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<mods:title id="F809E9A2631FC828F7A54B1CF7DF2591">Taxonomy of Middle Miocene foraminifera from the northern Namibian continental shelf</mods:title>
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<mods:namePart id="CD14DB48D2A5A4B97E4219CA90A537BF">Bergh, Eugene W.</mods:namePart>
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<mods:affiliation id="B7E355233FEC42D13D4B954EBA9715D7">Marine Research Institute and Department of Geological Sciences, University of Cape Town, Private Bag X 3, Rondebosch, 7701, South Africa. &amp; Department of Environmental and Geographical Sciences, University of Cape Town, Private Bag X 3, Rondebosch, 7701, South Africa; &amp; Invertebrate palaeontology and Geology, Iziko South African Museum, P. O. Box 61, Cape Town 8000, South Africa. john. compton @ uct. ac. za; https: // orcid. org / 0000 - 0002 - 0765 - 4141</mods:affiliation>
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<mods:namePart id="CFBED9D4C7E0C5DD2609E116E49CAF55">Compton, John S.</mods:namePart>
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<subSubSection id="D9C46527FFC493397FC99FD1FED1FA94" pageId="23" pageNumber="24" type="nomenclature">
<paragraph id="916136ACFFC493397FC99FD1FD0AFB7F" blockId="23.[151,718,1245,1308]" box="[151,718,1245,1272]" pageId="23" pageNumber="24">
<heading id="CA2981C0FFC493397FC99FD1FD0AFB7F" bold="true" box="[151,718,1245,1272]" fontSize="11" level="1" pageId="23" pageNumber="24" reason="1">
<taxonomicName id="56DE4D2FFFC493397FC99FD1FD0AFB7F" ID-CoL="835P3" authority="(Thalmann, 1937)" baseAuthorityName="Thalmann" baseAuthorityYear="1937" box="[151,718,1245,1272]" class="Nodosariata" family="Glandulonodosariidae" genus="Orthomorphina" kingdom="Chromista" order="Nodosariida" pageId="23" pageNumber="24" phylum="Foraminifera" rank="species" species="jedlitschkai">
<emphasis id="A3AAEABEFFC493397FC99FD1FD0AFB7F" bold="true" box="[151,718,1245,1272]" pageId="23" pageNumber="24">
<emphasis id="A3AAEABEFFC493397FC99FD1FE2CFB70" bold="true" box="[151,488,1245,1272]" italics="true" pageId="23" pageNumber="24">Orthomorphina jedlitschkai</emphasis>
(
<bibRefCitation id="F54F4B5DFFC493397EA99FD1FD02FB7F" author="Thalmann, H. E." box="[503,710,1245,1271]" pageId="23" pageNumber="24" pagination="337 - 356" refId="ref38224" refString="Thalmann, H. E. (1937) Mitteilungen uber Foraminiferen III. Eclogae Geologicae Helvetiae, 30, 337 - 356." type="journal article" year="1937">Thalmann, 1937</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="916136ACFFC493397FC99E0EFED1FA94" blockId="23.[151,718,1245,1308]" box="[151,277,1281,1308]" pageId="23" pageNumber="24">
<heading id="CA2981C0FFC493397FC99E0EFED1FA94" bold="true" box="[151,277,1281,1308]" fontSize="11" level="1" pageId="23" pageNumber="24" reason="1">
<emphasis id="A3AAEABEFFC493397FC99E0EFED1FA94" bold="true" box="[151,277,1281,1308]" pageId="23" pageNumber="24">Pl. 5, fig. 2</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D9C46527FFC493397FC99E4DFD5AFA1E" pageId="23" pageNumber="24" type="reference_group">
<paragraph id="916136ACFFC493397FC99E4DFD67FAD1" blockId="23.[151,1436,1345,1430]" box="[151,675,1345,1369]" pageId="23" pageNumber="24">
<treatmentCitationGroup id="B1CE1182FFC493397FC99E4DFD67FAD1" box="[151,675,1345,1369]" pageId="23" pageNumber="24">
<treatmentCitation id="107F10BDFFC493397FC99E4DFD5BFAD1" author="Thalmann, H. E." box="[151,671,1345,1369]" page="341" pageId="23" pageNumber="24" year="1937">
<taxonomicName id="56DE4D2FFFC493397FC99E4DFD5BFAD1" authority="Thalmann, 1937, p. 341" authorityName="Thalmann" authorityPageNumber="341" authorityYear="1937" box="[151,671,1345,1369]" class="Nodosariata" family="Stilostomellidae" genus="Nodogenerina" kingdom="Chromista" order="Nodosariida" pageId="23" pageNumber="24" phylum="Foraminifera" rank="species" species="jedlitschkai">
