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<document id="8B9D4E370A79DFA6BED6D28B66080E2F" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D2F926EFE46FC33CA852C94" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius heterosquamosus Heller 1910" docType="treatment" docVersion="10" lastPageNumber="54" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="52" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="DFABAB2B2837137CE4D11BEBBA5EB27D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5ABEF474B8C6894E92809F659A492D35">
<mods:title id="3024AC8289929D233A71D2BB11750A2D">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
</mods:titleInfo>
<mods:name id="08E9A8ADF71BA829A85A2BC83EA98583" type="personal">
<mods:role id="5041F5A71B0A30C847962EB9FCEFD8AB">
<mods:roleTerm id="8D60C33838FC405EE3D09E13C6ED42CB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="931C1AD2744722F6A6DC0621EA68C3D8">Telnov, Dmitry</mods:namePart>
</mods:name>
<mods:typeOfResource id="4CD952B30A704D93C9CDAFB227373E1B">text</mods:typeOfResource>
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<mods:titleInfo id="E4E1959B67457E18FC153005E8CAB53F">
<mods:title id="5DC785870C4431566CD2A84083AB670C">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="D8F64585AD29525D7D974F2A32DEE6AD">
<mods:date id="4130A1F27268B62EF059C01C46AD54B6">2020</mods:date>
<mods:detail id="F6506500859A399EE4A971AFF48CBAF1" type="pubDate">
<mods:number id="4B61201FEA34DA80FD72554EE92F7771">2020-10-19</mods:number>
</mods:detail>
<mods:detail id="8363811364A084E6E0408E259CC9E4A1" type="volume">
<mods:number id="9563ACAB2EB1B362D99F1799FC0A384E">721</mods:number>
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<mods:classification id="2CF4E79434F7E415D1C0B03A0B23911B">journal article</mods:classification>
<mods:identifier id="053F89D990EE45CF090514D70669CBBD" type="CLB-Dataset">9693</mods:identifier>
<mods:identifier id="0FAA79816B5324C3290E953305D3F864" type="DOI">10.5852/ejt.2020.721.1127</mods:identifier>
<mods:identifier id="7C6340B64C843A1698456BC09BD6CC72" type="GBIF-Dataset">0cd61b23-1089-4c1b-bfc0-487887abad84</mods:identifier>
<mods:identifier id="525DE151F6E6FE0EF574730DBD966EC6" type="Zenodo-Dep">4122118</mods:identifier>
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<treatment id="1A3787F69D2F926EFE46FC33CA852C94" ID-DOI="http://doi.org/10.5281/zenodo.4344495" ID-GBIF-Taxon="168648140" ID-Zenodo-Dep="4344495" LSID="urn:lsid:plazi:treatment:1A3787F69D2F926EFE46FC33CA852C94" httpUri="http://treatment.plazi.org/id/1A3787F69D2F926EFE46FC33CA852C94" lastPageId="53" lastPageNumber="54" pageId="51" pageNumber="52">
<subSubSection id="DA84656B9D2F9268FE46FC33CC092BCA" box="[469,1119,909,936]" pageId="51" pageNumber="52" type="nomenclature">
<paragraph id="922136E09D2F9268FE46FC33CC092BCA" blockId="51.[469,1119,909,970]" box="[469,1119,909,936]" pageId="51" pageNumber="52">
<heading id="C969818C9D2F9268FE46FC33CC092BCA" box="[469,1119,909,936]" centered="true" fontSize="11" level="2" pageId="51" pageNumber="52" reason="2">
<taxonomicName id="559E4D639D2F9268FE46FC33CB802BCA" ID-CoL="7X3WT" authority="Heller, 1910" authorityName="Heller" authorityYear="1910" box="[469,982,909,936]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="heterosquamosus">
<emphasis id="A0EAEAF29D2F9268FE46FC33CB6A2BCA" bold="true" box="[469,828,909,936]" italics="true" pageId="51" pageNumber="52">Maechidius heterosquamosus</emphasis>
<bibRefCitation id="F60F4B119D2F9268FCD7FC30CB802BCA" author="Heller K. M." box="[836,982,910,936]" pageId="51" pageNumber="52" pagination="1 - 42" refId="ref98170" refString="Heller K. M. 1910. Funfter Beitrag zur Papuanischen Kaferfauna hauptsachlich auf Grund der Ausbeute von Dr. Schlaginhaufen. Abhandlungen und Berichte des Koniglichen Zoologischen und Anthropologisch- Ethnographischen Museums zu Dresden 13 (3): 1 - 42." type="journal article" year="1910">Heller, 1910</bibRefCitation>
</taxonomicName>
comb. rest.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D2F9268FD98FC11CC7F2BA8" box="[523,1065,943,970]" pageId="51" pageNumber="52" type="description">
<paragraph id="922136E09D2F9268FD98FC11CC7F2BA8" blockId="51.[469,1119,909,970]" box="[523,1065,943,970]" pageId="51" pageNumber="52">
<figureCitation id="0AA52A659D2F9268FD98FC11CAC62BA8" box="[523,656,943,970]" captionStart="Figs 42–45" captionStartId="52.[189,241,1892,1918]" captionTargetBox="[334,1252,265,1842]" captionTargetId="figure-69@52.[334,1252,265,1861]" captionTargetPageId="52" captionText="Figs 42–45. Papuan Maechidius Macleay, 1819, habitus, dorsal view. 42. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 43. Same, ♂, holotype of M. clypeatus (Frey, 1969) comb. nov. (BPBM). 44. M. hirtipesArrow, 1941, lectotype, ♂ (BMNH). 45. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122144" httpUri="https://zenodo.org/record/4122144/files/figure.png" pageId="51" pageNumber="52">Figs 42–43</figureCitation>
,
