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<document id="DB0317501856490832DDA2DC19E5639B" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D02927BFDCEFEB6CA612A4D" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius boessnecki Telnov 2020, sp. nov." docType="treatment" docUuid="C4A68BE8-8247-4C13-93A5-DEB150FB8B09" docUuidSource="ZooBank" docVersion="10" lastPageNumber="33" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="31" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="81DD5429B989C4C76B57EE09A8144499" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A2786B54553D104A9E41E75288357730">
<mods:title id="70993BBD73585DAC45B1BCD9EB568F9D">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
</mods:titleInfo>
<mods:name id="418B74B4A689A68F07C1739698D1C558" type="personal">
<mods:role id="E5E05AA40469F33ABD30F1B8E64D907B">
<mods:roleTerm id="BD8C07FA45BBE58352B4DAD33FF7A0B4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4FD9F5E29C4BAE1B1D26C39D4E32EB7D">Telnov, Dmitry</mods:namePart>
</mods:name>
<mods:typeOfResource id="18028EF9A570130853EE5D757338D381">text</mods:typeOfResource>
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<mods:titleInfo id="6E55E1E53F00DCB95BFC01B480320A6E">
<mods:title id="F4D1939E6436CA66CA8C719D16E635A5">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="A1B3D690456F702A18E0207DC9DCE47B">
<mods:date id="DC9C8F1312A306969B78FFA591AC8C1B">2020</mods:date>
<mods:detail id="FC2556C73AB268ED943E47C8B972F0D3" type="pubDate">
<mods:number id="8A21E9CBFE78F7EE196A1B705257A956">2020-10-19</mods:number>
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<mods:detail id="87DB0EDCA44F8D6052F59349EFE323B6" type="volume">
<mods:number id="0C4D4CBA9DF3D171BF50289AC6BD3C5B">721</mods:number>
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<mods:classification id="723A156DA15687584F0446EEF0942157">journal article</mods:classification>
<mods:identifier id="55743FB519ACA071C16AD014FC9E3932" type="CLB-Dataset">9693</mods:identifier>
<mods:identifier id="617D2AD894758B97915D81CED5A54EC9" type="DOI">10.5852/ejt.2020.721.1127</mods:identifier>
<mods:identifier id="5D977C5C628949B3A75CAEFA1A116E0D" type="GBIF-Dataset">0cd61b23-1089-4c1b-bfc0-487887abad84</mods:identifier>
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</mods:mods>
<treatment id="1A3787F69D02927BFDCEFEB6CA612A4D" ID-DOI="http://doi.org/10.5281/zenodo.4344531" ID-GBIF-Taxon="168648186" ID-Zenodo-Dep="4344531" LSID="urn:lsid:zoobank.org:act:C4A68BE8-8247-4C13-93A5-DEB150FB8B09" httpUri="http://treatment.plazi.org/id/1A3787F69D02927BFDCEFEB6CA612A4D" lastPageId="32" lastPageNumber="33" pageId="30" pageNumber="31">
<subSubSection id="DA84656B9D029245FDCEFEB6CB812941" box="[605,983,264,291]" pageId="30" pageNumber="31" type="nomenclature">
<paragraph id="922136E09D029245FDCEFEB6CB812941" blockId="30.[364,1224,264,363]" box="[605,983,264,291]" pageId="30" pageNumber="31">
<heading id="C969818C9D029245FDCEFEB6CB812941" box="[605,983,264,291]" centered="true" fontSize="11" level="2" pageId="30" pageNumber="31" reason="2">
<taxonomicName id="559E4D639D029245FDCEFEB6CB222941" ID-CoL="7X3WD" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[605,884,264,291]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="boessnecki" status="sp. nov.">
<emphasis id="A0EAEAF29D029245FDCEFEB6CB222941" bold="true" box="[605,884,264,291]" italics="true" pageId="30" pageNumber="31">Maechidius boessnecki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D029245FCEFFEB7CB812941" box="[892,983,265,291]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D029245FEFFFE93CC442909" pageId="30" pageNumber="31" type="description">
<paragraph id="922136E09D029245FEFFFE93CC9E292A" blockId="30.[364,1224,264,363]" box="[364,1224,301,328]" pageId="30" pageNumber="31">
<uri id="E60F3AE29D029245FEFFFE93CC9E292A" box="[364,1224,301,328]" pageId="30" pageNumber="31">
urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
<paragraph id="922136E09D029245FDB2FEEFCC442909" blockId="30.[364,1224,264,363]" box="[545,1042,336,363]" pageId="30" pageNumber="31">
<figureCitation id="0AA52A659D029245FDB2FEEFCA2D2909" box="[545,635,337,363]" captionStart="Figs 28–30" captionStartId="31.[189,241,1928,1954]" captionTargetBox="[347,1240,269,1901]" captionTargetId="figure-52@31.[345,1242,265,1911]" captionTargetPageId="31" captionText="Figs 28–30. Papuan and Wallacean Maechidius Macleay, 1819, habitus, dorsal view. 28. M. boessnecki sp. nov., holotype, ♂ (SNSD). 29. M. brocki sp. nov., holotype, ♂ (BMNH). 30. M. caperatus sp. nov., holotype, ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122136" httpUri="https://zenodo.org/record/4122136/files/figure.png" pageId="30" pageNumber="31">Figs 28</figureCitation>
,
<figureCitation id="0AA52A659D029245FD14FEEECAE32909" box="[647,693,336,363]" captionStart="Figs 97–108" captionStartId="137.[189,241,1786,1812]" captionTargetBox="[351,1228,270,1756]" captionTargetId="figure-127@137.[351,1237,265,1772]" captionTargetPageId="137" captionText="Figs 97–108. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 97. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 98. M. aiyura sp. nov., holotype, ♂ (BMNH). 99. M. alesbezdeki sp. nov., holotype, ♂ (NME). 100. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 101. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 102. M. bintang sp. nov., holotype, ♂ (NME). 103. M. awu sp. nov., holotype, ♂ (IECA). 104. Same, paratype, ♀ (IECA). 105. M. boessnecki sp. nov., holotype, ♂ (SNSD). 106. M. babyrousa sp. nov., holotype, ♂ (BMNH). 107. Same, paratype, ♀ (BMNH). 108. M. caperatus sp. nov., holotype, ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122173" httpUri="https://zenodo.org/record/4122173/files/figure.png" pageId="30" pageNumber="31">105</figureCitation>
,
<figureCitation id="0AA52A659D029245FD52FEEECAA62909" box="[705,752,336,363]" captionStart="Figs 193–207" captionStartId="145.[189,241,1716,1742]" captionTargetBox="[385,1203,267,1681]" captionTargetId="figure-159@145.[385,1203,265,1694]" captionTargetPageId="145" captionText="Figs 193–207. Papuan and Wallacean Maechidius Macleay, 1819, head (193–196) and pronotum (197– 207), dorsal view. 193. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 194. Same, ♂ (SNSD). 195. M. yamdena sp. nov., paratype, ♂ (BMNH). 196. Same, paratype, ♀ (BMNH). 197. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 198. M. aiyura sp. nov., holotype, ♂ (BMNH). 199. M. alesbezdeki sp. nov., holotype, ♂ (NME). 200. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 201. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 202. M. awu sp. nov., holotype, ♂ (IECA). 203. M. babyrousa sp. nov., holotype, ♂ (BMNH). 204. M. bintang sp. nov., holotype, ♂ (NME). 205. M. boessnecki sp. nov., holotype, ♂ (SNSD). 206. M. brocki sp. nov., holotype, ♂ (BMNH). 207. M. crypticus sp. nov., paratype, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122189" httpUri="https://zenodo.org/record/4122189/files/figure.png" pageId="30" pageNumber="31">205</figureCitation>
,
<figureCitation id="0AA52A659D029245FD6FFEEECB7D2909" box="[764,811,336,363]" captionStart="Figs 286–301" captionStartId="152.[189,241,1716,1742]" captionTargetBox="[364,1224,266,1689]" captionTargetId="figure-172@152.[364,1224,266,1695]" captionTargetPageId="152" captionText="Figs 286–301. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 286. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 287. M. aiyura sp. nov., holotype, ♂ (BMNH). 288. M. alesbezdeki sp. nov., holotype, ♂ (NME). 289. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 290. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 291. M. paupianus Heller, 1910, ♂ (BMNH). 292. M. awu sp. nov., holotype, ♂ (IECA). 293. M. babyrousa sp. nov., holotype, ♂ (BMNH). 294. M. bintang sp. nov., holotype, ♂ (NME). 295. M. boessnecki sp. nov., holotype, ♂ (SNSD). 296. M. brocki sp. nov., holotype, ♂ (BMNH). 297. M. caperatus sp. nov., holotype, ♂ (NME). 298. M. ciliatus sp. nov., holotype, ♂ (IECA). 299. M. crypticus sp. nov., paratype, ♀ (BMNH). 300. M. dani sp. nov., holotype, ♂ (IECA). 301. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122206" httpUri="https://zenodo.org/record/4122206/files/figure.png" pageId="30" pageNumber="31">295</figureCitation>
