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<document id="1603F65AD8AEC1F60FD308DECFA3A760" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D329275FDACFEB6C9C72F8A" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius fraterculus Moser Determined 1920" docType="treatment" docVersion="10" lastPageNumber="47" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="47" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="ED6F6438210955C8F7D787A511B812BB">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
</mods:titleInfo>
<mods:name id="F376F450BCBE88CF74510CBB0F59003C" type="personal">
<mods:role id="7A0BE84BC4C7D1EE3CCAB0FC2BD20C17">
<mods:roleTerm id="737F7659A1746E55B2CB15B297134D2D">Author</mods:roleTerm>
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<mods:namePart id="C64EE205E00728E27A5A421872279426">Telnov, Dmitry</mods:namePart>
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<mods:typeOfResource id="19507F7F195BEC4C014F97298395A26E">text</mods:typeOfResource>
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<mods:title id="10E5AD5CC1D7ABD8C899B80C4814D032">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="949E033E9AD4B419E3A45A4B9BF07A52">
<mods:date id="D5FCCEB00ABB7FF80AADC5FB0EB92ED5">2020</mods:date>
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<mods:number id="34CF74B5B3B4890E6CDAE35F40D89916">2020-10-19</mods:number>
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<mods:number id="B9AFD00BE62135448B54105A10F7EE8C">721</mods:number>
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<mods:classification id="AD35E53FC649E914DA9E2BF800AB2CFE">journal article</mods:classification>
<mods:identifier id="49E404879BAD0CFAD7EDA5A56D88C570" type="CLB-Dataset">9693</mods:identifier>
<mods:identifier id="8E4547C50FA0F7592B787145F6539907" type="DOI">10.5852/ejt.2020.721.1127</mods:identifier>
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<treatment id="1A3787F69D329275FDACFEB6C9C72F8A" ID-DOI="http://doi.org/10.5281/zenodo.4344441" ID-GBIF-Taxon="168648142" ID-Zenodo-Dep="4344441" LSID="urn:lsid:plazi:treatment:1A3787F69D329275FDACFEB6C9C72F8A" httpUri="http://treatment.plazi.org/id/1A3787F69D329275FDACFEB6C9C72F8A" lastPageNumber="47" pageId="46" pageNumber="47">
<subSubSection id="DA84656B9D329275FDACFEB6CBA32941" box="[575,1013,264,291]" pageId="46" pageNumber="47" type="nomenclature">
<paragraph id="922136E09D329275FDACFEB6CBA32941" blockId="46.[575,1013,264,325]" box="[575,1013,264,291]" pageId="46" pageNumber="47">
<heading id="C969818C9D329275FDACFEB6CBA32941" box="[575,1013,264,291]" centered="true" fontSize="11" level="2" pageId="46" pageNumber="47" reason="2">
<taxonomicName id="559E4D639D329275FDACFEB6CBA32941" ID-CoL="3XFJ8" authority="Moser, 1920" authorityName="Moser Determined" authorityYear="1920" box="[575,1013,264,291]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="fraterculus">
<emphasis id="A0EAEAF29D329275FDACFEB6CB0C2941" bold="true" box="[575,858,264,291]" italics="true" pageId="46" pageNumber="47">Maechidius fraterculus</emphasis>
<bibRefCitation id="F60F4B119D329275FCF2FEB7CBA32941" author="Moser J." box="[865,1013,265,291]" pageId="46" pageNumber="47" pagination="3 - 28" refId="ref98398" refString="Moser J. 1920. Beitrag zur Kenntnis der Melolonthiden XI. Stettiner Entomologische Zeitung 81: 3 - 28." type="journal article" year="1920">Moser, 1920</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D329275FDCFFE95CB812927" box="[604,983,299,325]" pageId="46" pageNumber="47" type="description">