<figureCitation id="0AA52A659D2F9268FD0EFC11CB502BA8" box="[669,774,943,970]" captionStart="Figs 121–132" captionStartId="139.[189,241,1716,1742]" captionTargetBox="[357,1230,276,1689]" captionTargetId="figure-149@139.[357,1230,265,1697]" captionTargetPageId="139" captionText="Figs 121–132. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, head, dorsal (121, 123–124, 126–132) and frontal (122, 125) views. 121. M. hamatus sp. nov., holotype, ♂ (IECA). 122. M. helleri (Frey, 1969) comb. nov., holotype, ♂ (photograph courtesy BPBM) (BPBM). 123. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 124. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 125. Same, ♂, holotype of M. clypeatus (Frey, 1969) comb. nov. (BMNH). 126. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 127. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 128. Same, paralectotype, ♀ (SNSD). 129. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 130. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 131. M. konjo sp. nov., holotype, ♂ (NMPC). 132. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122177" httpUri="https://zenodo.org/record/4122177/files/figure.png" pageId="51" pageNumber="52">124–125</figureCitation>
,
<figureCitation id="0AA52A659D2F9268FC81FC11CB172BAB" box="[786,833,943,969]" captionStart="Figs 208–225" captionStartId="146.[189,241,1644,1670]" captionTargetBox="[414,1187,265,1610]" captionTargetId="figure-199@146.[414,1187,265,1618]" captionTargetPageId="146" captionText="Figs 208–225. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, pronotum, dorsal view. 208. M.caperatus sp. nov., holotype, ♂ (NME). 209. M.ciliatus sp. nov., holotype, ♂ (IECA). 210. M.dani sp. nov., holotype, ♂ (IECA). 211. M. deltouri sp. nov., holotype, ♂ (BMNH). 212. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 213. M. esau Heller, 1914 (lectotype, ♂ of M. setosus Moser, 1920) (ZMHB). 214. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 215. M. hamatus sp. nov., holotype, ♂ (IECA). 216. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 217. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 218. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 219. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 220. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 221. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 222. M. konjo sp. nov., holotype, ♂ (NMPC). 223. M. lapsus sp. nov., holotype, ♂ (BMNH). 224. M. legalovi sp. nov., holotype, ♂ (BMNH). 225. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122191" httpUri="https://zenodo.org/record/4122191/files/figure.png" pageId="51" pageNumber="52">216</figureCitation>
,
<figureCitation id="0AA52A659D2F9268FCDEFC11CB2A2BAB" box="[845,892,943,969]" captionStart="Figs 266–277" captionStartId="150.[189,241,1753,1779]" captionTargetBox="[366,1222,265,1687]" captionTargetId="figure-144@150.[366,1222,265,1703]" captionTargetPageId="150" captionText="Figs 266–277. Papuan and Wallacean Maechidius Macleay, 1819, pronotal hypomeron, ventrolateral view. 266. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 267. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 268. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 269. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 270. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 271. M. mailu sp. nov., holotype, ♂ (NHMB). 272. M. lobaticeps Frey, 1969, ♂ (BMNH). 273. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 274. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 275. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 276. M. peregrinus Lansberge, 1886, ♂ (DTC). 277. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122200" httpUri="https://zenodo.org/record/4122200/files/figure.png" pageId="51" pageNumber="52">267</figureCitation>
,
<figureCitation id="0AA52A659D2F9268FC14FC11CBE02BAB" box="[903,950,943,969]" captionStart="Figs 302–316" captionStartId="153.[189,241,1679,1705]" captionTargetBox="[393,1194,265,1649]" captionTargetId="figure-159@153.[393,1194,265,1658]" captionTargetPageId="153" captionText="Figs 302–316. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, elytron, dorsal view. 302. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 303. M. esau Heller, 1914, lectotype, ♂ (SNSD). 304. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 305. M. hamatus sp. nov., holotype, ♂ (IECA). 306. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 307. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 308. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 309. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 310. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 311. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 312. Same, enlarged. 313. M. konjo sp. nov., holotype, ♂ (NMPC). 314. M. lapsus sp. nov., holotype, ♂ (BMNH). 315. M. legalovi sp. nov., holotype, ♂ (BMNH). 316. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122208" httpUri="https://zenodo.org/record/4122208/files/figure.png" pageId="51" pageNumber="52">306</figureCitation>
, 380, 473
