treatments-xml/data/1A/37/87/1A3787F69D4E9208FDCFFC24CAF32AE8.xml
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<document id="4B51E980EAA7F54E6D4150A5CB17E667" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D4E9208FDCFFC24CAF32AE8" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius nanus Arrow 1941" docType="treatment" docVersion="10" lastPageNumber="84" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallaces Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="83" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="B850AC084C3FA8294A951622740EBCEB">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallaces Line</mods:title>
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<mods:title id="65FC8BEF49A0B4F27788B21E66D5EE3C">European Journal of Taxonomy</mods:title>
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<mods:date id="8394A97C7067E485E9AC0083963FDC2B">2020</mods:date>
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<treatment id="1A3787F69D4E9208FDCFFC24CAF32AE8" ID-DOI="http://doi.org/10.5281/zenodo.4344527" ID-GBIF-Taxon="168648162" ID-Zenodo-Dep="4344527" LSID="urn:lsid:plazi:treatment:1A3787F69D4E9208FDCFFC24CAF32AE8" httpUri="http://treatment.plazi.org/id/1A3787F69D4E9208FDCFFC24CAF32AE8" lastPageId="83" lastPageNumber="84" pageId="82" pageNumber="83">
<subSubSection id="DA84656B9D4E9209FDCFFC24CB8E2BD7" box="[604,984,922,949]" pageId="82" pageNumber="83" type="nomenclature">
<paragraph id="922136E09D4E9209FDCFFC24CB8E2BD7" blockId="82.[545,1042,922,985]" box="[604,984,922,949]" pageId="82" pageNumber="83">
<heading id="C969818C9D4E9209FDCFFC24CB8E2BD7" box="[604,984,922,949]" centered="true" fontSize="11" level="2" pageId="82" pageNumber="83" reason="2">
<taxonomicName id="559E4D639D4E9209FDCFFC24CB8E2BD7" ID-CoL="3XFKD" authority="Arrow, 1941" authorityName="Arrow" authorityYear="1941" box="[604,984,922,949]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="A0EAEAF29D4E9209FDCFFC24CB6A2BD7" bold="true" box="[604,828,922,949]" italics="true" pageId="82" pageNumber="83">Maechidius nanus</emphasis>
<bibRefCitation id="F60F4B119D4E9209FCD0FC25CB8E2BD7" author="Arrow G. E." box="[835,984,923,949]" pageId="82" pageNumber="83" pagination="448 - 464" refId="ref97558" refString="Arrow G. E. 1941. XXIX. - Some undescribed species of melolonthid Coleoptera from Western New Guinea and the adjacent islands of Waigeu and Japen. Annals and Magazine of Natural History, Series 11 7 (41): 448 - 464. https: // doi. org / 10.1080 / 03745481.1941.9727945" type="journal article" year="1941">Arrow, 1941</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="DA84656B9D4E9209FDB2FC00CC442BBA" box="[545,1042,958,985]" pageId="82" pageNumber="83" type="description">
<paragraph id="922136E09D4E9209FDB2FC00CC442BBA" blockId="82.[545,1042,922,985]" box="[545,1042,958,985]" pageId="82" pageNumber="83">
<figureCitation id="0AA52A659D4E9209FDB2FC00CA2D2BBA" box="[545,635,958,984]" captionStart="Figs 6265" captionStartId="81.[189,241,1926,1952]" captionTargetBox="[260,1331,260,1918]" captionTargetId="figure-59@81.[254,1332,254,1925]" captionTargetPageId="81" captionText="Figs 6265. Papuan Maechidius Macleay, 1819, habitus, dorsal view. 62. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 63. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 64. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 65. M. nepenthephilus sp. nov., holotype, ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122154" httpUri="https://zenodo.org/record/4122154/files/figure.png" pageId="82" pageNumber="83">Figs 64</figureCitation>
,
