267 lines
29 KiB
XML
267 lines
29 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1182.97346" ID-Pensoft-Pub="1313-2970-1182-165" ID-Pensoft-UUID="DCECB4AAD9E95ADB80188D91F386DD1C" ID-ZooBank="86F5AC45D35E4DDFAB042BCBDEEE8407" ModsDocID="1313-2970-1182-165" checkinTime="1697273694970" checkinUser="pensoft" docAuthor="Choi, Yeon-Jae, Park, Sun-Jae, Lee, Seung-Gyu & Park, Jong-Seok" docDate="2023" docId="1A296044ED595F88B5E6701BCF2B01F2" docLanguage="en" docName="ZooKeys 1182: 165-181" docOrigin="ZooKeys 1182" docPubDate="2023-10-13" docSource="http://dx.doi.org/10.3897/zookeys.1182.97346" docTitle="Bryaxis grandinodus Choi, Park, Lee & Park 2023, sp. nov." docType="treatment" docUuid="296C6B58-60CE-41D9-BD87-AAC45B5AC0C5" docUuidSource="ZooBank" docVersion="1" id="DCECB4AAD9E95ADB80188D91F386DD1C" lastPageNumber="165" masterDocId="DCECB4AAD9E95ADB80188D91F386DD1C" masterDocTitle="Five new species of Bryaxis Kugelann (Coleoptera, Staphylinidae, Pselaphinae) from Korea and a nomenclatural note on Bryaxis mahunkai Loebl" masterLastPageNumber="181" masterPageNumber="165" pageNumber="165" updateTime="1697273694970" updateUser="pensoft">
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<mods:mods id="9E94C8209FAFE4AB1CE6EBA7B1C9FAFA" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="A5062E0175BF1CB9DA967B7FB3293F03">
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<mods:title id="AA27FB693744BD403F886937A46157F5">Five new species of Bryaxis Kugelann (Coleoptera, Staphylinidae, Pselaphinae) from Korea and a nomenclatural note on Bryaxis mahunkai Loebl</mods:title>
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<mods:name id="B4A24AFE41F764ACB18DA69F21896FEF" type="personal">
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<mods:roleTerm id="C5E7385D3D8EF5D661706F5B2C44EA79">Author</mods:roleTerm>
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<mods:namePart id="DD2C21394CC606C7C1B024A3B0FF63A9">Choi, Yeon-Jae</mods:namePart>
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<mods:nameIdentifier id="BF03522BAD483A0370B5B1CDB475E353" type="ORCID">https://orcid.org/0000-0003-0172-1128</mods:nameIdentifier>
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<mods:affiliation id="AC55A8BD468A5CA23D77A1EAB6E7B422">Department of Biological Sciences and Biotechnology, Chungbuk National University, Cheongju 28644, Republic of Korea</mods:affiliation>
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</mods:name>
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<mods:name id="D420CFD551A5BC99FEC41967FE5DFD60" type="personal">
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<mods:roleTerm id="C3CCF47440CADD993C9391D34F0D222E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9E7F9D05E4A968EA9567BC5AE4B1134C">Park, Sun-Jae</mods:namePart>
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<mods:nameIdentifier id="3E1225BA4C3F85DC8739E4AD7F20ABEE" type="ORCID">https://orcid.org/0000-0003-3759-6517</mods:nameIdentifier>
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<mods:affiliation id="75F6840E0846F04DDD8B8AA4304708E0">Climate Change and Environmental Biology Research Division, National Institute of Biological Resources, Incheon 22689, Republic of Korea</mods:affiliation>
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</mods:name>
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<mods:name id="8B4CDA858F80B5A68973852CEF8549FD" type="personal">
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<mods:role id="D7150040C1B2022EE2ABF324D41D0B0F">
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<mods:roleTerm id="C7AA921404600669091CCAA48D6748E4">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7ABC6D975C921DE0AAD82D5E17B2DF0A">Lee, Seung-Gyu</mods:namePart>
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<mods:nameIdentifier id="486644E53A4478DC459B1AF2D21C26E4" type="ORCID">https://orcid.org/0000-0001-5576-2799</mods:nameIdentifier>
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<mods:affiliation id="5F26502AE35A6AEDF653945AA6C5E503">Climate Change and Environmental Biology Research Division, National Institute of Biological Resources, Incheon 22689, Republic of Korea</mods:affiliation>
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</mods:name>
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<mods:name id="FFB4002B8D2F284D0C45C5CEEC339E04" type="personal">
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<mods:roleTerm id="F45E9CF9799FBC3AE7B291FC7E928CA1">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="DF1BA505AEC5F71287F7B17FBEEBEAC1">Park, Jong-Seok</mods:namePart>
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<mods:nameIdentifier id="221DC401EA4BB12E0F053D61A2A56583" type="ORCID">https://orcid.org/0000-0003-2063-2534</mods:nameIdentifier>
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<mods:affiliation id="6644364B3C7FE322FCE3156FBEB79B9B">Department of Biological Sciences and Biotechnology, Chungbuk National University, Cheongju 28644, Republic of Korea</mods:affiliation>