<emphasis id="A3AAEABEFFC493397FC99E4DFE67FAD1" box="[151,419,1345,1369]" italics="true" pageId="23" pageNumber="24">Nodogenerina jedlitschkai</emphasis>
<bibRefCitation id="F54F4B5DFFC493397EF49E4DFD96FAD1" author="Thalmann, H. E." box="[426,594,1345,1369]" pageId="23" pageNumber="24" pagination="337 - 356" refId="ref38224" refString="Thalmann, H. E. (1937) Mitteilungen uber Foraminiferen III. Eclogae Geologicae Helvetiae, 30, 337 - 356." type="journal article" year="1937">Thalmann, 1937</bibRefCitation>
, p. 341
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="916136ACFFC493397FC99E6CFD5AFA1E" blockId="23.[151,1436,1345,1430]" pageId="23" pageNumber="24">
<taxonomicName id="56DE4D2FFFC493397FC99E6CFDFFFAFF" ID-CoL="835P3" authority="Barker, 1960" authorityName="Barker" authorityYear="1960" box="[151,571,1375,1400]" class="Nodosariata" family="Glandulonodosariidae" genus="Orthomorphina" kingdom="Chromista" order="Nodosariida" pageId="23" pageNumber="24" phylum="Foraminifera" rank="species" species="jedlitschkai">
<emphasis id="A3AAEABEFFC493397FC99E6CFE75FAF0" box="[151,433,1376,1400]" italics="true" pageId="23" pageNumber="24">Orthomorphina jedlitschkai</emphasis>
<bibRefCitation id="F54F4B5DFFC493397EE89E6CFDFFFAFF" author="Barker, R. W." box="[438,571,1375,1400]" pageId="23" pageNumber="24" pagination="1 - 238" refId="ref27766" refString="Barker, R. W. (1960) Taxonomic notes on the species figured by H. B. Brady in his report on the Foraminifera dredged by H. M. S. Challenger during the years 1873 - 1876. Society of Economic Palaeontologists and Mineralogists, Special Publication, 9, 1 - 238." type="journal article" year="1960">Barker, 1960</bibRefCitation>
</taxonomicName>
, pl. 62, fig. 12;
<bibRefCitation id="F54F4B5DFFC493397DBD9E6CFC99FAF0" author="Jones, R. W." box="[739,861,1376,1400]" pageId="23" pageNumber="24" refId="ref33343" refString="Jones, R. W. (1994) The Challenger Foraminifera. Oxford University Press, Oxford, 149 pp." type="book" year="1994">Jones, 1994</bibRefCitation>
, pl. 62, fig. 12, suppl. pl. 1, fig. 16, suppl. pl. 2, fig. 67;
<bibRefCitation id="F54F4B5DFFC493397F999E72FE5BFA1E" author="Hayward, B. W. &amp; Kawagata, S. &amp; Sabaa, A. T. &amp; Grenfell, H. R. &amp; van Kerckhoven, L. &amp; Johnson, K. &amp; Thomas, E." box="[199,415,1406,1430]" pageId="23" pageNumber="24" pagination="1 - 408" refId="ref32644" refString="Hayward, B. W., Kawagata, S., Sabaa, A. T., Grenfell, H. R., van Kerckhoven, L., Johnson, K. &amp; Thomas, E. (2012) The last global extinction (Mid-Pleistocene) of deep-sea benthic foraminifera (Chrysalogoniidae, Ellipsoidinidae, Glandulonodosariidae, Plectofrondiculariidae, Pleursostomellidae, Stilostomellidae), their Late Cretaceous-Cenozoic history and taxonomy. Cushman Foundation for Foraminiferal Research Special Publication, 43, 1 - 408." type="journal article" year="2012">
Hayward
<emphasis id="A3AAEABEFFC493397E749E73FE93FA1E" box="[298,343,1406,1430]" italics="true" pageId="23" pageNumber="24">et al</emphasis>
., 2012
</bibRefCitation>
, p. 136, pl. 8, fig. 2427.
</paragraph>
</subSubSection>
<subSubSection id="D9C46527FFC493397FC99EC8FF02F9C2" pageId="23" pageNumber="24" type="description">
<paragraph id="916136ACFFC493397FC99EC8FF02F9C2" blockId="23.[151,1437,1476,2043]" pageId="23" pageNumber="24">
<emphasis id="A3AAEABEFFC493397FC99EC8FEE5FA56" bold="true" box="[151,289,1476,1502]" pageId="23" pageNumber="24">Description</emphasis>
: The test wall is calcareous and generally smooth. The test is elongate, uniserial, with globular chambers and circular in cross-section. The chambers are of different diameters and separated from each other by deep incised sutures. The penultimate chamber is larger than the terminal chamber. The aperture is surrounded by a round thick rim.
