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<document id="CD1B8E227227BB28E32D7B9691D035B7" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D3C927AFDEBFDD3CBE02D90" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius brocki Telnov 2020, sp. nov." docType="treatment" docUuid="39227DD9-2D93-4AF8-BFF8-050C6B0EF97B" docUuidSource="ZooBank" docVersion="10" lastPageNumber="34" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="33" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="A836EF7F0510BA1F8CD6763E52EC5BD4">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
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<mods:name id="7CA072FE15493E097EFD1A892E87D433" type="personal">
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<mods:namePart id="9C9A5F3BC0AC8D3A9901019609B30C0C">Telnov, Dmitry</mods:namePart>
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<mods:title id="87A90199C58038BDFFB8258C1CC594E8">European Journal of Taxonomy</mods:title>
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<mods:date id="57AB6320EE68271D8F26BA14C42BB8A1">2020</mods:date>
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<mods:number id="29AF6E2B73DD5011AF5CDA22A763A5D3">2020-10-19</mods:number>
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<mods:identifier id="5790974517EA8B4D20B6D3DEA654CC25" type="DOI">10.5852/ejt.2020.721.1127</mods:identifier>
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<mods:identifier id="8A15A5C5A6D7243BB685642CCC0838D4" type="Zenodo-Dep">4122118</mods:identifier>
<mods:identifier id="DD0CF5406594B979162EB99F5A3260D2" type="ZooBank">89E62EF8-2E45-4C59-94B7-6A5603E8939B</mods:identifier>
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<treatment id="1A3787F69D3C927AFDEBFDD3CBE02D90" ID-DOI="http://doi.org/10.5281/zenodo.4344307" ID-GBIF-Taxon="168648150" ID-Zenodo-Dep="4344307" LSID="urn:lsid:zoobank.org:act:39227DD9-2D93-4AF8-BFF8-050C6B0EF97B" httpUri="http://treatment.plazi.org/id/1A3787F69D3C927AFDEBFDD3CBE02D90" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
<subSubSection id="DA84656B9D3C927BFDEBFDD3CBEA2AEA" box="[632,956,621,648]" pageId="32" pageNumber="33" type="nomenclature">
<paragraph id="922136E09D3C927BFDEBFDD3CBEA2AEA" blockId="32.[365,1223,621,718]" box="[632,956,621,648]" pageId="32" pageNumber="33">
<heading id="C969818C9D3C927BFDEBFDD3CBEA2AEA" box="[632,956,621,648]" centered="true" fontSize="11" level="2" pageId="32" pageNumber="33" reason="2">
<taxonomicName id="559E4D639D3C927BFDEBFDD3CB0C2AEA" ID-CoL="7X3WF" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[632,858,621,648]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="brocki" status="sp. nov.">
<emphasis id="A0EAEAF29D3C927BFDEBFDD3CB0C2AEA" bold="true" box="[632,858,621,648]" italics="true" pageId="32" pageNumber="33">Maechidius brocki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D3C927BFCF2FDD0CBEA2AEA" box="[865,956,622,648]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D3C927BFEFEFD2ECC1B2AAF" pageId="32" pageNumber="33" type="description">
<paragraph id="922136E09D3C927BFEFEFD2ECC912AC9" blockId="32.[365,1223,621,718]" box="[365,1223,656,683]" pageId="32" pageNumber="33">
<uri id="E60F3AE29D3C927BFEFEFD2ECC912AC9" box="[365,1223,656,683]" pageId="32" pageNumber="33">
urn:lsid:zoobank.org:act:
<uuid id="E6380C359D3C927BFD1DFD2ECC912AC9" box="[654,1223,656,683]" pageId="32" pageNumber="33">39227DD9-2D93-4AF8-BFF8-050C6B0EF97B</uuid>
</uri>
</paragraph>
<paragraph id="922136E09D3C927BFE74FD0DCC1B2AAF" blockId="32.[365,1223,621,718]" box="[487,1101,691,718]" pageId="32" pageNumber="33">
<figureCitation id="0AA52A659D3C927BFE74FD0DCA172AAF" box="[487,577,691,717]" captionStart="Figs 28–30" captionStartId="31.[189,241,1928,1954]" captionTargetBox="[347,1240,269,1901]" captionTargetId="figure-52@31.[345,1242,265,1911]" captionTargetPageId="31" captionText="Figs 28–30. Papuan and Wallacean Maechidius Macleay, 1819, habitus, dorsal view. 28. M. boessnecki sp. nov., holotype, ♂ (SNSD). 29. M. brocki sp. nov., holotype, ♂ (BMNH). 30. M. caperatus sp. nov., holotype, ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122136" httpUri="https://zenodo.org/record/4122136/files/figure.png" pageId="32" pageNumber="33">Figs 29</figureCitation>