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<mods:nameIdentifier id="579FAF049CA1D1DFFA2D71011AA8107B" type="email">jpark16@cbnu.ac.kr</mods:nameIdentifier>
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<mods:typeOfResource id="36C069D9A89D3301045ED9E72739AC0F">text</mods:typeOfResource>
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<mods:title id="3F8A0D27FCBD78D9A3B6D7BE8195A439">ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part id="E24E5DB84A30FBABF926AE175F754616">
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<mods:date id="C03B88B50F77B54F7562B2C5EA84A659">2023</mods:date>
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<mods:number id="613B1B68E6A2DDFAAC9B0910A0839210">2023-10-13</mods:number>
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<mods:number id="001C7CDECE5DCCA86031051E84A25956">1182</mods:number>
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<mods:start id="695844B2F8186DAB8131AE1F98075996">165</mods:start>
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<mods:end id="2D737991C272612BF792953B17C6EE0A">181</mods:end>
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<mods:url id="A093D3540956AA96F4F1DD9B3FE6C813">http://dx.doi.org/10.3897/zookeys.1182.97346</mods:url>
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</mods:location>
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<mods:classification id="B8D6D8D272D4F6B8B1435278229BEDB4">journal article</mods:classification>
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<mods:identifier id="96FFCD2DF6C9514C1FAD4BAC998887F4" type="DOI">http://dx.doi.org/10.3897/zookeys.1182.97346</mods:identifier>
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<mods:identifier id="DDC1F58EAB834531459BED1E47400B1F" type="Pensoft-Pub">1313-2970-1182-165</mods:identifier>
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<mods:identifier id="3E285E9A617256F70A693329AE4E7059" type="ZooBank">86F5AC45D35E4DDFAB042BCBDEEE8407</mods:identifier>
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<mods:identifier id="4EC8138304599E296C7D9936C01BD39A" type="Pensoft-UUID">DCECB4AAD9E95ADB80188D91F386DD1C</mods:identifier>
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<treatment id="1A296044ED595F88B5E6701BCF2B01F2" LSID="urn:lsid:zoobank.org:act:296C6B58-60CE-41D9-BD87-AAC45B5AC0C5" httpUri="http://treatment.plazi.org/id/1A296044ED595F88B5E6701BCF2B01F2" lastPageNumber="165" pageId="0" pageNumber="165">
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<subSubSection id="BB554562CF6315D6551C9528F2B49A31" pageId="0" pageNumber="165" type="nomenclature">
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<paragraph id="06024359615472302262BF5FA312494B" pageId="0" pageNumber="165">
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<taxonomicName id="7D9350A099E1FDA44B1DD5678FDBBE79" LSID="https://zoobank.org/296C6B58-60CE-41D9-BD87-AAC45B5AC0C5" authority="Choi, Park, Lee & Park" authorityName="Choi, Park, Lee & Park" authorityYear="2023" class="Insecta" family="Staphylinidae" genus="Bryaxis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bryaxis grandinodus" order="Coleoptera" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="grandinodus" status="sp. nov.">Bryaxis grandinodus Choi, Park, Lee & Park</taxonomicName>
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<taxonomicNameLabel id="012D7FB13B79335A3F3215B93DF75A22" pageId="0" pageNumber="165">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="5A8FA0B0E4760F13C618C05E5FC98BA8" pageId="0" pageNumber="165" type="description">
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<paragraph id="91CF80DC20B65068F076EC3BDECB3D45" pageId="0" pageNumber="165">
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<figureCitation id="A81F8FBB6BA7E18E74E356C651BAE9A7" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Bryaxis grandinodus Choi, Park, Lee & Park, sp. nov., male (A, B, E-I), female (C, D). A, C dorsal habitus B, D ventral habitus E head F antenna G fore leg H hind leg I aedeagus. Scale bars: 1 mm (A-D); 0.1 mm (E, I); 0.5 mm (F-H)." figureDoi="10.3897/zookeys.1182.97346.figure1" httpUri="https://binary.pensoft.net/fig/920944" pageId="0" pageNumber="165">Figs 1</figureCitation>
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<figureCitation id="E65671AF78DEAE0F6502D8520349CC4F" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Diagnostic characters of Bryaxis grandinodus sp. nov. (A, C) and B. koltzei (Reitter) (B, D). A, B antennal scapes and pedicels C, D protibiae. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1182.97346.figure2" httpUri="https://binary.pensoft.net/fig/920945" pageId="0" pageNumber="165">, 2A, C</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="9FDF0021422C2D42469B87DC110D504D" pageId="0" pageNumber="165" type="type material">
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<paragraph id="404113C2C5B01D5A08AC5AFAC629B70B" pageId="0" pageNumber="165">Type materials</paragraph>