<paragraph id="922136E09D329275FDCFFE95CB812927" blockId="46.[575,1013,264,325]" box="[604,983,299,325]" pageId="46" pageNumber="47">
<figureCitation id="0AA52A659D329275FDCFFE95CAE02927" box="[604,694,299,325]" captionStart="Figs 38–41" captionStartId="47.[189,241,1893,1919]" captionTargetBox="[360,1224,277,1874]" captionTargetId="figure-68@47.[360,1227,265,1875]" captionTargetPageId="47" captionText="Figs 38–41. Papuan Maechidius Macleay, 1819, habitus, dorsal view. 38. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 39. M. hamatus sp. nov., holotype, ♂ (IECA). 40. M. helleri (Frey, 1969) comb. nov., holotype, ♂ (BPBM) (photograph courtesy BPBM). 41. M. gressitti Frey, 1969, holotype, ♂ (photograph courtesy BPBM) (BPBM). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122142" httpUri="https://zenodo.org/record/4122142/files/figure.png" pageId="46" pageNumber="47">Figs 38</figureCitation>
,
<figureCitation id="0AA52A659D329275FD51FE95CAA62927" box="[706,752,299,325]" captionStart="Figs 109–120" captionStartId="138.[189,241,1750,1776]" captionTargetBox="[365,1214,275,1722]" captionTargetId="figure-15@138.[365,1222,265,1730]" captionTargetPageId="138" captionText="Figs 109–120. Papuan andWallacean MaechidiusMacleay, 1819,head, dorsal (109–119) and frontal (120) views. 109. M. brocki sp. nov., holotype, ♂ (BMNH). 110. Same, paratype, ♀ (BMNH). 111. M. ciliatus sp. nov., holotype, ♂ (IECA). 112. M. crypticus sp. nov., paratype, ♂ (BMNH). 113. Same, paratype, ♀ (BMNH). 114. M. dani sp. nov., holotype, ♂ (IECA). 115. M. deltouri sp. nov., holotype, ♂ (BMNH). 116. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 117. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 118. M. esau Heller, 1914, lectotype, ♂ (SNSD). 119. Same, ♂ (lectotype of M. setosus Moser, 1920) (ZMHB). 120. M. gressitti Frey, 1969, holotype, ♂ (photograph courtesy BPBM) (BPBM). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122175" httpUri="https://zenodo.org/record/4122175/files/figure.png" pageId="46" pageNumber="47">117</figureCitation>
,
<figureCitation id="0AA52A659D329275FD68FE95CB7C2927" box="[763,810,299,325]" captionStart="Figs 208–225" captionStartId="146.[189,241,1644,1670]" captionTargetBox="[414,1187,265,1610]" captionTargetId="figure-199@146.[414,1187,265,1618]" captionTargetPageId="146" captionText="Figs 208–225. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, pronotum, dorsal view. 208. M.caperatus sp. nov., holotype, ♂ (NME). 209. M.ciliatus sp. nov., holotype, ♂ (IECA). 210. M.dani sp. nov., holotype, ♂ (IECA). 211. M. deltouri sp. nov., holotype, ♂ (BMNH). 212. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 213. M. esau Heller, 1914 (lectotype, ♂ of M. setosus Moser, 1920) (ZMHB). 214. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 215. M. hamatus sp. nov., holotype, ♂ (IECA). 216. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 217. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 218. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 219. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 220. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 221. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 222. M. konjo sp. nov., holotype, ♂ (NMPC). 223. M. lapsus sp. nov., holotype, ♂ (BMNH). 224. M. legalovi sp. nov., holotype, ♂ (BMNH). 225. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122191" httpUri="https://zenodo.org/record/4122191/files/figure.png" pageId="46" pageNumber="47">214</figureCitation>