<figureCitation id="0AA52A659D4E9209FD14FC00CAE32BBB" box="[647,693,958,985]" captionStart="Figs 145156" captionStartId="141.[189,241,1787,1813]" captionTargetBox="[328,1254,265,1740]" captionTargetId="figure-124@141.[328,1260,265,1749]" captionTargetPageId="141" captionText="Figs 145156. Papuan Maechidius Macleay, 1819, head, dorsal view. 145. M. merdeka sp. nov., holotype, ♂ (BMNH). 146. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 147. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 148. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 149. Same, ♂ (SNSD). 150. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 151. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 152. M. nepenthephilus sp. nov., holotype, ♂ (NME). 153. Same, paratype, ♀ (DTC). 154. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 155. Same, paratype, ♀ (BMNH). 156. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122181" httpUri="https://zenodo.org/record/4122181/files/figure.png" pageId="82" pageNumber="83">151</figureCitation>
,
<figureCitation id="0AA52A659D4E9209FD52FC00CAA62BBA" box="[705,752,958,984]" captionStart="Figs 226241" captionStartId="147.[189,241,1681,1707]" captionTargetBox="[391,1192,268,1647]" captionTargetId="figure-170@147.[391,1196,265,1656]" captionTargetPageId="147" captionText="Figs 226241. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 226. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 227. M. lobaticeps Frey, 1969, ♂ (DTC). 228. M. longipes sp. nov., holotype, ♂ (BMNH). 229. M. luniceps Fairmaire, 1883, ♂ (BMNH). 230. M. mailu sp. nov., holotype, ♂ (NHMB). 231. M. maleo sp. nov., holotype, ♂ (IRSN). 232. M. merdeka sp. nov., holotype, ♂ (BMNH). 233. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 234. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 235. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 236. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 237. M. nepenthephilus sp. nov., holotype, ♂ (NME). 238. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 239. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 240. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 241. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122193" httpUri="https://zenodo.org/record/4122193/files/figure.png" pageId="82" pageNumber="83">236</figureCitation>
,
<figureCitation id="0AA52A659D4E9209FD6FFC00CB7D2BBA" box="[764,811,958,984]" captionStart="Figs 317332" captionStartId="154.[189,241,1681,1707]" captionTargetBox="[401,1187,265,1655]" captionTargetId="figure-172@154.[401,1187,265,1669]" captionTargetPageId="154" captionText="Figs 317332. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 317. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 318. M. lobaticeps Frey, 1969, ♂ (DTC). 319. M. longipes sp. nov., holotype, ♂ (BMNH). 320. M. luniceps Fairmaire, 1883, ♂ (BMNH). 321. M. mailu sp. nov., holotype, ♂ (NHMB). 322. M. maleo sp. nov., holotype, ♂ (IRSN). 323. M. merdeka sp. nov., holotype, ♂ (BMNH). 324. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 325. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 326. M. muticusArrow, 1941, syntype, ♂ (BMNH). 327. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 328. M. nepenthephilus sp. nov., holotype, ♂ (NME). 329. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 330. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 331. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 332. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122210" httpUri="https://zenodo.org/record/4122210/files/figure.png" pageId="82" pageNumber="83">327</figureCitation>
, 400, 496, 676678
</paragraph>
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<emphasis id="A0EAEAF29D4E9209FF2EFBBEC93A2C7B" bold="true" box="[189,364,1023,1050]" pageId="82" pageNumber="83">Type material</emphasis>
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<typeStatus id="4D2588429D4E9209FF79FB90C90C2C2A" box="[234,346,1070,1096]" pageId="82" pageNumber="83" type="syntype">Syntypes</typeStatus>
</emphasis>
</paragraph>
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<collectingCountry id="EA8976709D4E9209FF2EFBEFC90D2C09" box="[189,347,1105,1131]" name="Indonesia" pageId="82" pageNumber="83">INDONESIA</collectingCountry>
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; “
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[p, label circular, red frame] //
<typeStatus id="4D2588429D4E9209FCE4FBEFCB812C09" box="[887,983,1105,1131]" pageId="82" pageNumber="83" type="syntype">Syntype</typeStatus>
[p, label circular, blue frame] // N.