,
<figureCitation id="0AA52A659D3C927BFDDEFD0DCAE32AAF" box="[589,693,691,717]" captionStart="Figs 109–120" captionStartId="138.[189,241,1750,1776]" captionTargetBox="[365,1214,275,1722]" captionTargetId="figure-15@138.[365,1222,265,1730]" captionTargetPageId="138" captionText="Figs 109–120. Papuan andWallacean MaechidiusMacleay, 1819,head, dorsal (109–119) and frontal (120) views. 109. M. brocki sp. nov., holotype, ♂ (BMNH). 110. Same, paratype, ♀ (BMNH). 111. M. ciliatus sp. nov., holotype, ♂ (IECA). 112. M. crypticus sp. nov., paratype, ♂ (BMNH). 113. Same, paratype, ♀ (BMNH). 114. M. dani sp. nov., holotype, ♂ (IECA). 115. M. deltouri sp. nov., holotype, ♂ (BMNH). 116. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 117. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 118. M. esau Heller, 1914, lectotype, ♂ (SNSD). 119. Same, ♂ (lectotype of M. setosus Moser, 1920) (ZMHB). 120. M. gressitti Frey, 1969, holotype, ♂ (photograph courtesy BPBM) (BPBM). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122175" httpUri="https://zenodo.org/record/4122175/files/figure.png" pageId="32" pageNumber="33">109–110</figureCitation>
,
<figureCitation id="0AA52A659D3C927BFD52FD0DCAA62AAF" box="[705,752,691,717]" captionStart="Figs 193–207" captionStartId="145.[189,241,1716,1742]" captionTargetBox="[385,1203,267,1681]" captionTargetId="figure-159@145.[385,1203,265,1694]" captionTargetPageId="145" captionText="Figs 193–207. Papuan and Wallacean Maechidius Macleay, 1819, head (193–196) and pronotum (197– 207), dorsal view. 193. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 194. Same, ♂ (SNSD). 195. M. yamdena sp. nov., paratype, ♂ (BMNH). 196. Same, paratype, ♀ (BMNH). 197. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 198. M. aiyura sp. nov., holotype, ♂ (BMNH). 199. M. alesbezdeki sp. nov., holotype, ♂ (NME). 200. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 201. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 202. M. awu sp. nov., holotype, ♂ (IECA). 203. M. babyrousa sp. nov., holotype, ♂ (BMNH). 204. M. bintang sp. nov., holotype, ♂ (NME). 205. M. boessnecki sp. nov., holotype, ♂ (SNSD). 206. M. brocki sp. nov., holotype, ♂ (BMNH). 207. M. crypticus sp. nov., paratype, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122189" httpUri="https://zenodo.org/record/4122189/files/figure.png" pageId="32" pageNumber="33">206</figureCitation>
,
<figureCitation id="0AA52A659D3C927BFD68FD0DCB7C2AAF" box="[763,810,691,717]" captionStart="Figs 286–301" captionStartId="152.[189,241,1716,1742]" captionTargetBox="[364,1224,266,1689]" captionTargetId="figure-172@152.[364,1224,266,1695]" captionTargetPageId="152" captionText="Figs 286–301. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 286. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 287. M. aiyura sp. nov., holotype, ♂ (BMNH). 288. M. alesbezdeki sp. nov., holotype, ♂ (NME). 289. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 290. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 291. M. paupianus Heller, 1910, ♂ (BMNH). 292. M. awu sp. nov., holotype, ♂ (IECA). 293. M. babyrousa sp. nov., holotype, ♂ (BMNH). 294. M. bintang sp. nov., holotype, ♂ (NME). 295. M. boessnecki sp. nov., holotype, ♂ (SNSD). 296. M. brocki sp. nov., holotype, ♂ (BMNH). 297. M. caperatus sp. nov., holotype, ♂ (NME). 298. M. ciliatus sp. nov., holotype, ♂ (IECA). 299. M. crypticus sp. nov., paratype, ♀ (BMNH). 300. M. dani sp. nov., holotype, ♂ (IECA). 301. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122206" httpUri="https://zenodo.org/record/4122206/files/figure.png" pageId="32" pageNumber="33">296</figureCitation>
, 368, 462, 539, 601–603
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D3C927BFF2EFD4ACB1C2B89" pageId="32" pageNumber="33" type="diagnosis">
<paragraph id="922136E09D3C927BFF2EFD4AC99F2B6C" blockId="32.[189,457,756,782]" box="[189,457,756,782]" pageId="32" pageNumber="33">
<heading id="C969818C9D3C927BFF2EFD4AC99F2B6C" bold="true" box="[189,457,756,782]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
<emphasis id="A0EAEAF29D3C927BFF2EFD4AC99F2B6C" bold="true" box="[189,457,756,782]" pageId="32" pageNumber="33">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D3C927BFF2EFC9CCB1C2B89" blockId="32.[189,1399,802,1003]" pageId="32" pageNumber="33">