, 367, 461, 589–591
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D029245FF2EFE2CCB8B29BE" pageId="30" pageNumber="31" type="diagnosis">
<paragraph id="922136E09D029245FF2EFE2CC99F29CE" blockId="30.[189,457,402,428]" box="[189,457,402,428]" pageId="30" pageNumber="31">
<heading id="C969818C9D029245FF2EFE2CC99F29CE" bold="true" box="[189,457,402,428]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="A0EAEAF29D029245FF2EFE2CC99F29CE" bold="true" box="[189,457,402,428]" pageId="30" pageNumber="31">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D029245FF2EFE7CCB8B29BE" blockId="30.[189,989,450,476]" box="[189,989,450,476]" pageId="30" pageNumber="31">This species differs from all congeners in the shape of the aedeagus.</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D029245FF2EFDBACBA22A13" pageId="30" pageNumber="31" type="etymology">
<paragraph id="922136E09D029245FF2EFDBAC9122A7C" blockId="30.[189,324,516,542]" box="[189,324,516,542]" pageId="30" pageNumber="31">
<heading id="C969818C9D029245FF2EFDBAC9122A7C" bold="true" box="[189,324,516,542]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="A0EAEAF29D029245FF2EFDBAC9122A7C" bold="true" box="[189,324,516,542]" pageId="30" pageNumber="31">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D029245FF2EFD8DCBA22A13" blockId="30.[189,1398,563,625]" pageId="30" pageNumber="31">
Patronymic. This species is named in honour of Ulrich Bössneck (
<date id="E62010209D029245FC46FD8DCC3D2A2F" box="[981,1131,563,590]" pageId="30" pageNumber="31" value="1960-03-04">4 Mar. 1960</date>
–
<date id="E62010209D029245FBE8FD8DCD4D2A2F" box="[1147,1307,563,589]" pageId="30" pageNumber="31" value="2019-09-13">13 Sep. 2019</date>
; Erfurt,
<collectingCountry id="EA8976709D029245FF2EFDE9C9792A13" box="[189,303,599,625]" name="Germany" pageId="30" pageNumber="31">Germany</collectingCountry>
), a famous malacologist, respected colleague and my friend.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D029245FF2EFD27CA9B2B48" pageId="30" pageNumber="31" type="materials_examined">
<paragraph id="922136E09D029245FF2EFD27C93A2AD1" blockId="30.[189,364,665,691]" box="[189,364,665,691]" pageId="30" pageNumber="31">
<heading id="C969818C9D029245FF2EFD27C93A2AD1" bold="true" box="[189,364,665,691]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="A0EAEAF29D029245FF2EFD27C93A2AD1" bold="true" box="[189,364,665,691]" pageId="30" pageNumber="31">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D029245FF79FD76C90A2A80" blockId="30.[189,1399,712,810]" box="[234,348,712,738]" pageId="30" pageNumber="31">
<emphasis id="A0EAEAF29D029245FF79FD76C90A2A80" bold="true" box="[234,348,712,738]" pageId="30" pageNumber="31">
<typeStatus id="4D2588429D029245FF79FD76C90A2A80" box="[234,348,712,738]" pageId="30" pageNumber="31" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="922136E09D029245FF2EFD52CA9B2B48" blockId="30.[189,1399,712,810]" pageId="30" pageNumber="31">
<materialsCitation id="22F63CBD9D029245FF2EFD52CA9B2B48" ID-GBIF-Occurrence="2898878404" collectionCode="SNSD" country="Indonesia" location="Celebes" pageId="30" pageNumber="31" specimenCount="1" specimenCount-male="1" stateProvince="Celebes" typeStatus="holotype">
<collectingCountry id="EA8976709D029245FF2EFD52C90D2B64" box="[189,347,748,774]" name="Indonesia" pageId="30" pageNumber="31">INDONESIA</collectingCountry>
•
<specimenCount id="8498FD699D029245FEEEFD52C9C32B64" box="[381,405,748,774]" pageId="30" pageNumber="31" type="male">♂</specimenCount>
; “Drs. Sarasin N.
