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<mods:title id="CA5CC5E757DAA3C9FDE91966458F5FDD">Chirostyloidean squat lobsters (Crustacea: Decapoda: Anomura) from the Galapagos Islands</mods:title>
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<mods:namePart id="762B6B38838881E222D89A283935A18A">Baba, Keiji</mods:namePart>
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<heading id="CA1B8189FF9EF634FF24FE22DC17973E" bold="true" box="[151,620,508,534]" fontSize="11" level="1" pageId="27" pageNumber="418" reason="1">
<taxonomicName id="56EC4D66FF9EF634FF24FE22DC17973E" ID-CoL="c5212fcc-14ae-4f60-a8ff-d552f2a48d5f" authority="(Benedict, 1902)" baseAuthorityName="Benedict" baseAuthorityYear="1902" box="[151,620,508,534]" class="Malacostraca" family="Sternostylidae" genus="Sternostylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="418" phylum="Arthropoda" rank="species" species="defensus">
<emphasis id="A398EAF7FF9EF634FF24FE22DC17973E" bold="true" box="[151,620,508,534]" pageId="27" pageNumber="418">
<emphasis id="A398EAF7FF9EF634FF24FE22DFE0973E" bold="true" box="[151,411,508,534]" italics="true" pageId="27" pageNumber="418">Sternostylus defensus</emphasis>
(
<bibRefCitation id="F57D4B14FF9EF634FE19FE22DC1F973E" author="Benedict, J. E." box="[426,612,508,534]" pageId="27" pageNumber="418" pagination="243 - 334" refId="ref12321" refString="Benedict, J. E. (1902) Description of a new genus and forty-six new species of crustaceans of the family Galatheidae with a list of the known marine species. Proceedings of the Biological Society of Washington, 26, 243 - 334." type="journal article" year="1902">Benedict, 1902</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="915336E5FF9EF634FF24FDC0DE82971F" blockId="27.[151,620,508,567]" box="[151,249,542,567]" pageId="27" pageNumber="418">(Fig. 18)</paragraph>
</subSubSection>
<subSubSection id="D9F6656EFF9EF634FF24FDB8DDE29792" pageId="27" pageNumber="418" type="reference_group">
<paragraph id="915336E5FF9EF634FF24FDB8DDE29792" blockId="27.[151,1436,614,698]" pageId="27" pageNumber="418">
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<taxonomicName id="56EC4D66FF9EF634FF24FDB8DC349754" authority="Benedict, 1902: 299" authorityName="Benedict" authorityPageNumber="299" authorityYear="1902" box="[151,591,614,636]" class="Malacostraca" family="Chirostylidae" genus="Ptychogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="418" phylum="Arthropoda" rank="species" species="defensa">
<emphasis id="A398EAF7FF9EF634FF24FDB8DF099754" box="[151,370,614,636]" italics="true" pageId="27" pageNumber="418">Ptychogaster defensa</emphasis>
<treatmentCitation id="104D10F4FF9EF634FEC8FDB8DC349754" author="Benedict, J. E." box="[379,591,614,636]" page="299" pageId="27" pageNumber="418" year="1902">
<bibRefCitation id="F57D4B14FF9EF634FEC8FDB8DC619754" author="Benedict, J. E." box="[379,538,614,636]" pageId="27" pageNumber="418" pagination="243 - 334" refId="ref12321" refString="Benedict, J. E. (1902) Description of a new genus and forty-six new species of crustaceans of the family Galatheidae with a list of the known marine species. Proceedings of the Biological Society of Washington, 26, 243 - 334." type="journal article" year="1902">Benedict, 1902</bibRefCitation>
: 299
</treatmentCitation>
</taxonomicName>
, fig. 44 (type locality: Galapagos Islands, between Santa Cruz and
<collectingCountry id="E9FB7675FF9EF634FAA3FDB8DBE79754" box="[1296,1436,614,636]" name="Saint Kitts and Nevis" pageId="27" pageNumber="418">San Cristobal</collectingCountry>
Islands,
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,
<geoCoordinate id="F4D85022FF9EF634FE28FD5BDC6C97B3" box="[411,535,645,667]" direction="west" orientation="longitude" pageId="27" pageNumber="418" precision="15" value="-89.90833">89°54'30&quot;W</geoCoordinate>
,
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).—Baba
<emphasis id="A398EAF7FF9EF634FD77FD58DC8C97B3" box="[708,759,645,667]" italics="true" pageId="27" pageNumber="418">et al.</emphasis>
2018: 81.
