMNH). 489. M. luniceps Fairmaire, 1883, ♂ (BMNH). 490. M. mailu sp. nov., holotype, ♂ (NHMB). 491. M. maleo sp. nov., holotype, ♂ (IRSN). 492. M. merdeka sp. nov., holotype, ♂ (BMNH). 493. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 494. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 495. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 496. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 497. M. nepenthephilus sp. nov., holotype, ♂ (NME). 498. M. opatroidesArrow, 1941 comb. rest., lectotype, ♀ (BMNH). 499. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 500. Same, paratype, ♀ (BMNH). 501. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 502. Same, different aspect. 503. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122226" httpUri="https://zenodo.org/record/4122226/files/figure.png" pageId="84" pageNumber="85">Fig. 497</figureCitation>
|
||
). Setae of pygidium suberect, sparse, moderately long. Male and female protibia with two inconspicuous distal teeth (Fig. 401), female ones generally larger. Protibial terminal spur absent in both sexes. Male tarsal claws with large pulvilli. Male aedeagus as in
|
||
<figureCitation id="0AA52A659D48920FFD4FFB9BCBD62C5D" box="[732,896,1061,1087]" captionStart="Figs 667–681" captionStartId="176.[189,241,1892,1918]" captionTargetBox="[444,1142,267,1860]" captionTargetId="figure-72@176.[419,1168,265,1868]" captionTargetPageId="176" captionText="Figs 667–681. Papuan Maechidius Macleay, 1819, aedeagus, different aspects. 667– 669. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 670–672. M. milneanus Heller, 1914, ♂ (SNSD). 673–675. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 676–678. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 679-681. M. nepenthephilus sp. nov., holotype, ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122246" httpUri="https://zenodo.org/record/4122246/files/figure.png" pageId="84" pageNumber="85">Figs 679–681</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="922136E09D48920FFF2EFBD9C9E62CE3" blockId="84.[189,432,1127,1153]" box="[189,432,1127,1153]" pageId="84" pageNumber="85">
|
||
<heading id="C969818C9D48920FFF2EFBD9C9E62CE3" bold="true" box="[189,432,1127,1153]" fontSize="11" level="3" pageId="84" pageNumber="85" reason="3">
|
||
<emphasis id="A0EAEAF29D48920FFF2EFBD9C9E62CE3" bold="true" box="[189,432,1127,1153]" pageId="84" pageNumber="85">Sexual dimorphism</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="922136E09D48920FFF2EFB29CB432CB7" blockId="84.[189,1399,1175,1237]" pageId="84" pageNumber="85">Female lamellae of antennal club shorter; female labroclypeus comparatively shallower emarginate anteriorly with less protruding anterolateral angles.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DA84656B9D48920FFF2EFB40CAF62D0E" pageId="84" pageNumber="85" type="biology_ecology">
|
||
<paragraph id="922136E09D48920FFF2EFB40C9762D7A" blockId="84.[189,288,1278,1304]" box="[189,288,1278,1304]" pageId="84" pageNumber="85">
|
||
<heading id="C969818C9D48920FFF2EFB40C9762D7A" bold="true" box="[189,288,1278,1304]" fontSize="11" level="3" pageId="84" pageNumber="85" reason="3">
|
||
<emphasis id="A0EAEAF29D48920FFF2EFB40C9762D7A" bold="true" box="[189,288,1278,1304]" pageId="84" pageNumber="85">Ecology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="922136E09D48920FFF2EFA93CAF62D0E" blockId="84.[189,1398,1325,1388]" pageId="84" pageNumber="85">
|
||
Occurs on dry nutrient-poor semidry slopes in eucalypt stands at altitudes around
|
||
<specimenCount id="8498FD699D48920FFB37FA93CCAF2D2A" box="[1188,1273,1325,1352]" pageId="84" pageNumber="85" type="generic">
|
||
<quantity id="55669B059D48920FFB37FA93CCA32D2A" box="[1188,1269,1325,1352]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="84" pageNumber="85" unit="m" value="150.0">150 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
Found on flowers of
|
||
<taxonomicName id="559E4D639D48920FFEA9FAEFC9E22D09" authorityName="Linnaeus" authorityYear="1753" box="[314,436,1361,1387]" class="Magnoliopsida" family="Nepenthaceae" genus="Nepenthes" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="84" pageNumber="85" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="A0EAEAF29D48920FFEA9FAEFC9E22D09" box="[314,436,1361,1387]" italics="true" pageId="84" pageNumber="85">Nepenthes</emphasis>
|
||
</taxonomicName>
|
||
sp. (
|
||
<figureCitation id="0AA52A659D48920FFE7EFAEFCAC22D0E" box="[493,660,1361,1388]" captionStart="Figs 751–753" captionStartId="182.[189,241,1928,1954]" captionTargetBox="[216,1371,266,1899]" captionTargetId="figure-58@182.[216,1371,265,1899]" captionTargetPageId="182" captionText="Figs 751–753. Dry nutrient-poor semi-dry slopes in eucalypt stands near Doyo Lama village, Sentani env., North New Guinea. Numerous Maechidius lobaticeps Frey, 1969 and the type series of M. nepenthephilus sp. nov. were sampled from flowers and pitchers of Nepenthes sp. in this area." figureDoi="http://doi.org/10.5281/zenodo.4122258" httpUri="https://zenodo.org/record/4122258/files/figure.png" pageId="84" pageNumber="85">Figs 751–753</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DA84656B9D48920FFF2EFA2ACBA82DBF" box="[189,1022,1428,1502]" pageId="84" pageNumber="85" type="distribution">
|
||
<paragraph id="922136E09D48920FFF2EFA2AC9002DCC" blockId="84.[189,342,1428,1454]" box="[189,342,1428,1454]" pageId="84" pageNumber="85">
|
||
<heading id="C969818C9D48920FFF2EFA2AC9002DCC" bold="true" box="[189,342,1428,1454]" fontSize="11" level="3" pageId="84" pageNumber="85" reason="3">
|
||
<emphasis id="A0EAEAF29D48920FFF2EFA2AC9002DCC" bold="true" box="[189,342,1428,1454]" pageId="84" pageNumber="85">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="922136E09D48920FFF2EFA7DCBA82DBF" blockId="84.[189,1022,1475,1502]" box="[189,1022,1475,1502]" pageId="84" pageNumber="85">
|
||
Hitherto only known from Sentani surroundings in North New
|
||
<collectingCountry id="EA8976709D48920FFC31FA7DCBAD2DBF" box="[930,1019,1475,1501]" name="Guinea-Bissau" pageId="84" pageNumber="85">Guinea</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |