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<mods:title id="BD36593C59694A04154C26BD66EDFF2C">On the Palaearctic and Oriental species of Scymbalium and Micrillus (Coleoptera: Staphylinidae: Paederinae)</mods:title>
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Key to the Palaearctic and 
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species of 
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<emphasis id="A3A9EA9AFFD4FFF8FD5AEF1BFCB9FA83" bold="true" box="[748,849,1294,1317]" italics="true" pageId="33" pageNumber="1512">Micrillus</emphasis>
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and 
<taxonomicName id="56DD4D0BFFD4FFF8FC3FEF1BFBFAFA83" authority="ERICHSON." authorityName="ERICHSON." box="[905,1042,1294,1317]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="1512" phylum="Arthropoda" rank="genus">
<emphasis id="A3A9EA9AFFD4FFF8FC3FEF1BFBFAFA83" bold="true" box="[905,1042,1294,1317]" italics="true" pageId="33" pageNumber="1512">Scymbalium</emphasis>
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For convenience, 
<taxonomicName id="56DD4D0BFFD4FFF8FEDEEF5FFDA8FAC6" authorityName="Assing" authorityYear="2013" baseAuthorityName="Cameron" baseAuthorityYear="1947" box="[360,576,1354,1376]" class="Insecta" family="Staphylinidae" genus="Micrillus" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="1512" phylum="Arthropoda" rank="species" species="aethiopicus">
<emphasis id="A3A9EA9AFFD4FFF8FEDEEF5FFDA8FAC6" box="[360,576,1354,1376]" italics="true" pageId="33" pageNumber="1512">Micrillus aethiopicus</emphasis>
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, a close relative of 
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<emphasis id="A3A9EA9AFFD4FFF8FCBCEF5EFC4EFAC6" box="[778,934,1355,1376]" italics="true" pageId="33" pageNumber="1512">M. aegyptiacus</emphasis>
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, is included in the key, although this species has been recorded only from the Afrotropical region.
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1 Protarsomeres I-IV dilated in both sexes. Species of moderately large to large size; body length&gt; 7.0 mm; length of forebody&gt; 
<quantity id="56259B6DFFD4FFF8FD3EEFB4FD3CFA10" box="[648,724,1441,1462]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="33" pageNumber="1512" unit="mm" value="3.8">3.8 mm</quantity>
. Genus 
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ERICHSON
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........2
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- Protarsomeres I-IV not dilated. Small to moderately large species; body length 3.2-9.0 mm; length of forebody 1.8-4.0 mm. Genus 
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<emphasis id="A3A9EA9AFFD4FFF8FD34EFC3FD32FA4D" box="[642,730,1494,1515]" italics="true" pageId="33" pageNumber="1512">Micrillus</emphasis>
RAFFRAY
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........................................5
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2 Head and pronotum with distinct and moderately dense punctation (
<figureCitation id="09E62A0DFFD4FFF8FC2FEFE0FC30F9AC" box="[921,984,1525,1546]" captionStart="Figs 2-8" captionStartId="7.[178,218,1503,1523]" captionTargetBox="[182,1080,219,1485]" captionTargetId="figure-59@7.[178,1085,219,1487]" captionTargetPageId="7" captionText="Figs 2-8: Scymbalium persimile CAMERON, holotype (2-4) and S. nepalense nov.sp., paratype (5- 8): (2, 5) habitus; (3, 6) forebody; (4) abdomen; (7) aedeagus in lateral view; (8) aedeagus in ventral view. Scale bars: 2-6: 1.0 mm; 7-8: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5301293" httpUri="https://zenodo.org/record/5301293/files/figure.png" pageId="33" pageNumber="1512">Fig. 3</figureCitation>
). West Palaearctic region................................................................................................................3
</keyLead>
</paragraph>
<paragraph id="91623688FFD7FFFBFF04EAC3FBD5FEBE" blockId="34.[178,1086,214,1543]" pageId="34" pageNumber="1513">
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- Head and pronotum with very fine (sometimes except for some scattered macropunctures on frons), sparse or very dense punctation (
<figureCitation id="09E62A0DFFD7FFFBFCFEEAF8FC64FEA4" box="[840,908,237,258]" captionStart="Figs 2-8" captionStartId="7.[178,218,1503,1523]" captionTargetBox="[182,1080,219,1485]" captionTargetId="figure-59@7.[178,1085,219,1487]" captionTargetPageId="7" captionText="Figs 2-8: Scymbalium persimile CAMERON, holotype (2-4) and S. nepalense nov.sp., paratype (5- 8): (2, 5) habitus; (3, 6) forebody; (4) abdomen; (7) aedeagus in lateral view; (8) aedeagus in ventral view. Scale bars: 2-6: 1.0 mm; 7-8: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5301293" httpUri="https://zenodo.org/record/5301293/files/figure.png" pageId="34" pageNumber="1513">Figs 6</figureCitation>
, 
<figureCitation id="09E62A0DFFD7FFFBFC2BEAF8FC0AFEA4" box="[925,994,237,258]" captionStart="Figs 56-62" captionStartId="32.[178,218,1520,1540]" captionTargetBox="[190,1085,222,1489]" captionTargetId="figure-68@32.[178,1085,212,1497]" captionTargetPageId="32" captionText="Figs 56-62: Scymbalium badium (MOTSCHULSKY), lectotype (56-60) and S. nepalense nov.sp., holotype (61-62): (56) forebody; (57, 61) median dorsal portion of head; (58, 62) male sternite VIII; (59-60) aedeagus in lateral and in ventral view. Scale bars: 56: 1.0 mm; 58, 62: 0.5 mm; 59- 60: 0.2 mm; 57, 61: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5301311" httpUri="https://zenodo.org/record/5301311/files/figure.png" pageId="34" pageNumber="1513">57, 61</figureCitation>
). East Palaearctic and 
<collectingRegion id="5319F86AFFD7FFFBFEC7EB16FE28FEBE" box="[369,448,259,280]" country="Morocco" name="Oriental" pageId="34" pageNumber="1513">Oriental</collectingRegion>
regions..........................................................................................4
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<paragraph id="91623688FFD7FFFBFF04EB37FBD5FEEA" blockId="34.[178,1086,214,1543]" pageId="34" pageNumber="1513">
<keyLead id="2A2C95BDFFD7FFFBFF04EB37FBD5FEEA" pageId="34" pageNumber="1513">
3 Abdomen with very fine punctation (
<figureCitation id="09E62A0DFFD7FFFBFD8BEB37FD90FE91" box="[573,632,290,311]" captionStart="Figs 2-8" captionStartId="7.[178,218,1503,1523]" captionTargetBox="[182,1080,219,1485]" captionTargetId="figure-59@7.[178,1085,219,1487]" captionTargetPageId="7" captionText="Figs 2-8: Scymbalium persimile CAMERON, holotype (2-4) and S. nepalense nov.sp., paratype (5- 8): (2, 5) habitus; (3, 6) forebody; (4) abdomen; (7) aedeagus in lateral view; (8) aedeagus in ventral view. Scale bars: 2-6: 1.0 mm; 7-8: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5301293" httpUri="https://zenodo.org/record/5301293/files/figure.png" pageId="34" pageNumber="1513">Fig. 4</figureCitation>
). 
<collectingCountry id="E9CA7618FFD7FFFBFD3CEB37FD5BFE91" box="[650,691,290,311]" name="Iraq" pageId="34" pageNumber="1513">Iraq</collectingCountry>
, Russian South European territory.......... .......................................................................................................... 
<taxonomicName id="56DD4D0BFFD7FFFBFCD4EB2CFBD5FEEA" ID-CoL="9QRS5" authority="CAMERON" authorityName="Cameron" authorityYear="1940" box="[866,1085,313,334]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="1513" phylum="Arthropoda" rank="species" species="persimile">
<emphasis id="A3A9EA9AFFD7FFFBFCD4EB2CFC3EFEE8" bold="true" box="[866,982,313,334]" italics="true" pageId="34" pageNumber="1513">S. persimile</emphasis>
CAMERON
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<paragraph id="91623688FFD7FFFBFF04EB4DFBD5FE25" blockId="34.[178,1086,214,1543]" pageId="34" pageNumber="1513">
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- Abdomen with distinctly coarser punctation. Widespread, from 
<collectingCountry id="E9CA7618FFD7FFFBFCFAEB4DFC65FECB" box="[844,909,344,365]" name="France" pageId="34" pageNumber="1513">France</collectingCountry>
and 
<collectingCountry id="E9CA7618FFD7FFFBFC76EB4DFBE0FECB" box="[960,1032,344,365]" name="Algeria" pageId="34" pageNumber="1513">Algeria</collectingCountry>
to 
<collectingCountry id="E9CA7618FFD7FFFBFF6EEB7BFEB5FE25" box="[216,349,366,387]" name="Turkmenistan" pageId="34" pageNumber="1513">Turkmenistan</collectingCountry>
and 
<collectingCountry id="E9CA7618FFD7FFFBFE3AEB7BFE5BFE25" box="[396,435,366,387]" name="Iran" pageId="34" pageNumber="1513">Iran</collectingCountry>
...................................................................... 
<taxonomicName id="56DD4D0BFFD7FFFBFCD3EB7BFBD5FE25" ID-CoL="9N48F" authority="(NORDMANN)" baseAuthorityName="NORDMANN" baseAuthorityYear="1837" box="[869,1085,366,387]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="1513" phylum="Arthropoda" rank="species" species="anale">
<emphasis id="A3A9EA9AFFD7FFFBFCD3EB7BFC5EFE25" bold="true" box="[869,950,366,387]" italics="true" pageId="34" pageNumber="1513">S. anale</emphasis>
(NORDMANN)
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</keyStep>
<keyStep id="2A292E2DFFD7FFFBFF04EB98FBD6FDD1" pageId="34" pageNumber="1513">
<paragraph id="91623688FFD7FFFBFF04EB98FBD5FE5B" blockId="34.[178,1086,214,1543]" pageId="34" pageNumber="1513">
<keyLead id="2A2C95BDFFD7FFFBFF04EB98FBD5FE5B" pageId="34" pageNumber="1513">
4 On average larger species of more robust habitus (
<figureCitation id="09E62A0DFFD7FFFBFD78EB98FCF0FE04" box="[718,792,397,418]" captionStart="Figs 56-62" captionStartId="32.[178,218,1520,1540]" captionTargetBox="[190,1085,222,1489]" captionTargetId="figure-68@32.[178,1085,212,1497]" captionTargetPageId="32" captionText="Figs 56-62: Scymbalium badium (MOTSCHULSKY), lectotype (56-60) and S. nepalense nov.sp., holotype (61-62): (56) forebody; (57, 61) median dorsal portion of head; (58, 62) male sternite VIII; (59-60) aedeagus in lateral and in ventral view. Scale bars: 56: 1.0 mm; 58, 62: 0.5 mm; 59- 60: 0.2 mm; 57, 61: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5301311" httpUri="https://zenodo.org/record/5301311/files/figure.png" pageId="34" pageNumber="1513">Fig. 56</figureCitation>
); body length 
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; length of forebody 
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. Head with rather dense and moderately fine punctation (
<figureCitation id="09E62A0DFFD7FFFBFF69EBAFFEC0FE69" box="[223,296,442,463]" captionStart="Figs 56-62" captionStartId="32.[178,218,1520,1540]" captionTargetBox="[190,1085,222,1489]" captionTargetId="figure-68@32.[178,1085,212,1497]" captionTargetPageId="32" captionText="Figs 56-62: Scymbalium badium (MOTSCHULSKY), lectotype (56-60) and S. nepalense nov.sp., holotype (61-62): (56) forebody; (57, 61) median dorsal portion of head; (58, 62) male sternite VIII; (59-60) aedeagus in lateral and in ventral view. Scale bars: 56: 1.0 mm; 58, 62: 0.5 mm; 59- 60: 0.2 mm; 57, 61: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5301311" httpUri="https://zenodo.org/record/5301311/files/figure.png" pageId="34" pageNumber="1513">Fig. 57</figureCitation>
). Male sternite VIII with deeper posterior concavity (
<figureCitation id="09E62A0DFFD7FFFBFC87EBAFFC92FE69" box="[817,890,442,463]" captionStart="Figs 56-62" captionStartId="32.[178,218,1520,1540]" captionTargetBox="[190,1085,222,1489]" captionTargetId="figure-68@32.[178,1085,212,1497]" captionTargetPageId="32" captionText="Figs 56-62: Scymbalium badium (MOTSCHULSKY), lectotype (56-60) and S. nepalense nov.sp., holotype (61-62): (56) forebody; (57, 61) median dorsal portion of head; (58, 62) male sternite VIII; (59-60) aedeagus in lateral and in ventral view. Scale bars: 56: 1.0 mm; 58, 62: 0.5 mm; 59- 60: 0.2 mm; 57, 61: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5301311" httpUri="https://zenodo.org/record/5301311/files/figure.png" pageId="34" pageNumber="1513">Fig. 58</figureCitation>
). Ventral process of aedeagus apically more slender and more strongly bent dorsad in lateral view (
<figureCitation id="09E62A0DFFD7FFFBFC4EEBC4FEFCFE5B" captionStart="Figs 56-62" captionStartId="32.[178,218,1520,1540]" captionTargetBox="[190,1085,222,1489]" captionTargetId="figure-68@32.[178,1085,212,1497]" captionTargetPageId="32" captionText="Figs 56-62: Scymbalium badium (MOTSCHULSKY), lectotype (56-60) and S. nepalense nov.sp., holotype (61-62): (56) forebody; (57, 61) median dorsal portion of head; (58, 62) male sternite VIII; (59-60) aedeagus in lateral and in ventral view. Scale bars: 56: 1.0 mm; 58, 62: 0.5 mm; 59- 60: 0.2 mm; 57, 61: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5301311" httpUri="https://zenodo.org/record/5301311/files/figure.png" pageId="34" pageNumber="1513">Figs 59-60</figureCitation>
). Northeastern 
<collectingCountry id="E9CA7618FFD7FFFBFE11EBFDFE33FE5B" box="[423,475,488,509]" name="India" pageId="34" pageNumber="1513">India</collectingCountry>
; 
<collectingCountry id="E9CA7618FFD7FFFBFE53EBFDFDC6FE5B" box="[485,558,488,509]" name="Bhutan" pageId="34" pageNumber="1513">Bhutan</collectingCountry>
; 
<collectingCountry id="E9CA7618FFD7FFFBFD8EEBFDFD7EFE5B" box="[568,662,488,509]" name="Myanmar" pageId="34" pageNumber="1513">Myanmar</collectingCountry>
......................... 
