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<document id="230F1F375423167E1FB2C5F460BDD1C9" ID-CLB-Dataset="2636" ID-DOI="10.11646/zootaxa.5091.1.1" ID-GBIF-Dataset="51b42715-ddb1-4b4a-aad4-8be0f600a287" ID-ISSN="1175-5326" ID-Zenodo-Dep="5840434" ID-ZooBank="ABC8AF70-F691-4D07-8F20-70934642C8BC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1641986550580" checkinUser="plazi" docAuthor="Bergh, Eugene W. &amp; Compton, John S." docDate="2022" docId="197787BAFFC1933C7FC99887FE11F840" docLanguage="en" docName="zootaxa.5091.1.1.pdf" docOrigin="Zootaxa 5091 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Marginulina obesa Cushman 1923" docType="treatment" docVersion="6" lastPageNumber="19" masterDocId="E54EFFC2FFD3932E7F5E9B0CFFC4FF88" masterDocTitle="Taxonomy of Middle Miocene foraminifera from the northern Namibian continental shelf" masterLastPageNumber="55" masterPageNumber="1" pageNumber="19" updateTime="1699301522325" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="34E02FED0214F52A47D458963D241BC3">Taxonomy of Middle Miocene foraminifera from the northern Namibian continental shelf</mods:title>
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<mods:namePart id="F6198DDA4BDBCAA5A8D15D79096DE973">Bergh, Eugene W.</mods:namePart>
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<mods:affiliation id="3315695943F96ABADD2F2A4A3B92454A">Marine Research Institute and Department of Geological Sciences, University of Cape Town, Private Bag X 3, Rondebosch, 7701, South Africa. &amp; Department of Environmental and Geographical Sciences, University of Cape Town, Private Bag X 3, Rondebosch, 7701, South Africa; &amp; Invertebrate palaeontology and Geology, Iziko South African Museum, P. O. Box 61, Cape Town 8000, South Africa. john. compton @ uct. ac. za; https: // orcid. org / 0000 - 0002 - 0765 - 4141</mods:affiliation>
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<paragraph id="916136ACFFC1933C7FC99887FD91FC2D" blockId="18.[151,597,907,969]" box="[151,597,907,934]" pageId="18" pageNumber="19">
<heading id="CA2981C0FFC1933C7FC99887FD91FC2D" bold="true" box="[151,597,907,934]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
<taxonomicName id="56DE4D2FFFC1933C7FC99887FD91FC2D" ID-CoL="3Y5ZB" authority="Cushman, 1923 a" authorityName="Cushman" authorityYear="1923" box="[151,597,907,934]" class="Nodosariata" family="Vaginulinidae" genus="Marginulina" kingdom="Chromista" order="Vaginulinida" pageId="18" pageNumber="19" phylum="Foraminifera" rank="species" species="obesa">
<emphasis id="A3AAEABEFFC1933C7FC99887FD91FC2D" bold="true" box="[151,597,907,934]" pageId="18" pageNumber="19">
<emphasis id="A3AAEABEFFC1933C7FC99887FEB9FC2E" bold="true" box="[151,381,907,934]" italics="true" pageId="18" pageNumber="19">Marginulina obesa</emphasis>
<bibRefCitation id="F54F4B5DFFC1933C7EDB9887FD91FC2D" author="Cushman, J. A." box="[389,597,907,933]" pageId="18" pageNumber="19" pagination="1 - 228" refId="ref29789" refString="Cushman, J. A. (1923 a) The foraminifera of the Atlantic Ocean: part 4 - Lagenidae. Bulletin of the United States National Museum, 104 (4), 1 - 228. https: // doi. org / 10.5479 / si. 03629236.104.3" type="journal article" year="1923">Cushman, 1923a</bibRefCitation>
</emphasis>
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</paragraph>
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<heading id="CA2981C0FFC1933C7FC998A3FE0EFC41" bold="true" box="[151,458,943,969]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
<emphasis id="A3AAEABEFFC1933C7FC998A3FE0EFC41" bold="true" box="[151,458,943,969]" pageId="18" pageNumber="19">Pl. 3, figs. 89; Pl. 4, fig. 1</emphasis>
</heading>
</paragraph>
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<subSubSection id="D9C46527FFC1933C7FC998F8FD9CFBEF" pageId="18" pageNumber="19" type="reference_group">
<paragraph id="916136ACFFC1933C7FC998F8FBF5FB84" blockId="18.[151,1437,1011,1128]" box="[151,1073,1011,1036]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="B1CE1182FFC1933C7FC998F8FBF5FB84" box="[151,1073,1011,1036]" pageId="18" pageNumber="19">
<treatmentCitation id="107F10BDFFC1933C7FC998F8FDF4FB83" author="Brady, H. B." box="[151,560,1011,1036]" page="527" pageId="18" pageNumber="19" year="1884">
<taxonomicName id="56DE4D2FFFC1933C7FC998F8FDF4FB83" ID-CoL="7F7WR" authority="Brady, 1884, p. 527" authorityName="Brady" authorityPageNumber="527" authorityYear="1884" box="[151,560,1011,1036]" class="Nodosariata" family="Vaginulinidae" genus="Marginulina" kingdom="Chromista" order="Vaginulinida" pageId="18" pageNumber="19" phylum="Foraminifera" rank="species" species="glabra">
<emphasis id="A3AAEABEFFC1933C7FC998F8FEA5FB84" box="[151,353,1012,1036]" italics="true" pageId="18" pageNumber="19">Marginulina glabra</emphasis>
<bibRefCitation id="F54F4B5DFFC1933C7E3998F8FE20FB84" author="Brady, H. B." box="[359,484,1012,1036]" pageId="18" pageNumber="19" pagination="1 - 814" refId="ref28770" refString="Brady, H. B. (1884) Report of the foraminifera dredged by H. M. S. Challenger during the years 1873 - 1876. Zoology, 9, 1 - 814." type="journal article" year="1884">Brady, 1884</bibRefCitation>
, p. 527
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, pl. 65, figs. 57;
<treatmentCitation id="107F10BDFFC1933C7DB998F8FC60FB84" author="Flint, J. M." box="[743,932,1012,1036]" page="133" pageId="18" pageNumber="19" year="1899">
<bibRefCitation id="F54F4B5DFFC1933C7DB998F8FC9DFB84" author="Flint, J. M." box="[743,857,1012,1036]" pageId="18" pageNumber="19" pagination="249 - 349" refId="ref31613" refString="Flint, J. M. (1899) Recent foraminifera. A descriptive catalogue of specimens dredged by the US Fish Commission Steamer Albatross. Report of the United States National Museum for 1897, 1, 249 - 349. https: // doi. org / 10.5962 / bhl. title. 2153" type="journal article" year="1899">Flint, 1899</bibRefCitation>
, p. 133
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, pl. 60, fig. 1.
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</paragraph>
<paragraph id="916136ACFFC1933C7FC99F1EFC9CFBA2" blockId="18.[151,1437,1011,1128]" box="[151,856,1042,1066]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="B1CE1182FFC1933C7FC99F1EFC9CFBA2" box="[151,856,1042,1066]" pageId="18" pageNumber="19">
<treatmentCitation id="107F10BDFFC1933C7FC99F1EFD08FBA2" author="Cushman, J. A." box="[151,716,1042,1066]" page="128" pageId="18" pageNumber="19" year="1923">
<taxonomicName id="56DE4D2FFFC1933C7FC99F1EFD45FBA2" ID-CoL="3Y5ZB" authority="Cushman, 1923 a" authorityName="Cushman" authorityYear="1923" box="[151,641,1042,1066]" class="Nodosariata" family="Vaginulinidae" genus="Marginulina" kingdom="Chromista" order="Vaginulinida" pageId="18" pageNumber="19" phylum="Foraminifera" rank="variety" species="glabra" variety="obesa">
<emphasis id="A3AAEABEFFC1933C7FC99F1EFEA5FBA2" box="[151,353,1042,1066]" italics="true" pageId="18" pageNumber="19">Marginulina glabra</emphasis>
var.