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:
<location id="9741603B9D4E9209FD9CFBCACB442CEC" LSID="urn:lsid:plazi:treatment:1A3787F69D4E9208FDCFFC24CAF32AE8:9741603B9D4E9209FD9CFBCACB442CEC" box="[527,786,1140,1167]" country="Indonesia" name="Waigeu. Camp Nok." pageId="82" pageNumber="83">Waigeu. Camp Nok.</location>
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<date id="E62010209D4E9209FC1DFBCACBBF2CEC" box="[910,1001,1140,1166]" pageId="82" pageNumber="83" value="1938-04">
<collectingDate id="F664E9C89D4E9209FC1DFBCACBBF2CEC" box="[910,1001,1140,1166]" pageId="82" pageNumber="83" value="1938-04">iv.1938</collectingDate>
</date>
. L.E.Cheesman. B.M.1938-593. [p] //
<taxonomicName id="559E4D639D4E9209FF64FB26CAD92CD0" authority="Arrow Co-type" authorityName="Arrow Co-type" authorityYear="1942" box="[247,655,1176,1202]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="A0EAEAF29D4E9209FF64FB26C9862CD3" box="[247,464,1176,1202]" italics="true" pageId="82" pageNumber="83">Maechidius nanus</emphasis>
Arrow Co-type
</taxonomicName>
[h]”;
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; “
<typeStatus id="4D2588429D4E9209FC6DFB26CC372CD0" box="[1022,1121,1176,1202]" pageId="82" pageNumber="83" type="syntype">Syntype</typeStatus>
[p, label circular, blue frame] // N.
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:
<location id="9741603B9D4E9209FDE4FB05CB3E2CB7" LSID="urn:lsid:plazi:treatment:1A3787F69D4E9208FDCFFC24CAF32AE8:9741603B9D4E9209FDE4FB05CB3E2CB7" box="[631,872,1211,1237]" country="Guinea-Bissau" name="Waigeu. Camp Nok." pageId="82" pageNumber="83">Waigeu. Camp Nok.</location>
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<date id="E62010209D4E9209FC40FB05CC7B2CB7" box="[979,1069,1211,1237]" pageId="82" pageNumber="83" value="1938-04">
<collectingDate id="F664E9C89D4E9209FC40FB05CC7B2CB7" box="[979,1069,1211,1237]" pageId="82" pageNumber="83" value="1938-04">iv.1938</collectingDate>
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. L.E.Cheesman. B.M.1938- 593. [p]”;
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</materialsCitation>
<materialsCitation id="22F63CBD9D4E9209FE3FFB60CA912D7E" ID-GBIF-Occurrence="2898878322" collectingDate="1938-04" collectionCode="BMNH" collectorName="L. E. Cheesman." country="Guinea-Bissau" elevation="762" location="DUTCH NEW GUINEA" pageId="82" pageNumber="83" specimenCount="152" typeStatus="syntype">
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; “N.
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:
<location id="9741603B9D4E9209FC28FB60CCFA2C9A" LSID="urn:lsid:plazi:treatment:1A3787F69D4E9208FDCFFC24CAF32AE8:9741603B9D4E9209FC28FB60CCFA2C9A" box="[955,1196,1246,1272]" country="Guinea-Bissau" name="Waigeu. Camp Nok." pageId="82" pageNumber="83">Waigeu. Camp Nok.</location>
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<date id="E62010209D4E9209FA8BFB60CD252C9A" box="[1304,1395,1246,1272]" pageId="82" pageNumber="83" value="1938-04">
<collectingDate id="F664E9C89D4E9209FA8BFB60CD252C9A" box="[1304,1395,1246,1272]" pageId="82" pageNumber="83" value="1938-04">iv.1938</collectingDate>
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.
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B.M.1938-593. [p]”;
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</materialsCitation>
<materialsCitation id="22F63CBD9D4E9209FD4DFABFC94F2D00" ID-GBIF-Occurrence="2898878339" collectingDate="1938-04" collectionCode="BMNH" collectorName="L. E. Cheesman." country="Guinea-Bissau" elevation="762" location="DUTCH NEW GUINEA" pageId="82" pageNumber="83" specimenCount="2" specimenCount-male="2" typeStatus="syntype">
<specimenCount id="8498FD699D4E9209FD4DFABFCB5D2D7E" box="[734,779,1281,1308]" pageId="82" pageNumber="83" type="male">1 ♂</specimenCount>
; “
<specimenCount id="8498FD699D4E9209FCB6FABFCB6D2D79" box="[805,827,1281,1307]" pageId="82" pageNumber="83" type="male"></specimenCount>
[p] // N.