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D2F9268FF2EFC51CB712C49" pageId="51" pageNumber="52" type="reference_group">
<paragraph id="922136E09D2F9268FF2EFC51CAB22C6B" blockId="51.[189,807,1007,1068]" box="[189,740,1007,1034]" pageId="51" pageNumber="52">
<treatmentCitationGroup id="B28E11CE9D2F9268FF2EFC51CAB22C6B" box="[189,740,1007,1034]" pageId="51" pageNumber="52">
<treatmentCitation id="133F10F19D2F9268FF2EFC51CA8B2C68" author="Heller K. M." box="[189,733,1007,1034]" page="23" pageId="51" pageNumber="52" year="1910">
<taxonomicName id="559E4D639D2F9268FF2EFC51CA8B2C68" ID-CoL="7X3WT" authority="Heller, 1910: 23" authorityName="Heller" authorityPageNumber="23" authorityYear="1910" box="[189,733,1007,1034]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="heterosquamosus">
<emphasis id="A0EAEAF29D2F9268FF2EFC51CA4E2C6B" box="[189,536,1007,1033]" italics="true" pageId="51" pageNumber="52">Maechidius heterosquamosus</emphasis>
<bibRefCitation id="F60F4B119D2F9268FD8CFC51CA8B2C68" author="Heller K. M." box="[543,733,1007,1034]" pageId="51" pageNumber="52" pagination="1 - 42" refId="ref98170" refString="Heller K. M. 1910. Funfter Beitrag zur Papuanischen Kaferfauna hauptsachlich auf Grund der Ausbeute von Dr. Schlaginhaufen. Abhandlungen und Berichte des Koniglichen Zoologischen und Anthropologisch- Ethnographischen Museums zu Dresden 13 (3): 1 - 42." type="journal article" year="1910">Heller, 1910: 23</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="922136E09D2F9268FF2EFBAFCB712C49" blockId="51.[189,807,1007,1068]" box="[189,807,1041,1068]" pageId="51" pageNumber="52">
<treatmentCitationGroup id="B28E11CE9D2F9268FF2EFBAFCB712C49" box="[189,807,1041,1068]" pageId="51" pageNumber="52">
<treatmentCitation id="133F10F19D2F9268FF2EFBAFCAE52C4E" author="Frey G." box="[189,691,1041,1068]" page="508" pageId="51" pageNumber="52" year="1969">
<taxonomicName id="559E4D639D2F9268FF2EFBAFCAE52C4E" ID-CoL="7XWML" authority="Frey, 1969: 508" authorityName="Frey" authorityPageNumber="508" authorityYear="1969" box="[189,691,1041,1068]" class="Insecta" family="Scarabaeidae" genus="Paramaechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="clypeatus">
<emphasis id="A0EAEAF29D2F9268FF2EFBAFC9A22C49" box="[189,500,1041,1067]" italics="true" pageId="51" pageNumber="52">Paramaechidius clypeatus</emphasis>
<bibRefCitation id="F60F4B119D2F9268FE68FBACCAE52C4E" author="Frey G." box="[507,691,1041,1068]" pageId="51" pageNumber="52" pagination="494 - 509" refId="ref98001" refString="Frey G. 1969. Revision der Maechidius - Arten Neu-Guineas (Col., Melolonth.). Entomologische Arbeiten aus dem Museum G. Frey 20: 494 - 509." type="journal article" year="1969">Frey, 1969: 508</bibRefCitation>
</taxonomicName>
</treatmentCitation>
<emphasis id="A0EAEAF29D2F9268FD28FBAFCB712C49" bold="true" box="[699,807,1041,1067]" pageId="51" pageNumber="52">
<taxonomicNameLabel id="BBD957899D2F9268FD28FBAFCB712C49" box="[699,807,1041,1067]" pageId="51" pageNumber="52">syn. nov.</taxonomicNameLabel>
</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D2F9268FF2EFBEFCA102DA7" pageId="51" pageNumber="52" type="materials_examined">
<paragraph id="922136E09D2F9268FF2EFBEFC93A2C09" blockId="51.[189,364,1105,1131]" box="[189,364,1105,1131]" pageId="51" pageNumber="52">
<heading id="C969818C9D2F9268FF2EFBEFC93A2C09" bold="true" box="[189,364,1105,1131]" fontSize="11" level="3" pageId="51" pageNumber="52" reason="3">
<emphasis id="A0EAEAF29D2F9268FF2EFBEFC93A2C09" bold="true" box="[189,364,1105,1131]" pageId="51" pageNumber="52">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D2F9268FF79FBC0CBD92CFB" blockId="51.[189,1399,1150,1311]" box="[234,911,1150,1177]" pageId="51" pageNumber="52">
<emphasis id="A0EAEAF29D2F9268FF79FBC0CADE2CFB" bold="true" box="[234,648,1150,1177]" pageId="51" pageNumber="52">
<typeStatus id="4D2588429D2F9268FF79FBC0C9302CFA" box="[234,358,1150,1176]" pageId="51" pageNumber="52" type="lectotype">Lectotype</typeStatus>
of
<taxonomicName id="559E4D639D2F9268FE1EFBC0CADE2CFB" authorityName="Heller" authorityYear="1910" box="[397,648,1150,1177]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="heterosquamosus">
<emphasis id="A0EAEAF29D2F9268FE1EFBC0CADE2CFB" bold="true" box="[397,648,1150,1177]" italics="true" pageId="51" pageNumber="52">M. heterosquamosus</emphasis>
</taxonomicName>
</emphasis>
(herewith designated)
</paragraph>
<paragraph id="922136E09D2F9268FF2EFB1EC9462D7D" blockId="51.[189,1399,1150,1311]" pageId="51" pageNumber="52">
<materialsCitation id="22F63CBD9D2F9268FF2EFB1EC95A2D7D" ID-GBIF-Occurrence="2898878540" collectionCode="SNSD" country="Papua New Guinea" elevation="640" location="PAPUA NEW GUINEA" pageId="51" pageNumber="52" specimenCount="2" specimenCount-female="1" typeStatus="typus">
<collectingCountry id="EA8976709D2F9268FF2EFB1EC9B52CD8" box="[189,483,1184,1210]" name="Papua New Guinea" pageId="51" pageNumber="52">PAPUA NEW GUINEA</collectingCountry>
•