<emphasis id="A3AAEABEFFC1933C7ECC9F1EFE09FBA2" box="[402,461,1042,1066]" italics="true" pageId="18" pageNumber="19">obesa</emphasis>
<bibRefCitation id="F54F4B5DFFC1933C7E8A9F1EFD45FBA2" author="Cushman, J. A." box="[468,641,1042,1066]" pageId="18" pageNumber="19" pagination="1 - 228" refId="ref29789" refString="Cushman, J. A. (1923 a) The foraminifera of the Atlantic Ocean: part 4 - Lagenidae. Bulletin of the United States National Museum, 104 (4), 1 - 228. https: // doi. org / 10.5479 / si. 03629236.104.3" type="journal article" year="1923">Cushman, 1923a</bibRefCitation>
</taxonomicName>
, p. 128
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, pl. 37, fig. 1.
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</paragraph>
<paragraph id="916136ACFFC1933C7FC99F3DFD9CFBEF" blockId="18.[151,1437,1011,1128]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="B1CE1182FFC1933C7FC99F3DFD9CFBEF" pageId="18" pageNumber="19">
<taxonomicName id="56DE4D2FFFC1933C7FC99F3DFE26FBC1" ID-CoL="3Y5ZB" authority="Barker, 1960" authorityName="Barker" authorityYear="1960" box="[151,482,1073,1097]" class="Nodosariata" family="Vaginulinidae" genus="Marginulina" kingdom="Chromista" order="Vaginulinida" pageId="18" pageNumber="19" phylum="Foraminifera" rank="species" species="obesa">
<emphasis id="A3AAEABEFFC1933C7FC99F3DFE93FBC1" box="[151,343,1073,1097]" italics="true" pageId="18" pageNumber="19">Marginulina obesa</emphasis>
<bibRefCitation id="F54F4B5DFFC1933C7E039F3DFE26FBC1" author="Barker, R. W." box="[349,482,1073,1097]" pageId="18" pageNumber="19" pagination="1 - 238" refId="ref27766" refString="Barker, R. W. (1960) Taxonomic notes on the species figured by H. B. Brady in his report on the Foraminifera dredged by H. M. S. Challenger during the years 1873 - 1876. Society of Economic Palaeontologists and Mineralogists, Special Publication, 9, 1 - 238." type="journal article" year="1960">Barker, 1960</bibRefCitation>
</taxonomicName>
, pl. 65, figs. 56;
<treatmentCitation id="107F10BDFFC1933C7DC89F3DFC0FFBC1" author="LeRoy, D. O. &amp; Levinson, S. A." box="[662,971,1073,1097]" page="8" pageId="18" pageNumber="19" year="1974">
<bibRefCitation id="F54F4B5DFFC1933C7DC89F3DFC5EFBC1" author="LeRoy, D. O. &amp; Levinson, S. A." box="[662,922,1073,1097]" pageId="18" pageNumber="19" pagination="1 - 37" refId="ref34352" refString="LeRoy, D. O. &amp; Levinson, S. A. (1974) A deep-water Pleistocene microfossil assemblage from a well in the Northern Gulf of Mexico. Micropalaeontology, 20 (1), 1 - 37. https: // doi. org / 10.2307 / 1485098" type="journal article" year="1974">LeRoy &amp; Levinson, 1974</bibRefCitation>
, p. 8
</treatmentCitation>
, pl. 4, fig. 34;
<treatmentCitation id="107F10BDFFC1933C7B379F3DFAF3FBC1" author="Lowry, F. M. D." box="[1129,1335,1073,1097]" page="174" pageId="18" pageNumber="19" year="1987">
<bibRefCitation id="F54F4B5DFFC1933C7B379F3DFB28FBC1" author="Lowry, F. M. D." box="[1129,1260,1073,1097]" pageId="18" pageNumber="19" refId="ref34923" refString="Lowry, F. M. D. (1987) Foraminiferal thanatocoenoses from the continental shelf of southern Africa. Unpublished Ph. D. thesis, University College, London, 443 pp." type="book" year="1987">Lowry, 1987</bibRefCitation>
, p. 174
</treatmentCitation>
, pl. 9, fig. 12;
<treatmentCitation id="107F10BDFFC1933C7FB09F5CFE6CFBEF" author="Jones, R. W." box="[238,424,1104,1128]" page="77" pageId="18" pageNumber="19" year="1994">
<bibRefCitation id="F54F4B5DFFC1933C7FB09F5CFEACFBE0" author="Jones, R. W." box="[238,360,1104,1128]" pageId="18" pageNumber="19" refId="ref33343" refString="Jones, R. W. (1994) The Challenger Foraminifera. Oxford University Press, Oxford, 149 pp." type="book" year="1994">Jones, 1994</bibRefCitation>
, p. 77
</treatmentCitation>
, pl. 65, figs. 56.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="D9C46527FFC1933C7FC99F99FAD1FB70" pageId="18" pageNumber="19" type="description">
<paragraph id="916136ACFFC1933C7FC99F99FAD1FB70" blockId="18.[151,1437,1173,1992]" pageId="18" pageNumber="19">
<emphasis id="A3AAEABEFFC1933C7FC99F99FEE5FB27" bold="true" box="[151,289,1173,1199]" pageId="18" pageNumber="19">Description</emphasis>
: The test wall is calcareous and smooth. The test is elongate, circular in cross-section, initially curved, becoming rectilinear in the later stage. Three inflated chambers increase in size towards the terminal end. The chambers are separated by straight and slightly depressed sutures. The aperture is terminal and radiate.
