215 lines
30 KiB
XML
215 lines
30 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/jhr.84.68709" ID-GBIF-Dataset="b9e61bc3-4158-4bee-8d57-a65655588f70" ID-Pensoft-Pub="1314-2607-84-145" ID-Pensoft-UUID="AEDB6967BB17574BB6988187C56EE6CA" ID-Zenodo-Dep="5349694" ID-ZooBank="5F83EC637C7947FDBD29E7A2B9328518" ModsDocID="1314-2607-84-145" checkinTime="1629847073322" checkinUser="pensoft" docAuthor="Okayasu, Juriya" docDate="2021" docId="1A048A79BD0E579B8F8A615E811A9FDB" docLanguage="en" docName="JourHymenoptRes 84: 145-168" docOrigin="Journal of Hymenoptera Research 84" docPubDate="2021-08-24" docSource="http://dx.doi.org/10.3897/jhr.84.68709" docTitle="Arkaditilla leleji Okayasu 2021, sp. nov." docType="treatment" docUuid="1A4CD8F4-673F-40E8-8A9A-D12E0DBFFE0A" docUuidSource="ZooBank" docVersion="4" id="AEDB6967BB17574BB6988187C56EE6CA" lastPageNumber="145" masterDocId="AEDB6967BB17574BB6988187C56EE6CA" masterDocTitle="A new velvet ant genus Arkaditilla (Hymenoptera, Mutillidae, Trogaspidiini) from the Oriental Region, with review of species" masterLastPageNumber="168" masterPageNumber="145" pageNumber="145" updateTime="1678756233000" updateUser="pensoft">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>A new velvet ant genus Arkaditilla (Hymenoptera, Mutillidae, Trogaspidiini) from the Oriental Region, with review of species</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Okayasu, Juriya</mods:namePart>
|
||
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6307-2939</mods:nameIdentifier>
|
||
<mods:affiliation>Entomological Laboratory, Faculty of Agriculture, Ehime University, Tarumi 3 - 5 - 7, Matsuyama, 790 - 8566, Japan & Systematic Entomology, Graduate School of Agriculture, Hokkaido University, Sapporo, 060 - 8589, Japan</mods:affiliation>
|
||
<mods:nameIdentifier type="email">mutiphiidae@gmail.com</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>Journal of Hymenoptera Research</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2021</mods:date>
|
||
<mods:detail type="pubDate">
|
||
<mods:number>2021-08-24</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="volume">
|
||
<mods:number>84</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>145</mods:start>
|
||
<mods:end>168</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location>
|
||
<mods:url>http://dx.doi.org/10.3897/jhr.84.68709</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">http://dx.doi.org/10.3897/jhr.84.68709</mods:identifier>
|
||
<mods:identifier type="Pensoft-Pub">1314-2607-84-145</mods:identifier>
|
||
<mods:identifier type="ZooBank">5F83EC637C7947FDBD29E7A2B9328518</mods:identifier>
|
||
<mods:identifier type="Pensoft-UUID">AEDB6967BB17574BB6988187C56EE6CA</mods:identifier>
|
||
<mods:identifier type="Zenodo-Dep">5349694</mods:identifier>
|
||
</mods:mods>
|
||
<treatment ID-GBIF-Taxon="184941298" LSID="urn:lsid:zoobank.org:act:1A4CD8F4-673F-40E8-8A9A-D12E0DBFFE0A" httpUri="http://treatment.plazi.org/id/1A048A79BD0E579B8F8A615E811A9FDB" lastPageNumber="145" pageId="0" pageNumber="145">
|
||
<subSubSection pageId="0" pageNumber="145" type="nomenclature">
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<taxonomicName LSID="http://zoobank.org/1A4CD8F4-673F-40E8-8A9A-D12E0DBFFE0A" authority="Okayasu, 2021" authorityName="Okayasu" authorityYear="2021" class="Insecta" family="Mutillidae" genus="Arkaditilla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Arkaditilla leleji" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="leleji" status="sp. nov.">Arkaditilla leleji</taxonomicName>
|
||
<taxonomicNameLabel pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="description">
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<figureCitation captionStart="Figures 7–12" captionStartId="F2" captionText="Figures 7 - 12. Arkaditilla spp., ♂, habitus 7, 9, 11 dorsal view 8, 10, 12 lateral view 7, 8 A. gibba sp. nov., holotype 9, 10 A. leleji sp. nov., holotype 11, 12 A. nallinia (Zavattari). Scale bars: 3 mm." figureDoi="10.3897/jhr.84.68709.figures7-12" httpUri="https://binary.pensoft.net/fig/579481" pageId="0" pageNumber="145">Figures 9</figureCitation>
|
||