<taxonomicName id="56DD4D0BFFD7FFFBFC84EBFDFBD5FE5B" authority="(MOTSCHULSKY)" baseAuthorityName="Motschulsky" baseAuthorityYear="1858" box="[818,1085,488,509]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="1513" phylum="Arthropoda" rank="species" species="badium">
<emphasis id="A3A9EA9AFFD7FFFBFC84EBFDFC7FFE5B" bold="true" box="[818,919,488,509]" italics="true" pageId="34" pageNumber="1513">S. badium</emphasis>
(MOTSCHULSKY)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="91623688FFD7FFFBFF04E812FBD6FDD1" blockId="34.[178,1086,214,1543]" pageId="34" pageNumber="1513">
<keyLead id="2A2C95BDFFD7FFFBFF04E812FBD6FDD1" pageId="34" pageNumber="1513">
- On average smaller species of more slender habitus (
<figureCitation id="09E62A0DFFD7FFFBFD6FE812FCC3FDBA" box="[729,811,519,540]" captionStart="Figs 2-8" captionStartId="7.[178,218,1503,1523]" captionTargetBox="[182,1080,219,1485]" captionTargetId="figure-59@7.[178,1085,219,1487]" captionTargetPageId="7" captionText="Figs 2-8: Scymbalium persimile CAMERON, holotype (2-4) and S. nepalense nov.sp., paratype (5- 8): (2, 5) habitus; (3, 6) forebody; (4) abdomen; (7) aedeagus in lateral view; (8) aedeagus in ventral view. Scale bars: 2-6: 1.0 mm; 7-8: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5301293" httpUri="https://zenodo.org/record/5301293/files/figure.png" pageId="34" pageNumber="1513">Figs 5-6</figureCitation>
, 
<figureCitation id="09E62A0DFFD7FFFBFC81E812FCBAFDBA" box="[823,850,519,540]" captionStart="Figs 63-66" captionStartId="33.[178,218,834,854]" captionTargetBox="[191,1085,214,809]" captionTargetId="figure-296@33.[178,1085,212,809]" captionTargetPageId="33" captionText="Figs 63-66: Scymbalium nepalense nov.sp., holotype: (63) habitus; (64) male sternite VIII; (65-66) aedeagus in lateral and in ventral view. Scale bars: 63: 1.0 mm; 64: 0.5 mm; 65-66: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5301317" httpUri="https://zenodo.org/record/5301317/files/figure.png" pageId="34" pageNumber="1513">63</figureCitation>
); body length 
<quantity id="56259B6DFFD7FFFBFC68E812FEE8FD94" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="8.5" metricValueMin="7.5" pageId="34" pageNumber="1513" unit="mm" value="8.0" valueMax="8.5" valueMin="7.5">7.5-8.5 mm</quantity>
; length of forebody barely 
<quantity id="56259B6DFFD7FFFBFD96E808FD70FD94" box="[544,664,541,562]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" metricValueMax="4.7" metricValueMin="3.9" pageId="34" pageNumber="1513" unit="mm" value="4.3" valueMax="4.7" valueMin="3.9">3.9-4.7 mm</quantity>
. Head with extremely fine and sparse punctation (
<figureCitation id="09E62A0DFFD7FFFBFEFDE821FE7CFDEF" box="[331,404,564,585]" captionStart="Figs 56-62" captionStartId="32.[178,218,1520,1540]" captionTargetBox="[190,1085,222,1489]" captionTargetId="figure-68@32.[178,1085,212,1497]" captionTargetPageId="32" captionText="Figs 56-62: Scymbalium badium (MOTSCHULSKY), lectotype (56-60) and S. nepalense nov.sp., holotype (61-62): (56) forebody; (57, 61) median dorsal portion of head; (58, 62) male sternite VIII; (59-60) aedeagus in lateral and in ventral view. Scale bars: 56: 1.0 mm; 58, 62: 0.5 mm; 59- 60: 0.2 mm; 57, 61: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5301311" httpUri="https://zenodo.org/record/5301311/files/figure.png" pageId="34" pageNumber="1513">Fig. 61</figureCitation>
). Male sternite VIII with shallowly concave posterior margin (
<figureCitation id="09E62A0DFFD7FFFBFC4EE821FF1AFDC6" captionStart="Figs 56-62" captionStartId="32.[178,218,1520,1540]" captionTargetBox="[190,1085,222,1489]" captionTargetId="figure-68@32.[178,1085,212,1497]" captionTargetPageId="32" captionText="Figs 56-62: Scymbalium badium (MOTSCHULSKY), lectotype (56-60) and S. nepalense nov.sp., holotype (61-62): (56) forebody; (57, 61) median dorsal portion of head; (58, 62) male sternite VIII; (59-60) aedeagus in lateral and in ventral view. Scale bars: 56: 1.0 mm; 58, 62: 0.5 mm; 59- 60: 0.2 mm; 57, 61: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5301311" httpUri="https://zenodo.org/record/5301311/files/figure.png" pageId="34" pageNumber="1513">Figs 62</figureCitation>
, 
<figureCitation id="09E62A0DFFD7FFFBFF48E85EFEF1FDC6" box="[254,281,587,608]" captionStart="Figs 63-66" captionStartId="33.[178,218,834,854]" captionTargetBox="[191,1085,214,809]" captionTargetId="figure-296@33.[178,1085,212,809]" captionTargetPageId="33" captionText="Figs 63-66: Scymbalium nepalense nov.sp., holotype: (63) habitus; (64) male sternite VIII; (65-66) aedeagus in lateral and in ventral view. Scale bars: 63: 1.0 mm; 64: 0.5 mm; 65-66: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5301317" httpUri="https://zenodo.org/record/5301317/files/figure.png" pageId="34" pageNumber="1513">64</figureCitation>
). Aedeagus 
<quantity id="56259B6DFFD7FFFBFE24E85EFDEBFDC6" box="[402,515,587,608]" metricMagnitude="-4" metricUnit="m" metricValue="6.499999999999999" metricValueMax="7.0" metricValueMin="6.0" pageId="34" pageNumber="1513" unit="mm" value="0.6499999999999999" valueMax="0.7" valueMin="0.6">0.6-0.7 mm</quantity>
long, ventral process apically stouter and less strongly bent dorsad in lateral view (
<figureCitation id="09E62A0DFFD7FFFBFE52E877FDDFFDD1" box="[484,567,610,631]" captionStart="Figs 2-8" captionStartId="7.[178,218,1503,1523]" captionTargetBox="[182,1080,219,1485]" captionTargetId="figure-59@7.[178,1085,219,1487]" captionTargetPageId="7" captionText="Figs 2-8: Scymbalium persimile CAMERON, holotype (2-4) and S. nepalense nov.sp., paratype (5- 8): (2, 5) habitus; (3, 6) forebody; (4) abdomen; (7) aedeagus in lateral view; (8) aedeagus in ventral view. Scale bars: 2-6: 1.0 mm; 7-8: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5301293" httpUri="https://zenodo.org/record/5301293/files/figure.png" pageId="34" pageNumber="1513">Figs 7-8</figureCitation>
, 
<figureCitation id="09E62A0DFFD7FFFBFDF4E877FD96FDD1" box="[578,638,610,631]" captionStart="Figs 63-66" captionStartId="33.[178,218,834,854]" captionTargetBox="[191,1085,214,809]" captionTargetId="figure-296@33.[178,1085,212,809]" captionTargetPageId="33" captionText="Figs 63-66: Scymbalium nepalense nov.sp., holotype: (63) habitus; (64) male sternite VIII; (65-66) aedeagus in lateral and in ventral view. Scale bars: 63: 1.0 mm; 64: 0.5 mm; 65-66: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5301317" httpUri="https://zenodo.org/record/5301317/files/figure.png" pageId="34" pageNumber="1513">65-66</figureCitation>
). 
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............................ 
<taxonomicName id="56DD4D0BFFD7FFFBFCC0E877FC1AFDD1" ID-CoL="9QRRW" authority="Assing, 2013" authorityName="Assing" authorityYear="2013" box="[886,1010,610,631]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="1513" phylum="Arthropoda" rank="species" species="nepalense" status="sp. nov.">
<emphasis id="A3A9EA9AFFD7FFFBFCC0E877FC1AFDD1" bold="true" box="[886,1010,610,631]" italics="true" pageId="34" pageNumber="1513">S. nepalense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B89A57E1FFD7FFFBFC4EE877FBD6FDD1" box="[1016,1086,610,631]" pageId="34" pageNumber="1513" rank="species">nov.sp.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="2A292E2DFFD7FFFBFF04E895FBD5FC26" pageId="34" pageNumber="1513">
<paragraph id="91623688FFD7FFFBFF04E895FBD5FD56" blockId="34.[178,1086,214,1543]" pageId="34" pageNumber="1513">
<keyLead id="2A2C95BDFFD7FFFBFF04E895FBD5FD56" pageId="34" pageNumber="1513">
5 Apical joint of maxillary palpus long and slender, at least half the length of preapical palpomere (
<bibRefCitation id="F54C4B79FFD7FFFBFEF9E882FE3FFD0A" author="ASSING V" box="[335,471,663,684]" pageId="34" pageNumber="1513" pagination="301 - 344" refId="ref22718" refString="ASSING V. (2008): A revision of the Micrillus species of the Palaearctic region, with notes on two species from adjacent parts of the Afrotropical and Oriental regions (Coleoptera: Staphylinidae: Paederinae). - Stuttgarter Beitrage zur Naturkunde Serie A, Neue Serie 1: 301 - 344." type="journal article" year="2008">ASSING 2008</bibRefCitation>
: figure 39). Aedeagus of simple morphology, symmetric, usually slender with ventral process apically more or less bent dorsad; internal structures amorphous and weakly sclerotised. West Palaearctic region (including Middle Asia); absent from 
<collectingCountry id="E9CA7618FFD7FFFBFE6BE8CEFDFFFD56" box="[477,535,731,752]" name="Egypt" pageId="34" pageNumber="1513">Egypt</collectingCountry>
(
<bibRefCitation id="F54C4B79FFD7FFFBFD95E8CEFD4EFD56" author="ASSING V" box="[547,678,731,752]" pageId="34" pageNumber="1513" pagination="301 - 344" refId="ref22718" refString="ASSING V. (2008): A revision of the Micrillus species of the Palaearctic region, with notes on two species from adjacent parts of the Afrotropical and Oriental regions (Coleoptera: Staphylinidae: Paederinae). - Stuttgarter Beitrage zur Naturkunde Serie A, Neue Serie 1: 301 - 344." type="journal article" year="2008">ASSING 2008</bibRefCitation>
: figure 2). 