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<emphasis id="A398EAF7FF9EF634FF24FD7ADF069792" box="[151,381,676,698]" italics="true" pageId="27" pageNumber="418">Gastroptychus defensa</emphasis>
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.—
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<bibRefCitation id="F57D4B14FF9EF634FE28FD7ADC249792" author="Baba, K. &amp; Haig, J." box="[411,607,676,698]" pageId="27" pageNumber="418" pagination="854 - 860" refId="ref11789" refString="Baba, K. &amp; Haig, J. (1990) A new species of chirostylid crustacean (Decapoda: Anomura) from off the west coast of north America. Proceedings of the Biological Society of Washington, 103, 854 - 860." type="journal article" year="1990">Baba &amp; Haig 1990</bibRefCitation>
: 856
</treatmentCitation>
, fig. 3 (note on
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).
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</paragraph>
</subSubSection>
<subSubSection id="D9F6656EFF99F633FF74FF49DCD29715" pageId="28" pageNumber="419" type="discussion">
<paragraph id="915336E5FF99F633FF74FF49DAEA9435" blockId="28.[151,1436,151,645]" pageId="28" pageNumber="419">
<emphasis id="A398EAF7FF99F633FF74FF49DF4F9598" bold="true" box="[199,308,151,176]" pageId="28" pageNumber="419">Remarks</emphasis>
. The specimen examined matches both the original description and the note provided by
<bibRefCitation id="F57D4B14FF99F633FAF0FF49DF6495FC" author="Baba, K. &amp; Haig, J." pageId="28" pageNumber="419" pagination="854 - 860" refId="ref11789" refString="Baba, K. &amp; Haig, J. (1990) A new species of chirostylid crustacean (Decapoda: Anomura) from off the west coast of north America. Proceedings of the Biological Society of Washington, 103, 854 - 860." type="journal article" year="1990">Baba &amp; Haig (1990)</bibRefCitation>
on the
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, although the carapace spines are much sharper in this specimen. The spines on the P1 carpus also are more numerous and sharper than in the
<typeStatus id="4E578847FF99F633FCA5FF3EDD0795D0" box="[790,892,224,248]" pageId="28" pageNumber="419" type="syntype">syntypes</typeStatus>
. These differences in the spination between the present male and the
<typeStatus id="4E578847FF99F633FE36FEDBDFA79435" box="[389,476,261,285]" pageId="28" pageNumber="419" type="syntype">syntype</typeStatus>
male (CL
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vs.
<quantity id="56149B00FF99F633FD55FEDADD309434" box="[742,843,260,284]" metricMagnitude="-2" metricUnit="m" metricValue="1.29" pageId="28" pageNumber="419" unit="mm" value="12.9">12.9 mm</quantity>
) are likely to be size-related.
</paragraph>
<paragraph id="915336E5FF99F633FF74FEF9DD7394DD" blockId="28.[151,1436,151,645]" pageId="28" pageNumber="419">
The specimen was collected together with
<taxonomicName id="56EC4D66FF99F633FD17FEF9DDCF9468" authorityName="Faxon" authorityYear="1893" box="[676,948,295,320]" class="Malacostraca" family="Chirostylidae" genus="Uroptychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="28" pageNumber="419" phylum="Arthropoda" rank="species" species="occidentalis">
<emphasis id="A398EAF7FF99F633FD17FEF9DDCF9468" box="[676,948,295,320]" italics="true" pageId="28" pageNumber="419">Uroptychus occidentalis</emphasis>
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(see above). The photo (Fig. 19) shows the collected specimen on an antipatharian (black coral) next to a gorgonian,
<taxonomicName id="56EC4D66FF99F633FC6BFE92DA08944D" box="[984,1139,332,357]" class="Anthozoa" family="Chrysogorgiidae" genus="Chrysogorgia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="28" pageNumber="419" phylum="Cnidaria" rank="genus">
<emphasis id="A398EAF7FF99F633FC6BFE92DA08944D" box="[984,1139,332,357]" italics="true" pageId="28" pageNumber="419">Chrysogorgia</emphasis>
</taxonomicName>
sp. but it is not certain on which of the two corals the squat lobsters were originally resting. The in-situ photo in
<bibRefCitation id="F57D4B14FF99F633FBE6FEB1DB7694A0" author="Bell, K. L. C. &amp; Brennan, M. L. &amp; Flanders, J. &amp; Raineault, N. A. &amp; Wagner, K." box="[1109,1293,367,392]" pageId="28" pageNumber="419" pagination="1 - 84" refId="ref12234" refString="Bell, K. L. C., Brennan, M. L., Flanders, J., Raineault, N. A. &amp; Wagner, K. (Eds.) (2016) New frontiers in ocean exploration: The E / V Nautilus and NOAA Ship Okeanos Explorer 2015 field season. Oceanography, 29 (Supplement 1), 1 - 84. https: // doi. org / 10.5670 / oceanog. 2016. supplement. 01" type="journal article" year="2016">