<figureCitation captionStart="Figures 7–12" captionStartId="F2" captionText="Figures 7 - 12. Arkaditilla spp., ♂, habitus 7, 9, 11 dorsal view 8, 10, 12 lateral view 7, 8 A. gibba sp. nov., holotype 9, 10 A. leleji sp. nov., holotype 11, 12 A. nallinia (Zavattari). Scale bars: 3 mm." figureDoi="10.3897/jhr.84.68709.figures7-12" httpUri="https://binary.pensoft.net/fig/579481" pageId="0" pageNumber="145">, 10</figureCitation>
|
||
<figureCitation captionStart="Figures 13–18" captionStartId="F3" captionText="Figures 13 - 18. Arkaditilla spp., ♂, face 13 A. bagrada (Cameron) 14 A. depressicornis (Mickel) 15 A. frim sp. nov., holotype 16 A. gibba sp. nov., paratype 17 A. leleji sp. nov., holotype 18 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures13-18" httpUri="https://binary.pensoft.net/fig/579482" pageId="0" pageNumber="145">, 17</figureCitation>
|
||
<figureCitation captionStart="Figures 19–36" captionStartId="F4" captionText="Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures19-36" httpUri="https://binary.pensoft.net/fig/579483" pageId="0" pageNumber="145">, 23</figureCitation>
|
||
<figureCitation captionStart="Figures 19–36" captionStartId="F4" captionText="Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures19-36" httpUri="https://binary.pensoft.net/fig/579483" pageId="0" pageNumber="145">, 33</figureCitation>
|
||
<figureCitation captionStart="Figures 19–36" captionStartId="F4" captionText="Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures19-36" httpUri="https://binary.pensoft.net/fig/579483" pageId="0" pageNumber="145">, 34</figureCitation>
|
||
<figureCitation captionStart="Figures 37–48" captionStartId="F5" captionText="Figures 37 - 48. Arkaditilla spp., ♂, genitalia 37, 39, 41, 43, 45, 47 dorsal view 38, 40, 42, 44, 46, 48 ventral view 37, 38 A. bagrada (Cameron) 39, 40 A. depressicornis (Mickel) 41, 42 A. frim sp. nov., holotype 43, 44 A. gibba sp. nov., holotype 45, 46 A. leleji sp. nov., holotype 47, 48 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures37-48" httpUri="https://binary.pensoft.net/fig/579484" pageId="0" pageNumber="145">, 45</figureCitation>
|
||
<figureCitation captionStart="Figures 37–48" captionStartId="F5" captionText="Figures 37 - 48. Arkaditilla spp., ♂, genitalia 37, 39, 41, 43, 45, 47 dorsal view 38, 40, 42, 44, 46, 48 ventral view 37, 38 A. bagrada (Cameron) 39, 40 A. depressicornis (Mickel) 41, 42 A. frim sp. nov., holotype 43, 44 A. gibba sp. nov., holotype 45, 46 A. leleji sp. nov., holotype 47, 48 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures37-48" httpUri="https://binary.pensoft.net/fig/579484" pageId="0" pageNumber="145">, 46</figureCitation>
|
||
<figureCitation captionStart="Figures 49–66" captionStartId="F6" captionText="Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures49-66" httpUri="https://binary.pensoft.net/fig/579485" pageId="0" pageNumber="145">, 61-63</figureCitation>
|
||
<figureCitation captionStart="Figures 67–72" captionStartId="F7" captionText="Figures 67 - 72. Arkaditilla spp., ♂, genital volsella, ventro-inner view 67 A. bagrada (Cameron) 68 A. depressicornis (Mickel) 69 A. frim sp. nov., holotype 70 A. gibba sp. nov., holotype 71 A. leleji sp. nov., holotype 72 A. nallinia (Zavattari). White arrows indicate the paracuspis." figureDoi="10.3897/jhr.84.68709.figures67-72" httpUri="https://binary.pensoft.net/fig/579486" pageId="0" pageNumber="145">, 71</figureCitation>
|
||
<figureCitation captionStart="Figure 73–76" captionStartId="F8" captionText="Figure 73 - 76. Distributional map of Arkaditilla spp. based on specimens examined herein and the literature (Cameron 1902; Lelej 1996 b; Mickel 1935, 1937; Zavattari 1914) 73 A. bagrada (Cameron) (red circle) 74 A. depressicornis (Mickel) (blue circle) 75 A. frim sp. nov. (yellow circle), A. gibba sp. nov. (purple triangle), and A. leleji sp. nov. (light blue square) 76 A. nallinia (Zavattari) (orange circle)." figureDoi="10.3897/jhr.84.68709.figure73-76" httpUri="https://binary.pensoft.net/fig/579487" pageId="0" pageNumber="145">, 75</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="diagnosis">
|
||
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" pageId="0" pageNumber="145">Male.</emphasis>
|
||
Clypeus dorsally elevated, sloping ventrally. Mandible with sharp subapical and large inner subbasal teeth lamellately expanded near apex (Fig.