<taxonomicName id="56DD4D0BFFD7FFFBFCB8E8CEFC2CFD56" ID-CoL="9LW5Z" baseAuthorityName="ERICHSON" baseAuthorityYear="1840" box="[782,964,731,752]" class="Insecta" family="Staphylinidae" genus="Micrillus" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="1513" phylum="Arthropoda" rank="species" species="testaceus">
<emphasis id="A3A9EA9AFFD7FFFBFCB8E8CEFC2CFD56" box="[782,964,731,752]" italics="true" pageId="34" pageNumber="1513">Micrillus testaceus</emphasis>
</taxonomicName>
group.......6
</keyLead>
</paragraph>
<paragraph id="91623688FFD7FFFBFF04E8EFFBD5FC26" blockId="34.[178,1086,214,1543]" pageId="34" pageNumber="1513">
<keyLead id="2A2C95BDFFD7FFFBFF04E8EFFBD5FC26" pageId="34" pageNumber="1513">
- Apical joint of maxillary palpus of conical shape and usually shorter than half the length of preapical palpomere, only in one species from the 
<collectingRegion id="5319F86AFFD7FFFBFC89E904FC3DFC80" box="[831,981,785,806]" country="Egypt" name="Ash Sharqiyah" pageId="34" pageNumber="1513">Oriental region</collectingRegion>
longer. Aedeagus of complex morphology, usually asymmetric and/or with more or less distinctly sclerotised internal structures. Afrotropical, East Palaearctic, and Oriental regions, in the West Palaearctic confined to 
<collectingCountry id="E9CA7618FFD7FFFBFD39E940FD21FCCC" box="[655,713,853,874]" name="Egypt" pageId="34" pageNumber="1513">Egypt</collectingCountry>
and 
<collectingCountry id="E9CA7618FFD7FFFBFD4AE940FCADFCCC" box="[764,837,853,874]" name="Yemen" pageId="34" pageNumber="1513">Yemen</collectingCountry>
. 
<taxonomicName id="56DD4D0BFFD7FFFBFCF9E940FBCBFCCC" ID-CoL="9PLYR" box="[847,1059,853,874]" class="Insecta" family="Staphylinidae" genus="Micrillus" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="1513" phylum="Arthropoda" rank="species" species="aegyptiacus">
<emphasis id="A3A9EA9AFFD7FFFBFCF9E940FBCBFCCC" box="[847,1059,853,874]" italics="true" pageId="34" pageNumber="1513">Micrillus aegyptiacus</emphasis>
</taxonomicName>
and 
<taxonomicName id="56DD4D0BFFD7FFFBFEB7E97EFE9DFC26" ID-CoL="9PLZJ" authorityName="Assing" authorityYear="2013" baseAuthorityName="Cameron" baseAuthorityYear="1930" box="[257,373,875,896]" class="Insecta" family="Staphylinidae" genus="Micrillus" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="1513" phylum="Arthropoda" rank="species" species="suturalis">
<emphasis id="A3A9EA9AFFD7FFFBFEB7E97EFE9DFC26" box="[257,373,875,896]" italics="true" pageId="34" pageNumber="1513">M. suturalis</emphasis>
</taxonomicName>
groups.....................................................................................................13
</keyLead>
</paragraph>
</keyStep>
<keyStep id="2A292E2DFFD7FFFBFF04E99FFBD5FC4D" pageId="34" pageNumber="1513">
<paragraph id="91623688FFD7FFFBFF04E99FFBD5FC10" blockId="34.[178,1086,214,1543]" pageId="34" pageNumber="1513">
<keyLead id="2A2C95BDFFD7FFFBFF04E99FFBD5FC10" pageId="34" pageNumber="1513">6 Eyes fully developed or of reduced size, but always composed of ommatidia. Posterior margin of abdominal tergite VII at least with narrow rudiment of a palisade fringe..........7</keyLead>
</paragraph>
<paragraph id="91623688FFD7FFFBFF04E9D5FBD5FC4D" blockId="34.[178,1086,214,1543]" pageId="34" pageNumber="1513">
<keyLead id="2A2C95BDFFD7FFFBFF04E9D5FBD5FC4D" pageId="34" pageNumber="1513">- Eyes extremely reduced, without ommatidia. Abdominal tergite VII without palisade fringe. Distribution confined to North Africa....................................................................11</keyLead>
</paragraph>
</keyStep>
<keyStep id="2A292E2DFFD7FFFBFF04E9E0FBD5FB3C" pageId="34" pageNumber="1513">
<paragraph id="91623688FFD7FFFBFF04E9E0FBD5FBC3" blockId="34.[178,1086,214,1543]" pageId="34" pageNumber="1513">
<keyLead id="2A2C95BDFFD7FFFBFF04E9E0FBD5FBC3" pageId="34" pageNumber="1513">
7 Posterior margin of head distinctly concave in the middle (
<bibRefCitation id="F54C4B79FFD7FFFBFC9FE9E0FC46FBAC" author="ASSING V" box="[809,942,1013,1034]" pageId="34" pageNumber="1513" pagination="301 - 344" refId="ref22718" refString="ASSING V. (2008): A revision of the Micrillus species of the Palaearctic region, with notes on two species from adjacent parts of the Afrotropical and Oriental regions (Coleoptera: Staphylinidae: Paederinae). - Stuttgarter Beitrage zur Naturkunde Serie A, Neue Serie 1: 301 - 344." type="journal article" year="2008">ASSING 2008</bibRefCitation>
: figure 61). Eyes, elytra, and hind wings not reduced (
<bibRefCitation id="F54C4B79FFD7FFFBFDCAEE19FCE9FB87" author="ASSING V" box="[636,769,1036,1057]" pageId="34" pageNumber="1513" pagination="301 - 344" refId="ref22718" refString="ASSING V. (2008): A revision of the Micrillus species of the Palaearctic region, with notes on two species from adjacent parts of the Afrotropical and Oriental regions (Coleoptera: Staphylinidae: Paederinae). - Stuttgarter Beitrage zur Naturkunde Serie A, Neue Serie 1: 301 - 344." type="journal article" year="2008">ASSING 2008</bibRefCitation>
: figures 59-60, 62). Posterior margin of male sternite VIII with deeply and broadly triangular excision (
<bibRefCitation id="F54C4B79FFD7FFFBFC6CEE37FEE4FBE8" author="ASSING V" pageId="34" pageNumber="1513" pagination="301 - 344" refId="ref22718" refString="ASSING V. (2008): A revision of the Micrillus species of the Palaearctic region, with notes on two species from adjacent parts of the Afrotropical and Oriental regions (Coleoptera: Staphylinidae: Paederinae). - Stuttgarter Beitrage zur Naturkunde Serie A, Neue Serie 1: 301 - 344." type="journal article" year="2008">ASSING 2008</bibRefCitation>
: figure 64). Aedeagus as in 
<bibRefCitation id="F54C4B79FFD7FFFBFD80EE2CFD55FBE8" author="ASSING V" box="[566,701,1081,1102]" pageId="34" pageNumber="1513" pagination="301 - 344" refId="ref22718" refString="ASSING V. (2008): A revision of the Micrillus species of the Palaearctic region, with notes on two species from adjacent parts of the Afrotropical and Oriental regions (Coleoptera: Staphylinidae: Paederinae). - Stuttgarter Beitrage zur Naturkunde Serie A, Neue Serie 1: 301 - 344." type="journal article" year="2008">ASSING 2008</bibRefCitation>
: figures 66-74. Known distribution confined to 
<collectingCountry id="E9CA7618FFD7FFFBFEFDEE45FE9CFBC3" box="[331,372,1104,1125]" name="Iraq" pageId="34" pageNumber="1513">Iraq</collectingCountry>
, 
<collectingCountry id="E9CA7618FFD7FFFBFEC8EE45FE5EFBC3" box="[382,438,1104,1125]" name="Israel" pageId="34" pageNumber="1513">Israel</collectingCountry>
, and 
<collectingCountry id="E9CA7618FFD7FFFBFE5FEE45FDF8FBC3" box="[489,528,1104,1125]" name="Iran" pageId="34" pageNumber="1513">Iran</collectingCountry>
................................................... 
<taxonomicName id="56DD4D0BFFD7FFFBFCFAEE45FBD5FBC3" ID-CoL="9PLZ5" authority="(CAMERON)" baseAuthorityName="CAMERON" baseAuthorityYear="1940" box="[844,1085,1104,1125]" class="Insecta" family="Staphylinidae" genus="Micrillus" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="1513" phylum="Arthropoda" rank="species" species="irakensis">
<emphasis id="A3A9EA9AFFD7FFFBFCFAEE45FC2EFBC3" bold="true" box="[844,966,1104,1125]" italics="true" pageId="34" pageNumber="1513">M. irakensis</emphasis>
(CAMERON)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="91623688FFD7FFFBFF04EE7AFBD5FB3C" blockId="34.[178,1086,214,1543]" pageId="34" pageNumber="1513">
<keyLead id="2A2C95BDFFD7FFFBFF04EE7AFBD5FB3C" pageId="34" pageNumber="1513">- Posterior margin of head not distinctly concave, usually truncate. Aedeagus of different morphology..........................................................................................................8</keyLead>
</paragraph>
</keyStep>
<keyStep id="2A292E2DFFD7FFFBFF04EEB1FBD5FA28" pageId="34" pageNumber="1513">
<paragraph id="91623688FFD7FFFBFF04EEB1FBD5FA8D" blockId="34.[178,1086,214,1543]" pageId="34" pageNumber="1513">
<keyLead id="2A2C95BDFFD7FFFBFF04EEB1FBD5FA8D" pageId="34" pageNumber="1513">
8 Species of pale (yellowish, reddish or pale brown) coloration. Eyes, elytra, and hind wings not reduced (
<bibRefCitation id="F54C4B79FFD7FFFBFE23EEAEFDF0FB76" author="ASSING V" box="[405,536,1211,1232]" pageId="34" pageNumber="1513" pagination="301 - 344" refId="ref22718" refString="ASSING V. (2008): A revision of the Micrillus species of the Palaearctic region, with notes on two species from adjacent parts of the Afrotropical and Oriental regions (Coleoptera: Staphylinidae: Paederinae). - Stuttgarter Beitrage zur Naturkunde Serie A, Neue Serie 1: 301 - 344." type="journal article" year="2008">ASSING 2008</bibRefCitation>
: figures 48-49). Posterior margin of male sternite VIII with shallow emargination (
<bibRefCitation id="F54C4B79FFD7FFFBFE59EEC4FD9DFB40" author="ASSING V" box="[495,629,1233,1254]" pageId="34" pageNumber="1513" pagination="301 - 344" refId="ref22718" refString="ASSING V. (2008): A revision of the Micrillus species of the Palaearctic region, with notes on two species from adjacent parts of the Afrotropical and Oriental regions (Coleoptera: Staphylinidae: Paederinae). - Stuttgarter Beitrage zur Naturkunde Serie A, Neue Serie 1: 301 - 344." type="journal article" year="2008">ASSING 2008</bibRefCitation>
: figure 50). Aedeagus of distinctive shape, rather broad in ventral view, ventral process apically not bent dorsad, and with two clusters of moderately sclerotised internal structures (
<bibRefCitation id="F54C4B79FFD7FFFBFD4FEEEAFC69FAB2" author="ASSING V" box="[761,897,1279,1300]" pageId="34" pageNumber="1513" pagination="301 - 344" refId="ref22718" refString="ASSING V. (2008): A revision of the Micrillus species of the Palaearctic region, with notes on two species from adjacent parts of the Afrotropical and Oriental regions (Coleoptera: Staphylinidae: Paederinae). - Stuttgarter Beitrage zur Naturkunde Serie A, Neue Serie 1: 301 - 344." type="journal article" year="2008">ASSING 2008</bibRefCitation>
: figures 51-55). Caucasus region and Middle Asia ..................................................... 
<taxonomicName id="56DD4D0BFFD7FFFBFCD1EF03FBD5FA8D" ID-CoL="9PLZ9" authority="(REITTER)" baseAuthorityName="Reitter" baseAuthorityYear="1887" box="[871,1085,1302,1323]" class="Insecta" family="Staphylinidae" genus="Micrillus" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="1513" phylum="Arthropoda" rank="species" species="pallidus">
<emphasis id="A3A9EA9AFFD7FFFBFCD1EF03FC3FFA8D" bold="true" box="[871,983,1302,1323]" italics="true" pageId="34" pageNumber="1513">M. pallidus</emphasis>
(REITTER)
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<keyLead id="2A2C95BDFFD7FFFBFF04EF21FBD5FA28" pageId="34" pageNumber="1513">- Mature specimens usually (but not always!) of darker coloration, reddish to dark brown. Eyes, elytra, and hind wings often of reduced size or length, respectively. Aedeagus more slender, with one cluster of internal structures and with ventral process apically bent dorsad............................................................................................................9</keyLead>
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9 Wing-dimorphic, on average smaller species; eyes on average larger. Elytra and abdomen with finer punctation (
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: figure 11). Posterior margin of male sternite VIII with deeper triangular emargination (
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: figure 13). Widespread species with slender aedeagus with more weakly sclerotised ventral process (
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: figures 16-26), or distribution confined to 
<collectingCountry id="E9CA7618FFD7FFFBFCF9EFE7FC71F9A1" box="[847,921,1522,1543]" name="Algeria" pageId="34" pageNumber="1513">Algeria</collectingCountry>
.......................10
</keyLead>
</paragraph>
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- Monomorphic brachypterous species (
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: figures 34-35); eyes (
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: figure 38) usually smaller. Elytra and abdomen with coarser and denser punctation (
<bibRefCitation id="F54C4B79FFD6FFFAFEFAEB16FE39FEBE" author="ASSING V" box="[332,465,259,280]" pageId="35" pageNumber="1514" pagination="301 - 344" refId="ref22718" refString="ASSING V. (2008): A revision of the Micrillus species of the Palaearctic region, with notes on two species from adjacent parts of the Afrotropical and Oriental regions (Coleoptera: Staphylinidae: Paederinae). - Stuttgarter Beitrage zur Naturkunde Serie A, Neue Serie 1: 301 - 344." type="journal article" year="2008">ASSING 2008</bibRefCitation>
: figures 35, 37). Posterior margin of male sternite VIII with shallow emargination (
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: figure 56). Aedeagus relatively broad and with rather strongly sclerotised ventral process (
<bibRefCitation id="F54C4B79FFD6FFFAFD33EB24FCE2FEE0" author="ASSING V" box="[645,778,305,326]" pageId="35" pageNumber="1514" pagination="301 - 344" refId="ref22718" refString="ASSING V. (2008): A revision of the Micrillus species of the Palaearctic region, with notes on two species from adjacent parts of the Afrotropical and Oriental regions (Coleoptera: Staphylinidae: Paederinae). - Stuttgarter Beitrage zur Naturkunde Serie A, Neue Serie 1: 301 - 344." type="journal article" year="2008">ASSING 2008</bibRefCitation>
: figures 41-46). Endemic to northern 
<collectingCountry id="E9CA7618FFD6FFFAFE99EB5DFE6EFEFB" box="[303,390,328,349]" name="Morocco" pageId="35" pageNumber="1514">Morocco</collectingCountry>
............................................................................ 