Bell
<emphasis id="A398EAF7FF99F633FB39FEAFDABA94A0" box="[1162,1217,367,392]" italics="true" pageId="28" pageNumber="419">et al.</emphasis>
(2016
</bibRefCitation>
: 2) shows an unidentified squat lobster, which is very much like this species, on a black coral. According to
<bibRefCitation id="F57D4B14FF99F633FB62FE4BDE9F94F8" author="Le Guilloux, E. &amp; Hall-Spencer, J. M. &amp; Soffker, M. K. &amp; Olu, K." pageId="28" pageNumber="419" pagination="1363 - 1369" refId="ref13436" refString="Le Guilloux, E., Hall-Spencer, J. M., Soffker, M. K. &amp; Olu, K. (2010) Association between the squat lobster Gastroptychus formosus and cold-water corals in the North Atlantic. Journal of the Marine Biological Association of the United Kingdom, 90, 1363 - 1369. https: // doi. org / 10.1017 / S 0025315410000524" type="journal article" year="2010">
Le Guilloux
<emphasis id="A398EAF7FF99F633FAD0FE4BDBE79485" box="[1379,1436,404,429]" italics="true" pageId="28" pageNumber="419">et al.</emphasis>
(2010)
</bibRefCitation>
, adults of
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<emphasis id="A398EAF7FF99F633FEEDFE69DC2C94F8" box="[350,599,439,464]" italics="true" pageId="28" pageNumber="419">Sternostylus formosus</emphasis>
</taxonomicName>
(as
<taxonomicName id="56EC4D66FF99F633FD3AFE69DDE094F8" baseAuthorityName="Filhol" baseAuthorityYear="1884" box="[649,923,439,464]" class="Malacostraca" family="Chirostylidae" genus="Gastroptychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="28" pageNumber="419" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="A398EAF7FF99F633FD3AFE69DDE094F8" box="[649,923,439,464]" italics="true" pageId="28" pageNumber="419">Gastroptychus formosus</emphasis>
</taxonomicName>
) were found on gorgonian and antipatharian corals, but the black coral
<taxonomicName id="56EC4D66FF99F633FE0DFE02DC4294DD" box="[446,569,476,501]" class="Anthozoa" family="Leiopathidae" genus="Leiopathes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Antipatharia" pageId="28" pageNumber="419" phylum="Cnidaria" rank="genus">
<emphasis id="A398EAF7FF99F633FE0DFE02DC4294DD" box="[446,569,476,501]" italics="true" pageId="28" pageNumber="419">Leiopathes</emphasis>
</taxonomicName>
sp. was preferred.
</paragraph>
<paragraph id="915336E5FF99F633FF74FE21DCD29715" blockId="28.[151,1436,151,645]" pageId="28" pageNumber="419">
Five species of
<taxonomicName id="56EC4D66FF99F633FEC7FE21DF879730" authorityName="Baba, Ahyong &amp; Schnabel" authorityYear="2018" box="[372,508,511,536]" class="Malacostraca" family="Sternostylidae" genus="Sternostylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="28" pageNumber="419" phylum="Arthropoda" rank="genus">
<emphasis id="A398EAF7FF99F633FEC7FE21DF879730" box="[372,508,511,536]" italics="true" pageId="28" pageNumber="419">Sternostylus</emphasis>
</taxonomicName>
are known from the eastern Pacific. Distinguishing characters among these species are shown in the key to species in
<bibRefCitation id="F57D4B14FF99F633FDA4FDFADCDE9715" author="Baba, K." box="[535,677,548,573]" pageId="28" pageNumber="419" pagination="1 - 317" refId="ref11700" refString="Baba, K. (2005) Deep-sea chirostylid and galatheid crustaceans (Decapoda: Anomura) from the Indo-West Pacific, with a list of species. Galathea Report, 20, 1 - 317." type="journal article" year="2005">Baba (2005)</bibRefCitation>
.
</paragraph>
</subSubSection>
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<emphasis id="A398EAF7FF99F633FF74FD99DF229748" bold="true" box="[199,345,583,608]" pageId="28" pageNumber="419">Distribution</emphasis>
. Galapagos Islands between Santa Cruz and
<collectingCountry id="E9FB7675FF99F633FC33FD96DA649748" box="[896,1055,583,608]" name="Saint Kitts and Nevis" pageId="28" pageNumber="419">San Cristobal</collectingCountry>
Islands, and now at East Wolf Seamount; at depths of
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717
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.
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