|
||
<figureCitation captionStart="Figures 13–18" captionStartId="F3" captionText="Figures 13 - 18. Arkaditilla spp., ♂, face 13 A. bagrada (Cameron) 14 A. depressicornis (Mickel) 15 A. frim sp. nov., holotype 16 A. gibba sp. nov., paratype 17 A. leleji sp. nov., holotype 18 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures13-18" httpUri="https://binary.pensoft.net/fig/579482" pageId="0" pageNumber="145">17</figureCitation>
|
||
), ventrally slightly excised with small basal tooth (Fig.
|
||
<figureCitation captionStart="Figures 19–36" captionStartId="F4" captionText="Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures19-36" httpUri="https://binary.pensoft.net/fig/579483" pageId="0" pageNumber="145">23</figureCitation>
|
||
). Scape ventral carinae divergent apically (Fig.
|
||
<figureCitation captionStart="Figures 19–36" captionStartId="F4" captionText="Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures19-36" httpUri="https://binary.pensoft.net/fig/579483" pageId="0" pageNumber="145">33</figureCitation>
|
||
), densely punctate between them; F1 apical half and F2-F6 whitish yellow on ventral half, F7-F11 with ventral dark brown patch; F1 weakly depressed, F1 height subequal to pedicel diameter (Fig.
|
||
<figureCitation captionStart="Figures 19–36" captionStartId="F4" captionText="Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures19-36" httpUri="https://binary.pensoft.net/fig/579483" pageId="0" pageNumber="145">34</figureCitation>
|
||
). T3 black (Fig.
|
||
<figureCitation captionStart="Figures 7–12" captionStartId="F2" captionText="Figures 7 - 12. Arkaditilla spp., ♂, habitus 7, 9, 11 dorsal view 8, 10, 12 lateral view 7, 8 A. gibba sp. nov., holotype 9, 10 A. leleji sp. nov., holotype 11, 12 A. nallinia (Zavattari). Scale bars: 3 mm." figureDoi="10.3897/jhr.84.68709.figures7-12" httpUri="https://binary.pensoft.net/fig/579481" pageId="0" pageNumber="145">10</figureCitation>
|
||
); cuspis weakly widened on posterior 1/3 with rounded apex (Figs
|
||
<figureCitation captionStart="Figures 49–66" captionStartId="F6" captionText="Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures49-66" httpUri="https://binary.pensoft.net/fig/579485" pageId="0" pageNumber="145">61</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 67–72" captionStartId="F7" captionText="Figures 67 - 72. Arkaditilla spp., ♂, genital volsella, ventro-inner view 67 A. bagrada (Cameron) 68 A. depressicornis (Mickel) 69 A. frim sp. nov., holotype 70 A. gibba sp. nov., holotype 71 A. leleji sp. nov., holotype 72 A. nallinia (Zavattari). White arrows indicate the paracuspis." figureDoi="10.3897/jhr.84.68709.figures67-72" httpUri="https://binary.pensoft.net/fig/579486" pageId="0" pageNumber="145">71</figureCitation>
|
||
); digitus posteriorly slender stick-like (Fig.