</paragraph>
</subSubSection>
<subSubSection id="D9C46527FFC1933C7F999E0EFE11F840" pageId="18" pageNumber="19" type="discussion">
<paragraph id="916136ACFFC1933C7F999E0EFEF0FA00" blockId="18.[151,1437,1173,1992]" pageId="18" pageNumber="19">
<emphasis id="A3AAEABEFFC1933C7F999E0EFEF9FA94" bold="true" box="[199,317,1282,1308]" pageId="18" pageNumber="19">Remarks:</emphasis>
The relative abundance of this species is generally low, forming trace components (&lt;1%) in some of the samples of core 2670. The length of the tests in
<bibRefCitation id="F54F4B5DFFC1933C7D909E2AFC31FAC8" author="LeRoy, D. O. &amp; Levinson, S. A." box="[718,1013,1318,1344]" pageId="18" pageNumber="19" pagination="1 - 37" refId="ref34352" refString="LeRoy, D. O. &amp; Levinson, S. A. (1974) A deep-water Pleistocene microfossil assemblage from a well in the Northern Gulf of Mexico. Micropalaeontology, 20 (1), 1 - 37. https: // doi. org / 10.2307 / 1485098" type="journal article" year="1974">LeRoy &amp; Levinson (1974)</bibRefCitation>
is reported to be
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and a diameter of
<quantity id="56269B49FFC1933C7FE89E46FEDAFAEC" box="[182,286,1354,1380]" metricMagnitude="-4" metricUnit="m" metricValue="6.2" pageId="18" pageNumber="19" unit="mm" value="0.62">0.62 mm</quantity>
. The tests in this study are slightly smaller, measuring up to
<quantity id="56269B49FFC1933C7C959E46FBE6FAEC" box="[971,1058,1354,1380]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="18" pageNumber="19" unit="mm" value="0.4">0.4 mm</quantity>
in cross section diameter and
<quantity id="56269B49FFC1933C7A279E46FF07FA00" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="18" pageNumber="19" unit="mm" value="0.7">0.7 mm</quantity>
in length.