<collectingRegion id="505AF8029D029245FDEDFD52CA8D2B64" box="[638,731,748,774]" country="Indonesia" name="Sulawesi Utara" pageId="30" pageNumber="31">Celebes</collectingRegion>
[p] Matinang [h]// 10575. [h, label yellowish]// Staatl. Museum für Tierkunde Dresden [p]”;
<collectionCode id="F48FAE259D029245FDE8FCAECA9C2B48" box="[635,714,784,810]" collectionName="SNSD" pageId="30" pageNumber="31">SNSD</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D02927BFF2EFCECC9FD2933" lastPageId="32" lastPageNumber="33" pageId="30" pageNumber="31" type="description">
<paragraph id="922136E09D029245FF2EFCECC9182B0E" blockId="30.[189,334,850,876]" box="[189,334,850,876]" pageId="30" pageNumber="31">
<heading id="C969818C9D029245FF2EFCECC9182B0E" bold="true" box="[189,334,850,876]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="A0EAEAF29D029245FF2EFCECC9182B0E" bold="true" box="[189,334,850,876]" pageId="30" pageNumber="31">Description</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D029245FF2EFC3CC97B2B81" blockId="30.[189,1399,897,995]" pageId="30" pageNumber="31">
MEASUREMENTS.
<typeStatus id="4D2588429D029245FEEFFC3FC9BA2BF9" box="[380,492,897,923]" pageId="30" pageNumber="31" type="holotype">Holotype</typeStatus>
, total body length
<quantity id="55669B059D029245FD54FC3FCB652BFE" box="[711,819,897,924]" metricMagnitude="-3" metricUnit="m" metricValue="8.45" pageId="30" pageNumber="31" unit="mm" value="8.45">8.45 mm</quantity>
. Head
<quantity id="55669B059D029245FC16FC3FCBB82BFE" box="[901,1006,897,924]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="30" pageNumber="31" unit="mm" value="1.7">1.70 mm</quantity>
long, across eyes
<quantity id="55669B059D029245FB55FC3FCD792BFE" box="[1222,1327,897,924]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="30" pageNumber="31" unit="mm" value="1.8">1.80 mm</quantity>
wide. Pronotum
<quantity id="55669B059D029245FEA4FC1BC9C92BDD" box="[311,415,933,959]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="30" pageNumber="31" unit="mm" value="1.8">1.80 mm</quantity>
long, maximum width
<quantity id="55669B059D029245FD3EFC1BCB412BDD" box="[685,791,933,959]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="30" pageNumber="31" unit="mm" value="2.9">2.90 mm</quantity>
. Elytral length
<quantity id="55669B059D029245FC54FC1BCC642BDD" box="[967,1074,933,960]" metricMagnitude="-3" metricUnit="m" metricValue="4.95" pageId="30" pageNumber="31" unit="mm" value="4.95">4.95 mm</quantity>
, maximum combined width
<quantity id="55669B059D029245FF2EFC76C97F2B81" box="[189,297,968,995]" metricMagnitude="-3" metricUnit="m" metricValue="3.15" pageId="30" pageNumber="31" unit="mm" value="3.15">3.15 mm</quantity>
.
