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<document id="B2B26A2CF4C25A4145260173CDF3189C" ID-CLB-Dataset="2636" ID-DOI="10.11646/zootaxa.5091.1.1" ID-GBIF-Dataset="51b42715-ddb1-4b4a-aad4-8be0f600a287" ID-ISSN="1175-5326" ID-Zenodo-Dep="5840434" ID-ZooBank="ABC8AF70-F691-4D07-8F20-70934642C8BC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1641986550580" checkinUser="plazi" docAuthor="Bergh, Eugene W. &amp; Compton, John S." docDate="2022" docId="197787BAFFC6933B7FC99B9BFCA6FCAD" docLanguage="en" docName="zootaxa.5091.1.1.pdf" docOrigin="Zootaxa 5091 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bolivina reticulata Hantken 1875" docType="treatment" docVersion="6" lastPageNumber="22" masterDocId="E54EFFC2FFD3932E7F5E9B0CFFC4FF88" masterDocTitle="Taxonomy of Middle Miocene foraminifera from the northern Namibian continental shelf" masterLastPageNumber="55" masterPageNumber="1" pageNumber="22" updateTime="1699301522325" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="EF5A9E1BDF8215CA07F8B42959D4C48A">Taxonomy of Middle Miocene foraminifera from the northern Namibian continental shelf</mods:title>
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<mods:namePart id="7B47456510E33F7DD785E8AB744DFAE7">Bergh, Eugene W.</mods:namePart>
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<mods:affiliation id="00FE7DD959762154492650FE0BBD26E3">Marine Research Institute and Department of Geological Sciences, University of Cape Town, Private Bag X 3, Rondebosch, 7701, South Africa. &amp; Department of Environmental and Geographical Sciences, University of Cape Town, Private Bag X 3, Rondebosch, 7701, South Africa; &amp; Invertebrate palaeontology and Geology, Iziko South African Museum, P. O. Box 61, Cape Town 8000, South Africa. john. compton @ uct. ac. za; https: // orcid. org / 0000 - 0002 - 0765 - 4141</mods:affiliation>
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<treatment id="197787BAFFC6933B7FC99B9BFCA6FCAD" ID-DOI="http://doi.org/10.5281/zenodo.5840517" ID-GBIF-Taxon="192044069" ID-Zenodo-Dep="5840517" LSID="urn:lsid:plazi:treatment:197787BAFFC6933B7FC99B9BFCA6FCAD" httpUri="http://treatment.plazi.org/id/197787BAFFC6933B7FC99B9BFCA6FCAD" lastPageNumber="22" pageId="21" pageNumber="22">
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<paragraph id="916136ACFFC6933B7FC99B9BFDF3FF39" blockId="21.[151,567,151,213]" box="[151,567,151,178]" pageId="21" pageNumber="22">
<heading id="CA2981C0FFC6933B7FC99B9BFDF3FF39" bold="true" box="[151,567,151,178]" fontSize="11" level="1" pageId="21" pageNumber="22" reason="1">
<taxonomicName id="56DE4D2FFFC6933B7FC99B9BFDF3FF39" ID-CoL="5WMN9" authority="Hantken, 1875" authorityName="Hantken" authorityYear="1875" box="[151,567,151,178]" class="Globothalamea" family="Bolivinitidae" genus="Bolivina" kingdom="Chromista" order="Rotaliida" pageId="21" pageNumber="22" phylum="Foraminifera" rank="species" species="reticulata">
<emphasis id="A3AAEABEFFC6933B7FC99B9BFDF3FF39" bold="true" box="[151,567,151,178]" pageId="21" pageNumber="22">
<emphasis id="A3AAEABEFFC6933B7FC99B9BFEBDFF3A" bold="true" box="[151,377,151,178]" italics="true" pageId="21" pageNumber="22">Bolivina reticulata</emphasis>
<bibRefCitation id="F54F4B5DFFC6933B7EDE9B9BFDF3FF39" author="Hantken, M." box="[384,567,151,177]" pageId="21" pageNumber="22" pagination="1 - 93" refId="ref32414" refString="Hantken, M. (1875) A Clavulina szaboi retegek Faunaja. I. Foraminiferak. Foldtani Inezet Evkonyve, 4, 1 - 93." type="journal article" year="1875">Hantken, 1875</bibRefCitation>
</emphasis>
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</paragraph>