<taxonomicName id="56DD4D0BFFD6FFFAFCEEEB5DFBD5FEFB" ID-CoL="9PLZB" authority="(FAUVEL)" baseAuthorityName="FAUVEL" baseAuthorityYear="1875" box="[856,1085,328,349]" class="Insecta" family="Staphylinidae" genus="Micrillus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="1514" phylum="Arthropoda" rank="species" species="scabrosus">
<emphasis id="A3A9EA9AFFD6FFFAFCEEEB5DFC33FEFB" bold="true" box="[856,987,328,349]" italics="true" pageId="35" pageNumber="1514">M. scabrosus</emphasis>
(FAUVEL)
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</keyStep>
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10 Aedeagus longer (&gt; 
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), slender, and with weakly sclerotised internal structures (
<bibRefCitation id="F54C4B79FFD6FFFAFF69EB68FE8BFE34" author="ASSING V" box="[223,355,381,402]" pageId="35" pageNumber="1514" pagination="301 - 344" refId="ref22718" refString="ASSING V. (2008): A revision of the Micrillus species of the Palaearctic region, with notes on two species from adjacent parts of the Afrotropical and Oriental regions (Coleoptera: Staphylinidae: Paederinae). - Stuttgarter Beitrage zur Naturkunde Serie A, Neue Serie 1: 301 - 344." type="journal article" year="2008">ASSING 2008</bibRefCitation>
: figures 16-26). Widespread in the south of the West Palaearctic region (
<figureCitation id="09E62A0DFFD6FFFAFF68EB81FEF1FE0F" box="[222,281,404,425]" captionStart="Map 1" captionStartId="11.[178,223,1571,1591]" captionTargetBox="[178,1058,212,1558]" captionTargetId="figure-16@11.[178,1058,212,1558]" captionTargetPageId="11" captionText="Map 1: Distribution of Micrillus testaceus (ERICHSON), based on revised records." figureDoi="http://doi.org/10.5281/zenodo.6511852" httpUri="https://zenodo.org/record/6511852/files/figure.png" pageId="35" pageNumber="1514">Map 1</figureCitation>
).......................................................................................... 
<taxonomicName id="56DD4D0BFFD6FFFAFCFAEB81FBD5FE0F" ID-CoL="9LW5Z" authority="(ERICHSON)" baseAuthorityName="ERICHSON" baseAuthorityYear="1840" box="[844,1085,404,425]" class="Insecta" family="Staphylinidae" genus="Micrillus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="1514" phylum="Arthropoda" rank="species" species="testaceus">
<emphasis id="A3A9EA9AFFD6FFFAFCFAEB81FC2FFE0F" bold="true" box="[844,967,404,425]" italics="true" pageId="35" pageNumber="1514">M. testaceus</emphasis>
(ERICHSON)
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</paragraph>
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- Aedeagus shorter (&lt;
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), broader, and with distinctly sclerotised internal structures (
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: figures 106-110). Known only from the surroundings of 
<collectingRegion id="5319F86AFFD6FFFAFF6EEBF5FEF1FE53" box="[216,281,480,501]" country="Algeria" name="Biskra" pageId="35" pageNumber="1514">Biskra</collectingRegion>
, 
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................................................................................ 
<taxonomicName id="56DD4D0BFFD6FFFAFCE1EBF5FBD5FE53" ID-CoL="9PLYT" authority="(FAUVEL)" baseAuthorityName="Fauvel" baseAuthorityYear="1898" box="[855,1085,480,501]" class="Insecta" family="Staphylinidae" genus="Micrillus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="1514" phylum="Arthropoda" rank="species" species="biskrensis">
<emphasis id="A3A9EA9AFFD6FFFAFCE1EBF5FC33FE53" bold="true" box="[855,987,480,501]" italics="true" pageId="35" pageNumber="1514">M. biskrensis</emphasis>
(FAUVEL)
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11 Elytra slightly narrower than pronotum (
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: figure 98). Eye rudiments (
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: figure 99) slightly larger, 
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. Aedeagus apically pointed in ventral view (
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: figures 102-103). Known only from two localities in Taza district, 
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............................................................................. 
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<emphasis id="A3A9EA9AFFD6FFFAFCECE856FC18FDFE" bold="true" box="[858,1008,579,600]" italics="true" pageId="35" pageNumber="1514">M. tenuipennis</emphasis>
ASSING
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- Elytra approximately as wide as pronotum. Eye rudiments smaller (
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: figures 77, 88), eye length 
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. Aedeagus apically weakly to distinctly concave in ventral view. Distribution confined to 
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................................................12
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12 Smaller species; head length &lt;
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, head width &lt;
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, width of pronotum &lt;
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. Antennae shorter (
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: figure 89), &lt;
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long. Eye rudiments smaller (
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: figure 88), eye length 
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. Pronotum on average more oblong (
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: figure 87). Head approximately as broad as long (
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: figure 87) and smaller in relation to pronotum, approximately as wide as pronotum (
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: figure 87). Head with coarser and denser punctation and with more pronounced microsculpture (
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: figure 87). Aedeagus apically weakly concave in ventral view and less strongly bent dorsad in lateral view (
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: figures 93-95). 
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: surroundings of 
<collectingCountry id="E9CA7618FFD6FFFAFDD6E976FD76FCDE" box="[608,670,867,888]" name="Algeria" pageId="35" pageNumber="1514">Algier</collectingCountry>
......................... 
<taxonomicName id="56DD4D0BFFD6FFFAFC8DE976FBD6FCD1" ID-CoL="9PLZD" authority="RAFFRAY" authorityName="RAFFRAY" authorityYear="1873" box="[827,1086,867,888]" class="Insecta" family="Staphylinidae" genus="Micrillus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="1514" phylum="Arthropoda" rank="species" species="subterraneus">
<emphasis id="A3A9EA9AFFD6FFFAFC8DE976FC35FCDE" bold="true" box="[827,989,867,888]" italics="true" pageId="35" pageNumber="1514">M. subterraneus</emphasis>
RAFFRAY
</taxonomicName>
</keyLead>
</paragraph>
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<keyLead id="2A2C95BDFFD6FFFAFF04E997FBD5FBEB" pageId="35" pageNumber="1514">
- Larger species; head length 
<quantity id="56259B6DFFD6FFFAFDB4E997FDB4FC31" box="[514,604,898,919]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="35" pageNumber="1514" unit="mm" value="0.9">0.90 mm</quantity>
, head width&gt; 
<quantity id="56259B6DFFD6FFFAFD46E997FCA2FC31" box="[752,842,898,919]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" pageId="35" pageNumber="1514" unit="mm" value="0.85">0.85 mm</quantity>
, width of pronotum&gt; 
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. Antennae longer (Fig. 78),&gt; 
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long. Eye rudiments slightly larger (
<bibRefCitation id="F54C4B79FFD6FFFAFF69E9A5FE8CFC63" author="ASSING V" box="[223,356,944,965]" pageId="35" pageNumber="1514" pagination="301 - 344" refId="ref22718" refString="ASSING V. (2008): A revision of the Micrillus species of the Palaearctic region, with notes on two species from adjacent parts of the Afrotropical and Oriental regions (Coleoptera: Staphylinidae: Paederinae). - Stuttgarter Beitrage zur Naturkunde Serie A, Neue Serie 1: 301 - 344." type="journal article" year="2008">ASSING 2008</bibRefCitation>
: figure 77), 
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long. Pronotum on average less oblong (
<bibRefCitation id="F54C4B79FFD6FFFAFC6CE9A5FEE4FC7D" author="ASSING V" pageId="35" pageNumber="1514" pagination="301 - 344" refId="ref22718" refString="ASSING V. (2008): A revision of the Micrillus species of the Palaearctic region, with notes on two species from adjacent parts of the Afrotropical and Oriental regions (Coleoptera: Staphylinidae: Paederinae). - Stuttgarter Beitrage zur Naturkunde Serie A, Neue Serie 1: 301 - 344." type="journal article" year="2008">ASSING 2008</bibRefCitation>
: figure 76). Head oblong (
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: figure 76), larger in relation to pronotum, and wider than pronotum (
<bibRefCitation id="F54C4B79FFD6FFFAFDE4E9C8FD30FC54" author="ASSING V" box="[594,728,989,1010]" pageId="35" pageNumber="1514" pagination="301 - 344" refId="ref22718" refString="ASSING V. (2008): A revision of the Micrillus species of the Palaearctic region, with notes on two species from adjacent parts of the Afrotropical and Oriental regions (Coleoptera: Staphylinidae: Paederinae). - Stuttgarter Beitrage zur Naturkunde Serie A, Neue Serie 1: 301 - 344." type="journal article" year="2008">ASSING 2008</bibRefCitation>
: figure 76). Head with finer and sparser punctation and with weaker microsculpture (
<bibRefCitation id="F54C4B79FFD6FFFAFCA7E9E1FC76FBAF" author="ASSING V" box="[785,926,1012,1033]" pageId="35" pageNumber="1514" pagination="301 - 344" refId="ref22718" refString="ASSING V. (2008): A revision of the Micrillus species of the Palaearctic region, with notes on two species from adjacent parts of the Afrotropical and Oriental regions (Coleoptera: Staphylinidae: Paederinae). - Stuttgarter Beitrage zur Naturkunde Serie A, Neue Serie 1: 301 - 344." type="journal article" year="2008">ASSING 2008</bibRefCitation>
: figure 76). Aedeagus apically distinctly concave in ventral view and strongly bent dorsad in lateral view (
<bibRefCitation id="F54C4B79FFD6FFFAFEE0EE34FE30FB90" author="ASSING V" box="[342,472,1057,1078]" pageId="35" pageNumber="1514" pagination="301 - 344" refId="ref22718" refString="ASSING V. (2008): A revision of the Micrillus species of the Palaearctic region, with notes on two species from adjacent parts of the Afrotropical and Oriental regions (Coleoptera: Staphylinidae: Paederinae). - Stuttgarter Beitrage zur Naturkunde Serie A, Neue Serie 1: 301 - 344." type="journal article" year="2008">ASSING 2008</bibRefCitation>
: figures 81-85). 
<collectingCountry id="E9CA7618FFD6FFFAFDC0EE34FD2AFB90" box="[630,706,1057,1078]" name="Algeria" pageId="35" pageNumber="1514">Algeria</collectingCountry>
: Djebel Bou Zegz ................................. ............................................................................................. 