<location id="9741603B9D4E9209FC07FABFCC962D79" LSID="urn:lsid:plazi:treatment:1A3787F69D4E9208FDCFFC24CAF32AE8:9741603B9D4E9209FC07FABFCC962D79" box="[916,1216,1281,1308]" country="Guinea-Bissau" name="DUTCH NEW GUINEA" pageId="82" pageNumber="83">DUTCH NEW GUINEA</location>
:
<location id="9741603B9D4E9209FB59FABFC8A12D5C" LSID="urn:lsid:plazi:treatment:1A3787F69D4E9208FDCFFC24CAF32AE8:9741603B9D4E9209FB59FABFC8A12D5C" country="Guinea-Bissau" name="Waigeu. Camp Nok." pageId="82" pageNumber="83">Waigeu. Camp Nok.</location>
<quantity id="55669B059D4E9209FE90FA9AC9092D5D" box="[259,351,1316,1343]" metricMagnitude="2" metricUnit="m" metricValue="7.62" pageId="82" pageNumber="83" unit="ft" value="2500.0">
<elevation id="19B3D1D39D4E9209FE90FA9AC9092D5D" box="[259,351,1316,1343]" metricMagnitude="2" metricUnit="m" metricValue="7.62" pageId="82" pageNumber="83" unit="ft" value="2500.0">2,500ft.</elevation>
</quantity>
<date id="E62010209D4E9209FEF9FA9AC9932D5C" box="[362,453,1316,1342]" pageId="82" pageNumber="83" value="1938-04">
<collectingDate id="F664E9C89D4E9209FEF9FA9AC9932D5C" box="[362,453,1316,1342]" pageId="82" pageNumber="83" value="1938-04">iv.1938</collectingDate>
</date>
.
<collectorName id="3F6B53369D4E9209FE46FA9ACADF2D5C" box="[469,649,1316,1342]" pageId="82" pageNumber="83">L.E.Cheesman.</collectorName>
B.M.1938-593. [p] //
<taxonomicName id="559E4D639D4E9209FC1CFA9ACD652D5D" authority="Arrow Co-Type" authorityName="Arrow Co-type" authorityYear="1942" box="[911,1331,1316,1343]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="A0EAEAF29D4E9209FC1CFA9ACC3C2D5C" box="[911,1130,1316,1342]" italics="true" pageId="82" pageNumber="83">
<location id="9741603B9D4E9209FC1CFA9ACC4E2D5C" LSID="urn:lsid:plazi:treatment:1A3787F69D4E9208FDCFFC24CAF32AE8:9741603B9D4E9209FC1CFA9ACC4E2D5C" box="[911,1048,1316,1342]" country="Guinea-Bissau" name="Maechidius" pageId="82" pageNumber="83">Maechidius</location>
nanus
</emphasis>
<location id="9741603B9D4E9209FBE7FA9BCD652D5D" LSID="urn:lsid:plazi:treatment:1A3787F69D4E9208FDCFFC24CAF32AE8:9741603B9D4E9209FBE7FA9BCD652D5D" box="[1140,1331,1317,1343]" country="Guinea-Bissau" name="Arrow Co-Type" pageId="82" pageNumber="83">Arrow Co-Type</location>
</taxonomicName>
[h]”;
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</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D4E9209FF2EFA37CB542D97" pageId="82" pageNumber="83" type="discussion">
<paragraph id="922136E09D4E9209FF2EFA37C9792DC1" blockId="82.[189,303,1417,1443]" box="[189,303,1417,1443]" pageId="82" pageNumber="83">
<heading id="C969818C9D4E9209FF2EFA37C9792DC1" bold="true" box="[189,303,1417,1443]" fontSize="11" level="3" pageId="82" pageNumber="83" reason="3">
<emphasis id="A0EAEAF29D4E9209FF2EFA37C9792DC1" bold="true" box="[189,303,1417,1443]" pageId="82" pageNumber="83">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D4E9209FF2EFA06CB542D97" blockId="82.[189,1399,1464,1526]" pageId="82" pageNumber="83">
There are several male
<typeStatus id="4D2588429D4E9209FE5AFA06CA662DB0" box="[457,560,1464,1490]" pageId="82" pageNumber="83" type="syntype">syntypes</typeStatus>
from the same locality available for this species. A
<typeStatus id="4D2588429D4E9209FB11FA06CCB82DB0" box="[1154,1262,1464,1490]" pageId="82" pageNumber="83" type="lectotype">lectotype</typeStatus>
is therefore not designated since there is no risk of confusion.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D4E9208FF2EF9A2CAF32AE8" lastPageId="83" lastPageNumber="84" pageId="82" pageNumber="83" type="description">
<paragraph id="922136E09D4E9209FF2EF9A2C93D2E54" blockId="82.[189,363,1564,1590]" box="[189,363,1564,1590]" pageId="82" pageNumber="83">
<heading id="C969818C9D4E9209FF2EF9A2C93D2E54" bold="true" box="[189,363,1564,1590]" fontSize="11" level="3" pageId="82" pageNumber="83" reason="3">
<emphasis id="A0EAEAF29D4E9209FF2EF9A2C93D2E54" bold="true" box="[189,363,1564,1590]" pageId="82" pageNumber="83">Redescription</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D4E9208FF2EF9F5CB6D2A78" blockId="82.[189,1399,1611,2025]" lastBlockId="83.[189,1399,265,539]" lastPageId="83" lastPageNumber="84" pageId="82" pageNumber="83">