This species most closely resembles
<taxonomicName id="559E4D639D3C927BFDE8FC9CCC5C2B5E" authority="Arrow, 1941" authorityName="Arrow" authorityYear="1941" box="[635,1034,802,828]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="hirtipes">
<emphasis id="A0EAEAF29D3C927BFDE8FC9CCB3E2B5E" box="[635,872,802,828]" italics="true" pageId="32" pageNumber="33">Maechidius hirtipes</emphasis>
<bibRefCitation id="F60F4B119D3C927BFCE1FC9CCC5C2B5E" author="Arrow G. E." box="[882,1034,802,828]" pageId="32" pageNumber="33" pagination="448 - 464" refId="ref97558" refString="Arrow G. E. 1941. XXIX. - Some undescribed species of melolonthid Coleoptera from Western New Guinea and the adjacent islands of Waigeu and Japen. Annals and Magazine of Natural History, Series 11 7 (41): 448 - 464. https: // doi. org / 10.1080 / 03745481.1941.9727945" type="journal article" year="1941">Arrow, 1941</bibRefCitation>
</taxonomicName>
(Papuan Peninsula, East New
<collectingCountry id="EA8976709D3C927BFF2EFCFBC9422B02" box="[189,276,837,864]" name="Guinea-Bissau" pageId="32" pageNumber="33">Guinea</collectingCountry>
) and
<taxonomicName id="559E4D639D3C927BFEC5FCF8CAF52B02" authority="Heller, 1895" authorityName=", Hell. Determined" authorityYear="1895" box="[342,675,837,864]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="subcostatus">
<emphasis id="A0EAEAF29D3C927BFEC5FCF8CA5E2B3D" box="[342,520,837,863]" italics="true" pageId="32" pageNumber="33">M. subcostatus</emphasis>
<bibRefCitation id="F60F4B119D3C927BFD83FCFBCAF52B02" author="Heller K. M." box="[528,675,837,864]" pageId="32" pageNumber="33" pagination="1 - 17" refId="ref98131" refString="Heller K. M. 1895. Erster Beitrag zur Papuaniscen [sic!] Kaferfauna. Abhandlungen und Berichte des Koniglichen Zoologischen und Anthropologisch-Ethnographischen Museums zu Dresden 1894 / 1895 (16): 1 - 17." type="journal article" year="1895">Heller, 1895</bibRefCitation>
</taxonomicName>
(Finisterre Mountains, East New
<collectingCountry id="EA8976709D3C927BFBA6FCFBCCD82B02" box="[1077,1166,837,864]" name="Guinea-Bissau" pageId="32" pageNumber="33">Guinea</collectingCountry>
), but readily differs from both primarily in the shape of the male genital organs.Additionally, in
<taxonomicName id="559E4D639D3C927BFBB4FCD7CCF02BE0" authorityName="Arrow" authorityYear="1941" box="[1063,1190,872,898]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="hirtipes">
<emphasis id="A0EAEAF29D3C927BFBB4FCD7CCF02BE0" box="[1063,1190,872,898]" italics="true" pageId="32" pageNumber="33">M. hirtipes</emphasis>
</taxonomicName>
the lateral margin of the pronotum is denticulate (tuberculate in
<taxonomicName id="559E4D639D3C927BFD61FC35CB312BC7" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[754,871,907,933]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="brocki" status="sp. nov.">
<emphasis id="A0EAEAF29D3C927BFD61FC35CB312BC7" box="[754,871,907,933]" italics="true" pageId="32" pageNumber="33">M. brocki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D3C927BFCE0FC35CB852BC7" box="[883,979,907,933]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
) and slightly emarginate (strongly emarginate in
<taxonomicName id="559E4D639D3C927BFEF5FC10C98E2BA5" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[358,472,941,967]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="brocki" status="sp. nov.">
<emphasis id="A0EAEAF29D3C927BFEF5FC10C98E2BA5" box="[358,472,941,967]" italics="true" pageId="32" pageNumber="33">M. brocki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D3C927BFE73FC10CA6A2BAA" box="[480,572,942,968]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
) and the basal metatarsomere has a brush of very long, dense sinuous setae on the inner margin (absent in
<taxonomicName id="559E4D639D3C927BFDF5FC6FCA8E2B88" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[614,728,976,1002]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="brocki" status="sp. nov.">
<emphasis id="A0EAEAF29D3C927BFDF5FC6FCA8E2B88" box="[614,728,976,1002]" italics="true" pageId="32" pageNumber="33">M. brocki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D3C927BFD4CFC6FCB6B2B89" box="[735,829,977,1003]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
).