</paragraph>
<paragraph id="916136ACFFC1933C7F999E9EFE2CF944" blockId="18.[151,1437,1173,1992]" pageId="18" pageNumber="19">
<taxonomicName id="56DE4D2FFFC1933C7F999E9EFE67FA24" authorityName="Brady" authorityYear="1884" box="[199,419,1426,1452]" class="Nodosariata" family="Vaginulinidae" genus="Marginulina" kingdom="Chromista" order="Vaginulinida" pageId="18" pageNumber="19" phylum="Foraminifera" rank="species" species="glabra">
<emphasis id="A3AAEABEFFC1933C7F999E9EFE67FA24" box="[199,419,1426,1452]" italics="true" pageId="18" pageNumber="19">Marginulina glabra</emphasis>
</taxonomicName>
has been synonymised with
<taxonomicName id="56DE4D2FFFC1933C7D879E9EFC6FFA24" authorityName="Barker" authorityYear="1960" box="[729,939,1426,1452]" class="Nodosariata" family="Vaginulinidae" genus="Marginulina" kingdom="Chromista" order="Vaginulinida" pageId="18" pageNumber="19" phylum="Foraminifera" rank="species" species="obesa">
<emphasis id="A3AAEABEFFC1933C7D879E9EFC6FFA24" box="[729,939,1426,1452]" italics="true" pageId="18" pageNumber="19">Marginulina obesa</emphasis>
</taxonomicName>
(e.g.,
<bibRefCitation id="F54F4B5DFFC1933C7CB39E9EFB1DFA24" author="Holbourn, A. &amp; Henderson, A. S. &amp; MacLeod, N." box="[1005,1241,1426,1452]" pageId="18" pageNumber="19" refId="ref32986" refString="Holbourn, A., Henderson, A. S. &amp; MacLeod, N. (2013) Atlas of Benthic Foraminifera. Wiley-Blackwell, Oxford, 654 pp. https: // doi. org / 10.1002 / 9781118452493" type="book" year="2013">Holbourn et al., 2013</bibRefCitation>
), based on similar descriptions between
<bibRefCitation id="F54F4B5DFFC1933C7ECF9EBAFD8FFA59" author="Terquem, M. O." box="[401,587,1462,1489]" pageId="18" pageNumber="19" refId="ref38178" refString="Terquem, M. O. (1866) Cinquieme memoire sur les foraminiferes du lias des departements de la Moselle, de la Cote-d'Or et de l'Indre avec un apercu stratigraphique et petrologique des environs de Nohant. Lorette, Metz, 144 pp." type="book" year="1866">Terquem (1866)</bibRefCitation>
,
<bibRefCitation id="F54F4B5DFFC1933C7D059EBAFD31FA58" author="Brady, H. B." box="[603,757,1462,1488]" pageId="18" pageNumber="19" pagination="1 - 814" refId="ref28770" refString="Brady, H. B. (1884) Report of the foraminifera dredged by H. M. S. Challenger during the years 1873 - 1876. Zoology, 9, 1 - 814." type="journal article" year="1884">Brady (1884)</bibRefCitation>
and
<bibRefCitation id="F54F4B5DFFC1933C7C6D9EBAFBC5FA58" author="Cushman, J. A." box="[819,1025,1462,1488]" pageId="18" pageNumber="19" pagination="1 - 228" refId="ref29789" refString="Cushman, J. A. (1923 a) The foraminifera of the Atlantic Ocean: part 4 - Lagenidae. Bulletin of the United States National Museum, 104 (4), 1 - 228. https: // doi. org / 10.5479 / si. 03629236.104.3" type="journal article" year="1923">Cushman (1923a)</bibRefCitation>
.
<bibRefCitation id="F54F4B5DFFC1933C7B4E9EBAFB6EFA58" author="Brady, H. B." box="[1040,1194,1462,1488]" pageId="18" pageNumber="19" pagination="1 - 814" refId="ref28770" refString="Brady, H. B. (1884) Report of the foraminifera dredged by H. M. S. Challenger during the years 1873 - 1876. Zoology, 9, 1 - 814." type="journal article" year="1884">Brady (1884)</bibRefCitation>
does not distinguish between the variations within the species, where the length and stoutness is concerned. The test should remain inflated and the final chamber as large as at least one third of the test. Confusion might arise with