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<paragraph id="2729C77CF4FC7B2C910359A29E174453" pageId="0" pageNumber="165">
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<emphasis id="B9072B7AFE5FFBAF9261E658ADAFA70E" bold="true" pageId="0" pageNumber="165">
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(
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<emphasis id="C6F173945F99CBBB4051DF948DA04CDA" italics="true" pageId="0" pageNumber="165">N</emphasis>
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= 11, 6
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</emphasis>
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♂♂,
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<emphasis id="2D13C8BF94C7D1F939A84B2BBBB15AC5" bold="true" pageId="0" pageNumber="165">5</emphasis>
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♀♀)
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<emphasis id="741B698B05926D4C83AFC4C7EB8D676D" bold="true" pageId="0" pageNumber="165">
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.
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<emphasis id="976491D049E6F039D96A38C46AE87D33" italics="true" pageId="0" pageNumber="165">Holotype male</emphasis>
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.
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</emphasis>
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"Korea: Jeonnam Prov. Dangsan-ri, Gyegok-myeon, Haenam-gun, 18 May 2019,
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<geoCoordinate id="C07434F9F23B7CF90A9C386E0D48F341" degrees="34" direction="north" minutes="40" orientation="latitude" precision="1" seconds="53.0" value="34.68139">34°40'53.0"N</geoCoordinate>
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,
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<geoCoordinate id="4C033631871294EE23E95066ED628033" degrees="126" direction="east" minutes="38" orientation="longitude" precision="1" seconds="56.3" value="126.64897">126°38'56.3"E</geoCoordinate>
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, 211 m, sifting leaf litter & deadwood debris, J.-S. Park, M.-H. Song" (NIBR).
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<emphasis id="6083FAD32AD49492267AB99226B777A9" bold="true" pageId="0" pageNumber="165">
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<emphasis id="0C3F763CA22DD44A97347B15EC288A64" italics="true" pageId="0" pageNumber="165">Paratypes</emphasis>
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.
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</emphasis>
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2♂♂, 2♀♀ (CBNUIC, 1♂, 1♀ slide mounted, 1♂, 1♀ dried). "Korea: Jeonbuk Prov. Sinsi island. Sinsido-gil, Okdo-myeon, Gunsan-si, 4 Jul 2022,
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<geoCoordinate id="A767FD5BF1AEDD10C011C50BC27FE9B6" degrees="35" direction="north" minutes="49" orientation="latitude" precision="1" seconds="12.2" value="35.820053">35°49'12.2"N</geoCoordinate>
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,
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<geoCoordinate id="6F0BAFAA886B021C9C3206AEF0172D51" degrees="126" direction="east" minutes="27" orientation="longitude" precision="1" seconds="35.1" value="126.45975">126°27'35.1"E</geoCoordinate>
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, 36 m, sifting leaf & soil litter, M.-H. Song, U.-J. Byeon, J.-W. Kang, T.-Y. Jang". 2♂♂ (CBNUIC, dried) "Korea: Jeonbuk Prov. Seonyu island. Seonyubuk-gil, Okdo-myeon, Gunsan-si, 16 Jun 2021,
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<geoCoordinate id="21255ECF7CC411843372FCBA1887E5AA" degrees="35" direction="north" minutes="48" orientation="latitude" precision="1" seconds="36.5" value="35.81014">35°48'36.5"N</geoCoordinate>
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,
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<geoCoordinate id="D5BE63853FD7E39C46469BF6B8FCD98F" degrees="126" direction="east" minutes="24" orientation="longitude" precision="1" seconds="57.4" value="126.41595">126°24'57.4"E</geoCoordinate>
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, 25 m, sifting leaf, soil litter & fungi, J.-W. Kang, J.-I. Shin". 1♂, 3♀♀ (CBNUIC, DNA grade). "Korea: Jeonbuk Prov. Seonyu island. 5-1, Seonyunam-gil, Okdo-myeon, Gunsan-si, 4 Jul 2022,
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<geoCoordinate id="86D8AA5FF40C2436B906DE84993F95EC" degrees="35" direction="north" minutes="48" orientation="latitude" precision="1" seconds="24.7" value="35.80686">35°48'24.7"N</geoCoordinate>
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,
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<geoCoordinate id="956399FB0819813D4824C6056636504C" degrees="126" direction="east" minutes="24" orientation="longitude" precision="1" seconds="40.3" value="126.411194">126°24'40.3"E</geoCoordinate>
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, 9 m, sifting leaf & soil litter, M.-H. Song, U.-J. Byeon, J.-W. Kang, T.-Y. Jang".