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D3C927BFF2EFBACCC612C1E" pageId="32" pageNumber="33" type="etymology">
<paragraph id="922136E09D3C927BFF2EFBACC9122C4E" blockId="32.[189,324,1042,1068]" box="[189,324,1042,1068]" pageId="32" pageNumber="33">
<heading id="C969818C9D3C927BFF2EFBACC9122C4E" bold="true" box="[189,324,1042,1068]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
<emphasis id="A0EAEAF29D3C927BFF2EFBACC9122C4E" bold="true" box="[189,324,1042,1068]" pageId="32" pageNumber="33">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D3C927BFF2EFBFECC612C1E" blockId="32.[189,1398,1088,1149]" pageId="32" pageNumber="33">
Patronymic. This species is dedicated to Paul Brock (Brockenhurst,
<collectingCountry id="EA8976709D3C927BFC59FBFECCC22C39" box="[970,1172,1088,1115]" name="United Kingdom" pageId="32" pageNumber="33">United Kingdom</collectingCountry>
), a famous phasmid expert, experienced macrophotographer, respected colleague and my friend.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D3C927BFF2EFB1ACB442DBE" pageId="32" pageNumber="33" type="materials_examined">
<paragraph id="922136E09D3C927BFF2EFB1AC93A2CDF" blockId="32.[189,364,1187,1214]" box="[189,364,1187,1214]" pageId="32" pageNumber="33">
<heading id="C969818C9D3C927BFF2EFB1AC93A2CDF" bold="true" box="[189,364,1187,1214]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
<emphasis id="A0EAEAF29D3C927BFF2EFB1AC93A2CDF" bold="true" box="[189,364,1187,1214]" pageId="32" pageNumber="33">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D3C927BFF79FB6CC90A2C8E" blockId="32.[189,1399,1234,1330]" box="[234,348,1234,1260]" pageId="32" pageNumber="33">
<emphasis id="A0EAEAF29D3C927BFF79FB6CC90A2C8E" bold="true" box="[234,348,1234,1260]" pageId="32" pageNumber="33">
<typeStatus id="4D2588429D3C927BFF79FB6CC90A2C8E" box="[234,348,1234,1260]" pageId="32" pageNumber="33" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="922136E09D3C927BFF2EFB4BC9C02D50" blockId="32.[189,1399,1234,1330]" pageId="32" pageNumber="33">
<materialsCitation id="22F63CBD9D3C927BFF2EFB4BC9C02D50" ID-GBIF-Occurrence="2898878478" collectingDate="1933-12" collectionCode="PAPUA, BMNH" collectorName="L. E. Cheesman." country="Papua New Guinea" elevation="1219" location="PAPUA NEW GUINEA" pageId="32" pageNumber="33" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
<collectingCountry id="EA8976709D3C927BFF2EFB4BC9882D6D" box="[189,478,1269,1295]" name="Papua New Guinea" pageId="32" pageNumber="33">PAPUA NEW GUINEA</collectingCountry>
•
<specimenCount id="8498FD699D3C927BFE68FB4BCA452D6D" box="[507,531,1269,1295]" pageId="32" pageNumber="33" type="male">♂</specimenCount>
; “
<specimenCount id="8498FD699D3C927BFDA3FB4BCA102D6D" box="[560,582,1269,1295]" pageId="32" pageNumber="33" type="male">♂</specimenCount>
[p] //
<collectionCode id="F48FAE259D3C927BFD19FB4BCABA2D6D" box="[650,748,1269,1295]" pageId="32" pageNumber="33">PAPUA</collectionCode>
: Mafulu.
<quantity id="55669B059D3C927BFCF2FB4BCB912D6D" box="[865,967,1269,1295]" metricMagnitude="3" metricUnit="m" metricValue="1.2192" pageId="32" pageNumber="33" unit="ft" value="4000.0">
<elevation id="19B3D1D39D3C927BFCF2FB4BCB912D6D" box="[865,967,1269,1295]" metricMagnitude="3" metricUnit="m" metricValue="1.2192" pageId="32" pageNumber="33" unit="ft" value="4000.0">4,000 ft.</elevation>
</quantity>
<date id="E62010209D3C927BFC55FB4BCC7C2D6D" box="[966,1066,1269,1295]" pageId="32" pageNumber="33" value="1933-12">
<collectingDate id="F664E9C89D3C927BFC55FB4BCC7C2D6D" box="[966,1066,1269,1295]" pageId="32" pageNumber="33" value="1933-12">xii.1933</collectingDate>
</date>
.
<collectorName id="3F6B53369D3C927BFBABFB4BCCBA2D6D" box="[1080,1260,1269,1295]" pageId="32" pageNumber="33">L.E.Cheesman.</collectorName>
B.M.1933- 321. [p]”;
<collectionCode id="F48FAE259D3C927BFEA5FAA6C9C42D50" box="[310,402,1304,1330]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="32" pageNumber="33">BMNH</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="922136E09D3C927BFF79FAE7CA452D16" blockId="32.[189,1399,1369,1500]" box="[234,531,1369,1396]" pageId="32" pageNumber="33">
<emphasis id="A0EAEAF29D3C927BFF79FAE7C9312D11" bold="true" box="[234,359,1369,1395]" pageId="32" pageNumber="33">