|
||
<figureCitation captionStart="Figures 37–48" captionStartId="F5" captionText="Figures 37 - 48. Arkaditilla spp., ♂, genitalia 37, 39, 41, 43, 45, 47 dorsal view 38, 40, 42, 44, 46, 48 ventral view 37, 38 A. bagrada (Cameron) 39, 40 A. depressicornis (Mickel) 41, 42 A. frim sp. nov., holotype 43, 44 A. gibba sp. nov., holotype 45, 46 A. leleji sp. nov., holotype 47, 48 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures37-48" httpUri="https://binary.pensoft.net/fig/579484" pageId="0" pageNumber="145">45</figureCitation>
|
||
); paracuspis slightly tuberculate (Fig.
|
||
<figureCitation captionStart="Figures 67–72" captionStartId="F7" captionText="Figures 67 - 72. Arkaditilla spp., ♂, genital volsella, ventro-inner view 67 A. bagrada (Cameron) 68 A. depressicornis (Mickel) 69 A. frim sp. nov., holotype 70 A. gibba sp. nov., holotype 71 A. leleji sp. nov., holotype 72 A. nallinia (Zavattari). White arrows indicate the paracuspis." figureDoi="10.3897/jhr.84.68709.figures67-72" httpUri="https://binary.pensoft.net/fig/579486" pageId="0" pageNumber="145">71</figureCitation>
|
||
); penis valve strongly expanded ventrally on posterior half (Figs
|
||
<figureCitation captionStart="Figures 49–66" captionStartId="F6" captionText="Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures49-66" httpUri="https://binary.pensoft.net/fig/579485" pageId="0" pageNumber="145">62</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 49–66" captionStartId="F6" captionText="Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures49-66" httpUri="https://binary.pensoft.net/fig/579485" pageId="0" pageNumber="145">63</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="145">Female.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
<caption ID-Zenodo-Dep="5349706" doi="10.3897/jhr.84.68709.figures49-66" httpUri="https://binary.pensoft.net/fig/579485" pageId="0" pageNumber="145" start="Figures 49–66" startId="F6">
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" pageId="0" pageNumber="145">Figures 49-66.</emphasis>
|
||
<taxonomicName authorityName="Okayasu" authorityYear="2021" class="Insecta" family="Mutillidae" genus="Arkaditilla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Arkaditilla" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Arkaditilla</emphasis>
|
||
</taxonomicName>
|
||
spp., ♂, genitalia
|
||
<emphasis bold="true" pageId="0" pageNumber="145">49, 52, 55, 58, 61, 64</emphasis>
|
||
genital capsule and volsella, inner view
|
||
<emphasis bold="true" pageId="0" pageNumber="145">50, 53, 56, 59, 62, 65</emphasis>
|
||
left penis valve
|
||
<emphasis bold="true" pageId="0" pageNumber="145">51, 54, 57, 60, 63, 66</emphasis>
|
||
right penis valve
|
||
<emphasis bold="true" pageId="0" pageNumber="145">49-51</emphasis>
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. bagrada" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="bagrada">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">A. bagrada</emphasis>
|
||
</taxonomicName>
|
||
(Cameron)
|
||
<emphasis bold="true" pageId="0" pageNumber="145">52-54</emphasis>
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. depressicornis" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="depressicornis">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">A. depressicornis</emphasis>
|
||
</taxonomicName>
|
||
(Mickel)
|
||
<emphasis bold="true" pageId="0" pageNumber="145">55-57</emphasis>
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. frim" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="frim">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">A. frim</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype
|
||
<emphasis bold="true" pageId="0" pageNumber="145">58-60</emphasis>
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. gibba" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="gibba">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">A. gibba</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype
|
||
<emphasis bold="true" pageId="0" pageNumber="145">61-63</emphasis>
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. leleji" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="leleji">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">A. leleji</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype
|
||
<emphasis bold="true" pageId="0" pageNumber="145">64-66</emphasis>
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. nallinia" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="nallinia">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">A. nallinia</emphasis>
|
||
</taxonomicName>
|
||
(Zavattari).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="description">
|
||
<paragraph pageId="0" pageNumber="145">Description.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" pageId="0" pageNumber="145">Male.</emphasis>
|
||
Body length: 15.9 mm; forewing length: 13.0 mm.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" pageId="0" pageNumber="145">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Color and setae</emphasis>
|
||
.