,
<figureCitation id="0AA52A659D329275FCA5FE95CB332927" box="[822,869,299,325]" captionStart="Figs 302–316" captionStartId="153.[189,241,1679,1705]" captionTargetBox="[393,1194,265,1649]" captionTargetId="figure-159@153.[393,1194,265,1658]" captionTargetPageId="153" captionText="Figs 302–316. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, elytron, dorsal view. 302. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 303. M. esau Heller, 1914, lectotype, ♂ (SNSD). 304. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 305. M. hamatus sp. nov., holotype, ♂ (IECA). 306. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 307. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 308. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 309. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 310. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 311. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 312. Same, enlarged. 313. M. konjo sp. nov., holotype, ♂ (NMPC). 314. M. lapsus sp. nov., holotype, ♂ (BMNH). 315. M. legalovi sp. nov., holotype, ♂ (BMNH). 316. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122208" httpUri="https://zenodo.org/record/4122208/files/figure.png" pageId="46" pageNumber="47">304</figureCitation>
, 377, 471
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D329275FF2EFED5CA592A78" pageId="46" pageNumber="47" type="materials_examined">
<paragraph id="922136E09D329275FF2EFED5C93A29E7" blockId="46.[189,364,363,389]" box="[189,364,363,389]" pageId="46" pageNumber="47">
<heading id="C969818C9D329275FF2EFED5C93A29E7" bold="true" box="[189,364,363,389]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
<emphasis id="A0EAEAF29D329275FF2EFED5C93A29E7" bold="true" box="[189,364,363,389]" pageId="46" pageNumber="47">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D329275FF79FE27CA3A29D1" blockId="46.[189,1399,408,538]" box="[234,620,408,435]" pageId="46" pageNumber="47">
<emphasis id="A0EAEAF29D329275FF79FE27C93029D1" bold="true" box="[234,358,409,435]" pageId="46" pageNumber="47">
<typeStatus id="4D2588429D329275FF79FE27C93029D1" box="[234,358,409,435]" pageId="46" pageNumber="47" type="lectotype">Lectotype</typeStatus>
</emphasis>
[herewith designated]
</paragraph>
<paragraph id="922136E09D329275FF2EFE05CA592A78" blockId="46.[189,1399,408,538]" pageId="46" pageNumber="47">
<materialsCitation id="22F63CBD9D329275FF2EFE05CA592A78" ID-GBIF-Occurrence="2898878342" collectionCode="ZMHB" country="Guinea-Bissau" location="Island of New Guinea" pageId="46" pageNumber="47" specimenCode="MFNB 2019" specimenCount="1" specimenCount-female="1" typeStatus="syntype">
COUNTRY UNKNOWN (
<location id="9741603B9D329275FD97FE05CB4C29B4" LSID="urn:lsid:plazi:treatment:1A3787F69D329275FDACFEB6C9C72F8A:9741603B9D329275FD97FE05CB4C29B4" box="[516,794,443,470]" country="Guinea-Bissau" name="Island of New Guinea" pageId="46" pageNumber="47">Island of New Guinea</location>
) •
<specimenCount id="8498FD699D329275FCD6FE05CB0A29B7" box="[837,860,443,469]" pageId="46" pageNumber="47" type="female">♀</specimenCount>
; “
<taxonomicName id="559E4D639D329275FCEEFE05CD4529B7" authority="Mos. Typ" authorityName="Mos. Typ" box="[893,1299,443,469]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="fraterculus">
<emphasis id="A0EAEAF29D329275FCEEFE05CCC429B7" box="[893,1170,443,469]" italics="true" pageId="46" pageNumber="47">Maechidius fraterculus</emphasis>
Mos. Typ
</taxonomicName>
[h] // N.