<typeStatus id="4D2588429D3C927BFF79FAE7C9312D11" box="[234,359,1369,1395]" pageId="32" pageNumber="33" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="8498FD699D3C927BFEEAFAE7CA5D2D16" box="[377,523,1369,1396]" pageId="32" pageNumber="33" type="generic">8 specimens</specimenCount>
)
</paragraph>
<paragraph id="922136E09D3C927BFF2EFAC2CB442DBE" blockId="32.[189,1399,1369,1500]" pageId="32" pageNumber="33">
<materialsCitation id="22F63CBD9D3C927BFF2EFAC2CD7D2DF4" ID-GBIF-Occurrence="2898878403" box="[189,1323,1403,1430]" collectionCode="BMNH" country="Papua New Guinea" location="PAPUA NEW GUINEA" pageId="32" pageNumber="33" specimenCount="7" specimenCount-female="6" specimenCount-male="1" typeStatus="holotype">
<collectingCountry id="EA8976709D3C927BFF2EFAC2C9812DF4" box="[189,471,1404,1430]" name="Papua New Guinea" pageId="32" pageNumber="33">PAPUA NEW GUINEA</collectingCountry>
•
<specimenCount id="8498FD699D3C927BFE7CFAC2CA642DF4" box="[495,562,1404,1430]" pageId="32" pageNumber="33" type="male">1 ♂♂</specimenCount>
,
<specimenCount id="8498FD699D3C927BFDADFAC5CAD62DF4" box="[574,640,1403,1430]" pageId="32" pageNumber="33" type="female">6 ♀♀</specimenCount>
; same labels as for holotype but without sex label;
<collectionCode id="F48FAE259D3C927BFB40FAC2CD7D2DF4" box="[1235,1323,1404,1430]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="32" pageNumber="33">BMNH</collectionCode>
</materialsCitation>
•
<materialsCitation id="22F63CBD9D3C927BFAD0FAC2CB582DBE" ID-GBIF-Occurrence="2898878490" collectingDate="1933-12" collectionCode="PAPUA, BMNH" collectorName="L. E. Cheesman." country="Papua New Guinea" elevation="1219" location="Mafulu" municipality="Mafulu" pageId="32" pageNumber="33" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<specimenCount id="8498FD699D3C927BFAD0FAC2CD392DF4" box="[1347,1391,1404,1430]" pageId="32" pageNumber="33" type="female">1 ♀</specimenCount>
; “
<collectionCode id="F48FAE259D3C927BFF54FA21C97A2DDB" box="[199,300,1439,1465]" pageId="32" pageNumber="33">PAPUA</collectionCode>
:
<collectingMunicipality id="7245AC9A9D3C927BFEA5FA20C9C62DDA" box="[310,400,1438,1464]" pageId="32" pageNumber="33">Mafulu</collectingMunicipality>
.
<quantity id="55669B059D3C927BFE0AFA21C9AA2DDA" box="[409,508,1438,1465]" metricMagnitude="3" metricUnit="m" metricValue="1.2192" pageId="32" pageNumber="33" unit="ft" value="4000.0">
<elevation id="19B3D1D39D3C927BFE0AFA21C9AA2DDA" box="[409,508,1438,1465]" metricMagnitude="3" metricUnit="m" metricValue="1.2192" pageId="32" pageNumber="33" unit="ft" value="4000.0">4,000 ft.</elevation>
</quantity>
<date id="E62010209D3C927BFE68FA20CA092DDA" box="[507,607,1438,1464]" pageId="32" pageNumber="33" value="1933-12">
<collectingDate id="F664E9C89D3C927BFE68FA20CA092DDA" box="[507,607,1438,1464]" pageId="32" pageNumber="33" value="1933-12">xii.1933</collectingDate>
</date>
.
<collectorName id="3F6B53369D3C927BFDFAFA20CB482DDA" box="[617,798,1438,1464]" pageId="32" pageNumber="33">L.E.Cheesman.</collectorName>
B.M.1933-321. [p]//
<taxonomicName id="559E4D639D3C927BFB84FA20C9112DB9" authority=", Hell. Determined" authorityName=", Hell. Determined" authorityYear="1895" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="subcostatus">
<emphasis id="A0EAEAF29D3C927BFB84FA20CD792DDA" box="[1047,1327,1438,1464]" italics="true" pageId="32" pageNumber="33">Maechidius subcostatus</emphasis>
, Hell. Determined
</taxonomicName>
from description. G.J.A. [p]”;
<collectionCode id="F48FAE259D3C927BFD21FA7CCB582DBE" box="[690,782,1474,1500]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="32" pageNumber="33">BMNH</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D3C927AFF2EF9BCC9112D6D" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="description">
<paragraph id="922136E09D3C927BFF2EF9BCC9182E7E" blockId="32.[189,334,1538,1564]" box="[189,334,1538,1564]" pageId="32" pageNumber="33">
<heading id="C969818C9D3C927BFF2EF9BCC9182E7E" bold="true" box="[189,334,1538,1564]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
<emphasis id="A0EAEAF29D3C927BFF2EF9BCC9182E7E" bold="true" box="[189,334,1538,1564]" pageId="32" pageNumber="33">Description</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D3C927BFF2EF98FCB502EF2" blockId="32.[189,1399,1584,1681]" pageId="32" pageNumber="33">
MEASUREMENTS.