<taxonomicName id="56DD4D0BFFD6FFFAFCAEEE2DFBD5FBEB" ID-CoL="9PLZ2" authority="(PEYERIMHOFF)" baseAuthorityName="PEYERIMHOFF" baseAuthorityYear="1907" box="[792,1085,1080,1101]" class="Insecta" family="Staphylinidae" genus="Micrillus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="1514" phylum="Arthropoda" rank="species" species="hypogaeus">
<emphasis id="A3A9EA9AFFD6FFFAFCAEEE2DFC4AFBEB" bold="true" box="[792,930,1080,1101]" italics="true" pageId="35" pageNumber="1514">M. hypogaeus</emphasis>
(PEYERIMHOFF)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="2A292E2DFFD6FFFAFF04EE42FBD5FB2C" pageId="35" pageNumber="1514">
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<keyLead id="2A2C95BDFFD6FFFAFF04EE42FBD5FBCA" box="[178,1085,1111,1132]" pageId="35" pageNumber="1514">13 Species from North Africa................................................................................................14</keyLead>
</paragraph>
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<keyLead id="2A2C95BDFFD6FFFAFF04EE60FBD5FB2C" box="[178,1085,1141,1162]" pageId="35" pageNumber="1514">
- Species distributed in the East Palaearctic and 
<collectingRegion id="5319F86AFFD6FFFAFD2FEE60FD00FB2C" box="[665,744,1141,1162]" country="Morocco" name="Oriental" pageId="35" pageNumber="1514">Oriental</collectingRegion>
regions.......................................17
</keyLead>
</paragraph>
</keyStep>
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<keyLead id="2A2C95BDFFD6FFFAFF04EE81FBD5FAEE" pageId="35" pageNumber="1514">
14 Smallest species of the genus; head length &lt;
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, head width &lt;
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, length of pronotum &lt;
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, width of pronotum &lt;
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. Wing-dimorphic (
<bibRefCitation id="F54C4B79FFD6FFFAFC6CEEBEFEE4FB70" author="ASSING V" pageId="35" pageNumber="1514" pagination="301 - 344" refId="ref22718" refString="ASSING V. (2008): A revision of the Micrillus species of the Palaearctic region, with notes on two species from adjacent parts of the Afrotropical and Oriental regions (Coleoptera: Staphylinidae: Paederinae). - Stuttgarter Beitrage zur Naturkunde Serie A, Neue Serie 1: 301 - 344." type="journal article" year="2008">ASSING 2008</bibRefCitation>
: figures 128-129); brachypterous morph with very small eyes (
<bibRefCitation id="F54C4B79FFD6FFFAFC2EEED4FBC8FB70" author="ASSING V" box="[920,1056,1217,1238]" pageId="35" pageNumber="1514" pagination="301 - 344" refId="ref22718" refString="ASSING V. (2008): A revision of the Micrillus species of the Palaearctic region, with notes on two species from adjacent parts of the Afrotropical and Oriental regions (Coleoptera: Staphylinidae: Paederinae). - Stuttgarter Beitrage zur Naturkunde Serie A, Neue Serie 1: 301 - 344." type="journal article" year="2008">ASSING 2008</bibRefCitation>
: figure 130), short elytra, and reduced hind wings (
<bibRefCitation id="F54C4B79FFD6FFFAFD72EECDFCA0FB4B" author="ASSING V" box="[708,840,1240,1261]" pageId="35" pageNumber="1514" pagination="301 - 344" refId="ref22718" refString="ASSING V. (2008): A revision of the Micrillus species of the Palaearctic region, with notes on two species from adjacent parts of the Afrotropical and Oriental regions (Coleoptera: Staphylinidae: Paederinae). - Stuttgarter Beitrage zur Naturkunde Serie A, Neue Serie 1: 301 - 344." type="journal article" year="2008">ASSING 2008</bibRefCitation>
: figure 129). Valvulae of abdominal segment IX relatively short (
<bibRefCitation id="F54C4B79FFD6FFFAFDC7EEFAFD1CFAA2" author="ASSING V" box="[625,756,1263,1284]" pageId="35" pageNumber="1514" pagination="301 - 344" refId="ref22718" refString="ASSING V. (2008): A revision of the Micrillus species of the Palaearctic region, with notes on two species from adjacent parts of the Afrotropical and Oriental regions (Coleoptera: Staphylinidae: Paederinae). - Stuttgarter Beitrage zur Naturkunde Serie A, Neue Serie 1: 301 - 344." type="journal article" year="2008">ASSING 2008</bibRefCitation>
: figure 136). Male sternite VIII posteriorly very deeply and very narrowly incised (
<bibRefCitation id="F54C4B79FFD6FFFAFD55EF13FC82FABD" author="ASSING V" box="[739,874,1286,1307]" pageId="35" pageNumber="1514" pagination="301 - 344" refId="ref22718" refString="ASSING V. (2008): A revision of the Micrillus species of the Palaearctic region, with notes on two species from adjacent parts of the Afrotropical and Oriental regions (Coleoptera: Staphylinidae: Paederinae). - Stuttgarter Beitrage zur Naturkunde Serie A, Neue Serie 1: 301 - 344." type="journal article" year="2008">ASSING 2008</bibRefCitation>
: figures 134-135). Aedeagus of distinctive morphology (
<bibRefCitation id="F54C4B79FFD6FFFAFDF2EF09FD2EFA97" author="ASSING V" box="[580,710,1308,1329]" pageId="35" pageNumber="1514" pagination="301 - 344" refId="ref22718" refString="ASSING V. (2008): A revision of the Micrillus species of the Palaearctic region, with notes on two species from adjacent parts of the Afrotropical and Oriental regions (Coleoptera: Staphylinidae: Paederinae). - Stuttgarter Beitrage zur Naturkunde Serie A, Neue Serie 1: 301 - 344." type="journal article" year="2008">ASSING 2008</bibRefCitation>
: figure 130-134). 
<collectingCountry id="E9CA7618FFD6FFFAFCC5EF09FC45FA97" box="[883,941,1308,1329]" name="Egypt" pageId="35" pageNumber="1514">Egypt</collectingCountry>
and 
<collectingCountry id="E9CA7618FFD6FFFAFC6AEF09FBF2FA97" box="[988,1050,1308,1329]" name="Sudan" pageId="35" pageNumber="1514">Sudan</collectingCountry>
...... .......................................................................................................... 
<taxonomicName id="56DD4D0BFFD6FFFAFCD3EF26FBD5FAEE" ID-CoL="9PLZM" authority="(KOCH)" baseAuthorityName="KOCH" box="[869,1085,1331,1352]" class="Insecta" family="Staphylinidae" genus="Micrillus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="1514" phylum="Arthropoda" rank="species" species="torretassoi">
<emphasis id="A3A9EA9AFFD6FFFAFCD3EF26FC06FAEE" bold="true" box="[869,1006,1331,1352]" italics="true" pageId="35" pageNumber="1514">M. torretassoi</emphasis>
(KOCH)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="91623688FFD6FFFAFF04EF47FBD6FA32" blockId="35.[178,1086,214,1573]" pageId="35" pageNumber="1514">
<keyLead id="2A2C95BDFFD6FFFAFF04EF47FBD6FA32" pageId="35" pageNumber="1514">- Distinctly larger, monomorphic macropterous species with distinctly larger eyes. Valvulae of abdominal segment very long. Male sternite VIII posteriorly with much less deep and broader excision. Aedeagus of completely different morphology...............15</keyLead>
</paragraph>
</keyStep>
<keyStep id="2A292E2DFFD6FFFDFF04EF8BFBD5FE89" lastPageId="36" lastPageNumber="1515" pageId="35" pageNumber="1514">
<paragraph id="91623688FFD6FFFAFF04EF8BFBD5F983" blockId="35.[178,1086,214,1573]" pageId="35" pageNumber="1514">
<keyLead id="2A2C95BDFFD6FFFAFF04EF8BFBD5F983" pageId="35" pageNumber="1514">
15 Abdomen with less dense and more distinct punctation and with less pronounced microsculpture. Head subparallel behind eyes. Eyes rather small and weakly convex (
<bibRefCitation id="F54C4B79FFD6FFFAFF69EFDEFE8DFA46" author="ASSING V" box="[223,357,1483,1504]" pageId="35" pageNumber="1514" pagination="301 - 344" refId="ref22718" refString="ASSING V. (2008): A revision of the Micrillus species of the Palaearctic region, with notes on two species from adjacent parts of the Afrotropical and Oriental regions (Coleoptera: Staphylinidae: Paederinae). - Stuttgarter Beitrage zur Naturkunde Serie A, Neue Serie 1: 301 - 344." type="journal article" year="2008">ASSING 2008</bibRefCitation>
: figures 112-113). Head oblong. Male sternite VIII without modified setae and with broader and less deep posterior excision (
<bibRefCitation id="F54C4B79FFD6FFFAFCACEFF7FC48FA51" author="ASSING V" box="[794,928,1506,1527]" pageId="35" pageNumber="1514" pagination="301 - 344" refId="ref22718" refString="ASSING V. (2008): A revision of the Micrillus species of the Palaearctic region, with notes on two species from adjacent parts of the Afrotropical and Oriental regions (Coleoptera: Staphylinidae: Paederinae). - Stuttgarter Beitrage zur Naturkunde Serie A, Neue Serie 1: 301 - 344." type="journal article" year="2008">ASSING 2008</bibRefCitation>
: figure 116). Aedeagus shaped as in 
<bibRefCitation id="F54C4B79FFD6FFFAFE0FEFECFDD2F9A8" author="ASSING V" box="[441,570,1529,1550]" pageId="35" pageNumber="1514" pagination="301 - 344" refId="ref22718" refString="ASSING V. (2008): A revision of the Micrillus species of the Palaearctic region, with notes on two species from adjacent parts of the Afrotropical and Oriental regions (Coleoptera: Staphylinidae: Paederinae). - Stuttgarter Beitrage zur Naturkunde Serie A, Neue Serie 1: 301 - 344." type="journal article" year="2008">ASSING 2008</bibRefCitation>
: figures 118-123. 
<collectingCountry id="E9CA7618FFD6FFFAFD5EEFECFCCCF9A8" box="[744,804,1529,1550]" name="Egypt" pageId="35" pageNumber="1514">Egypt</collectingCountry>
, 
<collectingCountry id="E9CA7618FFD6FFFAFC98EFECFC82F9A8" box="[814,874,1529,1550]" name="Sudan" pageId="35" pageNumber="1514">Sudan</collectingCountry>
(
<figureCitation id="09E62A0DFFD6FFFAFCC1EFECFC5FF9A8" box="[887,951,1529,1550]" captionStart="Map 2" captionStartId="12.[178,223,1191,1211]" captionTargetBox="[178,1085,212,1169]" captionTargetId="figure-224@12.[178,1085,212,1169]" captionTargetPageId="12" captionText="Map 2: Distributions of Micrillus aegyptiacus (BERNHAUER) (circles), M. sudanicus ASSING (triangles), and M. aethiopicus (CAMERON) (diamonds; open diamonds unrevised records; unidentified locality &quot;Nargi&quot; omitted)." figureDoi="http://doi.org/10.5281/zenodo.6511858" httpUri="https://zenodo.org/record/6511858/files/figure.png" pageId="35" pageNumber="1514">Map 2</figureCitation>
) .................... .............................................................................................. 
<taxonomicName id="56DD4D0BFFD6FFFAFCAAEC05FBD5F983" ID-CoL="9PLYR" authority="(BERNHAUER)" baseAuthorityName="BERNHAUER" box="[796,1085,1552,1573]" class="Insecta" family="Staphylinidae" genus="Micrillus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="1514" phylum="Arthropoda" rank="species" species="aegyptiacus">
<emphasis id="A3A9EA9AFFD6FFFAFCAAEC05FC5AF983" bold="true" box="[796,946,1552,1573]" italics="true" pageId="35" pageNumber="1514">M. aegyptiacus</emphasis>
(BERNHAUER)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="91623688FFD1FFFDFF04EAC3FBD5FE89" blockId="36.[178,1086,214,1559]" pageId="36" pageNumber="1515">
<keyLead id="2A2C95BDFFD1FFFDFF04EAC3FBD5FE89" pageId="36" pageNumber="1515">- Abdomen with dense and very fine punctation, and with pronounced microsculpture. Head tapering behind eyes, widest across the more convex eyes. Male sternite VIII with short stout modified setae in the middle and with deep and narrow posterior excision. Aedeagus of different morphology....................................................................16</keyLead>
</paragraph>
</keyStep>
<keyStep id="2A292E2DFFD1FFFDFF04EB2CFBD5FE61" pageId="36" pageNumber="1515">
<paragraph id="91623688FFD1FFFDFF04EB2CFBD5FE36" blockId="36.[178,1086,214,1559]" pageId="36" pageNumber="1515">
<keyLead id="2A2C95BDFFD1FFFDFF04EB2CFBD5FE36" pageId="36" pageNumber="1515">
16 Eyes larger and more bulging, strongly projecting from lateral contours of head (
<bibRefCitation id="F54C4B79FFD1FFFDFF69EB45FE8CFEC3" author="ASSING V" box="[223,356,336,357]" pageId="36" pageNumber="1515" pagination="301 - 344" refId="ref22718" refString="ASSING V. (2008): A revision of the Micrillus species of the Palaearctic region, with notes on two species from adjacent parts of the Afrotropical and Oriental regions (Coleoptera: Staphylinidae: Paederinae). - Stuttgarter Beitrage zur Naturkunde Serie A, Neue Serie 1: 301 - 344." type="journal article" year="2008">ASSING 2008</bibRefCitation>
: figures 155-156). Head as broad as long or indistinctly oblong. Male sternite VIII and aedeagus as in 
<bibRefCitation id="F54C4B79FFD1FFFDFD94EB73FD4EFEDD" author="ASSING V" box="[546,678,358,379]" pageId="36" pageNumber="1515" pagination="301 - 344" refId="ref22718" refString="ASSING V. (2008): A revision of the Micrillus species of the Palaearctic region, with notes on two species from adjacent parts of the Afrotropical and Oriental regions (Coleoptera: Staphylinidae: Paederinae). - Stuttgarter Beitrage zur Naturkunde Serie A, Neue Serie 1: 301 - 344." type="journal article" year="2008">ASSING 2008</bibRefCitation>
: figures 159, 161-163. 