<taxonomicName id="56DE4D2FFFC1933C7BA79EF3FAAEF990" authorityName="Brady" authorityYear="1884" box="[1273,1386,1534,1560]" class="Nodosariata" family="Vaginulinidae" genus="Marginulina" kingdom="Chromista" order="Vaginulinida" pageId="18" pageNumber="19" phylum="Foraminifera" rank="species" species="glabra">
<emphasis id="A3AAEABEFFC1933C7BA79EF3FAAEF990" box="[1273,1386,1534,1560]" italics="true" pageId="18" pageNumber="19">M. glabra</emphasis>
</taxonomicName>
that is also described by
<bibRefCitation id="F54F4B5DFFC1933C7EDE9D2EFDA4F9B5" author="Parker W. K. &amp; Jones T. R. &amp; Brady H. B." box="[384,608,1570,1597]" pageId="18" pageNumber="19" pagination="15 - 41" refId="ref36138" refString="Parker W. K., Jones T. R. &amp; Brady H. B. (1865) On the nomenclature of the Foraminifera. Part X. (continued). The species enumerated by D'Orbigny in the ' Annales des Sciences Naturelles, ' 1826, Vol. VII - III. The species illustrated by models. Annals and Magazine of Natural History, 16, 15 - 41. https: // doi. org / 10.1080 / 00222936508679369" type="journal article" year="1865">
Parker
<emphasis id="A3AAEABEFFC1933C7E8F9D2FFDC0F9B4" box="[465,516,1570,1596]" italics="true" pageId="18" pageNumber="19">et al</emphasis>
. (1865)
</bibRefCitation>
, but the test of that species is not inflated, is thin and is more elongated, compared to
<taxonomicName id="56DE4D2FFFC1933C7E749D4BFE55F9E8" authorityName="Barker" authorityYear="1960" box="[298,401,1606,1632]" class="Nodosariata" family="Vaginulinidae" genus="Marginulina" kingdom="Chromista" order="Vaginulinida" pageId="18" pageNumber="19" phylum="Foraminifera" rank="species" species="obesa">
<emphasis id="A3AAEABEFFC1933C7E749D4BFE55F9E8" box="[298,401,1606,1632]" italics="true" pageId="18" pageNumber="19">M. obesa</emphasis>
</taxonomicName>
and
<taxonomicName id="56DE4D2FFFC1933C7E979D4BFDFEF9E8" authorityName="Brady" authorityYear="1884" box="[457,570,1606,1632]" class="Nodosariata" family="Vaginulinidae" genus="Marginulina" kingdom="Chromista" order="Vaginulinida" pageId="18" pageNumber="19" phylum="Foraminifera" rank="species" species="glabra">
<emphasis id="A3AAEABEFFC1933C7E979D4BFDFEF9E8" box="[457,570,1606,1632]" italics="true" pageId="18" pageNumber="19">M. glabra</emphasis>
</taxonomicName>
described in
<bibRefCitation id="F54F4B5DFFC1933C7D8F9D4AFCAEF9E8" author="Brady, H. B." box="[721,874,1606,1632]" pageId="18" pageNumber="19" pagination="1 - 814" refId="ref28770" refString="Brady, H. B. (1884) Report of the foraminifera dredged by H. M. S. Challenger during the years 1873 - 1876. Zoology, 9, 1 - 814." type="journal article" year="1884">Brady (1884)</bibRefCitation>
.
<taxonomicName id="56DE4D2FFFC1933C7C289D4BFC23F9E8" authorityName="Brady" authorityYear="1884" box="[886,999,1606,1632]" class="Nodosariata" family="Vaginulinidae" genus="Marginulina" kingdom="Chromista" order="Vaginulinida" pageId="18" pageNumber="19" phylum="Foraminifera" rank="species" species="glabra">
<emphasis id="A3AAEABEFFC1933C7C289D4BFC23F9E8" box="[886,999,1606,1632]" italics="true" pageId="18" pageNumber="19">M. glabra</emphasis>
</taxonomicName>
is now not accepted as a species name and the more elongated form described in
<bibRefCitation id="F54F4B5DFFC1933C7D2E9D66FC8DF90D" author="Parker W. K. &amp; Jones T. R. &amp; Brady H. B." box="[624,841,1642,1669]" pageId="18" pageNumber="19" pagination="15 - 41" refId="ref36138" refString="Parker W. K., Jones T. R. &amp; Brady H. B. (1865) On the nomenclature of the Foraminifera. Part X. (continued). The species enumerated by D'Orbigny in the ' Annales des Sciences Naturelles, ' 1826, Vol. VII - III. The species illustrated by models. Annals and Magazine of Natural History, 16, 15 - 41. https: // doi. org / 10.1080 / 00222936508679369" type="journal article" year="1865">