</paragraph>
<paragraph id="922136E09D029245FF2EFBB5CA482F8B" blockId="30.[189,1399,1035,2025]" pageId="30" pageNumber="31">
Dorsum black-brown, labroclypeus, antennae, legs and venter brown. Head glossy dorsally, slightly convex on frons. Male labroclypeus (
<figureCitation id="0AA52A659D029245FDEBFB90CA882C2B" box="[632,734,1070,1097]" captionStart="Figs 97–108" captionStartId="137.[189,241,1786,1812]" captionTargetBox="[351,1228,270,1756]" captionTargetId="figure-127@137.[351,1237,265,1772]" captionTargetPageId="137" captionText="Figs 97–108. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 97. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 98. M. aiyura sp. nov., holotype, ♂ (BMNH). 99. M. alesbezdeki sp. nov., holotype, ♂ (NME). 100. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 101. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 102. M. bintang sp. nov., holotype, ♂ (NME). 103. M. awu sp. nov., holotype, ♂ (IECA). 104. Same, paratype, ♀ (IECA). 105. M. boessnecki sp. nov., holotype, ♂ (SNSD). 106. M. babyrousa sp. nov., holotype, ♂ (BMNH). 107. Same, paratype, ♀ (BMNH). 108. M. caperatus sp. nov., holotype, ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122173" httpUri="https://zenodo.org/record/4122173/files/figure.png" pageId="30" pageNumber="31">Fig. 105</figureCitation>
) very shallowly broadly emarginate on anterior margin, its lateral margins slightly sinuous in both dorsal and lateral views. Anterolateral angles of labroclypeus obtuse, hardly protruding anteriad. Compound eye large, occupy about half side of head. Canthus nearly straight in dorsal view. Head punctures ovoid, deep and dense, filled with microscopical velvety pubescence. Intervening spaces glossy, much smaller than punctures. Head setae suberect, thickened, moderately long, longer on frons than on labroclypeus; each seta rises from anterior margin of corresponding puncture. Antenna 9-segmented, club 3-lamellate. Pronotum transverse, glossy dorsally and laterally. Anterior margin of pronotum broadly emarginate, basal margin very broadly rounded. Lateral margin of pronotum in dorsal view broadly rounded, maximum width slightly postmedially, crenulate all along (
<figureCitation id="0AA52A659D029245FE32FACFCA502DEE" box="[417,518,1393,1420]" captionStart="Figs 193–207" captionStartId="145.[189,241,1716,1742]" captionTargetBox="[385,1203,267,1681]" captionTargetId="figure-159@145.[385,1203,265,1694]" captionTargetPageId="145" captionText="Figs 193–207. Papuan and Wallacean Maechidius Macleay, 1819, head (193–196) and pronotum (197– 207), dorsal view. 193. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 194. Same, ♂ (SNSD). 195. M. yamdena sp. nov., paratype, ♂ (BMNH). 196. Same, paratype, ♀ (BMNH). 197. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 198. M. aiyura sp. nov., holotype, ♂ (BMNH). 199. M. alesbezdeki sp. nov., holotype, ♂ (NME). 200. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 201. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 202. M. awu sp. nov., holotype, ♂ (IECA). 203. M. babyrousa sp. nov., holotype, ♂ (BMNH). 204. M. bintang sp. nov., holotype, ♂ (NME). 205. M. boessnecki sp. nov., holotype, ♂ (SNSD). 206. M. brocki sp. nov., holotype, ♂ (BMNH). 207. M. crypticus sp. nov., paratype, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122189" httpUri="https://zenodo.org/record/4122189/files/figure.png" pageId="30" pageNumber="31">Fig. 205</figureCitation>
). Moderately long curved suberect seta present between every two crenulae. Lateral margin of pronotum nearly straight in lateral view. Pronotal punctures ovoid, deep and dense, filled with microscopical velvety pubescence. Intervening spaces glossy, much smaller than punctures, in part wrinkled. Lateral and basal margins, antero- and posterolateral angles partly covered with microscopical velvety pubescence. Pronotal setae inconspicuous, suberect; each seta rises from anterior margin of corresponding puncture, surpassing its length. Hypomeron flanged-like protruding, slightly emarginate and long setose opposite to compound eye. Antennal pocket deep. Elytron glabrous, with vague track of one glabrous longitudinal carina. Sutural carina not indicated. Elytral punctures irregularly ovoid, deep and dense, filled with microscopical velvety pubescence (