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</paragraph>
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</subSubSection>
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<subSubSection id="BB4748A0A078EDFF301881C0E82ED61B" pageId="0" pageNumber="165" type="diagnosis">
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<paragraph id="15847FFD1658FB82C457BFF43998BEA3" pageId="0" pageNumber="165">Diagnosis.</paragraph>
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<paragraph id="D1DE6D6BB99A9104788DB1B29E527FE1" pageId="0" pageNumber="165">
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Antennal scapes robust, with bowl-like glandular nodule on inner margin (Figs
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<figureCitation id="B200D0A8BFC7E2D35F587B672C956B7D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Bryaxis grandinodus Choi, Park, Lee & Park, sp. nov., male (A, B, E-I), female (C, D). A, C dorsal habitus B, D ventral habitus E head F antenna G fore leg H hind leg I aedeagus. Scale bars: 1 mm (A-D); 0.1 mm (E, I); 0.5 mm (F-H)." figureDoi="10.3897/zookeys.1182.97346.figure1" httpUri="https://binary.pensoft.net/fig/920944" pageId="0" pageNumber="165">1E, F</figureCitation>
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,
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<figureCitation id="E30B3CB6564B3842967C97A8D805A983" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Diagnostic characters of Bryaxis grandinodus sp. nov. (A, C) and B. koltzei (Reitter) (B, D). A, B antennal scapes and pedicels C, D protibiae. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1182.97346.figure2" httpUri="https://binary.pensoft.net/fig/920945" pageId="0" pageNumber="165">2A</figureCitation>
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, arrows), 2.45 times as long as pedicels; endophallus of male genitalia with three bifid struts, joined at base (Fig.
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<figureCitation id="822548350D36F56DBB4780C8770D5BE8" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Bryaxis grandinodus Choi, Park, Lee & Park, sp. nov., male (A, B, E-I), female (C, D). A, C dorsal habitus B, D ventral habitus E head F antenna G fore leg H hind leg I aedeagus. Scale bars: 1 mm (A-D); 0.1 mm (E, I); 0.5 mm (F-H)." figureDoi="10.3897/zookeys.1182.97346.figure1" httpUri="https://binary.pensoft.net/fig/920944" pageId="0" pageNumber="165">1I</figureCitation>
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).
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</paragraph>
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<caption id="C444274A558AD02031886E2F7319B781" doi="10.3897/zookeys.1182.97346.figure1" httpUri="https://binary.pensoft.net/fig/920944" pageId="0" pageNumber="165" start="Figure 1" startId="F1">
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<paragraph id="21F53237DE11594DE0539EBEA63BDD0E" pageId="0" pageNumber="165">
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<emphasis id="284256A3B5F6F86C746BDDA3BB1ED744" bold="true" pageId="0" pageNumber="165">Figure 1.</emphasis>
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<taxonomicName id="F604D3FC6E5D458979E13DC87B1A8023" authorityName="Choi, Park, Lee & Park" authorityYear="2023" class="Insecta" family="Staphylinidae" genus="Bryaxis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bryaxis grandinodus" order="Coleoptera" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="grandinodus">
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<emphasis id="2320539DE0113F1F7C2AAE4856A45F4E" italics="true" pageId="0" pageNumber="165">Bryaxis grandinodus</emphasis>
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</taxonomicName>
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Choi, Park, Lee & Park, sp. nov., male (
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<emphasis id="66F51FD68397C407A7574A100BB53D4C" bold="true" pageId="0" pageNumber="165">A, B, E-I</emphasis>
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), female (
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<emphasis id="36DD11BE91FC4518DA03E668265A7892" bold="true" pageId="0" pageNumber="165">C, D</emphasis>
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).