Parker
<emphasis id="A3AAEABEFFC1933C7D9E9D67FD35F90C" box="[704,753,1642,1668]" italics="true" pageId="18" pageNumber="19">et al</emphasis>
. (1865)
</bibRefCitation>
should be synonymised with
<taxonomicName id="56DE4D2FFFC1933C7BC89D67FAC0F90C" box="[1174,1284,1642,1668]" class="Nodosariata" family="Vaginulinidae" genus="Marginulina" kingdom="Chromista" order="Vaginulinida" pageId="18" pageNumber="19" phylum="Foraminifera" rank="species" species="similis">
<emphasis id="A3AAEABEFFC1933C7BC89D67FAC0F90C" box="[1174,1284,1642,1668]" italics="true" pageId="18" pageNumber="19">M. similis</emphasis>
</taxonomicName>
in
<bibRefCitation id="F54F4B5DFFC1933C7A769D66FF21F921" author="d'Orbigny, A. D." pageId="18" pageNumber="19" refId="ref30601" refString="d'Orbigny, A. D. (1846) Foraminiferes Fossiles du Basin Tertiaire de Vienne (Austriche). Gide et Comp, Paris, 312 pp. https: // doi. org / 10.5962 / bhl. title. 145432" type="book" year="1846">dOrbigny (1846)</bibRefCitation>
. The terminal chamber in
<taxonomicName id="56DE4D2FFFC1933C7D509D83FDB9F920" box="[526,637,1678,1704]" class="Nodosariata" family="Vaginulinidae" genus="Marginulina" kingdom="Chromista" order="Vaginulinida" pageId="18" pageNumber="19" phylum="Foraminifera" rank="species" species="similis">
<emphasis id="A3AAEABEFFC1933C7D509D83FDB9F920" box="[526,637,1678,1704]" italics="true" pageId="18" pageNumber="19">M. similis</emphasis>
</taxonomicName>
is also much more reduced, approximately a quarter in size, compared to the more inflated
<taxonomicName id="56DE4D2FFFC1933C7E249DBFFE25F944" authorityName="Barker" authorityYear="1960" box="[378,481,1714,1740]" class="Nodosariata" family="Vaginulinidae" genus="Marginulina" kingdom="Chromista" order="Vaginulinida" pageId="18" pageNumber="19" phylum="Foraminifera" rank="species" species="obesa">
<emphasis id="A3AAEABEFFC1933C7E249DBFFE25F944" box="[378,481,1714,1740]" italics="true" pageId="18" pageNumber="19">M. obesa</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="916136ACFFC1933C7F999DD9FC8CF8B0" blockId="18.[151,1437,1173,1992]" pageId="18" pageNumber="19">
<emphasis id="A3AAEABEFFC1933C7F999DD9FEACF978" bold="true" box="[199,360,1749,1776]" pageId="18" pageNumber="19">Life strategy:</emphasis>
The bathymetric distribution of
<taxonomicName id="56DE4D2FFFC1933C7D8F9DDBFCF3F978" authorityName="Barker" authorityYear="1960" box="[721,823,1750,1776]" class="Nodosariata" family="Vaginulinidae" genus="Marginulina" kingdom="Chromista" order="Vaginulinida" pageId="18" pageNumber="19" phylum="Foraminifera" rank="species" species="obesa">
<emphasis id="A3AAEABEFFC1933C7D8F9DDBFCF3F978" box="[721,823,1750,1776]" italics="true" pageId="18" pageNumber="19">M. obesa</emphasis>
</taxonomicName>
is recorded as slope to abyssal (
<bibRefCitation id="F54F4B5DFFC1933C7BC59DDAFA55F978" author="Holbourn, A. &amp; Henderson, A. S. &amp; MacLeod, N." box="[1179,1425,1750,1776]" pageId="18" pageNumber="19" refId="ref32986" refString="Holbourn, A., Henderson, A. S. &amp; MacLeod, N. (2013) Atlas of Benthic Foraminifera. Wiley-Blackwell, Oxford, 654 pp. https: // doi. org / 10.1002 / 9781118452493" type="book" year="2013">
Holbourn
<emphasis id="A3AAEABEFFC1933C7A519DDBFA84F978" box="[1295,1344,1750,1776]" italics="true" pageId="18" pageNumber="19">et al</emphasis>
., 2013
</bibRefCitation>
). Species of the genus