<specimenCount id="8498FD699D2F9268FD96FB1ECA4A2CD8" box="[517,540,1184,1210]" pageId="51" pageNumber="52" type="female">♀</specimenCount>
; “[upper side of label] Kais.Wilhelmsland Toricelli [sic
<specimenCount id="8498FD699D2F9268FB42FB21CC832CD8" box="[1233,1237,1183,1210]" pageId="51" pageNumber="52" type="male">♂</specimenCount>
] Gebirge [p] [underside of label]
<quantity id="55669B059D2F9268FE27FB7FCA542CBE" box="[436,514,1217,1244]" metricMagnitude="2" metricUnit="m" metricValue="6.4" pageId="51" pageNumber="52" unit="m" value="640.0">
<elevation id="19B3D1D39D2F9268FE27FB7FCA542CBE" box="[436,514,1217,1244]" metricMagnitude="2" metricUnit="m" metricValue="6.4" pageId="51" pageNumber="52" unit="m" value="640.0">640 m</elevation>
</quantity>
[h] [blue label]// 1910 [p] 1 [h] [blue label]// [upper side of label]
<typeStatus id="4D2588429D2F9268FAA4FB7CCD212CBE" box="[1335,1399,1218,1244]" pageId="51" pageNumber="52">typus</typeStatus>
[p] [underside of label]
<taxonomicName id="559E4D639D2F9268FE4AFB5DCAF22C9F" authorityName="Heller" authorityYear="1910" box="[473,676,1251,1277]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="heterosquamosus">
<emphasis id="A0EAEAF29D2F9268FE4AFB5DCAF22C9F" box="[473,676,1251,1277]" italics="true" pageId="51" pageNumber="52">heterosquamosus</emphasis>
</taxonomicName>
[h] [red label]// Staatl. Museum für Tierkunde.Dresden [p]”;
<collectionCode id="F48FAE259D2F9268FF2EFABBC95A2D7D" box="[189,268,1285,1311]" collectionName="SNSD" pageId="51" pageNumber="52">SNSD</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="922136E09D2F9268FF79FAFBCA4A2D3D" blockId="51.[189,1398,1348,1477]" box="[234,540,1348,1375]" pageId="51" pageNumber="52">
<emphasis id="A0EAEAF29D2F9268FF79FAFBCA4A2D3D" bold="true" box="[234,540,1348,1375]" pageId="51" pageNumber="52">
<typeStatus id="4D2588429D2F9268FF79FAFBC90A2D3D" box="[234,348,1349,1375]" pageId="51" pageNumber="52" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="559E4D639D2F9268FE17FAFBCA4A2D3D" authorityName="Frey" authorityYear="1969" box="[388,540,1348,1375]" class="Insecta" family="Scarabaeidae" genus="Paramaechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="clypeatus">
<emphasis id="A0EAEAF29D2F9268FE17FAFBCA4A2D3D" bold="true" box="[388,540,1348,1375]" italics="true" pageId="51" pageNumber="52">M. clypeatus</emphasis>
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="922136E09D2F9268FF2EFAD9CA102DA7" blockId="51.[189,1398,1348,1477]" pageId="51" pageNumber="52">
<materialsCitation id="22F63CBD9D2F9268FF2EFAD9CA102DA7" ID-GBIF-Occurrence="2898878440" collectingDate="II- 28 - III- 4" collectionCode="NHMB" collectorName="W. W. Brandt Collector Bishop" country="Papua New Guinea" determinerName="G. Frey" elevation="750" location="NEW GUINEA" municipality="Mts. Mobitei" pageId="51" pageNumber="52" specimenCount="751" typeStatus="type">
<collectingCountry id="EA8976709D2F9268FF2EFAD9C9B72DE3" box="[189,481,1383,1409]" name="Papua New Guinea" pageId="51" pageNumber="52">PAPUA NEW GUINEA</collectingCountry>
•
<specimenCount id="8498FD699D2F9268FD92FAD9CA4F2DE3" box="[513,537,1383,1409]" pageId="51" pageNumber="52" type="male">♂</specimenCount>
; “
<location id="9741603B9D2F9268FDA6FAD9CAAD2DE3" LSID="urn:lsid:plazi:treatment:1A3787F69D2F926EFE46FC33CA852C94:9741603B9D2F9268FDA6FAD9CAAD2DE3" box="[565,763,1383,1409]" country="Papua New Guinea" municipality="Mts. Mobitei" name="NEW GUINEA" pageId="51" pageNumber="52">NEW GUINEA</location>
:
<locationDeviation id="83932FC29D2F9268FC9AFAD9CBE62DE3" box="[777,944,1383,1409]" pageId="51" pageNumber="52">NE. Torricelli</locationDeviation>
Mts. Mobitei,
<specimenCount id="8498FD699D2F9268FBF8FAD8CC962DE3" box="[1131,1216,1382,1409]" pageId="51" pageNumber="52" type="generic">
<quantity id="55669B059D2F9268FBF8FAD8CCEA2DE3" box="[1131,1212,1382,1409]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="51" pageNumber="52" unit="m" value="750.0">750 m</quantity>
.
</specimenCount>
<date id="E62010209D2F9268FB58FAD8CD1C2DE3" box="[1227,1354,1382,1409]" pageId="51" pageNumber="52">
<collectingDate id="F664E9C89D2F9268FB58FAD8CD1C2DE3" box="[1227,1354,1382,1409]" pageId="51" pageNumber="52">II-28-III-4</collectingDate>
</date>
-’59 [p] //
<collectorName id="3F6B53369D2F9268FF65FA37CA3F2DC0" box="[246,617,1416,1443]" pageId="51" pageNumber="52">W.W. Brandt Collector Bishop</collectorName>
[p] //
<typeStatus id="4D2588429D2F9268FD3EFA37CAB12DC1" box="[685,743,1417,1443]" pageId="51" pageNumber="52">Type</typeStatus>
[h, red label]//
<typeStatus id="4D2588429D2F9268FC3BFA37CBA72DC1" box="[936,1009,1417,1443]" pageId="51" pageNumber="52">TYPE</typeStatus>
<taxonomicName id="559E4D639D2F9268FC6EFA36CD612DC0" authorityName="Frey" authorityYear="1969" box="[1021,1335,1416,1442]" class="Insecta" family="Scarabaeidae" genus="Paramaechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="clypeatus">
<emphasis id="A0EAEAF29D2F9268FC6EFA36CD612DC0" box="[1021,1335,1416,1442]" italics="true" pageId="51" pageNumber="52">Paramaechidius clypeatus</emphasis>
</taxonomicName>
n sp det.