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<emphasis id="625BFE2827615D9CE23D480D65F66C62" bold="true" pageId="0" pageNumber="165">A, C</emphasis>
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dorsal habitus
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<emphasis id="BB92475CEF32E16D11F1D00330940FB4" bold="true" pageId="0" pageNumber="165">B, D</emphasis>
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ventral habitus
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<emphasis id="37ECBD7B0A1CFEE30C25A793A63B0623" bold="true" pageId="0" pageNumber="165">E</emphasis>
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head
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<emphasis id="21DEE0FD388114FDD339495D64996CF8" bold="true" pageId="0" pageNumber="165">F</emphasis>
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antenna
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<emphasis id="9EFF8D374EC085783118B0F0B08DAA78" bold="true" pageId="0" pageNumber="165">G</emphasis>
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fore leg
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<emphasis id="39F95DD7B058FC11667A80DC4949CAE5" bold="true" pageId="0" pageNumber="165">H</emphasis>
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hind leg
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<emphasis id="D510CE936626CA81D7889FB133FB40BF" bold="true" pageId="0" pageNumber="165">I</emphasis>
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aedeagus. Scale bars: 1 mm (
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<emphasis id="8731086CE358A6E91D8CC4610216303B" bold="true" pageId="0" pageNumber="165">A-D</emphasis>
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); 0.1 mm (
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<emphasis id="6ACA8C50B3D250B47E6B2AEB6FB8C804" bold="true" pageId="0" pageNumber="165">E, I</emphasis>
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); 0.5 mm (
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<emphasis id="52E26EEC1E165E364C293255BAC58BB4" bold="true" pageId="0" pageNumber="165">F-H</emphasis>
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).
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</paragraph>
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</caption>
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<caption id="AD02FBA01C943132F36806F7B4168FB0" doi="10.3897/zookeys.1182.97346.figure2" httpUri="https://binary.pensoft.net/fig/920945" pageId="0" pageNumber="165" start="Figure 2" startId="F2">
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<paragraph id="B029A60439F4BB7F17AAF54ABEB0DE97" pageId="0" pageNumber="165">
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<emphasis id="ACB197E18198A0523BE55DE46E8EA209" bold="true" pageId="0" pageNumber="165">Figure 2.</emphasis>
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Diagnostic characters of
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<taxonomicName id="D0B70804340D831214866ACEA738C88F" authorityName="Choi, Park, Lee & Park" authorityYear="2023" class="Insecta" family="Staphylinidae" genus="Bryaxis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bryaxis grandinodus" order="Coleoptera" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="grandinodus">
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<emphasis id="E740B653C86E7E5C7D021EF516690EC6" italics="true" pageId="0" pageNumber="165">Bryaxis grandinodus</emphasis>
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</taxonomicName>
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sp. nov. (
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<emphasis id="704218137695648DE4103A2BFF206343" bold="true" pageId="0" pageNumber="165">A, C</emphasis>
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) and
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<taxonomicName id="773C6146117E5E9D17874DCB3B60A065" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="B. koltzei" order="Coleoptera" pageId="0" pageNumber="165" rank="species" species="koltzei">
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<emphasis id="3BE90B8C78C4DAC2FD7E08D0DDB984DE" italics="true" pageId="0" pageNumber="165">B. koltzei</emphasis>
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</taxonomicName>
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(Reitter) (
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|
<emphasis id="E1999CC798C679306BFC159576CD75EA" bold="true" pageId="0" pageNumber="165">B, D</emphasis>
|
|
).
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|
<emphasis id="F9186291AFA693D75046836A831C2D9E" bold="true" pageId="0" pageNumber="165">A, B</emphasis>
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|
antennal scapes and pedicels
|
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<emphasis id="B0A1F71F59A5DA0ACD7B9E0106B658FA" bold="true" pageId="0" pageNumber="165">C, D</emphasis>
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|
protibiae. Scale bars: 0.1 mm.