<collectingCountry id="E9CA7618FFD1FFFDFC24EB73FC38FEDD" box="[914,976,358,379]" name="Sudan" pageId="36" pageNumber="1515">Sudan</collectingCountry>
, 
<collectingCountry id="E9CA7618FFD1FFFDFC6BEB73FBCAFEDD" box="[989,1058,358,379]" name="Yemen" pageId="36" pageNumber="1515">Yemen</collectingCountry>
(
<figureCitation id="09E62A0DFFD1FFFDFF68EB68FEF5FE34" box="[222,285,381,402]" captionStart="Map 2" captionStartId="12.[178,223,1191,1211]" captionTargetBox="[178,1085,212,1169]" captionTargetId="figure-224@12.[178,1085,212,1169]" captionTargetPageId="12" captionText="Map 2: Distributions of Micrillus aegyptiacus (BERNHAUER) (circles), M. sudanicus ASSING (triangles), and M. aethiopicus (CAMERON) (diamonds; open diamonds unrevised records; unidentified locality &quot;Nargi&quot; omitted)." figureDoi="http://doi.org/10.5281/zenodo.6511858" httpUri="https://zenodo.org/record/6511858/files/figure.png" pageId="36" pageNumber="1515">Map 2</figureCitation>
)............................................................................................... 
<taxonomicName id="56DD4D0BFFD1FFFDFCDDEB68FBD5FE36" ID-CoL="9PLZF" authority="ASSING" authorityName="ASSING" box="[875,1085,381,402]" class="Insecta" family="Staphylinidae" genus="Micrillus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="1515" phylum="Arthropoda" rank="species" species="sudanicus">
<emphasis id="A3A9EA9AFFD1FFFDFCDDEB68FC18FE34" bold="true" box="[875,1008,381,402]" italics="true" pageId="36" pageNumber="1515">M. sudanicus</emphasis>
ASSING
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="91623688FFD1FFFDFF04EB89FBD5FE61" blockId="36.[178,1086,214,1559]" pageId="36" pageNumber="1515">
<keyLead id="2A2C95BDFFD1FFFDFF04EB89FBD5FE61" pageId="36" pageNumber="1515">
- Eyes somewhat smaller and less bulging (
<figureCitation id="09E62A0DFFD1FFFDFDDDEB89FD5BFE17" box="[619,691,412,433]" captionStart="Figs 9-16" captionStartId="14.[178,218,1520,1540]" captionTargetBox="[182,1076,213,1487]" captionTargetId="figure-58@14.[178,1085,212,1497]" captionTargetPageId="14" captionText="Figs 9-16: Micrillus aethiopicus (CAMERON), paratype (9-13) and M. nigriceps (CAMERON), lectotype (14-16): (9) habitus; (10) forebody; (11, 14) male sternite VIII; (12-13, 15-16) aedeagus in lateral and in ventral view. Scale bars: 9-10: 1.0 mm; 11-16: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5301295" httpUri="https://zenodo.org/record/5301295/files/figure.png" pageId="36" pageNumber="1515">Fig. 10</figureCitation>
). Head oblong. Male sternite VIII and aedeagus as illustrated in 
<figureCitation id="09E62A0DFFD1FFFDFE66EBA7FDD2FE61" box="[464,570,434,455]" captionStart="Figs 9-16" captionStartId="14.[178,218,1520,1540]" captionTargetBox="[182,1076,213,1487]" captionTargetId="figure-58@14.[178,1085,212,1497]" captionTargetPageId="14" captionText="Figs 9-16: Micrillus aethiopicus (CAMERON), paratype (9-13) and M. nigriceps (CAMERON), lectotype (14-16): (9) habitus; (10) forebody; (11, 14) male sternite VIII; (12-13, 15-16) aedeagus in lateral and in ventral view. Scale bars: 9-10: 1.0 mm; 11-16: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5301295" httpUri="https://zenodo.org/record/5301295/files/figure.png" pageId="36" pageNumber="1515">Figs 11-13</figureCitation>
. 
<collectingCountry id="E9CA7618FFD1FFFDFDF2EBA7FD7EFE61" box="[580,662,434,455]" name="Ethiopia" pageId="36" pageNumber="1515">Ethiopia</collectingCountry>
(
<figureCitation id="09E62A0DFFD1FFFDFD14EBA7FD0BFE61" box="[674,739,434,455]" captionStart="Map 2" captionStartId="12.[178,223,1191,1211]" captionTargetBox="[178,1085,212,1169]" captionTargetId="figure-224@12.[178,1085,212,1169]" captionTargetPageId="12" captionText="Map 2: Distributions of Micrillus aegyptiacus (BERNHAUER) (circles), M. sudanicus ASSING (triangles), and M. aethiopicus (CAMERON) (diamonds; open diamonds unrevised records; unidentified locality &quot;Nargi&quot; omitted)." figureDoi="http://doi.org/10.5281/zenodo.6511858" httpUri="https://zenodo.org/record/6511858/files/figure.png" pageId="36" pageNumber="1515">Map 2</figureCitation>
) ........... 
<taxonomicName id="56DD4D0BFFD1FFFDFC85EBA7FBD5FE61" ID-CoL="9PLYS" authority="(CAMERON)" authorityName="Assing" authorityYear="2013" baseAuthorityName="Cameron" baseAuthorityYear="1947" box="[819,1085,434,455]" class="Insecta" family="Staphylinidae" genus="Micrillus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="1515" phylum="Arthropoda" rank="species" species="aethiopicus">
<emphasis id="A3A9EA9AFFD1FFFDFC85EBA7FC2FFE61" bold="true" box="[819,967,434,455]" italics="true" pageId="36" pageNumber="1515">M. aethiopicus</emphasis>
(CAMERON)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="2A292E2DFFD1FFFDFF04EBC4FBD5FD2B" pageId="36" pageNumber="1515">
<paragraph id="91623688FFD1FFFDFF04EBC4FBD5FDE7" blockId="36.[178,1086,214,1559]" pageId="36" pageNumber="1515">
<keyLead id="2A2C95BDFFD1FFFDFF04EBC4FBD5FDE7" pageId="36" pageNumber="1515">
17 Larger species; body length 6.5-7.0 mm; length of forebody 
<quantity id="56259B6DFFD1FFFDFC92EBC4FC7FFE40" box="[804,919,465,486]" metricMagnitude="-3" metricUnit="m" metricValue="3.65" metricValueMax="3.8" metricValueMin="3.5" pageId="36" pageNumber="1515" unit="mm" value="3.65" valueMax="3.8" valueMin="3.5">3.5-3.8 mm</quantity>
. Coloration of body dark-brown to blackish-brown. Posterior excision of male sternite VIII broad and moderately deep (
<figureCitation id="09E62A0DFFD1FFFDFE33EBEAFE25FDB2" box="[389,461,511,532]" captionStart="Figs 48-55" captionStartId="29.[178,218,1520,1540]" captionTargetBox="[182,1085,213,1495]" captionTargetId="figure-61@29.[178,1085,212,1497]" captionTargetPageId="29" captionText="Figs 48-55: Micrillus suturalis (CAMERON) (holotypes of Scymbalium borneense CAMERON (48- 49, 51, 54-55) and S. javanicum CAMERON (50-53)): (48) habitus; (49-50) forebody; (51) male sternite VIII; (52-55) aedeagus in lateral and in ventral view. Scale bars: 48-50: 1.0 mm; 51-55: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5301309" httpUri="https://zenodo.org/record/5301309/files/figure.png" pageId="36" pageNumber="1515">Fig. 51</figureCitation>
). Aedeagus 
<quantity id="56259B6DFFD1FFFDFDF5EBEAFD64FDB2" box="[579,652,511,532]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="36" pageNumber="1515" unit="mm" value="1.3">1.3 mm</quantity>
long and slender, with conspicuous series of long and strongly sclerotized spines in internal sac (
<figureCitation id="09E62A0DFFD1FFFDFD53E800FCBAFD8C" box="[741,850,533,554]" captionStart="Figs 48-55" captionStartId="29.[178,218,1520,1540]" captionTargetBox="[182,1085,213,1495]" captionTargetId="figure-61@29.[178,1085,212,1497]" captionTargetPageId="29" captionText="Figs 48-55: Micrillus suturalis (CAMERON) (holotypes of Scymbalium borneense CAMERON (48- 49, 51, 54-55) and S. javanicum CAMERON (50-53)): (48) habitus; (49-50) forebody; (51) male sternite VIII; (52-55) aedeagus in lateral and in ventral view. Scale bars: 48-50: 1.0 mm; 51-55: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5301309" httpUri="https://zenodo.org/record/5301309/files/figure.png" pageId="36" pageNumber="1515">Figs 52-55</figureCitation>
). 
<collectingCountry id="E9CA7618FFD1FFFDFCD4E800FC55FD8C" box="[866,957,533,554]" name="Malaysia" pageId="36" pageNumber="1515">Malaysia</collectingCountry>
, 
<collectingCountry id="E9CA7618FFD1FFFDFC71E800FBF7FD8C" box="[967,1055,533,554]" name="Thailand" pageId="36" pageNumber="1515">Thailand</collectingCountry>
, Borneo, Java (
<figureCitation id="09E62A0DFFD1FFFDFED4E839FE4BFDE7" box="[354,419,556,577]" captionStart="Map 4" captionStartId="27.[178,223,1091,1111]" captionTargetBox="[178,1085,212,1066]" captionTargetId="figure-174@27.[178,1085,212,1066]" captionTargetPageId="27" captionText="Map 4: Distributions of Micrillus distortus nov.sp. (triangles) and M. suturalis (CAMERON) (circles) in the Oriental region, based on examined records." figureDoi="http://doi.org/10.5281/zenodo.5301305" httpUri="https://zenodo.org/record/5301305/files/figure.png" pageId="36" pageNumber="1515">Map 4</figureCitation>
).................................................................... 
<taxonomicName id="56DD4D0BFFD1FFFDFCF8E839FBD5FDE7" ID-CoL="9PLZJ" authority="(CAMERON)" authorityName="Assing" authorityYear="2013" baseAuthorityName="Cameron" baseAuthorityYear="1930" box="[846,1085,556,577]" class="Insecta" family="Staphylinidae" genus="Micrillus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="1515" phylum="Arthropoda" rank="species" species="suturalis">
<emphasis id="A3A9EA9AFFD1FFFDFCF8E839FC2FFDE7" bold="true" box="[846,967,556,577]" italics="true" pageId="36" pageNumber="1515">M. suturalis</emphasis>
(CAMERON)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="91623688FFD1FFFDFF04E85EFBD5FD2B" blockId="36.[178,1086,214,1559]" pageId="36" pageNumber="1515">
<keyLead id="2A2C95BDFFD1FFFDFF04E85EFBD5FD2B" pageId="36" pageNumber="1515">- Distinctly smaller and paler species. Posterior excision of male sternite VIII deep and narrow. Aedeagus much smaller, compact, and with completely different internal structures...........................................................................................................................18</keyLead>
</paragraph>
</keyStep>
<keyStep id="2A292E2DFFD1FFFDFF04E882FBD5FC80" pageId="36" pageNumber="1515">
<paragraph id="91623688FFD1FFFDFF04E882FBD5FD56" blockId="36.[178,1086,214,1559]" pageId="36" pageNumber="1515">
<keyLead id="2A2C95BDFFD1FFFDFF04E882FBD5FD56" pageId="36" pageNumber="1515">
18 Preapical antennomeres (VII-IX) transverse (
<figureCitation id="09E62A0DFFD1FFFDFD0CE882FCE6FD0A" box="[698,782,663,684]" captionStart="Figs 29-35" captionStartId="22.[178,218,1520,1540]" captionTargetBox="[182,1076,212,1495]" captionTargetId="figure-79@22.[178,1085,212,1497]" captionTargetPageId="22" captionText="Figs 29-35: Micrillus sutteri (SCHEERPELTZ), holotype (29-32) and M. flavescens (MOTSCHULSKY) (lectotype of M. asperiventris (FAUVEL)) (33-35): (29) male sternite VIII; (30-31) aedeagus in lateral and in ventral view; (32) internal and dorso-apical structures of aedeagus in lateral view; (33) habitus; (34) forebody; (35) spermatheca. Scale bars: 33-34: 1.0 mm; 29-33: 0.2 mm; 35: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5301301" httpUri="https://zenodo.org/record/5301301/files/figure.png" pageId="36" pageNumber="1515">Fig. 33</figureCitation>
). Pronotum with broadly impunctate midline and laterally with coarser (though shallow) and dense punctation (
<figureCitation id="09E62A0DFFD1FFFDFF69E8D1FEC7FD7F" box="[223,303,708,729]" captionStart="Figs 29-35" captionStartId="22.[178,218,1520,1540]" captionTargetBox="[182,1076,212,1495]" captionTargetId="figure-79@22.[178,1085,212,1497]" captionTargetPageId="22" captionText="Figs 29-35: Micrillus sutteri (SCHEERPELTZ), holotype (29-32) and M. flavescens (MOTSCHULSKY) (lectotype of M. asperiventris (FAUVEL)) (33-35): (29) male sternite VIII; (30-31) aedeagus in lateral and in ventral view; (32) internal and dorso-apical structures of aedeagus in lateral view; (33) habitus; (34) forebody; (35) spermatheca. Scale bars: 33-34: 1.0 mm; 29-33: 0.2 mm; 35: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5301301" httpUri="https://zenodo.org/record/5301301/files/figure.png" pageId="36" pageNumber="1515">Fig. 34</figureCitation>
). Head with fine but transverse microsculpture. Male sexual characters unknown. 