</paragraph>
</subSubSection>
<subSubSection id="D9C46527FFC493397F999D59FA81F873" pageId="23" pageNumber="24" type="discussion">
<paragraph id="916136ACFFC493397F999D59FE9AF93E" blockId="23.[151,1437,1476,2043]" pageId="23" pageNumber="24">
<emphasis id="A3AAEABEFFC493397F999D59FEF9F9E7" bold="true" box="[199,317,1621,1647]" pageId="23" pageNumber="24">Remarks:</emphasis>
Few tests of this species were recorded in this study, forming a trace component (&lt;1%) in core 2670. The tests are moderate in size. The broadest chamber measures
<quantity id="56269B49FFC493397C019D75FC70F91B" box="[863,948,1657,1683]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="23" pageNumber="24" unit="mm" value="0.3">0.3 mm</quantity>
in cross section and the test measures up to
<quantity id="56269B49FFC493397FC99D91FF29F93F" box="[151,237,1693,1719]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="23" pageNumber="24" unit="mm" value="0.8">0.8 mm</quantity>
in length.
</paragraph>
<paragraph id="916136ACFFC493397F999DCDFB54F8AA" blockId="23.[151,1437,1476,2043]" pageId="23" pageNumber="24">
<bibRefCitation id="F54F4B5DFFC493397F999DCDFE93F953" author="Jones, R. W." box="[199,343,1729,1755]" pageId="23" pageNumber="24" refId="ref33343" refString="Jones, R. W. (1994) The Challenger Foraminifera. Oxford University Press, Oxford, 149 pp." type="book" year="1994">Jones (1994)</bibRefCitation>
illustrated the similarity in test morphology between
<taxonomicName id="56DE4D2FFFC493397CEF9DCDFB91F952" authorityName="Barker" authorityYear="1960" box="[945,1109,1728,1755]" class="Nodosariata" family="Glandulonodosariidae" genus="Orthomorphina" kingdom="Chromista" order="Nodosariida" pageId="23" pageNumber="24" phylum="Foraminifera" rank="species" species="jedlitschkai">
<emphasis id="A3AAEABEFFC493397CEF9DCDFB91F952" box="[945,1109,1728,1755]" italics="true" pageId="23" pageNumber="24">O. jedlitschkai</emphasis>
</taxonomicName>
and
<taxonomicName id="56DE4D2FFFC493397BD29DCCFA59F953" box="[1164,1437,1728,1755]" class="Nodosariata" family="Glandulonodosariidae" genus="Glandulonodosaria" kingdom="Chromista" order="Nodosariida" pageId="23" pageNumber="24" phylum="Foraminifera" rank="species" species="undetermined">
<emphasis id="A3AAEABEFFC493397BD29DCCFAA3F952" box="[1164,1383,1728,1754]" italics="true" pageId="23" pageNumber="24">Glandulonodosaria</emphasis>
spp.
</taxonomicName>
Based on the shape of the aperture (slightly protruding and rounded) and resemblance to the test morphology displayed in
<bibRefCitation id="F54F4B5DFFC493397E789C05FE7CF8AB" author="Jones, R. W." box="[294,440,1801,1827]" pageId="23" pageNumber="24" refId="ref33343" refString="Jones, R. W. (1994) The Challenger Foraminifera. Oxford University Press, Oxford, 149 pp." type="book" year="1994">Jones (1994)</bibRefCitation>
, this specimen has been assigned to
<emphasis id="A3AAEABEFFC493397C0F9C04FB54F8AA" box="[849,1168,1800,1826]" italics="true" pageId="23" pageNumber="24">
<taxonomicName id="56DE4D2FFFC493397C0F9C04FB49F8AA" authorityName="Barker" authorityYear="1960" box="[849,1165,1800,1826]" class="Nodosariata" family="Glandulonodosariidae" genus="Orthomorphina" kingdom="Chromista" order="Nodosariida" pageId="23" pageNumber="24" phylum="Foraminifera" rank="species" species="jedlitschkai">Orthomorphina jedlitschkai</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="916136ACFFC493397F999C20FDE0F8E3" blockId="23.[151,1437,1476,2043]" pageId="23" pageNumber="24">
<emphasis id="A3AAEABEFFC493397F999C20FEA3F8CF" bold="true" box="[199,359,1836,1863]" pageId="23" pageNumber="24">Life strategy:</emphasis>
Species of the genus
<taxonomicName id="56DE4D2FFFC493397D0C9C20FCC6F8CE" authorityName="Stainforth" authorityYear="1952" box="[594,770,1836,1862]" class="Nodosariata" family="Glandulonodosariidae" genus="Orthomorphina" kingdom="Chromista" order="Nodosariida" pageId="23" pageNumber="24" phylum="Foraminifera" rank="genus">
<emphasis id="A3AAEABEFFC493397D0C9C20FCC6F8CE" box="[594,770,1836,1862]" italics="true" pageId="23" pageNumber="24">Orthomorphina</emphasis>
</taxonomicName>
are recorded to be infaunal under suboxic conditions (
<bibRefCitation id="F54F4B5DFFC493397A089C20FED1F8E3" author="Pezelj, D &amp; Mandic, O. &amp; Coric, S." pageId="23" pageNumber="24" pagination="81 - 100" refId="ref36415" refString="Pezelj, D, Mandic, O. &amp; Coric, S. (2013) Paleoenvironmental dynamics in the southern Pannonian Basin during initial Middle Miocene marine flooding. Geologica Carpathica, 64, 81 - 100." type="journal article" year="2013">
Pezelj
<emphasis id="A3AAEABEFFC493397FC99C5DFF12F8E2" box="[151,214,1872,1898]" italics="true" pageId="23" pageNumber="24">et al.,</emphasis>
2013
</bibRefCitation>
and references therein).