<determinerName id="67FDDA249D2F9268FF74FA14C96C2DA6" box="[231,314,1450,1476]" pageId="51" pageNumber="52">G.Frey</determinerName>
,1967/68 [p]”;
<collectionCode id="F48FAE259D2F9268FE75FA15CA142DA7" box="[486,578,1451,1477]" collectionName="Switzerland, Basel, Naturhistorisches Museum" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="51" pageNumber="52">NHMB</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D2F9268FF2EFA54CC712EB4" pageId="51" pageNumber="52" type="discussion">
<paragraph id="922136E09D2F9268FF2EFA54C9792E66" blockId="51.[189,303,1514,1540]" box="[189,303,1514,1540]" pageId="51" pageNumber="52">
<heading id="C969818C9D2F9268FF2EFA54C9792E66" bold="true" box="[189,303,1514,1540]" fontSize="11" level="3" pageId="51" pageNumber="52" reason="3">
<emphasis id="A0EAEAF29D2F9268FF2EFA54C9792E66" bold="true" box="[189,303,1514,1540]" pageId="51" pageNumber="52">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D2F9268FF2EF9A9CD112E50" blockId="51.[189,1351,1559,1586]" box="[189,1351,1559,1586]" pageId="51" pageNumber="52">
The
<typeStatus id="4D2588429D2F9268FF61F9A6C9012E50" box="[242,343,1560,1586]" pageId="51" pageNumber="52" type="paratype">paratype</typeStatus>
of
<taxonomicName id="559E4D639D2F9268FEEDF9A6CA422E53" authorityName="Frey" authorityYear="1969" box="[382,532,1559,1585]" class="Insecta" family="Scarabaeidae" genus="Paramaechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="clypeatus">
<emphasis id="A0EAEAF29D2F9268FEEDF9A6CA422E53" box="[382,532,1559,1585]" italics="true" pageId="51" pageNumber="52">M. clypeatus</emphasis>
</taxonomicName>
from NHMB (
<bibRefCitation id="F60F4B119D2F9268FD50F9A9CB2C2E50" author="Frey G." box="[707,890,1559,1586]" pageId="51" pageNumber="52" pagination="494 - 509" refId="ref98001" refString="Frey G. 1969. Revision der Maechidius - Arten Neu-Guineas (Col., Melolonth.). Entomologische Arbeiten aus dem Museum G. Frey 20: 494 - 509." type="journal article" year="1969">Frey 1969: 508</bibRefCitation>
) belongs to
<taxonomicName id="559E4D639D2F9268FB99F9A9CCB32E53" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1034,1253,1559,1585]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="lapsus" status="sp. nov.">
<emphasis id="A0EAEAF29D2F9268FB99F9A9CCB32E53" box="[1034,1253,1559,1585]" italics="true" pageId="51" pageNumber="52">Maechidius lapsus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D2F9268FB7FF9A6CD112E50" box="[1260,1351,1560,1586]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="922136E09D2F9268FF2EF9E9CC712EB4" blockId="51.[189,1398,1623,1751]" pageId="51" pageNumber="52">
<bibRefCitation id="F60F4B119D2F9268FF2EF9E9C9DF2E13" author="Heller K. M." box="[189,393,1623,1649]" pageId="51" pageNumber="52" pagination="1 - 42" refId="ref98170" refString="Heller K. M. 1910. Funfter Beitrag zur Papuanischen Kaferfauna hauptsachlich auf Grund der Ausbeute von Dr. Schlaginhaufen. Abhandlungen und Berichte des Koniglichen Zoologischen und Anthropologisch- Ethnographischen Museums zu Dresden 13 (3): 1 - 42." type="journal article" year="1910">Heller (1910: 23)</bibRefCitation>
likely based his description of
<taxonomicName id="559E4D639D2F9268FD6BF9E6CBBC2E13" authorityName="Heller" authorityYear="1910" box="[760,1002,1623,1649]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="heterosquamosus">
<emphasis id="A0EAEAF29D2F9268FD6BF9E6CBBC2E13" box="[760,1002,1623,1649]" italics="true" pageId="51" pageNumber="52">M. heterosquamosus</emphasis>
</taxonomicName>
comb. rest. on a single specimen, although not explicitly stated. The single SNSD specimen labelled “typus” is herewith designated as
<typeStatus id="4D2588429D2F9268FF2EF924C97F2ED6" box="[189,297,1690,1716]" pageId="51" pageNumber="52" type="lectotype">lectotype</typeStatus>
in order to enhance the stability of nomenclature and fix the specimen I based my redescription and new synonymy on, in case other
<typeStatus id="4D2588429D2F9268FDE3F903CA812EB5" box="[624,727,1725,1751]" pageId="51" pageNumber="52" type="syntype">syntypes</typeStatus>
are discovered in the future.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D2F926EFF2EF942CA852C94" lastPageId="53" lastPageNumber="54" pageId="51" pageNumber="52" type="description">
<paragraph id="922136E09D2F9268FF2EF942C9182F74" blockId="51.[189,334,1788,1814]" box="[189,334,1788,1814]" pageId="51" pageNumber="52">
<heading id="C969818C9D2F9268FF2EF942C9182F74" bold="true" box="[189,334,1788,1814]" fontSize="11" level="3" pageId="51" pageNumber="52" reason="3">
<emphasis id="A0EAEAF29D2F9268FF2EF942C9182F74" bold="true" box="[189,334,1788,1814]" pageId="51" pageNumber="52">Description</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D2F9268FF2EF894CD7C2F26" blockId="51.[189,1322,1833,1860]" box="[189,1322,1833,1860]" pageId="51" pageNumber="52">
MEASUREMENTS. Total body length 7.00–
<quantity id="55669B059D2F9268FD00F897CAAD2F26" box="[659,763,1833,1860]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="51" pageNumber="52" unit="mm" value="8.1">8.10 mm</quantity>
(
<typeStatus id="4D2588429D2F9268FC99F897CB212F21" box="[778,887,1833,1859]" pageId="51" pageNumber="52" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="559E4D639D2F9268FC0DF894CCC72F21" authorityName="Heller" authorityYear="1910" box="[926,1169,1833,1859]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="heterosquamosus">
<emphasis id="A0EAEAF29D2F9268FC0DF894CCC72F21" box="[926,1169,1833,1859]" italics="true" pageId="51" pageNumber="52">M. heterosquamosus</emphasis>
</taxonomicName>
comb. rest.).