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<heading id="CA2981C0FFC6933B7FC99BB7FE88FF5D" bold="true" box="[151,332,187,213]" fontSize="11" level="1" pageId="21" pageNumber="22" reason="1">
<emphasis id="A3AAEABEFFC6933B7FC99BB7FE88FF5D" bold="true" box="[151,332,187,213]" pageId="21" pageNumber="22">Pl. 4, figs. 8ab</emphasis>
</heading>
</paragraph>
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<subSubSection id="D9C46527FFC6933B7FC99BF6FD7EFED8" pageId="21" pageNumber="22" type="reference_group">
<paragraph id="916136ACFFC6933B7FC99BF6FF23FEB9" blockId="21.[151,1437,250,336]" pageId="21" pageNumber="22">
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<treatmentCitation id="107F10BDFFC6933B7FC99BF6FDF4FE9A" author="Hantken, M." box="[151,560,250,274]" page="65" pageId="21" pageNumber="22" year="1875">
<taxonomicName id="56DE4D2FFFC6933B7FC99BF6FDF4FE9A" ID-CoL="5WMN9" authority="Hantken, 1875, p. 65" authorityName="Hantken" authorityPageNumber="65" authorityYear="1875" box="[151,560,250,274]" class="Globothalamea" family="Bolivinitidae" genus="Bolivina" kingdom="Chromista" order="Rotaliida" pageId="21" pageNumber="22" phylum="Foraminifera" rank="species" species="reticulata">
<emphasis id="A3AAEABEFFC6933B7FC99BF6FE91FE9A" box="[151,341,250,274]" italics="true" pageId="21" pageNumber="22">Bolivina reticulata</emphasis>
<bibRefCitation id="F54F4B5DFFC6933B7E059BF6FE36FE9A" author="Hantken, M." box="[347,498,250,274]" pageId="21" pageNumber="22" pagination="1 - 93" refId="ref32414" refString="Hantken, M. (1875) A Clavulina szaboi retegek Faunaja. I. Foraminiferak. Foldtani Inezet Evkonyve, 4, 1 - 93." type="journal article" year="1875">Hantken, 1875</bibRefCitation>
, p. 65
</taxonomicName>
</treatmentCitation>
, pl. 15, fig. 6;
<treatmentCitation id="107F10BDFFC6933B7D9C9BF7FC0AFE9A" author="Fenero, R. &amp; Thomas, E. &amp; Alegret, L. &amp; Molina, E." box="[706,974,250,274]" page="292" pageId="21" pageNumber="22" year="2012">
<bibRefCitation id="F54F4B5DFFC6933B7D9C9BF7FC46FE9A" author="Fenero, R. &amp; Thomas, E. &amp; Alegret, L. &amp; Molina, E." box="[706,898,250,274]" pageId="21" pageNumber="22" pagination="286 - 304" refId="ref31452" refString="Fenero, R., Thomas, E., Alegret, L. &amp; Molina, E. (2012) Oligocene benthic foraminifera from the Fuente Caldera section (Spain, western Tethys): taxonomy and paleoenvironmental inferences. Journal of Foraminiferal Research, 42 (4), 286 - 304. https: // doi. org / 10.2113 / gsjfr. 42.4.286" type="journal article" year="2012">
Fenero
<emphasis id="A3AAEABEFFC6933B7C519BF7FCFFFE9A" box="[783,827,250,274]" italics="true" pageId="21" pageNumber="22">et al</emphasis>
., 2012
</bibRefCitation>
, p. 292
</treatmentCitation>
, fig. 5.7;
<treatmentCitation id="107F10BDFFC6933B7B739BF6FAF2FE9A" author="Valchev, B. &amp; Stojanova, V. &amp; Juranov, S." box="[1069,1334,250,274]" page="95" pageId="21" pageNumber="22" year="2013">
<bibRefCitation id="F54F4B5DFFC6933B7B739BF6FB3CFE9A" author="Valchev, B. &amp; Stojanova, V. &amp; Juranov, S." box="[1069,1272,250,274]" pageId="21" pageNumber="22" pagination="81 - 110" refId="ref38624" refString="Valchev, B., Stojanova, V. &amp; Juranov, S. (2013) Paleogene hyaline benthic foraminifera (LAGENINA and ROTALIINA) from the Republic of Macedonia. Review of the Bulgarian Geological Society, 74, 81 - 110." type="journal article" year="2013">
Valchev
<emphasis id="A3AAEABEFFC6933B7BDA9BF7FB78FE9A" box="[1156,1212,250,274]" italics="true" pageId="21" pageNumber="22">et al.,</emphasis>
2013
</bibRefCitation>
, p. 95
</treatmentCitation>
, pl. 1, fig. 16.