<figureCitation id="0AA52A659D029245FC33F92ECC5E2EC8" box="[928,1032,1679,1706]" captionStart="Figs 286–301" captionStartId="152.[189,241,1716,1742]" captionTargetBox="[364,1224,266,1689]" captionTargetId="figure-172@152.[364,1224,266,1695]" captionTargetPageId="152" captionText="Figs 286–301. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 286. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 287. M. aiyura sp. nov., holotype, ♂ (BMNH). 288. M. alesbezdeki sp. nov., holotype, ♂ (NME). 289. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 290. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 291. M. paupianus Heller, 1910, ♂ (BMNH). 292. M. awu sp. nov., holotype, ♂ (IECA). 293. M. babyrousa sp. nov., holotype, ♂ (BMNH). 294. M. bintang sp. nov., holotype, ♂ (NME). 295. M. boessnecki sp. nov., holotype, ♂ (SNSD). 296. M. brocki sp. nov., holotype, ♂ (BMNH). 297. M. caperatus sp. nov., holotype, ♂ (NME). 298. M. ciliatus sp. nov., holotype, ♂ (IECA). 299. M. crypticus sp. nov., paratype, ♀ (BMNH). 300. M. dani sp. nov., holotype, ♂ (IECA). 301. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122206" httpUri="https://zenodo.org/record/4122206/files/figure.png" pageId="30" pageNumber="31">Fig. 295</figureCitation>
), those on elytral disc arranged in
<quantity id="55669B059D029245FF4FF90AC94F2EAF" box="[220,281,1715,1742]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="30" pageNumber="31" unit="in" value="12.0">12 in</quantity>
part irregular longitudinal rows, those along lateral sides confused. Intervening spaces glossy, generally larger than punctures. Setae of elytra inconspicuous, appressed; minute seta rises from anterior margin of each puncture, not surpassing its length (
<figureCitation id="0AA52A659D029245FCBAF945CBC42F74" box="[809,914,1787,1814]" captionStart="Figs 286–301" captionStartId="152.[189,241,1716,1742]" captionTargetBox="[364,1224,266,1689]" captionTargetId="figure-172@152.[364,1224,266,1695]" captionTargetPageId="152" captionText="Figs 286–301. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 286. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 287. M. aiyura sp. nov., holotype, ♂ (BMNH). 288. M. alesbezdeki sp. nov., holotype, ♂ (NME). 289. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 290. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 291. M. paupianus Heller, 1910, ♂ (BMNH). 292. M. awu sp. nov., holotype, ♂ (IECA). 293. M. babyrousa sp. nov., holotype, ♂ (BMNH). 294. M. bintang sp. nov., holotype, ♂ (NME). 295. M. boessnecki sp. nov., holotype, ♂ (SNSD). 296. M. brocki sp. nov., holotype, ♂ (BMNH). 297. M. caperatus sp. nov., holotype, ♂ (NME). 298. M. ciliatus sp. nov., holotype, ♂ (IECA). 299. M. crypticus sp. nov., paratype, ♀ (BMNH). 300. M. dani sp. nov., holotype, ♂ (IECA). 301. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122206" httpUri="https://zenodo.org/record/4122206/files/figure.png" pageId="30" pageNumber="31">Fig. 295</figureCitation>
). Several longer suberect setae arranged in 5–6 inconspicuous longitudinal rows. Lateral margin of elytron with row of short suberect setae. Male pygidium flattened dorsally, with moderately large dense annular punctures (
<figureCitation id="0AA52A659D029245FB05F8FDCCA92F3C" box="[1174,1279,1859,1886]" captionStart="Figs 451–468" captionStartId="163.[189,241,1681,1707]" captionTargetBox="[456,1130,262,1650]" captionTargetId="figure-178@163.[456,1131,262,1657]" captionTargetPageId="163" captionText="Figs 451–468. Papuan and Wallacean Maechidius Macleay, 1819, pygidium, dorsal view. 451. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 452. M. aiyura sp. nov., holotype, ♂ (BMNH). 453. Same, different aspect (BMNH). 454. M. alesbezdeki sp. nov., holotype, ♂ (NME). 455. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 456. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 457. M. awu sp. nov., holotype, ♂ (IECA). 458. M. babyrousa sp. nov., holotype, ♂ (BMNH). 459. Same, paratype, ♀ (BMNH). 460. M. bintang sp. nov., holotype, ♂ (NME). 461. M. boessnecki sp. nov., holotype, ♂ (SNSD). 462. M. brocki sp. nov., holotype, ♂ (BMNH). 463. M. caperatus sp. nov., holotype, ♂ (NME). 464. M. ciliatus sp. nov., holotype, ♂ (IECA). 465. M. crypticus sp. nov., paratype, ♂ (BMNH). 466. Same, paratype, ♀ (BMNH). 467. M. dani sp. nov., holotype, ♂ (IECA). 468. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122222" httpUri="https://zenodo.org/record/4122222/files/figure.png" pageId="30" pageNumber="31">Fig. 461</figureCitation>