<collectingCountry id="E9CA7618FFD1FFFDFE8BE8CEFE76FD56" box="[317,414,731,752]" name="Myanmar" pageId="36" pageNumber="1515">Myanmar</collectingCountry>
. ......................................................... 
<taxonomicName id="56DD4D0BFFD1FFFDFCA6E8CEFBD5FD56" authority="(MOTSCHULSKY)" authorityName="Assing" authorityYear="2013" baseAuthorityName="Motschulsky" baseAuthorityYear="2013" box="[784,1085,731,752]" class="Insecta" family="Staphylinidae" genus="Lithocharis" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="1515" phylum="Arthropoda" rank="species" species="flavescens">
<emphasis id="A3A9EA9AFFD1FFFDFCA6E8CEFC7FFD56" bold="true" box="[784,919,731,752]" italics="true" pageId="36" pageNumber="1515">M. flavescens</emphasis>
(MOTSCHULSKY)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="91623688FFD1FFFDFF04E8EFFBD5FC80" blockId="36.[178,1086,214,1559]" pageId="36" pageNumber="1515">
<keyLead id="2A2C95BDFFD1FFFDFF04E8EFFBD5FC80" pageId="36" pageNumber="1515">- Preapical antennomeres not transverse, usually oblong. Punctation of pronotum different.............................................................................................................................19</keyLead>
</paragraph>
</keyStep>
<keyStep id="2A292E2DFFD1FFFDFF04E93AFBD5FBA4" pageId="36" pageNumber="1515">
<paragraph id="91623688FFD1FFFDFF04E93AFBD6FC39" blockId="36.[178,1086,214,1559]" pageId="36" pageNumber="1515">
<keyLead id="2A2C95BDFFD1FFFDFF04E93AFBD6FC39" pageId="36" pageNumber="1515">
19 Head (
<figureCitation id="09E62A0DFFD1FFFDFEAAE93AFE8FFCE2" box="[284,359,815,836]" captionStart="Figs 36-43" captionStartId="26.[178,218,1520,1540]" captionTargetBox="[186,1078,212,1484]" captionTargetId="figure-65@26.[178,1085,212,1497]" captionTargetPageId="26" captionText="Figs 36-43: Micrillus distortus nov.sp.: (36) habitus; (37) forebody; (38) male sternite VII; (39) male sternite VIII; (40-41) aedeagus in lateral and in ventral view; (42) ventral process of aedeagus in ventral view; (43) internal structure of aedeagus in dorsal view. Scale bars: 36-37: 1.0 mm; 38- 43: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5301303" httpUri="https://zenodo.org/record/5301303/files/figure.png" pageId="36" pageNumber="1515">Fig. 37</figureCitation>
) at least as broad as long, with marked posterior angles, subparallel behind eyes. Head and pronotum somewhat flattened (cross-section) and distinctly glossy, with sparse and rather distinct punctation. Posterior excision of male sternite VIII very narrow. Aedeagus with long, basally strongly curved, strongly sclerotized, spine-like internal structure in dorsal position..................................................................20
</keyLead>
</paragraph>
<paragraph id="91623688FFD1FFFDFF04E9BCFBD5FBA4" blockId="36.[178,1086,214,1559]" pageId="36" pageNumber="1515">
<keyLead id="2A2C95BDFFD1FFFDFF04E9BCFBD5FBA4" pageId="36" pageNumber="1515">- Head usually oblong, rarely as broad as long, mostly more or less distinctly tapering behind eyes and with more broadly convex posterior angles. Head and pronotum not distinctly flattened, less glossy and with different punctation. Posterior excision of male sternite VIII less narrow. Aedeagus of different morphology..................................21</keyLead>
</paragraph>
</keyStep>
<keyStep id="2A292E2DFFD1FFFDFF04EE19FBD5FB3C" pageId="36" pageNumber="1515">
<paragraph id="91623688FFD1FFFDFF04EE19FBD6FBE8" blockId="36.[178,1086,214,1559]" pageId="36" pageNumber="1515">
<keyLead id="2A2C95BDFFD1FFFDFF04EE19FBD6FBE8" pageId="36" pageNumber="1515">
<specimenCount id="87DBFD01FFD1FFFDFF04EE19FEE1FB87" box="[178,265,1036,1057]" pageId="36" pageNumber="1515" type="male">20 Male</specimenCount>
sternite VIII distinctly oblong; posterior excision distinctly less than one third the length of sternite (
<figureCitation id="09E62A0DFFD1FFFDFE28EE36FE06FB9E" box="[414,494,1059,1080]" captionStart="Figs 36-43" captionStartId="26.[178,218,1520,1540]" captionTargetBox="[186,1078,212,1484]" captionTargetId="figure-65@26.[178,1085,212,1497]" captionTargetPageId="26" captionText="Figs 36-43: Micrillus distortus nov.sp.: (36) habitus; (37) forebody; (38) male sternite VII; (39) male sternite VIII; (40-41) aedeagus in lateral and in ventral view; (42) ventral process of aedeagus in ventral view; (43) internal structure of aedeagus in dorsal view. Scale bars: 36-37: 1.0 mm; 38- 43: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5301303" httpUri="https://zenodo.org/record/5301303/files/figure.png" pageId="36" pageNumber="1515">Fig. 39</figureCitation>
). Aedeagus shaped as in 
<figureCitation id="09E62A0DFFD1FFFDFCB3EE36FC9FFB9E" box="[773,887,1059,1080]" captionStart="Figs 36-43" captionStartId="26.[178,218,1520,1540]" captionTargetBox="[186,1078,212,1484]" captionTargetId="figure-65@26.[178,1085,212,1497]" captionTargetPageId="26" captionText="Figs 36-43: Micrillus distortus nov.sp.: (36) habitus; (37) forebody; (38) male sternite VII; (39) male sternite VIII; (40-41) aedeagus in lateral and in ventral view; (42) ventral process of aedeagus in ventral view; (43) internal structure of aedeagus in dorsal view. Scale bars: 36-37: 1.0 mm; 38- 43: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5301303" httpUri="https://zenodo.org/record/5301303/files/figure.png" pageId="36" pageNumber="1515">Figs 40-42</figureCitation>
. 
<collectingCountry id="E9CA7618FFD1FFFDFC31EE36FBF7FB9E" box="[903,1055,1059,1080]" isEnumeration="true" pageId="36" pageNumber="1515">Laos, Vietnam</collectingCountry>
, 
<collectingCountry id="E9CA7618FFD1FFFDFF6EEE2CFEC6FBE8" box="[216,302,1081,1102]" name="Thailand" pageId="36" pageNumber="1515">Thailand</collectingCountry>
(
<figureCitation id="09E62A0DFFD1FFFDFE8CEE2CFE93FBE8" box="[314,379,1081,1102]" captionStart="Map 4" captionStartId="27.[178,223,1091,1111]" captionTargetBox="[178,1085,212,1066]" captionTargetId="figure-174@27.[178,1085,212,1066]" captionTargetPageId="27" captionText="Map 4: Distributions of Micrillus distortus nov.sp. (triangles) and M. suturalis (CAMERON) (circles) in the Oriental region, based on examined records." figureDoi="http://doi.org/10.5281/zenodo.5301305" httpUri="https://zenodo.org/record/5301305/files/figure.png" pageId="36" pageNumber="1515">Map 4</figureCitation>
) .................................................................................. 
<taxonomicName id="56DD4D0BFFD1FFFDFCCDEE2CFC19FBE8" ID-CoL="9PLYY" authority="Assing, 2013" authorityName="Assing" authorityYear="2013" box="[891,1009,1081,1102]" class="Insecta" family="Staphylinidae" genus="Micrillus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="1515" phylum="Arthropoda" rank="species" species="distortus" status="sp. nov.">
<emphasis id="A3A9EA9AFFD1FFFDFCCDEE2CFC19FBE8" bold="true" box="[891,1009,1081,1102]" italics="true" pageId="36" pageNumber="1515">M. distortus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B89A57E1FFD1FFFDFC4EEE2CFBD6FBE8" box="[1016,1086,1081,1102]" pageId="36" pageNumber="1515" rank="species">nov.sp.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="91623688FFD1FFFDFF04EE4DFBD5FB3C" blockId="36.[178,1086,214,1559]" pageId="36" pageNumber="1515">
<keyLead id="2A2C95BDFFD1FFFDFF04EE4DFBD5FB3C" pageId="36" pageNumber="1515">
- Male sternite VIII very weakly oblong; posterior excision deeper, approximately one third the length of sternite (
<figureCitation id="09E62A0DFFD1FFFDFE44EE7AFDD7FB22" box="[498,575,1135,1156]" captionStart="Figs 44-47" captionStartId="28.[178,218,1334,1354]" captionTargetBox="[191,1076,578,1315]" captionTargetId="figure-220@28.[178,1085,578,1316]" captionTargetPageId="28" captionText="Figs 44-47: Micrillus sumbaensis (SCHEERPELTZ), holotype: (44) male sternite VIII; (45-46) aedeagus in lateral and in ventral view; (47) internal structure of aedeagus in dorsal view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5301307" httpUri="https://zenodo.org/record/5301307/files/figure.png" pageId="36" pageNumber="1515">Fig. 44</figureCitation>
). Aedeagus shaped as in 
<figureCitation id="09E62A0DFFD1FFFDFCF5EE7AFC59FB22" box="[835,945,1135,1156]" captionStart="Figs 44-47" captionStartId="28.[178,218,1334,1354]" captionTargetBox="[191,1076,578,1315]" captionTargetId="figure-220@28.[178,1085,578,1316]" captionTargetPageId="28" captionText="Figs 44-47: Micrillus sumbaensis (SCHEERPELTZ), holotype: (44) male sternite VIII; (45-46) aedeagus in lateral and in ventral view; (47) internal structure of aedeagus in dorsal view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5301307" httpUri="https://zenodo.org/record/5301307/files/figure.png" pageId="36" pageNumber="1515">Figs 45-47</figureCitation>
. 
<collectingCountry id="E9CA7618FFD1FFFDFC08EE7AFBF7FB22" box="[958,1055,1135,1156]" name="Indonesia" pageId="36" pageNumber="1515">Indonesia</collectingCountry>
: Sumba. ............................................................................... 
<taxonomicName id="56DD4D0BFFD1FFFDFCA7EE90FBD5FB3C" ID-CoL="9PLZG" authority="(SCHEERPELTZ)" authorityName="Assing" authorityYear="2013" baseAuthorityName="Scheerpeltz" baseAuthorityYear="1957" box="[785,1085,1157,1178]" class="Insecta" family="Staphylinidae" genus="Micrillus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="1515" phylum="Arthropoda" rank="species" species="sumbaensis">
<emphasis id="A3A9EA9AFFD1FFFDFCA7EE90FC4DFB3C" bold="true" box="[785,933,1157,1178]" italics="true" pageId="36" pageNumber="1515">M. sumbaensis</emphasis>
(SCHEERPELTZ)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="2A292E2DFFD1FFFDFF04EEB1FBD6FAA3" pageId="36" pageNumber="1515">
<paragraph id="91623688FFD1FFFDFF04EEB1FBD6FB76" blockId="36.[178,1086,214,1559]" pageId="36" pageNumber="1515">
<keyLead id="2A2C95BDFFD1FFFDFF04EEB1FBD6FB76" pageId="36" pageNumber="1515">
21 Larger species, length of forebody 
<quantity id="56259B6DFFD1FFFDFD9BEEB1FD92FB1F" box="[557,634,1188,1209]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="36" pageNumber="1515" unit="mm" value="3.2">3.2 mm</quantity>
. Male sternite VIII and aedeagus as in 
<figureCitation id="09E62A0DFFD1FFFDFC4FEEB1FEFBFB76" captionStart="Figs 17-22" captionStartId="19.[178,218,1468,1488]" captionTargetBox="[178,1084,212,1435]" captionTargetId="figure-81@19.[178,1085,212,1445]" captionTargetPageId="19" captionText="Figs 17-22: Micrillus siamensis nov.sp.: (17) habitus; (18) forebody; (19) male sternite VIII; (20) aedeagus in lateral view; (21) aedeagus in ventral view; (22) aedeagus in dorsal view. Scale bars: 17-18: 1.0 mm; 19-22: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5301297" httpUri="https://zenodo.org/record/5301297/files/figure.png" pageId="36" pageNumber="1515">Figs 19-22</figureCitation>
. 