</treatmentCitationGroup>
</paragraph>
<paragraph id="916136ACFFC6933B7FC99A34FD7EFED8" blockId="21.[151,1437,250,336]" box="[151,698,311,336]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="B1CE1182FFC6933B7FC99A34FD7EFED8" box="[151,698,311,336]" pageId="21" pageNumber="22">
<treatmentCitation id="107F10BDFFC6933B7FC99A34FD85FEC7" author="Jones, R. W." box="[151,577,311,336]" page="59" pageId="21" pageNumber="22" year="1994">
<taxonomicName id="56DE4D2FFFC6933B7FC99A34FD85FEC7" ID-CoL="NC2X" authority="Jones, 1994, p. 59" authorityName="Jones" authorityPageNumber="59" authorityYear="1994" box="[151,577,311,336]" class="Globothalamea" family="Bolivinitidae" genus="Brizalina" kingdom="Chromista" order="Rotaliida" pageId="21" pageNumber="22" phylum="Foraminifera" rank="species" species="subreticulata">
<emphasis id="A3AAEABEFFC6933B7FC99A34FE45FED8" box="[151,385,312,336]" italics="true" pageId="21" pageNumber="22">Brizalina subreticulata</emphasis>
<bibRefCitation id="F54F4B5DFFC6933B7ED99A34FDC5FED8" author="Jones, R. W." box="[391,513,312,336]" pageId="21" pageNumber="22" refId="ref33343" refString="Jones, R. W. (1994) The Challenger Foraminifera. Oxford University Press, Oxford, 149 pp." type="book" year="1994">Jones, 1994</bibRefCitation>
, p. 59
</taxonomicName>
</treatmentCitation>
, fig. 3031.
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</paragraph>
</subSubSection>
<subSubSection id="D9C46527FFC6933B7FC99A71FE6CFDF9" pageId="21" pageNumber="22" type="description">
<paragraph id="916136ACFFC6933B7FC99A71FE6EFD8C" blockId="21.[151,1437,381,805]" pageId="21" pageNumber="22">
<emphasis id="A3AAEABEFFC6933B7FC99A71FEE5FE1F" bold="true" box="[151,289,381,407]" pageId="21" pageNumber="22">Description</emphasis>
: The test wall is calcareous and microperforate. The test is small, broad, biserial, triangular-ovoid in shape and elliptical in cross-section. The test is less than twice as long as it is wide. Irregular, reticulate, interwoven costae cover the entire test, giving a reticulate appearance to the test. The aperture is slit-like and situated at the base of the terminal chamber.
</paragraph>
<paragraph id="916136ACFFC6933B7F999901FE6CFDF9" blockId="21.[151,1437,381,805]" pageId="21" pageNumber="22">
<emphasis id="A3AAEABEFFC6933B7F999901FEAEFDA0" bold="true" box="[199,362,525,552]" pageId="21" pageNumber="22">Life strategy:</emphasis>
This species is infaunal (
<bibRefCitation id="F54F4B5DFFC6933B7DCC9902FCAFFDA0" author="Drinia, H. &amp; Antonarakou, A. &amp; Tsaparas, N. &amp; Dermitzakis, M. D." box="[658,875,526,552]" pageId="21" pageNumber="22" pagination="145 - 158" refId="ref31154" refString="Drinia, H., Antonarakou, A., Tsaparas, N. &amp; Dermitzakis, M. D. (2007) Foraminiferal stratigraphy and palaeoecological implications in turbidite-like deposits from the Early Tortonian (Late Miocene) of Greece. Journal of Micropalaeontology, 26, 145 - 158. https: // doi. org / 10.1144 / jm. 26.2.145" type="journal article" year="2007">
Drinia
<emphasis id="A3AAEABEFFC6933B7DBA9903FCE2FDA0" box="[740,806,526,552]" italics="true" pageId="21" pageNumber="22">et al.,</emphasis>
2007
</bibRefCitation>
), unattached and prefers muddy sediments under dysoxic conditions (
<bibRefCitation id="F54F4B5DFFC6933B7E26993EFDC3FDC4" author="Kaiho, K." box="[376,519,562,588]" pageId="21" pageNumber="22" pagination="719 - 722" refId="ref33365" refString="Kaiho, K. (1994) Benthic foraminiferal dissolved-oxygen index and dissolved-oxygen levels in the modern ocean. Geology, 22 (8), 719 - 722. https: // doi. org / 10.1130 / 0091 - 7613 (1994) 022 &lt;0719: BFDOIA&gt; 2.3. CO; 2" type="journal article" year="1994">Kaiho, 1994</bibRefCitation>
). The bathymetric distribution of this species is recorded to be from the inner shelf to slope (