<collectingCountry id="EA8976709D329275FF2EFE63C9442995" box="[189,274,477,503]" name="Guinea-Bissau" pageId="46" pageNumber="47">Guinea</collectingCountry>
… [h, in part unreadable] //
<typeStatus id="4D2588429D329275FDC8FE63CA892995" box="[603,735,477,503]" pageId="46" pageNumber="47" type="syntype">SYNTYPE</typeStatus>
<taxonomicName id="559E4D639D329275FD74FE63CCC52995" authority="Moser, 1920" authorityName="Moser Determined" authorityYear="1920" box="[743,1171,477,504]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="fraterculus">
<emphasis id="A0EAEAF29D329275FD74FE63CBAE2995" box="[743,1016,477,503]" italics="true" pageId="46" pageNumber="47">Maechidius fraterculus</emphasis>
<bibRefCitation id="F60F4B119D329275FC6CFE60CCC52995" author="Moser J." box="[1023,1171,477,504]" pageId="46" pageNumber="47" pagination="3 - 28" refId="ref98398" refString="Moser J. 1920. Beitrag zur Kenntnis der Melolonthiden XI. Stettiner Entomologische Zeitung 81: 3 - 28." type="journal article" year="1920">Moser, 1920</bibRefCitation>
</taxonomicName>
labelled by
<specimenCode id="C2389E9B9D329275FAB0FE60C8AE2A7B" collectionCode="MFNB" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="46" pageNumber="47">MFNB 2019</specimenCode>
[p, red label]”;
<collectionCode id="F48FAE259D329275FE21FDBECA5D2A78" box="[434,523,512,538]" collectionName="Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet" pageId="46" pageNumber="47">ZMHB</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D329275FF2EFDFECD162AA9" pageId="46" pageNumber="47" type="discussion">
<paragraph id="922136E09D329275FF2EFDFEC9792A38" blockId="46.[189,303,576,602]" box="[189,303,576,602]" pageId="46" pageNumber="47">
<heading id="C969818C9D329275FF2EFDFEC9792A38" bold="true" box="[189,303,576,602]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
<emphasis id="A0EAEAF29D329275FF2EFDFEC9792A38" bold="true" box="[189,303,576,602]" pageId="46" pageNumber="47">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D329275FF2EFDD3CD162AA9" blockId="46.[189,1398,621,716]" pageId="46" pageNumber="47">
<bibRefCitation id="F60F4B119D329275FF2EFDD3C9D72AEA" author="Moser J." box="[189,385,621,648]" pageId="46" pageNumber="47" pagination="3 - 28" refId="ref98398" refString="Moser J. 1920. Beitrag zur Kenntnis der Melolonthiden XI. Stettiner Entomologische Zeitung 81: 3 - 28." type="journal article" year="1920">Moser (1920: 15)</bibRefCitation>
based his description of
<taxonomicName id="559E4D639D329275FD07FDD0CB622AE5" authorityName="Moser Determined" authorityYear="1920" box="[660,820,621,647]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="fraterculus">
<emphasis id="A0EAEAF29D329275FD07FDD0CB622AE5" box="[660,820,621,647]" italics="true" pageId="46" pageNumber="47">M. fraterculus</emphasis>
</taxonomicName>
on an unstated number of specimens. The
<typeStatus id="4D2588429D329275FA9EFDD3CD202AE5" box="[1293,1398,621,647]" pageId="46" pageNumber="47" type="lectotype">lectotype</typeStatus>
is therefore designated for the single
<typeStatus id="4D2588429D329275FDF9FD31CA952ACB" box="[618,707,655,681]" pageId="46" pageNumber="47" type="syntype">syntype</typeStatus>
in ZMHB in order to enhance the stability of nomenclature and fix the specimen I based my redescription on, in case other
<typeStatus id="4D2588429D329275FC07FD0CCBAE2AAE" box="[916,1016,690,716]" pageId="46" pageNumber="47" type="syntype">syntypes</typeStatus>
are discovered in the future.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D329275FF2EFD4FC9C72F8A" pageId="46" pageNumber="47" type="description">
<paragraph id="922136E09D329275FF2EFD4FC9182B69" blockId="46.[189,334,753,779]" box="[189,334,753,779]" pageId="46" pageNumber="47">
<heading id="C969818C9D329275FF2EFD4FC9182B69" bold="true" box="[189,334,753,779]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
<emphasis id="A0EAEAF29D329275FF2EFD4FC9182B69" bold="true" box="[189,334,753,779]" pageId="46" pageNumber="47">Description</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D329275FF2EFCA1CB112B58" blockId="46.[189,839,799,826]" box="[189,839,799,826]" pageId="46" pageNumber="47">
MEASUREMENTS. Total body length
<quantity id="55669B059D329275FDC2FCA1CAEF2B5B" box="[593,697,799,825]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="46" pageNumber="47" unit="mm" value="9.1">9.10 mm</quantity>
(
<typeStatus id="4D2588429D329275FD5BFCA1CB6D2B58" box="[712,827,799,826]" pageId="46" pageNumber="47" type="lectotype">lectotype</typeStatus>
).