). Setae of pygidium rise from anterior margin of corresponding punctures, generally suberect to erect and longer than corresponding punctures. Male protibia with two distal teeth on external margin, of which distal one acute (Fig. 367). Male protibial terminal spur strongly curved. Spiculum gastrale as in
<figureCitation id="0AA52A659D029245FA98F815CD252FA7" box="[1291,1395,1963,1989]" captionStart="Figs 451–468" captionStartId="163.[189,241,1681,1707]" captionTargetBox="[456,1130,262,1650]" captionTargetId="figure-178@163.[456,1131,262,1657]" captionTargetPageId="163" captionText="Figs 451–468. Papuan and Wallacean Maechidius Macleay, 1819, pygidium, dorsal view. 451. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 452. M. aiyura sp. nov., holotype, ♂ (BMNH). 453. Same, different aspect (BMNH). 454. M. alesbezdeki sp. nov., holotype, ♂ (NME). 455. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 456. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 457. M. awu sp. nov., holotype, ♂ (IECA). 458. M. babyrousa sp. nov., holotype, ♂ (BMNH). 459. Same, paratype, ♀ (BMNH). 460. M. bintang sp. nov., holotype, ♂ (NME). 461. M. boessnecki sp. nov., holotype, ♂ (SNSD). 462. M. brocki sp. nov., holotype, ♂ (BMNH). 463. M. caperatus sp. nov., holotype, ♂ (NME). 464. M. ciliatus sp. nov., holotype, ♂ (IECA). 465. M. crypticus sp. nov., paratype, ♂ (BMNH). 466. Same, paratype, ♀ (BMNH). 467. M. dani sp. nov., holotype, ♂ (IECA). 468. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122222" httpUri="https://zenodo.org/record/4122222/files/figure.png" pageId="30" pageNumber="31">Fig. 461</figureCitation>
. Aedeagus as in
<figureCitation id="0AA52A659D029245FEE5F870CA4C2F8B" box="[374,538,1998,2025]" captionStart="Figs 586–600" captionStartId="171.[189,241,1892,1918]" captionTargetBox="[400,1175,268,1865]" captionTargetId="figure-72@171.[391,1196,265,1868]" captionTargetPageId="171" captionText="Figs 586–600. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 586– 588. M. bintang sp. nov., holotype, ♂ (NME). 589–591. M. boessnecki sp. nov., holotype, ♂ (SNSD). 592–594. M. caperatus sp. nov., holotype, ♂ (NME). 595–597. M. ciliatus sp. nov., holotype, ♂ (IECA). 598–600. M. crypticus sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122234" httpUri="https://zenodo.org/record/4122234/files/figure.png" pageId="30" pageNumber="31">Figs 589–591</figureCitation>
.
</paragraph>
<caption id="C6E166689D039244FF2EF836CA112F8A" ID-DOI="http://doi.org/10.5281/zenodo.4122136" ID-Zenodo-Dep="4122136" httpUri="https://zenodo.org/record/4122136/files/figure.png" pageId="31" pageNumber="32" startId="31.[189,241,1928,1954]" targetBox="[347,1240,269,1901]" targetPageId="31">
<paragraph id="922136E09D039244FF2EF836CA112F8A" blockId="31.[189,1399,1928,2025]" pageId="31" pageNumber="32">
<emphasis id="A0EAEAF29D039244FF2EF836C91F2FC0" bold="true" box="[189,329,1928,1954]" pageId="31" pageNumber="32">Figs 28–30.</emphasis>
Papuan and Wallacean
<taxonomicName id="559E4D639D039244FDF3F836CBC92FC0" authority="Macleay, 1819" authorityName="Macleay" authorityYear="1819" box="[608,927,1928,1954]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="A0EAEAF29D039244FDF3F836CABF2FC0" box="[608,745,1928,1954]" italics="true" pageId="31" pageNumber="32">Maechidius</emphasis>
<bibRefCitation id="F60F4B119D039244FD62F836CBC92FC0" author="Macleay W. S." box="[753,927,1928,1954]" pageId="31" pageNumber="32" refId="ref98319" refString="Macleay W. S. 1819. Horae Entomologicae: or Essays on the Annulose Animals. Volume I, Part I. Containing General Observations on the Geography, Manners, and the Affinities of the Insects which compose the Genus Scarabaeus of Linnaeus; to which are added a few incidental Remarks on the Genera Lucanus and Hister of the same Author. With Appendix and Plates. R. and A. Taylor, London." type="book" year="1819">Macleay, 1819</bibRefCitation>
</taxonomicName>
, habitus, dorsal view.