<bibRefCitation id="F54F4B5DFFC6933B7FA0995AFE59FDF9" author="Murray, J. W." box="[254,413,598,625]" pageId="21" pageNumber="22" refId="ref35617" refString="Murray, J. W. (2006) Ecology and Applications of Benthic Foraminifera. Cambridge University Press, Cambridge, 426 pp. https: // doi. org / 10.1017 / CBO 9780511535529" type="book" year="2006">Murray, 2006</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="D9C46527FFC6933B7F999976FCA6FCAD" pageId="21" pageNumber="22" type="discussion">
<paragraph id="916136ACFFC6933B7F999976FC38FD30" blockId="21.[151,1437,381,805]" pageId="21" pageNumber="22">
<emphasis id="A3AAEABEFFC6933B7F999976FEF9FD1C" bold="true" box="[199,317,634,660]" pageId="21" pageNumber="22">Remarks:</emphasis>
The relative abundance is generally low, comprising trace components (&lt;1%) in the samples of core 2670. The tests are small, measuring
<quantity id="56269B49FFC6933B7D669992FD4AFD30" box="[568,654,670,696]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="21" pageNumber="22" unit="mm" value="0.2">0.2 mm</quantity>
in width and
<quantity id="56269B49FFC6933B7C799992FC4FFD30" box="[807,907,670,697]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="21" pageNumber="22" unit="mm" value="0.35">0.35 mm</quantity>
in length.
</paragraph>
<paragraph id="916136ACFFC6933B7F9999CDFAF2FD54" blockId="21.[151,1437,381,805]" box="[199,1334,705,732]" pageId="21" pageNumber="22">
<emphasis id="A3AAEABEFFC6933B7F9999CDFDCBFD54" bold="true" box="[199,527,705,732]" pageId="21" pageNumber="22">Global stratigraphic range:</emphasis>
This species occurs from the Oligocene to Recent (
<bibRefCitation id="F54F4B5DFFC6933B7B0F99CEFAEEFD54" author="Fenero, R. &amp; Thomas, E. &amp; Alegret, L. &amp; Molina, E." box="[1105,1322,706,732]" pageId="21" pageNumber="22" pagination="286 - 304" refId="ref31452" refString="Fenero, R., Thomas, E., Alegret, L. &amp; Molina, E. (2012) Oligocene benthic foraminifera from the Fuente Caldera section (Spain, western Tethys): taxonomy and paleoenvironmental inferences. Journal of Foraminiferal Research, 42 (4), 286 - 304. https: // doi. org / 10.2113 / gsjfr. 42.4.286" type="journal article" year="2012">
Fenero
<emphasis id="A3AAEABEFFC6933B7BF999CFFB1DFD54" box="[1191,1241,706,732]" italics="true" pageId="21" pageNumber="22">et al</emphasis>
., 2012
</bibRefCitation>
).
</paragraph>
<paragraph id="916136ACFFC6933B7F9999E9FCA6FCAD" blockId="21.[151,1437,381,805]" pageId="21" pageNumber="22">
<emphasis id="A3AAEABEFFC6933B7F9999E9FE7BFC88" bold="true" box="[199,447,741,768]" pageId="21" pageNumber="22">Regional occurrence</emphasis>
:
<taxonomicName id="56DE4D2FFFC6933B7E8F99EAFD62FC88" authorityName="Hantken" authorityYear="1875" box="[465,678,742,768]" class="Globothalamea" family="Bolivinitidae" genus="Bolivina" kingdom="Chromista" order="Rotaliida" pageId="21" pageNumber="22" phylum="Foraminifera" rank="species" species="reticulata">
<emphasis id="A3AAEABEFFC6933B7E8F99EAFD62FC88" box="[465,678,742,768]" italics="true" pageId="21" pageNumber="22">Bolivina reticulata</emphasis>
</taxonomicName>
is recorded in middle Miocene sediments on the Namibian outer continental shelf, south of the
<collectingRegion id="531AF84EFFC6933B7EB49806FD42FCAC" box="[490,646,778,804]" isEnumeration="true" pageId="21" pageNumber="22">
<collectingRegion id="531AF84EFFC6933B7EB49806FD85FCAC" box="[490,577,778,804]" country="Namibia" name="Kunene" pageId="21" pageNumber="22">Kunene</collectingRegion>
<collectingRegion id="531AF84EFFC6933B7D169806FD42FCAC" box="[584,646,778,804]" country="Nigeria" name="Rivers" pageId="21" pageNumber="22">River</collectingRegion>
</collectingRegion>
mouth (this study).
</paragraph>
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