</paragraph>
<paragraph id="922136E09D2F926EFF2EF8D4C9592CE8" blockId="51.[189,1399,1897,2025]" lastBlockId="53.[189,1399,265,1162]" lastPageId="53" lastPageNumber="54" pageId="51" pageNumber="52">
Dorsum and venter uniformly brown. Head strongly convex dorsally between eyes, glossy dorsally and ventrally. Female labroclypeus (
<figureCitation id="0AA52A659D2F9268FDA5F835CA8F2FC4" box="[566,729,1931,1958]" captionStart="Figs 121–132" captionStartId="139.[189,241,1716,1742]" captionTargetBox="[357,1230,276,1689]" captionTargetId="figure-149@139.[357,1230,265,1697]" captionTargetPageId="139" captionText="Figs 121–132. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, head, dorsal (121, 123–124, 126–132) and frontal (122, 125) views. 121. M. hamatus sp. nov., holotype, ♂ (IECA). 122. M. helleri (Frey, 1969) comb. nov., holotype, ♂ (photograph courtesy BPBM) (BPBM). 123. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 124. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 125. Same, ♂, holotype of M. clypeatus (Frey, 1969) comb. nov. (BMNH). 126. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 127. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 128. Same, paralectotype, ♀ (SNSD). 129. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 130. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 131. M. konjo sp. nov., holotype, ♂ (NMPC). 132. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122177" httpUri="https://zenodo.org/record/4122177/files/figure.png" pageId="51" pageNumber="52">Figs 124–125</figureCitation>
) broadly emarginate anteriorly, its lateral margins nearly straight in dorsal, slightly sinuous in lateral view.Anterolateral angles acute, protruding, raised up at angle of nearly 80-90° to frons in lateral view. Anterior and lateral margins of labroclypeus smooth. Underside of labroclypeus with sparse long setae, upper side with clavate short setae along lateral margins. Canthus straight in dorsal view. Punctures of frons ovoid, very deep and dense, on periphery encircled with dense microscopical velvety pubescence. Intervening spaces glossy, generally smaller than punctures. Setae moderately long, scale-like, suberect to erect, variably strongly clavate to fusiform. Female antenna 9-segmented, club 3-lamellate. Pronotum transverse, glossy dorsally and laterally. Anterior margin of pronotum broadly emarginate with protruding anterolateral angles, basal margin broadly rounded. Lateral margin of pronotum sinuous in lateral view, with obtuse median protruding in dorsal view, slightly emarginate anteriorly and somewhat stronger towards base, delicately (indistinctly) crenulate all along (
<figureCitation id="0AA52A659D29926EFEA1FDABC9CF2A52" box="[306,409,533,560]" captionStart="Figs 208–225" captionStartId="146.[189,241,1644,1670]" captionTargetBox="[414,1187,265,1610]" captionTargetId="figure-199@146.[414,1187,265,1618]" captionTargetPageId="146" captionText="Figs 208–225. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, pronotum, dorsal view. 208. M.caperatus sp. nov., holotype, ♂ (NME). 209. M.ciliatus sp. nov., holotype, ♂ (IECA). 210. M.dani sp. nov., holotype, ♂ (IECA). 211. M. deltouri sp. nov., holotype, ♂ (BMNH). 212. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 213. M. esau Heller, 1914 (lectotype, ♂ of M. setosus Moser, 1920) (ZMHB). 214. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 215. M. hamatus sp. nov., holotype, ♂ (IECA). 216. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 217. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 218. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 219. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 220. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 221. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 222. M. konjo sp. nov., holotype, ♂ (NMPC). 223. M. lapsus sp. nov., holotype, ♂ (BMNH). 224. M. legalovi sp. nov., holotype, ♂ (BMNH). 225. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122191" httpUri="https://zenodo.org/record/4122191/files/figure.png" pageId="53" pageNumber="54">Fig. 216</figureCitation>
). Clavate erect seta present between every two crenulae. Lateral margin of pronotum sinuous in lateral view. Hypomeron emarginate and long setose on anterior margin which is flange-like (
<figureCitation id="0AA52A659D29926EFF55FDE6C97A2A11" box="[198,300,600,627]" captionStart="Figs 266–277" captionStartId="150.[189,241,1753,1779]" captionTargetBox="[366,1222,265,1687]" captionTargetId="figure-144@150.[366,1222,265,1703]" captionTargetPageId="150" captionText="Figs 266–277. Papuan and Wallacean Maechidius Macleay, 1819, pronotal hypomeron, ventrolateral view. 266. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 267. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 268. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 269. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 270. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 271. M. mailu sp. nov., holotype, ♂ (NHMB). 272. M. lobaticeps Frey, 1969, ♂ (BMNH). 273. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 274. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 275. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 276. M. peregrinus Lansberge, 1886, ♂ (DTC). 277. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122200" httpUri="https://zenodo.org/record/4122200/files/figure.png" pageId="53" pageNumber="54">Fig. 267</figureCitation>
). Antennal pocket deep. Pronotal punctures deeper than those on head, of variable shape (ovoid to hexagonal). Scale-like clavate seta rises from anterior margin of each puncture. Lateral, basal margins and hypomeron covered with dense microscopical velvety (fur-like) pubescence (