Male labroclypeus (
<figureCitation id="0AA52A659D4E9209FE31F9F5CA522E04" box="[418,516,1611,1638]" captionStart="Figs 145156" captionStartId="141.[189,241,1787,1813]" captionTargetBox="[328,1254,265,1740]" captionTargetId="figure-124@141.[328,1260,265,1749]" captionTargetPageId="141" captionText="Figs 145156. Papuan Maechidius Macleay, 1819, head, dorsal view. 145. M. merdeka sp. nov., holotype, ♂ (BMNH). 146. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 147. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 148. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 149. Same, ♂ (SNSD). 150. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 151. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 152. M. nepenthephilus sp. nov., holotype, ♂ (NME). 153. Same, paratype, ♀ (DTC). 154. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 155. Same, paratype, ♀ (BMNH). 156. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122181" httpUri="https://zenodo.org/record/4122181/files/figure.png" pageId="82" pageNumber="83">Fig. 151</figureCitation>
) broadly V-shaped emarginate on anterior margin, its lateral margins sinuous in both dorsal and lateral views. Head dorsal punctures circular, variably large, shallow, annular, with shagreened background. A long suberect seta rises from anterior margin of each puncture. Intervening spaces glossy, larger on labroclypeus, generally much narrower than punctures on frons. Pronotum broadly emarginate on anterior margin, anterolateral angles slightly protruding, basal margin broadly rounded. Lateral margin of pronotum broadly rounded in dorsal view, with large crenulae all along (
<figureCitation id="0AA52A659D4E9209FF55F8A0C9642F5B" box="[198,306,1822,1849]" captionStart="Figs 226241" captionStartId="147.[189,241,1681,1707]" captionTargetBox="[391,1192,268,1647]" captionTargetId="figure-170@147.[391,1196,265,1656]" captionTargetPageId="147" captionText="Figs 226241. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 226. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 227. M. lobaticeps Frey, 1969, ♂ (DTC). 228. M. longipes sp. nov., holotype, ♂ (BMNH). 229. M. luniceps Fairmaire, 1883, ♂ (BMNH). 230. M. mailu sp. nov., holotype, ♂ (NHMB). 231. M. maleo sp. nov., holotype, ♂ (IRSN). 232. M. merdeka sp. nov., holotype, ♂ (BMNH). 233. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 234. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 235. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 236. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 237. M. nepenthephilus sp. nov., holotype, ♂ (NME). 238. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 239. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 240. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 241. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122193" httpUri="https://zenodo.org/record/4122193/files/figure.png" pageId="82" pageNumber="83">Fig. 236</figureCitation>
). A moderately long suberect seta present between every two crenulae. Lateral margin of pronotum nearly straight in lateral view. Hypomeron separated from prosternum by low nearly straight carina, with moderately long setae on its anterolateral margin opposite to compound eye. Antennal pocket shallow. Pronotal punctures circular, annular, large, shallow. Intervening spaces glossy, variably large. Minute appressed seta rises from anterior margin of each puncture, not surpassing midlength of corresponding puncture. Elytral punctures irregularly ovoid, annular, shallow, arranged in double longitudinal rows (