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|
</paragraph>
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|
</caption>
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</subSubSection>
|
|
<subSubSection id="41691A6D07E84B115357337AA3929E5E" pageId="0" pageNumber="165" type="description">
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<paragraph id="8E46208A94A48FD93B053DAC1DAD322F" pageId="0" pageNumber="165">Description.</paragraph>
|
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<paragraph id="3FCCCBBA49BBA49ED63A6400E2133A3B" pageId="0" pageNumber="165">
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|
Body reddish brown, antennae, maxillary palpi, and tarsi slightly lighter, length 1.2-1.32 mm, maximum width 0.51-0.58 mm (Fig.
|
|
<figureCitation id="51364A583B2859BB4BADEA72ACBE919F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Bryaxis grandinodus Choi, Park, Lee & Park, sp. nov., male (A, B, E-I), female (C, D). A, C dorsal habitus B, D ventral habitus E head F antenna G fore leg H hind leg I aedeagus. Scale bars: 1 mm (A-D); 0.1 mm (E, I); 0.5 mm (F-H)." figureDoi="10.3897/zookeys.1182.97346.figure1" httpUri="https://binary.pensoft.net/fig/920944" pageId="0" pageNumber="165">1A-D</figureCitation>
|
|
). Setae on body yellowish, long. Head 0.92 times as long as wide (Fig.
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|
<figureCitation id="DB98715B00205ACF2C0E83CFBF010E79" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Bryaxis grandinodus Choi, Park, Lee & Park, sp. nov., male (A, B, E-I), female (C, D). A, C dorsal habitus B, D ventral habitus E head F antenna G fore leg H hind leg I aedeagus. Scale bars: 1 mm (A-D); 0.1 mm (E, I); 0.5 mm (F-H)." figureDoi="10.3897/zookeys.1182.97346.figure1" httpUri="https://binary.pensoft.net/fig/920944" pageId="0" pageNumber="165">1E</figureCitation>
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|
). Frons with U-shaped impression between antennal tubercles; frontal foveae absent; frontal rostrum distinct anteriorly. Vertex slightly convex; longitudinal carina weak; vertexal foveae large. Eyes as long as tempora, with 23-26 facets. Maxillary palpi moderately developed; palpomeres II-III with tubercles; palpomere IV 0.23 mm long and about 3.15 times as long as wide, subcylindrical pseudosegment at apex. Antennae about 0.54 mm long; pedicels subglobose with long setae, 0.89 times as long as wide; antennomere III 1.12 times as long as wide; IV-VIII subequal in length; IX-X transverse, IX 0.64 times as long as wide and X 0.63 times as long as wide; XI largest, pointed at apex, 1.67 times as long as wide (Fig.
|
|
<figureCitation id="6210B8BE6D5A51D233D86D79E35B751F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Bryaxis grandinodus Choi, Park, Lee & Park, sp. nov., male (A, B, E-I), female (C, D). A, C dorsal habitus B, D ventral habitus E head F antenna G fore leg H hind leg I aedeagus. Scale bars: 1 mm (A-D); 0.1 mm (E, I); 0.5 mm (F-H)." figureDoi="10.3897/zookeys.1182.97346.figure1" httpUri="https://binary.pensoft.net/fig/920944" pageId="0" pageNumber="165">1F</figureCitation>
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|
). Pronotum 0.84 times as long as wide and widest at basal 2/3, lateral antebasal foveae connected by antebasal sulcus. Elytra slightly convex, 0.89 times as long as wide and widest at basal 1/4, each elytron with two basal foveae and subhumeral fovea. Legs slender; protibiae without spine (Figs
|
|
<figureCitation id="BEFC333131E329BF1F8FF3556EED56AF" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Bryaxis grandinodus Choi, Park, Lee & Park, sp. nov., male (A, B, E-I), female (C, D). A, C dorsal habitus B, D ventral habitus E head F antenna G fore leg H hind leg I aedeagus. Scale bars: 1 mm (A-D); 0.1 mm (E, I); 0.5 mm (F-H)." figureDoi="10.3897/zookeys.1182.97346.figure1" httpUri="https://binary.pensoft.net/fig/920944" pageId="0" pageNumber="165">1G</figureCitation>
|
|
,
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|
<figureCitation id="175EB6CB1F646AF5D14BFAA38BEB3321" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Diagnostic characters of Bryaxis grandinodus sp. nov. (A, C) and B. koltzei (Reitter) (B, D). A, B antennal scapes and pedicels C, D protibiae. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1182.97346.figure2" httpUri="https://binary.pensoft.net/fig/920945" pageId="0" pageNumber="165">2C</figureCitation>
|
|
); metatibiae with spine at apex (Fig.