<figureCitation id="0AA52A659D29926EFB1DFD25CCAE2AD7" box="[1166,1272,666,693]" captionStart="Figs 266–277" captionStartId="150.[189,241,1753,1779]" captionTargetBox="[366,1222,265,1687]" captionTargetId="figure-144@150.[366,1222,265,1703]" captionTargetPageId="150" captionText="Figs 266–277. Papuan and Wallacean Maechidius Macleay, 1819, pronotal hypomeron, ventrolateral view. 266. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 267. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 268. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 269. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 270. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 271. M. mailu sp. nov., holotype, ♂ (NHMB). 272. M. lobaticeps Frey, 1969, ♂ (BMNH). 273. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 274. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 275. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 276. M. peregrinus Lansberge, 1886, ♂ (DTC). 277. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122200" httpUri="https://zenodo.org/record/4122200/files/figure.png" pageId="53" pageNumber="54">Fig. 267</figureCitation>
). Scutellar shield pointed apically. Elytron opaque, with five raised glabrous partly interrupted longitudinal carinae, including sutural one. Punctures of elytral disc irregular, small to rather large, deep, of variable shape (
<figureCitation id="0AA52A659D29926EFF55FD41C9672B78" box="[198,305,767,794]" captionStart="Figs 302–316" captionStartId="153.[189,241,1679,1705]" captionTargetBox="[393,1194,265,1649]" captionTargetId="figure-159@153.[393,1194,265,1658]" captionTargetPageId="153" captionText="Figs 302–316. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, elytron, dorsal view. 302. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 303. M. esau Heller, 1914, lectotype, ♂ (SNSD). 304. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 305. M. hamatus sp. nov., holotype, ♂ (IECA). 306. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 307. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 308. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 309. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 310. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 311. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 312. Same, enlarged. 313. M. konjo sp. nov., holotype, ♂ (NMPC). 314. M. lapsus sp. nov., holotype, ♂ (BMNH). 315. M. legalovi sp. nov., holotype, ♂ (BMNH). 316. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122208" httpUri="https://zenodo.org/record/4122208/files/figure.png" pageId="53" pageNumber="54">Fig. 306</figureCitation>
). Intervening spaces hidden under dense microscopical velvety pubescence. Pronotal setae scale-like, short to rather long, strongly to less strong clavate, suberect to erect. Obtuse partially glossy hump near apex. Female pygidium slightly longitudinally impressed dorsally, with dense deep ovoid punctures (except on anterior margin) and long dense erect scale-like clavate (medially) to ordinary (along posterior margin) setae. Elytral setae generally pointing obliquely to middle of pygidium. Midline of female pygidium generally sparser punctured, in part glabrous (
<figureCitation id="0AA52A659D29926EFC5BFC18CC662BA2" box="[968,1072,934,960]" captionStart="Figs 469–486" captionStartId="164.[189,241,1681,1707]" captionTargetBox="[450,1137,265,1633]" captionTargetId="figure-184@164.[450,1138,265,1647]" captionTargetPageId="164" captionText="Figs 469–486. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, pygidium, dorsal view. 469. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 470. M. esau Heller, 1914, ♂ (BMNH). 471. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 472. M. hamatus sp. nov., holotype, ♂ (IECA). 473. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 474. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 475. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 476. Same, paralectotype, ♀ (SNSD). 477. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 478. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 479. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 480. M. konjo sp. nov., holotype, ♂ (NMPC). 481. M. lapsus sp. nov., holotype, ♂ (BMNH). 482. Same, paratype, ♀ (BMNH). 483. M. legalovi sp. nov., holotype, ♂ (BMNH). 484. Same, paratype, ♀ (BMNH). 485. M. leucopsar sp. nov., holotype, ♂ (IECA). 486. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122224" httpUri="https://zenodo.org/record/4122224/files/figure.png" pageId="53" pageNumber="54">Fig. 473</figureCitation>
). Intervening spaces glossy, covered with fur-like microscopical pubescence except along midline. Venter covered with extremely delicate fur-like microscopical pubescence. Protibia slightly widened distally, dorsally with delicate longitudinal carina, in female with two large distal external teeth of which distal one is stronger and narrower (Fig. 380). Female protibial terminal spur short, nearly straight. Female metatibial terminal spurs strongly unequal, lower one about twice as long as upper. Tarsal claws without pulvilli (in females only?).
</paragraph>
<caption id="C6E166689D28926FFF2EF8DAC9E82F85" ID-DOI="http://doi.org/10.5281/zenodo.4122144" ID-Zenodo-Dep="4122144" httpUri="https://zenodo.org/record/4122144/files/figure.png" pageId="52" pageNumber="53" startId="52.[189,241,1892,1918]" targetBox="[334,1252,265,1842]" targetPageId="52">
<paragraph id="922136E09D28926FFF2EF8DAC9E82F85" blockId="52.[189,1399,1892,2024]" pageId="52" pageNumber="53">
<emphasis id="A0EAEAF29D28926FFF2EF8DAC91C2F1C" bold="true" box="[189,330,1892,1918]" pageId="52" pageNumber="53">Figs 42–45.</emphasis>
Papuan
<taxonomicName id="559E4D639D28926FFE21F8DACAA22F1C" authority="Macleay, 1819" authorityName="Macleay" authorityYear="1819" box="[434,756,1892,1918]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="A0EAEAF29D28926FFE21F8DACA6D2F1C" box="[434,571,1892,1918]" italics="true" pageId="52" pageNumber="53">Maechidius</emphasis>
<bibRefCitation id="F60F4B119D28926FFDD7F8DACAA22F1C" author="Macleay W. S." box="[580,756,1892,1918]" pageId="52" pageNumber="53" refId="ref98319" refString="Macleay W. S. 1819. Horae Entomologicae: or Essays on the Annulose Animals. Volume I, Part I. Containing General Observations on the Geography, Manners, and the Affinities of the Insects which compose the Genus Scarabaeus of Linnaeus; to which are added a few incidental Remarks on the Genera Lucanus and Hister of the same Author. With Appendix and Plates. R. and A. Taylor, London." type="book" year="1819">Macleay, 1819</bibRefCitation>
</taxonomicName>
, habitus, dorsal view.
<emphasis id="A0EAEAF29D28926FFC6DF8DACC4D2F1C" bold="true" box="[1022,1051,1892,1918]" pageId="52" pageNumber="53">42</emphasis>
.