<typeStatus id="4D2588429D3C927BFEEFF98EC9BA2E28" box="[380,492,1584,1610]" pageId="32" pageNumber="33" type="holotype">Holotype</typeStatus>
, total body length
<quantity id="55669B059D3C927BFD54F98ECB652E29" box="[711,819,1584,1611]" metricMagnitude="-3" metricUnit="m" metricValue="8.25" pageId="32" pageNumber="33" unit="mm" value="8.25">8.25 mm</quantity>
. Head
<quantity id="55669B059D3C927BFC16F98ECBB82E29" box="[901,1006,1584,1611]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" pageId="32" pageNumber="33" unit="mm" value="1.55">1.55 mm</quantity>
long, across eyes
<quantity id="55669B059D3C927BFB55F98ECD792E29" box="[1222,1327,1584,1611]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" pageId="32" pageNumber="33" unit="mm" value="1.85">1.85 mm</quantity>
wide. Pronotum
<quantity id="55669B059D3C927BFEA4F9EAC9C92E0C" box="[311,415,1620,1646]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="32" pageNumber="33" unit="mm" value="1.7">1.70 mm</quantity>
long, maximum width
<quantity id="55669B059D3C927BFD3EF9EDCB412E0C" box="[685,791,1619,1646]" metricMagnitude="-3" metricUnit="m" metricValue="2.85" pageId="32" pageNumber="33" unit="mm" value="2.85">2.85 mm</quantity>
. Elytral length 5.00 mm, maximum combined width
<quantity id="55669B059D3C927BFF2EF9C8C97F2EF3" box="[189,297,1654,1681]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="32" pageNumber="33" unit="mm" value="3.9">3.90 mm</quantity>
. The largest
<typeStatus id="4D2588429D3C927BFE2EF9C9CA742EF3" box="[445,546,1655,1681]" pageId="32" pageNumber="33" type="paratype">paratype</typeStatus>
(
<specimenCount id="8498FD699D3C927BFDA6F9C8CA1E2EF2" box="[565,584,1654,1680]" pageId="32" pageNumber="33" type="female">♀</specimenCount>
)
<quantity id="55669B059D3C927BFDC8F9C8CA952EF3" box="[603,707,1654,1681]" metricMagnitude="-3" metricUnit="m" metricValue="9.25" pageId="32" pageNumber="33" unit="mm" value="9.25">9.25 mm</quantity>
long.
</paragraph>
<paragraph id="922136E09D3C927AFF2EF906CC952C19" blockId="32.[189,1399,1719,2025]" lastBlockId="33.[189,1399,265,1148]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
Dorsum and venter uniformly black-brown with castaneous labroclypeus, mouthparts, and legs. Head transverse, glossy dorsally and ventrally, flattened between eyes and vaguely impressed on labroclypeus anterior to each compound eye. Compound eye large, occupying over about half side of head. Male labroclypeus (
<figureCitation id="0AA52A659D3C927BFEF4F89EC9992F58" box="[359,463,1824,1850]" captionStart="Figs 109–120" captionStartId="138.[189,241,1750,1776]" captionTargetBox="[365,1214,275,1722]" captionTargetId="figure-15@138.[365,1222,265,1730]" captionTargetPageId="138" captionText="Figs 109–120. Papuan andWallacean MaechidiusMacleay, 1819,head, dorsal (109–119) and frontal (120) views. 109. M. brocki sp. nov., holotype, ♂ (BMNH). 110. Same, paratype, ♀ (BMNH). 111. M. ciliatus sp. nov., holotype, ♂ (IECA). 112. M. crypticus sp. nov., paratype, ♂ (BMNH). 113. Same, paratype, ♀ (BMNH). 114. M. dani sp. nov., holotype, ♂ (IECA). 115. M. deltouri sp. nov., holotype, ♂ (BMNH). 116. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 117. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 118. M. esau Heller, 1914, lectotype, ♂ (SNSD). 119. Same, ♂ (lectotype of M. setosus Moser, 1920) (ZMHB). 120. M. gressitti Frey, 1969, holotype, ♂ (photograph courtesy BPBM) (BPBM). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122175" httpUri="https://zenodo.org/record/4122175/files/figure.png" pageId="32" pageNumber="33">Fig. 109</figureCitation>
) broadly emarginate on anterior margin, its lateral margins strongly sinuous in both dorsal and lateral views, anterolateral angles protruding, bent up at ~90° to frons in lateral view. Female labroclypeus (
<figureCitation id="0AA52A659D3C927BFD9FF8D8CA262FE2" box="[524,624,1894,1920]" captionStart="Figs 109–120" captionStartId="138.[189,241,1750,1776]" captionTargetBox="[365,1214,275,1722]" captionTargetId="figure-15@138.[365,1222,265,1730]" captionTargetPageId="138" captionText="Figs 109–120. Papuan andWallacean MaechidiusMacleay, 1819,head, dorsal (109–119) and frontal (120) views. 109. M. brocki sp. nov., holotype, ♂ (BMNH). 110. Same, paratype, ♀ (BMNH). 111. M. ciliatus sp. nov., holotype, ♂ (IECA). 112. M. crypticus sp. nov., paratype, ♂ (BMNH). 113. Same, paratype, ♀ (BMNH). 114. M. dani sp. nov., holotype, ♂ (IECA). 115. M. deltouri sp. nov., holotype, ♂ (BMNH). 116. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 117. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 118. M. esau Heller, 1914, lectotype, ♂ (SNSD). 119. Same, ♂ (lectotype of M. setosus Moser, 1920) (ZMHB). 120. M. gressitti Frey, 1969, holotype, ♂ (photograph courtesy BPBM) (BPBM). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122175" httpUri="https://zenodo.org/record/4122175/files/figure.png" pageId="32" pageNumber="33">Fig. 110</figureCitation>