|
|
<figureCitation id="E6F1B0B075D2297647AC7AD65BC0593B" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Bryaxis grandinodus Choi, Park, Lee & Park, sp. nov., male (A, B, E-I), female (C, D). A, C dorsal habitus B, D ventral habitus E head F antenna G fore leg H hind leg I aedeagus. Scale bars: 1 mm (A-D); 0.1 mm (E, I); 0.5 mm (F-H)." figureDoi="10.3897/zookeys.1182.97346.figure1" httpUri="https://binary.pensoft.net/fig/920944" pageId="0" pageNumber="165">1H</figureCitation>
|
|
, arrow). Aedeagus large, 0.37 mm long and 1.69 times as long as wide; penis bulbous and dorsal diaphragm ovoid; parameres short and symmetrical, apices almost encountered, one robust seta and three fine setae on each apex; endophallus divided into three large struts, left dorsal strut branched at basal 1/3 and bifid at apex, right dorsal strut weakly branched basally and bifid dorso-ventrally at apex, ventral strut robust and bifid at apex (Fig.
|
|
<figureCitation id="4CACA9D9C1DD03900F86DBCFEACA2904" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Bryaxis grandinodus Choi, Park, Lee & Park, sp. nov., male (A, B, E-I), female (C, D). A, C dorsal habitus B, D ventral habitus E head F antenna G fore leg H hind leg I aedeagus. Scale bars: 1 mm (A-D); 0.1 mm (E, I); 0.5 mm (F-H)." figureDoi="10.3897/zookeys.1182.97346.figure1" httpUri="https://binary.pensoft.net/fig/920944" pageId="0" pageNumber="165">1I</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="3CAA5776BCA23DB8EAEF08EB82D68FA2" pageId="0" pageNumber="165">
|
|
<emphasis id="21AA2D8D226C1E956179A8DEA4C7CDBB" bold="true" pageId="0" pageNumber="165">
|
|
<emphasis id="C5ADCEE6A58F4C71CEBC5249D515DBC7" italics="true" pageId="0" pageNumber="165">Sexual dimorphism</emphasis>
|
|
.
|
|
</emphasis>
|
|
Female eyes shorter than tempora, composed of 11 facets; antennal scapes subcylindrical, without modification; metatibial spines absent.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="0E65614EF9A113A84D8AC578720A09E2" pageId="0" pageNumber="165" type="remarks">
|
|
<paragraph id="3771C1C972A92BC48F602BC9FDEACB17" pageId="0" pageNumber="165">Remarks.</paragraph>
|
|
<paragraph id="B99AD049DAFAC75848F21D97A272F632" pageId="0" pageNumber="165">
|
|
Adults of this species are very similar to
|
|
<taxonomicName id="3E129CFB7C27BC905F6859D2B86F615A" baseAuthorityName="Reitter" baseAuthorityYear="1887" class="Insecta" family="Staphylinidae" genus="Bryaxis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bryaxis koltzei" order="Coleoptera" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="koltzei">
|
|
<emphasis id="FB9A8EF0F1DA78C75DF4B7B4364B288A" italics="true" pageId="0" pageNumber="165">Bryaxis koltzei</emphasis>
|
|
</taxonomicName>
|
|
(Reitter, 1887) in the general body characters, but can be distinguished by the shape of the antennal scapes and its glandular nodule (Fig.
|
|
<figureCitation id="7BD5B46E086FDF98358BD1C82A9969E0" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Diagnostic characters of Bryaxis grandinodus sp. nov. (A, C) and B. koltzei (Reitter) (B, D). A, B antennal scapes and pedicels C, D protibiae. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1182.97346.figure2" httpUri="https://binary.pensoft.net/fig/920945" pageId="0" pageNumber="165">2A</figureCitation>
|
|
, arrow) and the spineless protibiae in the male (Fig.