<taxonomicName id="559E4D639D28926FFBB8F8DBC8AE2FC3" authority="Heller, 1910" authorityName="Heller" authorityYear="1910" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="heterosquamosus">
<emphasis id="A0EAEAF29D28926FFBB8F8DBCD492F1C" box="[1067,1311,1892,1918]" italics="true" pageId="52" pageNumber="53">M. heterosquamosus</emphasis>
<bibRefCitation id="F60F4B119D28926FFAB4F8DAC8AE2FC3" author="Heller K. M." pageId="52" pageNumber="53" pagination="1 - 42" refId="ref98170" refString="Heller K. M. 1910. Funfter Beitrag zur Papuanischen Kaferfauna hauptsachlich auf Grund der Ausbeute von Dr. Schlaginhaufen. Abhandlungen und Berichte des Koniglichen Zoologischen und Anthropologisch- Ethnographischen Museums zu Dresden 13 (3): 1 - 42." type="journal article" year="1910">Heller, 1910</bibRefCitation>
</taxonomicName>
comb. rest., lectotype, ♀ (SNSD).
<emphasis id="A0EAEAF29D28926FFD00F839CAE62FC3" bold="true" box="[659,688,1927,1953]" pageId="52" pageNumber="53">43</emphasis>
. Same, ♂, holotype of
<taxonomicName id="559E4D639D28926FFC52F836CCB82FC0" authority="(Frey, 1969)" authorityName="Telnov" authorityYear="2020" baseAuthorityName="Frey" baseAuthorityYear="1969" box="[961,1262,1927,1954]" class="Insecta" family="Scarabaeidae" genus="Paramaechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="clypeatus" status="comb. nov.">
<emphasis id="A0EAEAF29D28926FFC52F836CC012FC3" box="[961,1111,1927,1953]" italics="true" pageId="52" pageNumber="53">M. clypeatus</emphasis>
(
<bibRefCitation id="F60F4B119D28926FFBF5F839CCB02FC0" author="Frey G." box="[1126,1254,1927,1954]" pageId="52" pageNumber="53" pagination="494 - 509" refId="ref98001" refString="Frey G. 1969. Revision der Maechidius - Arten Neu-Guineas (Col., Melolonth.). Entomologische Arbeiten aus dem Museum G. Frey 20: 494 - 509." type="journal article" year="1969">Frey, 1969</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="BBD957899D28926FFB66F839CD212FC0" box="[1269,1399,1927,1954]" pageId="52" pageNumber="53" rank="species">comb. nov.</taxonomicNameLabel>
(BPBM).
<emphasis id="A0EAEAF29D28926FFEBDF814C91D2FA6" bold="true" box="[302,331,1962,1988]" pageId="52" pageNumber="53">44</emphasis>
.
<taxonomicName id="559E4D639D28926FFECBF815CA382FA6" authority="Arrow, 1941" authorityName="Arrow" authorityYear="1941" box="[344,622,1962,1988]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="hirtipes">
<emphasis id="A0EAEAF29D28926FFECBF815C9812FA6" box="[344,471,1962,1988]" italics="true" pageId="52" pageNumber="53">M. hirtipes</emphasis>
<bibRefCitation id="F60F4B119D28926FFE49F814CA382FA6" author="Arrow G. E." box="[474,622,1962,1988]" pageId="52" pageNumber="53" pagination="448 - 464" refId="ref97558" refString="Arrow G. E. 1941. XXIX. - Some undescribed species of melolonthid Coleoptera from Western New Guinea and the adjacent islands of Waigeu and Japen. Annals and Magazine of Natural History, Series 11 7 (41): 448 - 464. https: // doi. org / 10.1080 / 03745481.1941.9727945" type="journal article" year="1941">Arrow, 1941</bibRefCitation>
</taxonomicName>
, lectotype, ♂ (BMNH).
<emphasis id="A0EAEAF29D28926FFC17F814CBF72FA6" bold="true" box="[900,929,1962,1988]" pageId="52" pageNumber="53">45</emphasis>
.
<taxonomicName id="559E4D639D28926FFC3EF815CC882FA6" authority="Heller, 1914" authorityName="Heller" authorityYear="1914" box="[941,1246,1962,1988]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="humeralis">
<emphasis id="A0EAEAF29D28926FFC3EF815CC112FA6" box="[941,1095,1962,1988]" italics="true" pageId="52" pageNumber="53">M. humeralis</emphasis>
<bibRefCitation id="F60F4B119D28926FFBDEF814CC882FA6" author="Heller K. M." box="[1101,1246,1962,1988]" pageId="52" pageNumber="53" pagination="615 - 666" refId="ref98213" refString="Heller K. M. 1914. Coleoptera. Nova Guinea. Resultats de l'Expedition scientifique neerlandaise a la Nouvelle-Guinee en 1907 et 1909 sous les auspices du Dr. H. A. Lorentz. Zoologie 9: 615 - 666." type="journal article" year="1914">Heller, 1914</bibRefCitation>
</taxonomicName>
, lectotype, ♂ (SNSD). Not to scale.
</paragraph>
</caption>
<paragraph id="922136E09D29926EFF2EFB11C9E62CAB" blockId="53.[189,432,1199,1225]" box="[189,432,1199,1225]" pageId="53" pageNumber="54">
<heading id="C969818C9D29926EFF2EFB11C9E62CAB" bold="true" box="[189,432,1199,1225]" fontSize="11" level="3" pageId="53" pageNumber="54" reason="3">
<emphasis id="A0EAEAF29D29926EFF2EFB11C9E62CAB" bold="true" box="[189,432,1199,1225]" pageId="53" pageNumber="54">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D29926EFF2EFB62CA852C94" blockId="53.[189,723,1244,1270]" box="[189,723,1244,1270]" pageId="53" pageNumber="54">This species’ male genitalia were not studied.</paragraph>
</subSubSection>
</treatment>
</document>
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