</paragraph>
<paragraph id="922136E09D329275FF2EFCDECC5A2F18" blockId="46.[189,1399,863,1914]" pageId="46" pageNumber="47">
Dorsum and venter uniformly castaneous brown except for paler appendages and antennae. Head flattened dorsally on frons between eyes, glossy dorsally and ventrally. Female labroclypeus deeply and broadly V-shaped emarginate anteriorly, its lateral margins strongly sinuous in both dorsal and lateral views (
<figureCitation id="0AA52A659D329275FE87FC7BC92B2B82" box="[276,381,965,992]" captionStart="Figs 109–120" captionStartId="138.[189,241,1750,1776]" captionTargetBox="[365,1214,275,1722]" captionTargetId="figure-15@138.[365,1222,265,1730]" captionTargetPageId="138" captionText="Figs 109–120. Papuan andWallacean MaechidiusMacleay, 1819,head, dorsal (109–119) and frontal (120) views. 109. M. brocki sp. nov., holotype, ♂ (BMNH). 110. Same, paratype, ♀ (BMNH). 111. M. ciliatus sp. nov., holotype, ♂ (IECA). 112. M. crypticus sp. nov., paratype, ♂ (BMNH). 113. Same, paratype, ♀ (BMNH). 114. M. dani sp. nov., holotype, ♂ (IECA). 115. M. deltouri sp. nov., holotype, ♂ (BMNH). 116. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 117. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 118. M. esau Heller, 1914, lectotype, ♂ (SNSD). 119. Same, ♂ (lectotype of M. setosus Moser, 1920) (ZMHB). 120. M. gressitti Frey, 1969, holotype, ♂ (photograph courtesy BPBM) (BPBM). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122175" httpUri="https://zenodo.org/record/4122175/files/figure.png" pageId="46" pageNumber="47">Fig. 117</figureCitation>
). Anterolateral angles broadly rounded, moderately protruding, raised up at angle of nearly 80° to frons in lateral view. Anterior and lateral margins of labroclypeus smooth. Upper- and underside of labroclypeus with sparse moderately long setae along anterior and lateral margins. Canthus broadly rounded in dorsal view. Punctures of frons large, of irregular shape (circular to hexagonal), moderately deep and dense. Intervening spaces glossy, distinctly smaller than punctures, in part wrinkled. Setae very inconspicuous, not surpassing length of corresponding punctures; each seta rises from anterior margin of puncture. Few much longer suberect setae present along inner eye margin. Female antenna 9-segmented, club 3-lamellate. Pronotum transverse, glossy dorsally and laterally. Anterior margin of pronotum broadly emarginate with protruding anterolateral angles, basal margin broadly rounded. Lateral margin of pronotum broadly rounded and crenulate in anterior half, stronger constricted towards base and with 5–6 large denticles in basal third (
<figureCitation id="0AA52A659D329275FC75FAA4CC182D57" box="[998,1102,1306,1333]" captionStart="Figs 208–225" captionStartId="146.[189,241,1644,1670]" captionTargetBox="[414,1187,265,1610]" captionTargetId="figure-199@146.[414,1187,265,1618]" captionTargetPageId="146" captionText="Figs 208–225. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, pronotum, dorsal view. 208. M.caperatus sp. nov., holotype, ♂ (NME). 209. M.ciliatus sp. nov., holotype, ♂ (IECA). 210. M.dani sp. nov., holotype, ♂ (IECA). 211. M. deltouri sp. nov., holotype, ♂ (BMNH). 212. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 213. M. esau Heller, 1914 (lectotype, ♂ of M. setosus Moser, 1920) (ZMHB). 214. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 215. M. hamatus sp. nov., holotype, ♂ (IECA). 216. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 217. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 218. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 219. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 220. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 221. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 222. M. konjo sp. nov., holotype, ♂ (NMPC). 223. M. lapsus sp. nov., holotype, ♂ (BMNH). 224. M. legalovi sp. nov., holotype, ♂ (BMNH). 225. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122191" httpUri="https://zenodo.org/record/4122191/files/figure.png" pageId="46" pageNumber="47">Fig. 214</figureCitation>