</paragraph>
<paragraph id="916136ACFFC493397F999C78FD84F83B" blockId="23.[151,1437,1476,2043]" pageId="23" pageNumber="24">
<emphasis id="A3AAEABEFFC493397F999C78FDC3F807" bold="true" box="[199,519,1908,1935]" pageId="23" pageNumber="24">Global stratigraphic range</emphasis>
:
<taxonomicName id="56DE4D2FFFC493397D489C78FC8BF806" authorityName="Barker" authorityYear="1960" box="[534,847,1908,1934]" class="Nodosariata" family="Glandulonodosariidae" genus="Orthomorphina" kingdom="Chromista" order="Nodosariida" pageId="23" pageNumber="24" phylum="Foraminifera" rank="species" species="jedlitschkai">
<emphasis id="A3AAEABEFFC493397D489C78FC8BF806" box="[534,847,1908,1934]" italics="true" pageId="23" pageNumber="24">Orthomorphina jedlitschkai</emphasis>
</taxonomicName>
is an extinct species, occurring from the Cretaceous to Pleistocene (
<bibRefCitation id="F54F4B5DFFC493397E1C9C94FDF0F83B" author="Hayward, B. W. &amp; Kawagata, S. &amp; Sabaa, A. T. &amp; Grenfell, H. R. &amp; van Kerckhoven, L. &amp; Johnson, K. &amp; Thomas, E." box="[322,564,1944,1971]" pageId="23" pageNumber="24" pagination="1 - 408" refId="ref32644" refString="Hayward, B. W., Kawagata, S., Sabaa, A. T., Grenfell, H. R., van Kerckhoven, L., Johnson, K. &amp; Thomas, E. (2012) The last global extinction (Mid-Pleistocene) of deep-sea benthic foraminifera (Chrysalogoniidae, Ellipsoidinidae, Glandulonodosariidae, Plectofrondiculariidae, Pleursostomellidae, Stilostomellidae), their Late Cretaceous-Cenozoic history and taxonomy. Cushman Foundation for Foraminiferal Research Special Publication, 43, 1 - 408." type="journal article" year="2012">
Hayward
<emphasis id="A3AAEABEFFC493397EEF9C95FE27F83A" box="[433,483,1944,1970]" italics="true" pageId="23" pageNumber="24">et al</emphasis>
., 2012
</bibRefCitation>
).
</paragraph>
<paragraph id="916136ACFFC493397F999CB0FA81F873" blockId="23.[151,1437,1476,2043]" pageId="23" pageNumber="24">
<emphasis id="A3AAEABEFFC493397F999CB0FE04F85F" bold="true" box="[199,448,1980,2007]" pageId="23" pageNumber="24">Regional occurrence</emphasis>
: This is the first documentation of this species to occur in the region and in middle Miocene sediments on the Namibian outer continental shelf, south of the
<collectingRegion id="531AF84EFFC493397C929CEDFBADF872" box="[972,1129,2016,2043]" isEnumeration="true" pageId="23" pageNumber="24">
<collectingRegion id="531AF84EFFC493397C929CEDFBE7F873" box="[972,1059,2017,2043]" country="Namibia" name="Kunene" pageId="23" pageNumber="24">Kunene</collectingRegion>
<collectingRegion id="531AF84EFFC493397B759CECFBADF872" box="[1067,1129,2016,2042]" country="Nigeria" name="Rivers" pageId="23" pageNumber="24">River</collectingRegion>
</collectingRegion>
mouth (this study).
</paragraph>
</subSubSection>
</treatment>
</document>