<collectingCountry id="E9CA7618FFD1FFFDFEAAEEAEFE9AFB76" box="[284,370,1211,1232]" name="Thailand" pageId="36" pageNumber="1515">Thailand</collectingCountry>
(
<figureCitation id="09E62A0DFFD1FFFDFEC8EEAEFE57FB76" box="[382,447,1211,1232]" captionStart="Map 3" captionStartId="17.[178,223,1423,1443]" captionTargetBox="[178,1085,552,1406]" captionTargetId="figure-217@17.[178,1085,552,1406]" captionTargetPageId="17" captionText="Map 3: Distributions of Micrillus indicus (EPPELSHEIM) (triangles), M. nigriceps (CAMERON) (diamonds), M. intermedius (CAMERON) (open and filled circles), and M. siamensis nov.sp. (filled circle) in the Oriental region, based on examined records." figureDoi="http://doi.org/10.5281/zenodo.6512094" httpUri="https://zenodo.org/record/6512094/files/figure.png" pageId="36" pageNumber="1515">Map 3</figureCitation>
). .................................................................... 
<taxonomicName id="56DD4D0BFFD1FFFDFCC6EEAEFC1AFB76" ID-CoL="9PLZC" authority="Assing, 2013" authorityName="Assing" authorityYear="2013" box="[880,1010,1211,1232]" class="Insecta" family="Staphylinidae" genus="Micrillus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="1515" phylum="Arthropoda" rank="species" species="siamensis" status="sp. nov.">
<emphasis id="A3A9EA9AFFD1FFFDFCC6EEAEFC1AFB76" bold="true" box="[880,1010,1211,1232]" italics="true" pageId="36" pageNumber="1515">M. siamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B89A57E1FFD1FFFDFC4EEEAEFBD6FB76" box="[1016,1086,1211,1232]" pageId="36" pageNumber="1515" rank="species">nov.sp.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="91623688FFD1FFFDFF04EECCFBD6FAA3" blockId="36.[178,1086,214,1559]" pageId="36" pageNumber="1515">
<keyLead id="2A2C95BDFFD1FFFDFF04EECCFBD6FAA3" pageId="36" pageNumber="1515">- Smaller species; length of forebody &lt;3.0 mm. A reliable identification of the following species is possible only based on the morphology of the aedeagus..................22</keyLead>
</paragraph>
</keyStep>
<keyStep id="2A292E2DFFD1FFFDFF04EF1AFBD5FA6D" pageId="36" pageNumber="1515">
<paragraph id="91623688FFD1FFFDFF04EF1AFBD5FACE" blockId="36.[178,1086,214,1559]" pageId="36" pageNumber="1515">
<keyLead id="2A2C95BDFFD1FFFDFF04EF1AFBD5FACE" pageId="36" pageNumber="1515">
22 Depth of posterior excision of male sternite VIII distinctly less than one third the length of sternite; aedeagus minute, 
<quantity id="56259B6DFFD1FFFDFDD4EF33FD58FA9D" box="[610,688,1318,1339]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="36" pageNumber="1515" unit="mm" value="0.4">0.4 mm</quantity>
long, compact, without conspicuous dorso-apical structure, and with relatively small sclerotized internal structure. Species from 
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..........................................................................................................................23
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- Depth of posterior excision of male sternite VIII nearly one third the length of sternite; aedeagus&gt; 
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long, more slender, with conspicuous curved dorso-apical structure, and with large, long, and apically finger-shaped sclerotized internal structure. Species from 
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, 
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, 
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, and 
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....................................24
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23 Head usually reddish; slightly darker than pronotum at most. Aedeagus as illustrated in 
<bibRefCitation id="F54C4B79FFD1FFFDFF6EEFFEFEB1F9A6" author="ASSING V" box="[216,345,1515,1536]" pageId="36" pageNumber="1515" pagination="301 - 344" refId="ref22718" refString="ASSING V. (2008): A revision of the Micrillus species of the Palaearctic region, with notes on two species from adjacent parts of the Afrotropical and Oriental regions (Coleoptera: Staphylinidae: Paederinae). - Stuttgarter Beitrage zur Naturkunde Serie A, Neue Serie 1: 301 - 344." type="journal article" year="2008">ASSING 2008</bibRefCitation>
: figures 150-153. Central and northern 
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(
<figureCitation id="09E62A0DFFD1FFFDFCA4EFFEFCB8F9A6" box="[786,848,1515,1536]" captionStart="Map 3" captionStartId="17.[178,223,1423,1443]" captionTargetBox="[178,1085,552,1406]" captionTargetId="figure-217@17.[178,1085,552,1406]" captionTargetPageId="17" captionText="Map 3: Distributions of Micrillus indicus (EPPELSHEIM) (triangles), M. nigriceps (CAMERON) (diamonds), M. intermedius (CAMERON) (open and filled circles), and M. siamensis nov.sp. (filled circle) in the Oriental region, based on examined records." figureDoi="http://doi.org/10.5281/zenodo.6512094" httpUri="https://zenodo.org/record/6512094/files/figure.png" pageId="36" pageNumber="1515">Map 3</figureCitation>
)..................................... ..................................................................................................... 
<taxonomicName id="56DD4D0BFFD1FFFDFCFFEC17FBD5F9B1" ID-CoL="9PLZ3" authority="(EPPELSHEIM)" baseAuthorityName="Eppelsheim" baseAuthorityYear="1890" box="[841,1085,1538,1559]" class="Insecta" family="Staphylinidae" genus="Micrillus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="1515" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="A3A9EA9AFFD1FFFDFCFFEC17FC5BF9B1" bold="true" box="[841,947,1538,1559]" italics="true" pageId="36" pageNumber="1515">M. indicus</emphasis>
(EPPELSHEIM)
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</paragraph>
<paragraph id="91623688FFD0FFFCFF04EAC3FBD5FEA4" blockId="37.[178,1085,214,455]" pageId="37" pageNumber="1516">
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- Head brown to dark-brown, darker than the pale-reddish pronotum. Aedeagus as in 
<figureCitation id="09E62A0DFFD0FFFCFF6EEAF8FEAAFEA4" box="[216,322,237,258]" captionStart="Figs 9-16" captionStartId="14.[178,218,1520,1540]" captionTargetBox="[182,1076,213,1487]" captionTargetId="figure-58@14.[178,1085,212,1497]" captionTargetPageId="14" captionText="Figs 9-16: Micrillus aethiopicus (CAMERON), paratype (9-13) and M. nigriceps (CAMERON), lectotype (14-16): (9) habitus; (10) forebody; (11, 14) male sternite VIII; (12-13, 15-16) aedeagus in lateral and in ventral view. Scale bars: 9-10: 1.0 mm; 11-16: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5301295" httpUri="https://zenodo.org/record/5301295/files/figure.png" pageId="37" pageNumber="1516">Figs 15-16</figureCitation>
. Northeastern 
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(
<figureCitation id="09E62A0DFFD0FFFCFDBAEAF8FDA4FEA4" box="[524,588,237,258]" captionStart="Map 3" captionStartId="17.[178,223,1423,1443]" captionTargetBox="[178,1085,552,1406]" captionTargetId="figure-217@17.[178,1085,552,1406]" captionTargetPageId="17" captionText="Map 3: Distributions of Micrillus indicus (EPPELSHEIM) (triangles), M. nigriceps (CAMERON) (diamonds), M. intermedius (CAMERON) (open and filled circles), and M. siamensis nov.sp. (filled circle) in the Oriental region, based on examined records." figureDoi="http://doi.org/10.5281/zenodo.6512094" httpUri="https://zenodo.org/record/6512094/files/figure.png" pageId="37" pageNumber="1516">Map 3</figureCitation>
) ........................................ 
<taxonomicName id="56DD4D0BFFD0FFFCFCFCEAF8FBD5FEA4" authority="(CAMERON)" authorityName="Assing" authorityYear="2013" baseAuthorityName="Cameron" baseAuthorityYear="2013" box="[842,1085,237,258]" class="Insecta" family="Staphylinidae" genus="Scymbalium" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="1516" phylum="Arthropoda" rank="species" species="nigriceps">
<emphasis id="A3A9EA9AFFD0FFFCFCFCEAF8FC2FFEA4" bold="true" box="[842,967,237,258]" italics="true" pageId="37" pageNumber="1516">M. nigriceps</emphasis>
(CAMERON)
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24 Body smaller, length of forebody 
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. Head distinctly darker than remainder of body. Aedeagus with ventral process pointed and acute apically; dorso-apical structure short and strongly curved (
<figureCitation id="09E62A0DFFD0FFFCFE6CEB2CFDAFFEE8" box="[474,583,313,334]" captionStart="Figs 23-28" captionStartId="21.[178,218,1490,1510]" captionTargetBox="[183,1036,212,1453]" captionTargetId="figure-55@21.[178,1085,212,1467]" captionTargetPageId="21" captionText="Figs 23-28: Micrillus intermedius (CAMERON): (23) habitus; (24) forebody; (25) male sternite VIII; (26) aedeagus in lateral view; (27) aedeagus in ventral view; (28) internal structure of aedeagus in lateral view. Scale bars: 23-24: 1.0 mm; 25-28: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5301299" httpUri="https://zenodo.org/record/5301299/files/figure.png" pageId="37" pageNumber="1516">Figs 26-28</figureCitation>
). 
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, 
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, 
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(
<figureCitation id="09E62A0DFFD0FFFCFCEFEB2CFC71FEE8" box="[857,921,313,334]" captionStart="Map 3" captionStartId="17.[178,223,1423,1443]" captionTargetBox="[178,1085,552,1406]" captionTargetId="figure-217@17.[178,1085,552,1406]" captionTargetPageId="17" captionText="Map 3: Distributions of Micrillus indicus (EPPELSHEIM) (triangles), M. nigriceps (CAMERON) (diamonds), M. intermedius (CAMERON) (open and filled circles), and M. siamensis nov.sp. (filled circle) in the Oriental region, based on examined records." figureDoi="http://doi.org/10.5281/zenodo.6512094" httpUri="https://zenodo.org/record/6512094/files/figure.png" pageId="37" pageNumber="1516">Map 3</figureCitation>
) ......................... ................................................................................................. 
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<emphasis id="A3A9EA9AFFD0FFFCFC99EB45FC2FFEC3" bold="true" box="[815,967,336,357]" italics="true" pageId="37" pageNumber="1516">M. intermedius</emphasis>
(CAMERON)
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- Body larger, length of forebody 
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. Head mostly reddish, only indistinctly darker than pronotum at most. Aedeagus with ventral process apically bifid in ventral view; dorso-apical structure long and only apically curved (
<figureCitation id="09E62A0DFFD0FFFCFC8FEB89FC41FE17" box="[825,937,412,433]" captionStart="Figs 29-35" captionStartId="22.[178,218,1520,1540]" captionTargetBox="[182,1076,212,1495]" captionTargetId="figure-79@22.[178,1085,212,1497]" captionTargetPageId="22" captionText="Figs 29-35: Micrillus sutteri (SCHEERPELTZ), holotype (29-32) and M. flavescens (MOTSCHULSKY) (lectotype of M. asperiventris (FAUVEL)) (33-35): (29) male sternite VIII; (30-31) aedeagus in lateral and in ventral view; (32) internal and dorso-apical structures of aedeagus in lateral view; (33) habitus; (34) forebody; (35) spermatheca. Scale bars: 33-34: 1.0 mm; 29-33: 0.2 mm; 35: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5301301" httpUri="https://zenodo.org/record/5301301/files/figure.png" pageId="37" pageNumber="1516">Figs 30-32</figureCitation>
). 
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: Sumba .......................................................................................... 
<taxonomicName id="56DD4D0BFFD0FFFCFCF3EBA7FBD5FE61" ID-CoL="9PLZH" authority="(SCHEERPELTZ)" authorityName="Assing" authorityYear="2013" baseAuthorityName="Scheerpeltz" baseAuthorityYear="1957" box="[837,1085,434,455]" class="Insecta" family="Staphylinidae" genus="Micrillus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="1516" phylum="Arthropoda" rank="species" species="sutteri">
<emphasis id="A3A9EA9AFFD0FFFCFCF3EBA7FC4DFE61" bold="true" box="[837,933,434,455]" italics="true" pageId="37" pageNumber="1516">M. sutteri</emphasis>
(SCHEERPELTZ)
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