<figureCitation id="0AA52A659D4F9208FE0CFEB7CA5E2941" box="[415,520,265,291]" captionStart="Figs 317332" captionStartId="154.[189,241,1681,1707]" captionTargetBox="[401,1187,265,1655]" captionTargetId="figure-172@154.[401,1187,265,1669]" captionTargetPageId="154" captionText="Figs 317332. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 317. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 318. M. lobaticeps Frey, 1969, ♂ (DTC). 319. M. longipes sp. nov., holotype, ♂ (BMNH). 320. M. luniceps Fairmaire, 1883, ♂ (BMNH). 321. M. mailu sp. nov., holotype, ♂ (NHMB). 322. M. maleo sp. nov., holotype, ♂ (IRSN). 323. M. merdeka sp. nov., holotype, ♂ (BMNH). 324. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 325. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 326. M. muticusArrow, 1941, syntype, ♂ (BMNH). 327. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 328. M. nepenthephilus sp. nov., holotype, ♂ (NME). 329. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 330. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 331. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 332. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122210" httpUri="https://zenodo.org/record/4122210/files/figure.png" pageId="83" pageNumber="84">Fig. 327</figureCitation>
). Intervening spaces glossy, generally larger than punctures. Elytral setae minute, surface looks glabrous under 50 × magnification; minute seta rises from anterior margin of each puncture, not surpassing midlength of corresponding puncture. Pygidium in both sexes flattened dorsally, with large circular to ovoid shallow annular punctures (
<figureCitation id="0AA52A659D4F9208FC3DFECDCC4329EC" box="[942,1045,371,398]" captionStart="Figs487503" captionStartId="165.[189,241,1681,1707]" captionTargetBox="[422,1160,265,1655]" captionTargetId="figure-171@165.[421,1167,265,1658]" captionTargetPageId="165" captionText="Figs487503.Papuan andWallaceanMaechidius Macleay, 1819,pygidium, dorsalview.487.M.lobaticeps Frey, 1969, ♂ (DTC). 488. M. longipes sp. nov., holotype, ♂ (BMNH). 489. M. luniceps Fairmaire, 1883, ♂ (BMNH). 490. M. mailu sp. nov., holotype, ♂ (NHMB). 491. M. maleo sp. nov., holotype, ♂ (IRSN). 492. M. merdeka sp. nov., holotype, ♂ (BMNH). 493. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 494. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 495. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 496. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 497. M. nepenthephilus sp. nov., holotype, ♂ (NME). 498. M. opatroidesArrow, 1941 comb. rest., lectotype, ♀ (BMNH). 499. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 500. Same, paratype, ♀ (BMNH). 501. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 502. Same, different aspect. 503. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122226" httpUri="https://zenodo.org/record/4122226/files/figure.png" pageId="83" pageNumber="84">Fig. 496</figureCitation>
). Intervening spaces generally smaller than punctures, glossy. Setae inconspicuous and short in anterior part, becoming longer than corresponding punctures in distal part. Protibia in both sexes widened distally, with three distal teeth on external margin, of which two distal ones larger and stronger acute (Fig. 400). Protibia without terminal spur in both sexes. Male aedeagus as in
<figureCitation id="0AA52A659D4F9208FD00FDBECB612A78" box="[659,823,512,538]" captionStart="Figs 667681" captionStartId="176.[189,241,1892,1918]" captionTargetBox="[444,1142,267,1860]" captionTargetId="figure-72@176.[419,1168,265,1868]" captionTargetPageId="176" captionText="Figs 667681. Papuan Maechidius Macleay, 1819, aedeagus, different aspects. 667 669. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 670672. M. milneanus Heller, 1914, ♂ (SNSD). 673675. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 676678. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 679-681. M. nepenthephilus sp. nov., holotype, ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122246" httpUri="https://zenodo.org/record/4122246/files/figure.png" pageId="83" pageNumber="84">Figs 676678</figureCitation>
.
</paragraph>
<paragraph id="922136E09D4F9208FF2EFDFFC9E62A39" blockId="83.[189,432,577,603]" box="[189,432,577,603]" pageId="83" pageNumber="84">
<heading id="C969818C9D4F9208FF2EFDFFC9E62A39" bold="true" box="[189,432,577,603]" fontSize="11" level="3" pageId="83" pageNumber="84" reason="3">
<emphasis id="A0EAEAF29D4F9208FF2EFDFFC9E62A39" bold="true" box="[189,432,577,603]" pageId="83" pageNumber="84">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D4F9208FF2EFDCECAF32AE8" blockId="83.[189,677,624,650]" box="[189,677,624,650]" pageId="83" pageNumber="84">Lamellae of female antennal club shorter.</paragraph>
</subSubSection>
</treatment>
</document>