<taxonomicName id="56DE4D2FFFC1933C7ED69DF6FDD1F89C" authorityName="d'Orbigny" authorityYear="1826" box="[392,533,1786,1812]" class="Nodosariata" family="Vaginulinidae" genus="Marginulina" kingdom="Chromista" order="Vaginulinida" pageId="18" pageNumber="19" phylum="Foraminifera" rank="genus">
<emphasis id="A3AAEABEFFC1933C7ED69DF6FDD1F89C" box="[392,533,1786,1812]" italics="true" pageId="18" pageNumber="19">Marginulina</emphasis>
</taxonomicName>
are generally shallow-infaunal in environments with variable conditions of low (
<bibRefCitation id="F54F4B5DFFC1933C7FC19C12FE94F8B1" author="Bernhard, J. M." box="[159,336,1822,1849]" pageId="18" pageNumber="19" pagination="207 - 215" refId="ref28220" refString="Bernhard, J. M. (1986) Characteristic assemblages and morphologies from anoxic organic rich deposits: Jurassic through Holocene. Journal of Foraminiferal Research, 16 (3), 207 - 215. https: // doi. org / 10.2113 / gsjfr. 16.3.207" type="journal article" year="1986">Bernhard, 1986</bibRefCitation>
) to high oxygen (
<bibRefCitation id="F54F4B5DFFC1933C7D4B9C12FCF8F8B0" author="Milker, Y. &amp; Schmiedl, G." box="[533,828,1822,1848]" pageId="18" pageNumber="19" pagination="1 - 134" refId="ref35283" refString="Milker, Y. &amp; Schmiedl, G. (2012) A taxonomic guide to modern benthic shelf foraminifera of the western Mediterranean Sea. Palaeontologica Electronica, 15, 1 - 134. https: // doi. org / 10.26879 / 271" type="journal article" year="2012">Milker &amp; Schmiedl, 2012</bibRefCitation>
).
</paragraph>
<paragraph id="916136ACFFC1933C7F999C4DFB13F8D4" blockId="18.[151,1437,1173,1992]" box="[199,1239,1857,1884]" pageId="18" pageNumber="19">
<emphasis id="A3AAEABEFFC1933C7F999C4DFDCBF8D4" bold="true" box="[199,527,1857,1884]" pageId="18" pageNumber="19">Global stratigraphic range:</emphasis>
This species occurs from the Miocene to Recent (
<bibRefCitation id="F54F4B5DFFC1933C7B1E9C4EFB0EF8D4" author="Jones, R. W." box="[1088,1226,1858,1884]" pageId="18" pageNumber="19" refId="ref33343" refString="Jones, R. W. (1994) The Challenger Foraminifera. Oxford University Press, Oxford, 149 pp." type="book" year="1994">Jones, 1994</bibRefCitation>
).
</paragraph>
<paragraph id="916136ACFFC1933C7F999C69FE11F840" blockId="18.[151,1437,1173,1992]" pageId="18" pageNumber="19">
<emphasis id="A3AAEABEFFC1933C7F999C69FE79F808" bold="true" box="[199,445,1893,1920]" pageId="18" pageNumber="19">Regional occurrence</emphasis>
:
<taxonomicName id="56DE4D2FFFC1933C7E929C6BFDF7F808" authorityName="Barker" authorityYear="1960" box="[460,563,1894,1920]" class="Nodosariata" family="Vaginulinidae" genus="Marginulina" kingdom="Chromista" order="Vaginulinida" pageId="18" pageNumber="19" phylum="Foraminifera" rank="species" species="obesa">
<emphasis id="A3AAEABEFFC1933C7E929C6BFDF7F808" box="[460,563,1894,1920]" italics="true" pageId="18" pageNumber="19">M. obesa</emphasis>
</taxonomicName>
is documented to occur in middle Miocene sediments on the Namibian outer continental shelf, south of the Kunene River mouth (this study) and in surface sediments along the south coast of
<collectingCountry id="E9C9763CFFC1933C7FC99CA2FEE2F840" box="[151,294,1966,1992]" name="South Africa" pageId="18" pageNumber="19">South Africa</collectingCountry>
(
<bibRefCitation id="F54F4B5DFFC1933C7E6A9CA2FE0DF840" author="Lowry, F. M. D." box="[308,457,1966,1992]" pageId="18" pageNumber="19" refId="ref34923" refString="Lowry, F. M. D. (1987) Foraminiferal thanatocoenoses from the continental shelf of southern Africa. Unpublished Ph. D. thesis, University College, London, 443 pp." type="book" year="1987">Lowry, 1987</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>