) shallower emarginate on anterior margin, with broader rounded anterolateral angles. Upper- and underside of labroclypeus with sparse long setae along anterior and lateral margins. Canthus sinuous in dorsal view. Anterior and lateral margins of labroclypeus smooth. Head dorsal punctures circular to irregularly hexagonal, large and moderately deep. Intervening spaces glossy, much smaller than punctures, in part wrinkled. Head setae suberect, generally surpassing length of corresponding punctures. Few much longer suberect setae along inner margin of either eye. Antenna 9-segmented, club 3-lamellate. Scape large, with bulbous predistal projection on dorsal side, provided with two very long setae near distal margin. Antennomere 2 trapezoid, transverse. Pronotum glossy dorsally and laterally. Anterior margin of pronotum sinuous with anterolateral angles (stronger) and mesal portion (slightly) protruding anteriad. Basal margin of pronotum broadly rounded. Lateral margin gradually widens towards postmedium, very deeply emarginate prebasally (
<figureCitation id="0AA52A659D3D927AFB02FE61CCAF2998" box="[1169,1273,479,506]" captionStart="Figs 193–207" captionStartId="145.[189,241,1716,1742]" captionTargetBox="[385,1203,267,1681]" captionTargetId="figure-159@145.[385,1203,265,1694]" captionTargetPageId="145" captionText="Figs 193–207. Papuan and Wallacean Maechidius Macleay, 1819, head (193–196) and pronotum (197– 207), dorsal view. 193. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 194. Same, ♂ (SNSD). 195. M. yamdena sp. nov., paratype, ♂ (BMNH). 196. Same, paratype, ♀ (BMNH). 197. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 198. M. aiyura sp. nov., holotype, ♂ (BMNH). 199. M. alesbezdeki sp. nov., holotype, ♂ (NME). 200. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 201. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 202. M. awu sp. nov., holotype, ♂ (IECA). 203. M. babyrousa sp. nov., holotype, ♂ (BMNH). 204. M. bintang sp. nov., holotype, ♂ (NME). 205. M. boessnecki sp. nov., holotype, ♂ (SNSD). 206. M. brocki sp. nov., holotype, ♂ (BMNH). 207. M. crypticus sp. nov., paratype, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122189" httpUri="https://zenodo.org/record/4122189/files/figure.png" pageId="33" pageNumber="34">Fig. 206</figureCitation>
). Crenulae of lateral margin long and large except for smooth emargination area; suberect seta present between every two crenulae. Lateral margin of pronotum sinuous in lateral view. Pronotal punctures irregularly hexagonal, dense and moderately deep, variably large. Intervening spaces glossy and glabrous, much smaller than punctures, in part wrinkled. Appressed seta rises from anterior margin of each puncture, not surpassing length of corresponding puncture. Hypomeron separated from prosternum by moderately high sinuous carina strongly acutely dentate medially, with long setae on anterolateral margin. Antennal pocket moderately deep. Median anterior process of prosternum long brushy setose, short, flange-like rised. Scutellar shield narrowly rounded apically. Elytra slightly widened in posterior half, maximum width post midlength, opaque dorsally, with distinct humeri. Vague tracks of 2–3 inconspicuous, very flat longitudinal carinae on disc of each elytron. Elytral punctures linear (long and narrow), incisionshaped, moderately deep and dense (
<figureCitation id="0AA52A659D3D927AFDE3FCD9CA8F2BE0" box="[624,729,871,898]" captionStart="Figs 286–301" captionStartId="152.[189,241,1716,1742]" captionTargetBox="[364,1224,266,1689]" captionTargetId="figure-172@152.[364,1224,266,1695]" captionTargetPageId="152" captionText="Figs 286–301. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 286. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 287. M. aiyura sp. nov., holotype, ♂ (BMNH). 288. M. alesbezdeki sp. nov., holotype, ♂ (NME). 289. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 290. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 291. M. paupianus Heller, 1910, ♂ (BMNH). 292. M. awu sp. nov., holotype, ♂ (IECA). 293. M. babyrousa sp. nov., holotype, ♂ (BMNH). 294. M. bintang sp. nov., holotype, ♂ (NME). 295. M. boessnecki sp. nov., holotype, ♂ (SNSD). 296. M. brocki sp. nov., holotype, ♂ (BMNH). 297. M. caperatus sp. nov., holotype, ♂ (NME). 298. M. ciliatus sp. nov., holotype, ♂ (IECA). 299. M. crypticus sp. nov., paratype, ♀ (BMNH). 300. M. dani sp. nov., holotype, ♂ (IECA). 301. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122206" httpUri="https://zenodo.org/record/4122206/files/figure.png" pageId="33" pageNumber="34">Fig. 296</figureCitation>
). Intervening spaces covered with microscopical velvety pubescence, larger than incisions. Setation inconspicuous, suberect; seta rises from anterior margin of each incision, not or hardly surpassing its length. Male pygidium flat dorsally, with large shallow annular punctures (