). An inconspicuous short erect seta present between every two crenulae / denticles. Lateral margin of pronotum in lateral view nearly straight, slightly angulate postmedially. Pronotal disc with vague circular impression on either side of dorsally hardly longitudinally impressed midline. Hypomeron slightly emarginate opposite of compound eye, separated from prosternum by low carina, with long setae on its anterolateral margin. Antennal pocket shallow. Punctures of pronotal disc generally smaller but denser than those on head, circular to hexagonal, inner margin with or without thin membrane. Punctures along lateral margins distinctly oblong. Intervening spaces glossy, much smaller than punctures, in part globose. Setae very inconspicuous, not surpassing length of corresponding punctures. Scutellar shield narrowly rounded apically. Elytra subopaque, with tracks of four vague longitudinal carinae, including sutural one. Punctures of elytral disc sinuous, long and narrow, incision-shaped, dense (
<figureCitation id="0AA52A659D329275FBB7F9D1CCDC2EEB" box="[1060,1162,1647,1673]" captionStart="Figs 302–316" captionStartId="153.[189,241,1679,1705]" captionTargetBox="[393,1194,265,1649]" captionTargetId="figure-159@153.[393,1194,265,1658]" captionTargetPageId="153" captionText="Figs 302–316. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, elytron, dorsal view. 302. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 303. M. esau Heller, 1914, lectotype, ♂ (SNSD). 304. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 305. M. hamatus sp. nov., holotype, ♂ (IECA). 306. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 307. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 308. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 309. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 310. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 311. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 312. Same, enlarged. 313. M. konjo sp. nov., holotype, ♂ (NMPC). 314. M. lapsus sp. nov., holotype, ♂ (BMNH). 315. M. legalovi sp. nov., holotype, ♂ (BMNH). 316. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122208" httpUri="https://zenodo.org/record/4122208/files/figure.png" pageId="46" pageNumber="47">Fig. 304</figureCitation>
). Intervening spaces subopaque, in part densely transverse microstriate. Setae inconspicuous, appressed to suberect; each seta rising from anterior margin of corresponding incision-shaped puncture and not surpassing its length. Female pygidium flattened dorsally, with shallow transverse impression at posterior margin, densely and shallowly annularly punctate and with sparse short to moderately suberect setae (
<figureCitation id="0AA52A659D329275FB36F946CD5A2F70" box="[1189,1292,1784,1810]" captionStart="Figs 469–486" captionStartId="164.[189,241,1681,1707]" captionTargetBox="[450,1137,265,1633]" captionTargetId="figure-184@164.[450,1138,265,1647]" captionTargetPageId="164" captionText="Figs 469–486. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, pygidium, dorsal view. 469. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 470. M. esau Heller, 1914, ♂ (BMNH). 471. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 472. M. hamatus sp. nov., holotype, ♂ (IECA). 473. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 474. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 475. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 476. Same, paralectotype, ♀ (SNSD). 477. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 478. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 479. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 480. M. konjo sp. nov., holotype, ♂ (NMPC). 481. M. lapsus sp. nov., holotype, ♂ (BMNH). 482. Same, paratype, ♀ (BMNH). 483. M. legalovi sp. nov., holotype, ♂ (BMNH). 484. Same, paratype, ♀ (BMNH). 485. M. leucopsar sp. nov., holotype, ♂ (IECA). 486. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122224" httpUri="https://zenodo.org/record/4122224/files/figure.png" pageId="46" pageNumber="47">Fig. 471</figureCitation>
). Female protibia with three distal teeth, two of which much larger and stronger acute (Fig. 377). Female protibial terminal spur short, slightly curved apically. Female metatibial terminal spurs long, rounded apically, strongly subequal in length. Tarsal claws with pulvilli (in female only?).
</paragraph>
<paragraph id="922136E09D329275FF2EF821C9E62FDB" blockId="46.[189,432,1951,1977]" box="[189,432,1951,1977]" pageId="46" pageNumber="47">
<heading id="C969818C9D329275FF2EF821C9E62FDB" bold="true" box="[189,432,1951,1977]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
<emphasis id="A0EAEAF29D329275FF2EF821C9E62FDB" bold="true" box="[189,432,1951,1977]" pageId="46" pageNumber="47">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D329275FF2EF870C9C72F8A" blockId="46.[189,401,1998,2024]" box="[189,401,1998,2024]" pageId="46" pageNumber="47">Male is unknown.</paragraph>
</subSubSection>
</treatment>
</document>
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