|
|
<figureCitation id="D40F33A07582AF64A942585841C15B88" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Diagnostic characters of Bryaxis grandinodus sp. nov. (A, C) and B. koltzei (Reitter) (B, D). A, B antennal scapes and pedicels C, D protibiae. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1182.97346.figure2" httpUri="https://binary.pensoft.net/fig/920945" pageId="0" pageNumber="165">2C</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="A2149A8BEDE6EF9DCB6B011CD2245E2C" pageId="0" pageNumber="165" type="comments">
|
|
<paragraph id="26F863EF0E055B2C7A53B26F867905D4" pageId="0" pageNumber="165">Comments.</paragraph>
|
|
<paragraph id="5B8938DD748FF3A41D9056A92A10BDB5" pageId="0" pageNumber="165">
|
|
The localities of
|
|
<taxonomicName id="482F5782FF71B4873F8057B2CBE324CA" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="B. grandinodus" order="Coleoptera" pageId="0" pageNumber="165" rank="species" species="grandinodus">
|
|
<emphasis id="BA3CFE51A7E847A3D775DF37F65E74ED" italics="true" pageId="0" pageNumber="165">B. grandinodus</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. probably overlap with those of
|
|
<taxonomicName id="A75883F87382AEC3184D7275E1D4952D" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="B. koltzei" order="Coleoptera" pageId="0" pageNumber="165" rank="species" species="koltzei">
|
|
<emphasis id="E0F062E4CA35C7F50932EAE4A06D8149" italics="true" pageId="0" pageNumber="165">B. koltzei</emphasis>
|
|
</taxonomicName>
|
|
(Reitter) given that the latter are distributed across the entire country (Fig.
|
|
<figureCitation id="58FBFF65F3781FEE501706A845FECB80" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Collection localities. Bryaxis grandinodus sp. nov. (blue pentagon); B. uljinensis sp. nov. (purple cross); B. fabaiformis sp. nov. (sky-blue star); B. girinensis sp. nov. (green diamond); B. nemorosus sp. nov. (navy hexagon); B. mahunkai Loebl (yellow square); B. koltzei (Reitter) (red circle); B. kimjongkuki Nomura & Lee (orange triangle)." figureDoi="10.3897/zookeys.1182.97346.figure11" httpUri="https://binary.pensoft.net/fig/920954" pageId="0" pageNumber="165">11</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="E58D8A61E979F9435364BA6B2FF2941D" pageId="0" pageNumber="165" type="etymology">
|
|
<paragraph id="B83671D52A2FAC10EE281F39573C19D5" pageId="0" pageNumber="165">Etymology.</paragraph>
|
|
<paragraph id="BEE45FCC1196427C86319C4577D8590E" pageId="0" pageNumber="165">
|
|
The specific epithet is a combination of the Latin words
|
|
<emphasis id="F299F68A937EBACA04CBAE34E03DCE7D" italics="true" pageId="0" pageNumber="165">grandis</emphasis>
|
|
(
|
|
<normalizedToken id="F5CA1499D35BC5EBB75A847A5CC25D91" originalValue="“large”">"large"</normalizedToken>
|
|
, masculine) and
|
|
<emphasis id="4FC5451B109C56BAFECB940BB4A23B86" italics="true" pageId="0" pageNumber="165">nodus</emphasis>
|
|
(
|
|
<normalizedToken id="B685F4CBBE10A0B0DCCC48A7A149837A" originalValue="“knob”">"knob"</normalizedToken>
|
|
, masculine) and refers to the shape of the glandular nodules on the male antennal scapes.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="501A672BE44728DB1D23614A1C92D664" pageId="0" pageNumber="165" type="habitat">
|
|
<paragraph id="2049D20C6AE89FF7EE1DE44CD165286C" pageId="0" pageNumber="165">Habitat.</paragraph>
|
|
<paragraph id="E16F5D23785CD7357099E036EF970713" pageId="0" pageNumber="165">The holotype was collected by sifting leaf litter in mixed forest. Paratypes were collected by sifting leaf litter and soil.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C325E82414E5FFA6BBDAF217801E469F" pageId="0" pageNumber="165" type="distribution">
|
|
<paragraph id="A86D9DDFDE644E371FD518F8EA2DFF9A" pageId="0" pageNumber="165">Distribution.</paragraph>
|
|
<paragraph id="56FA36839AEB7764FAFD40FF61098271" pageId="0" pageNumber="165">Korea (Haenam-gun, Jeollanam-do; Gunsan-si, Jeollabuk-do).</paragraph>
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</subSubSection>
|
|
</treatment>
|
|
</document> |