<figureCitation id="0AA52A659D3D927AFEACFC6CC9F02B8F" box="[319,422,978,1005]" captionStart="Figs 451–468" captionStartId="163.[189,241,1681,1707]" captionTargetBox="[456,1130,262,1650]" captionTargetId="figure-178@163.[456,1131,262,1657]" captionTargetPageId="163" captionText="Figs 451–468. Papuan and Wallacean Maechidius Macleay, 1819, pygidium, dorsal view. 451. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 452. M. aiyura sp. nov., holotype, ♂ (BMNH). 453. Same, different aspect (BMNH). 454. M. alesbezdeki sp. nov., holotype, ♂ (NME). 455. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 456. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 457. M. awu sp. nov., holotype, ♂ (IECA). 458. M. babyrousa sp. nov., holotype, ♂ (BMNH). 459. Same, paratype, ♀ (BMNH). 460. M. bintang sp. nov., holotype, ♂ (NME). 461. M. boessnecki sp. nov., holotype, ♂ (SNSD). 462. M. brocki sp. nov., holotype, ♂ (BMNH). 463. M. caperatus sp. nov., holotype, ♂ (NME). 464. M. ciliatus sp. nov., holotype, ♂ (IECA). 465. M. crypticus sp. nov., paratype, ♂ (BMNH). 466. Same, paratype, ♀ (BMNH). 467. M. dani sp. nov., holotype, ♂ (IECA). 468. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122222" httpUri="https://zenodo.org/record/4122222/files/figure.png" pageId="33" pageNumber="34">Fig. 462</figureCitation>
). Intervening spaces generally smaller than punctures, in part microreticulate. Setae of pygidium inconspicuous, suberect, about length of corresponding punctures. Male protibia slightly arched on external margin, with three obtuse distal teeth on external margin (Fig. 368). Male protibial terminal spur short, thick and pointed apically, female one longer, straight and acute. All tarsal claws with large pulvilli. Spiculum gastrale as in Fig. 539. Male aedeagus as in
<figureCitation id="0AA52A659D3D927AFB89FBDFCCE92C19" box="[1050,1215,1121,1147]" captionStart="Figs 601–616" captionStartId="172.[189,241,1892,1918]" captionTargetBox="[458,1119,274,1854]" captionTargetId="figure-73@172.[450,1138,265,1868]" captionTargetPageId="172" captionText="Figs 601–616. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 601– 603. M. brocki sp. nov., holotype, ♂ (BMNH). 604–606. M. dani sp. nov., holotype, ♂ (IECA). 607–610. M. deltouri sp. nov., holotype, ♂ (BMNH). 611–613. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 614–616. M. esau Heller, 1914, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122238" httpUri="https://zenodo.org/record/4122238/files/figure.png" pageId="33" pageNumber="34">Figs 601–603</figureCitation>
.
</paragraph>
<paragraph id="922136E09D3D927AFF2EFB1DC9E62CDF" blockId="33.[189,432,1187,1213]" box="[189,432,1187,1213]" pageId="33" pageNumber="34">
<heading id="C969818C9D3D927AFF2EFB1DC9E62CDF" bold="true" box="[189,432,1187,1213]" fontSize="11" level="3" pageId="33" pageNumber="34" reason="3">
<emphasis id="A0EAEAF29D3D927AFF2EFB1DC9E62CDF" bold="true" box="[189,432,1187,1213]" pageId="33" pageNumber="34">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D3D927AFF2EFB6CC9112D6D" blockId="33.[189,1398,1234,1295]" pageId="33" pageNumber="34">Female lamellae of antennal club shorter; anterior margin of labroclypeus comparatively less deeply emarginate.</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D3D927AFF2EFA86CA6A2DE2" box="[189,572,1336,1409]" pageId="33" pageNumber="34" type="biology_ecology">
<paragraph id="922136E09D3D927AFF2EFA86C9762D30" blockId="33.[189,288,1336,1362]" box="[189,288,1336,1362]" pageId="33" pageNumber="34">
<heading id="C969818C9D3D927AFF2EFA86C9762D30" bold="true" box="[189,288,1336,1362]" fontSize="11" level="3" pageId="33" pageNumber="34" reason="3">
<emphasis id="A0EAEAF29D3D927AFF2EFA86C9762D30" bold="true" box="[189,288,1336,1362]" pageId="33" pageNumber="34">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D3D927AFF2EFAD9CA6A2DE2" blockId="33.[189,572,1382,1409]" box="[189,572,1382,1409]" pageId="33" pageNumber="34">
Occurs at about
<quantity id="55669B059D3D927AFEEFFAD9C9832DE3" box="[380,469,1383,1409]" metricMagnitude="3" metricUnit="m" metricValue="1.22" pageId="33" pageNumber="34" unit="m" value="1220.0">1220 m</quantity>
altitude.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D3D927AFF2EFA16CBE02D90" box="[189,950,1448,1523]" pageId="33" pageNumber="34" type="distribution">
<paragraph id="922136E09D3D927AFF2EFA16C9002DA0" blockId="33.[189,342,1448,1474]" box="[189,342,1448,1474]" pageId="33" pageNumber="34">
<heading id="C969818C9D3D927AFF2EFA16C9002DA0" bold="true" box="[189,342,1448,1474]" fontSize="11" level="3" pageId="33" pageNumber="34" reason="3">
<emphasis id="A0EAEAF29D3D927AFF2EFA16C9002DA0" bold="true" box="[189,342,1448,1474]" pageId="33" pageNumber="34">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D3D927AFF2EFA66CBE02D90" blockId="33.[189,950,1496,1523]" box="[189,950,1496,1523]" pageId="33" pageNumber="34">
Hitherto only known from the Papuan Peninsula of New
<collectingCountry id="EA8976709D3D927AFCC9FA66CBE52D90" box="[858,947,1496,1522]" name="Guinea-Bissau" pageId="33" pageNumber="34">Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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