|
||
</emphasis>
|
||
Frons, vertex, gena, clypeus, mandible mostly, pedicel, flagellum mostly, prementum, stipes, mesosoma, meso- and metacoxae, T3-T7, S1 medial carina, S4-S8 black; mandible medial 1/3 brown; labrum, protibial spur yellow; scape, F7-F11 ventrally, trochanters dark brown; F1 apical half, F2-F6 whitish yellow on ventral half; tegula, procoxa, femora, tibiae, tarsi, S3 lateral portion brownish black; meso- and metatibial spurs pale yellow; T1-T2, S1 except medial carina, S2-S3 mostly orange; wings brown, with veins dark brown. Upper frons, vertex, T1, S1 with sparse long erect pale golden setae; lower frons, gena, pronotal dorsum, dorsal propodeal face, dorsal half of lateral propodeal face, coxae with dense short appressed and sparse long erect pale golden setae; eye dorsal margin with long erect black setae; clypeus with dense long recumbent pale golden setae; mandible with sparse short recumbent and sparse long erect pale golden setae; scape with sparse long recumbent and sparse long erect pale golden setae; pedicel, F1, trochanters, T2 lateral margin with sparse short recumbent pale golden setae; F2-F11 with sparse erect pale golden microsetae; pronotal collar anteriorly, T2 disc, T3-T5, S3-S6 with sparse short erect pale golden setae; pronotal collar posteriorly glabrous; pronotal lateral face with sparse long appressed pale golden setae; propleuron with sparse long appressed and sparse long erect pale golden setae; mesoscutum with sparse short recumbent and sparse short erect dark golden setae; meso- and metascutellum, femora with sparse short recumbent pale golden and sparse long erect golden setae; anterior and posterior parts of mesopleuron, metapleuron, ventral half of lateral propodeal face with sparse short appressed pale golden setae; axilla, metanotal trough, medial and ventral parts of mesopleuron with dense long appressed and sparse long erect pale golden setae; tegula anterior portion, tarsi with sparse long recumbent golden setae; tegula inner margin with sparse short recumbent and sparse short erect golden setae; tibiae with sparse short recumbent golden and sparse long erect golden setae; T6-T7, S7-S8 with sparse short erect black setae; S2 with sparse short recumbent and sparse short erect pale golden setae; T2 with lateral golden felt line; T1-T5, S2-S5 with posterior fringe of sparse pale golden setae; S6-S8 with posterior fringe of sparse black setae.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" pageId="0" pageNumber="145">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Structure</emphasis>
|
||
.
|
||
</emphasis>
|
||
Head 2.00
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as wide as long; lateral margins strongly convergent behind eye; vertex rounded posteriorly; POD/OOD = 0.54; OD:MOD:LOD = 1.00:1.29:1.00; ocellar region slightly convex, ocelli without posterior carina; antennal scrobe with weak inverted V-shaped dorsal carina not reaching inner eye margin; genal carina absent; postgenal bridge flattened; hypostomal carina weakly developed; clypeus dorsally elevated, sloping ventrally, without subventral transverse groove; anterior clypeal margin projecting medially; mandible robust, narrowed to apex with inner margin expanded forming large subbasal tooth; mandible apex bidentate; mandible ventral margin slightly excised into small basal tooth; mandible dorsal carina sharp; prementum flattened; scape curved medially, with two longitudinal ventral carinae divergent apically; F1 depressed, F1 height 0.91
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
pedicel diameter; relative length of pedicel, F1 and F2 = 1.00:3.00:2.37; F2-F11 almost equal in width; F11 as long as F2, conical.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
Mesosoma widest at mesonotum including tegulae; head width:humeral width:pronotal width:mesonotal width = 83:62:85:100; humeral angle rounded, with sharp humeral carina not reaching pronotal dorsum; mesoscutum strongly convex; mesoscutal width 1.11
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
its length; tegula short rounded, posteriorly reaching mesoscuto-scutellar articulation, 1.54
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as wide; notaulus evenly wide, reaching anterior 1/2 of mesoscutum, subparallel; parapsidal line length 0.3
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