<emphasis id="A0EAEAF29D039244FB37F836CC972FC0" bold="true" box="[1188,1217,1928,1954]" pageId="31" pageNumber="32">28</emphasis>
.
<taxonomicName id="559E4D639D039244FB43F837CD202FC0" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1232,1398,1928,1954]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="boessnecki" status="sp. nov.">
<emphasis id="A0EAEAF29D039244FB43F837CD202FC0" box="[1232,1398,1928,1954]" italics="true" pageId="31" pageNumber="32">M. boessnecki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D039244FF2EF812C94C2FA4" box="[189,282,1964,1990]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (SNSD).
<emphasis id="A0EAEAF29D039244FDBBF815CA132FA7" bold="true" box="[552,581,1963,1989]" pageId="31" pageNumber="32">29</emphasis>
.
<taxonomicName id="559E4D639D039244FDC7F812CA902FA7" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[596,710,1963,1989]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="brocki" status="sp. nov.">
<emphasis id="A0EAEAF29D039244FDC7F812CA902FA7" box="[596,710,1963,1989]" italics="true" pageId="31" pageNumber="32">M. brocki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D039244FD5EF812CB7D2FA4" box="[717,811,1964,1990]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (BMNH).
<emphasis id="A0EAEAF29D039244FBD6F815CC342FA7" bold="true" box="[1093,1122,1963,1989]" pageId="31" pageNumber="32">30</emphasis>
.
<taxonomicName id="559E4D639D039244FBE1F812CD5B2FA7" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1138,1293,1964,1989]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="caperatus" status="sp. nov.">
<emphasis id="A0EAEAF29D039244FBE1F812CD5B2FA7" box="[1138,1293,1964,1989]" italics="true" pageId="31" pageNumber="32">M. caperatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D039244FA87F812CD242FA4" box="[1300,1394,1964,1990]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (NME). Not to scale.
</paragraph>
</caption>
<paragraph id="922136E09D3C927BFF2EFEB6C9E62940" blockId="32.[189,432,264,290]" box="[189,432,264,290]" pageId="32" pageNumber="33">
<heading id="C969818C9D3C927BFF2EFEB6C9E62940" bold="true" box="[189,432,264,290]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
<emphasis id="A0EAEAF29D3C927BFF2EFEB6C9E62940" bold="true" box="[189,432,264,290]" pageId="32" pageNumber="33">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D3C927BFF2EFE89C9FD2933" blockId="32.[189,427,311,337]" box="[189,427,311,337]" pageId="32" pageNumber="33">Female is unknown.</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D3C927BFF2EFEC6C96F29A2" box="[189,313,376,448]" pageId="32" pageNumber="33" type="biology_ecology">
<paragraph id="922136E09D3C927BFF2EFEC6C97629F0" blockId="32.[189,288,376,402]" box="[189,288,376,402]" pageId="32" pageNumber="33">
<heading id="C969818C9D3C927BFF2EFEC6C97629F0" bold="true" box="[189,288,376,402]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
<emphasis id="A0EAEAF29D3C927BFF2EFEC6C97629F0" bold="true" box="[189,288,376,402]" pageId="32" pageNumber="33">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D3C927BFF2EFE18C96F29A2" blockId="32.[189,313,422,448]" box="[189,313,422,448]" pageId="32" pageNumber="33">Unknown.</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D3C927BFF2EFE59CA612A4D" box="[189,567,487,559]" pageId="32" pageNumber="33" type="distribution">
<paragraph id="922136E09D3C927BFF2EFE59C9002A63" blockId="32.[189,342,487,513]" box="[189,342,487,513]" pageId="32" pageNumber="33">
<heading id="C969818C9D3C927BFF2EFE59C9002A63" bold="true" box="[189,342,487,513]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
<emphasis id="A0EAEAF29D3C927BFF2EFE59C9002A63" bold="true" box="[189,342,487,513]" pageId="32" pageNumber="33">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D3C927BFF2EFDABCA612A4D" blockId="32.[189,567,533,559]" box="[189,567,533,559]" pageId="32" pageNumber="33">
Northern peninsula of
<collectingRegion id="505AF8029D3C927BFE55FDABCA622A4D" box="[454,564,533,559]" country="Indonesia" name="Sulawesi Utara" pageId="32" pageNumber="33">Sulawesi</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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