mesoscutal length; parascutal carina forming rounded, weakly elevated process on postero-lateral angle of mesoscutum; mesoscutellum convex without medial carina, tubercle or impunctate line; mesopleuron divided into dorsal and ventral convexities by distinct transverse mesopleural groove, ventrally with transverse precoxal ridge, without longitudinal impunctate line, or precoxal swelling; dorsal propodeal face without distinct dorsum and sublateral carina, with anterior inverted triangular cell and semicircular sublateral cell; lateral margins of propodeum gradually convergent posteriorly; dorsal and lateral propodeal faces not separated by carina.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">Wings well developed; distance between origin of RS on vein SC and base of stigmatic cell equal to stigmatic cell length and first abscissa of RS length; cell 2RS present.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">Meso- and metacoxae evenly convex; metacoxa with weak complete inner longitudinal carina; protibial spur slender; tibiae without outer spines.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
T1 without distinct dorsum; T1 spiracle weakly tuberculate; T2 evenly convex, 1.15
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as wide as long; S1 with straight medial longitudinal carina with its ventral margin convex; S2 swollen anteriorly; T7 medially with longitudinal impunctate convex line, posteriorly not forming tubercle; S6-S8 without lateral carina or tubercle; S7 entirely exposed, 0.75
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as S8; S8 mostly sclerotized with posterior membranous area evenly convex posteriorly. Paramere down-curved posteriorly, without inner protuberance on dorso-inner margin; inner margin of paramere dorsally with short erect golden setae; ventral lobe posteriorly projecting, pointed; parapenial lobe large, with posterior margin slightly concave; cuspis long slender, weakly widened on posterior 1/3 with rounded apex, apparently surpassing apex of penis valve, with dense long erect setae ventrally and posteriorly; digitus straight, posteriorly not expanded, with dense erect golden setae posteriorly; paracuspis slightly tuberculate with few erect golden setae; penis valves symmetrical slender, strongly expanded ventrally on posterior half, with posterior ventral hook, without setae.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">Frons, vertex, gena, pronotal dorsum, mesoscutum, medial part of mesopleuron with large dense punctures; postgenal bridge, trochanters, femora, tibiae, T1 medially, T2 disc with small sparse punctures; clypeus mostly, mandible outer and dorsal faces, scape between ventral carinae, flagellum, pronotal lateral face, propleuron, axilla, metanotal trough, anterior and posterior parts of mesopleuron, metapleuron, ventral half of lateral propodeal face, coxae, tarsi with minute dense punctures; clypeus anterior margin with few large punctures; scape mostly, pedicel, T1-T2 laterally, T3-T7, S1-S8 with small dense punctures; pronotal collar anteriorly with minute sparse punctures; pronotal collar posteriorly smooth; mesoscutellum with large confluent punctures; metascutellum with small confluent punctures; dorsal propodeal face, dorsal half of lateral propodeal face reticulate; tegula mostly microreticulate with few small punctures anteriorly, postero-inner portion with minute sparse punctures.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="type material">
|
||
<paragraph pageId="0" pageNumber="145">Type material.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
1♂.
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
|
||
♂:
|
||
<emphasis bold="true" pageId="0" pageNumber="145">Indonesia</emphasis>
|
||
: Kalimantan Timur: Bukit Soeharto, 13-17.III.1994, K. Matsumoto & K. Konishi leg. [NARO].
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="distribution">
|
||
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">Indonesia: Kalimantan Timur.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="etymology">
|
||
<paragraph pageId="0" pageNumber="145">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">This new species is dedicated to Arkady Lelej as is the genus itself.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="remarks">
|
||
<paragraph pageId="0" pageNumber="145">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
This new species is unique in
|
||
<taxonomicName authorityName="Okayasu" authorityYear="2021" class="Insecta" family="Mutillidae" genus="Arkaditilla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Arkaditilla" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Arkaditilla</emphasis>
|
||
</taxonomicName>
|
||
by having the mandible ventral margin slightly excised with small basal tooth, F1 weakly depressed, T3 black, and the penis valve strongly expanded ventrally on posterior half.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |