312 lines
38 KiB
XML
312 lines
38 KiB
XML
<document id="7C85AAD878FF2400C2E0BF88C63E1695" ID-DOI="10.5281/zenodo.5347271" ID-ISSN="2345-7600" ID-Zenodo-Dep="5347271" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630203671219" checkinUser="tatiana" docAuthor="Nguyen, Anh D." docDate="2012" docId="191D7C72CD23E81FFF7D90F4FF23FD10" docLanguage="en" docName="60rbz289-311.pdf" docOrigin="Raffles Bulletin of Zoology 60 (2)" docStyle="DocumentStyle:C6443723295B2E4C6F22B9B23F08089C.5:RafflesBZool.2006-2013.journal_article" docStyleId="C6443723295B2E4C6F22B9B23F08089C" docStyleName="RafflesBZool.2006-2013.journal_article" docStyleVersion="5" docTitle="Tylopus phanluongi Nguyen 2012, new species" docType="treatment" docVersion="4" lastPageNumber="293" masterDocId="E524040ACD22E81BFF859465FF88FFC5" masterDocTitle="Tylopus Millipedes In Vietnam (Diplopoda: Polydesmida: Paradoxosomatidae: Sulciferini), With Descriptions Of Five New Species" masterLastPageNumber="311" masterPageNumber="289" pageNumber="290" updateTime="1699183855207" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="850062B3665EB76752A0786E10FC08BF">
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<mods:title id="BC22740BA5D531269A2F734ABE72239F">Tylopus Millipedes In Vietnam (Diplopoda: Polydesmida: Paradoxosomatidae: Sulciferini), With Descriptions Of Five New Species</mods:title>
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<mods:name id="F18944BAE5C83B68F13B8A802EAA59B4" type="personal">
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<mods:namePart id="03E2CD3C464E4628C378F9F7C3E66788">Nguyen, Anh D.</mods:namePart>
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<mods:title id="4D0B72E825D6B6EC98350910F60A87F1">Raffles Bulletin of Zoology</mods:title>
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<mods:part id="2A38D6809F3705E7BECDA1C3763E5D36">
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<mods:date id="BD4A2FAE55EEA5525AE38311EA87E598">2012</mods:date>
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<mods:number id="58A387EF8ABBEC9DB72CCC8B9EC02109">2012-08-31</mods:number>
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<mods:detail id="2EF9A01393CB76AB4AF75A7DED3E4B27" type="volume">
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<mods:number id="223FD1554BEF8A2E572B71C93679EE14">60</mods:number>
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<mods:identifier id="8DF6AC79F717ABF006E8291423372623" type="DOI">10.5281/zenodo.5347271</mods:identifier>
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<treatment id="191D7C72CD23E81FFF7D90F4FF23FD10" ID-DOI="http://doi.org/10.5281/zenodo.5449261" ID-Zenodo-Dep="5449261" LSID="urn:lsid:plazi:treatment:191D7C72CD23E81FFF7D90F4FF23FD10" httpUri="http://treatment.plazi.org/id/191D7C72CD23E81FFF7D90F4FF23FD10" lastPageId="4" lastPageNumber="293" pageId="1" pageNumber="290">
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<subSubSection id="D9AE9EEFCD23E81AFF7D90F4FDE1FB6F" box="[248,617,1169,1195]" pageId="1" pageNumber="290" type="nomenclature">
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<heading id="CA437A08CD23E81AFF7D90F4FDE1FB6F" bold="true" box="[248,617,1169,1195]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="290" reason="2">
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<emphasis id="A3C01176CD23E81AFF7D90F4FDE1FB6F" bold="true" box="[248,617,1169,1195]" pageId="1" pageNumber="290">
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<taxonomicName id="56B4B6E7CD23E81AFF7D90F4FE5DFB6F" ID-CoL="7CZPN" authority="Nguyen, 2012" authorityName="Nguyen" authorityYear="2012" box="[248,469,1169,1194]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="290" phylum="Arthropoda" rank="species" species="phanluongi" status="sp. nov.">
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<emphasis id="A3C01176CD23E81AFF7D90F4FE5DFB6F" bold="true" box="[248,469,1169,1194]" italics="true" pageId="1" pageNumber="290">Tylopus phanluongi</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="B8F3AC0DCD23E81AFE6190F6FDE1FB6F" box="[484,617,1170,1195]" pageId="1" pageNumber="290" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="D9AE9EEFCD23E81AFEE590D7FD89FB0E" box="[352,513,1202,1227]" pageId="1" pageNumber="290" type="description">
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<paragraph id="910BCD64CD23E81AFEE590D7FD89FB0E" blockId="1.[248,617,1169,1227]" box="[352,513,1202,1227]" pageId="1" pageNumber="290">
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(
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<figureCitation id="098FD1E1CD23E81AFEED90D7FE3AFB0E" box="[360,434,1202,1227]" captionStart="Fig" captionStartId="2.[137,172,2059,2081]" captionTargetBox="[139,1480,356,1904]" captionTargetId="figure-8@2.[107,1493,337,1932]" captionTargetPageId="2" captionText="Fig. 1. Tylopus phanluongi, new species, holotype: body ring 10, dorsal view (A); hypoproct, ventral view (B); sternite 5, posterioventral view (C); leg 9 (D); right gonopod, mesal view (E)and lateral view (F). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5347273" httpUri="https://zenodo.org/record/5347273/files/figure.png" pageId="1" pageNumber="290">Figs. 1</figureCitation>
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,
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<figureCitation id="098FD1E1CD23E81AFE4490D6FE58FB0E" box="[449,464,1203,1227]" captionStart="Fig" captionStartId="3.[113,148,2092,2114]" captionTargetBox="[113,1452,549,2054]" captionTargetId="figure-80@3.[113,1453,548,2054]" captionTargetPageId="3" captionText="Fig. 2. Tylopus phanluongi, new species, holotype: right gonopod, mesal view (A); postfemoral region, mesal view (B)." figureDoi="http://doi.org/10.5281/zenodo.5347275" httpUri="https://zenodo.org/record/5347275/files/figure.png" pageId="1" pageNumber="290">2</figureCitation>
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,
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<figureCitation id="098FD1E1CD23E81AFE5890D6FE70FB0E" box="[477,504,1203,1227]" captionStart="Fig" captionStartId="21.[113,147,1802,1824]" captionTargetBox="[113,753,1083,1781]" captionTargetId="figure-736@21.[113,753,1083,1779]" captionTargetPageId="21" captionText="Fig. 18.Distribution map ofthe millipede genus Tylopus Jeekel, 1968 in Vietnam. 1: Sa Pa, Lao Cai Prov. (T. crassipes, T. procurvus, T. nodulipes, T. sigma, T. sapaensis, T. maculatus, T. magnicus); 2: Van Ban, Lao Cai Prov. (T. crassipes, T. nodulipes); 3: Xuan Son NP (T. strongylosomoides, T. golovatchi); 4: Tam Dao NP (T. tamdaoensis, T. roseiparaterga, T. strongylosomoides); 5: Cuc Phuong NP (T. granulatus, T. hilaroides, T. topali); 6: Pu Mat NP (T. nodulipes); 7: Huong Son, Ha Tinh prov. (T. nodulipes); 8: Bach Ma–Ba Na NP (T. hilaris, T. multilatus); 9: Ngoc Linh Mts., Kon Tum prov. (T. hilaris, T. phanluongi); 10: Bi Doup–Nui Ba NP (T. spinisterna)." figureDoi="http://doi.org/10.5281/zenodo.5347309" httpUri="https://zenodo.org/record/5347309/files/figure.png" pageId="1" pageNumber="290">18</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="D9AE9EEFCD23E81AFFF49095FE39F981" pageId="1" pageNumber="290" type="materials_examined">
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<paragraph id="910BCD64CD23E81AFFF49095FD14FA98" blockId="1.[113,752,1264,1604]" pageId="1" pageNumber="290">
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<emphasis id="A3C01176CD23E81AFFF49095FE9EFAC3" bold="true" box="[113,278,1264,1286]" italics="true" pageId="1" pageNumber="290">Material studied.</emphasis>
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—
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<materialsCitation id="21DCC739CD23E81AFEB29094FD14FA98" collectingDate="2006-03-11" collectingDateMax="2006-04-09" collectingDateMin="2006-03-11" collectorName="Nguyen Duc Anh" county="Male" elevation="1850" latitude="15.3" location="Ngoc Linh Mountain" longLatPrecision="1285" longitude="8.016666" municipality="Dak Glei District" pageId="1" pageNumber="290" specimenCount="1" stateProvince="Kon Tum" typeStatus="holotype">
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<typeStatus id="4E0F73C6CD23E81AFEB29094FE1BFAC2" box="[311,403,1265,1287]" pageId="1" pageNumber="290" type="holotype">Holotype</typeStatus>
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:
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<collectingCounty id="786AB5E8CD23E81AFE1F9094FE43FAC2" box="[410,459,1265,1287]" pageId="1" pageNumber="290">Male</collectingCounty>
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in the bottle
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||
<emphasis id="A3C01176CD23E81AFDC69094FD33FAC3" bold="true" box="[579,699,1265,1286]" pageId="1" pageNumber="290">IEBR-100H</emphasis>
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, with a label “
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<collectingRegion id="53700386CD23E81AFF38916BFEFBFAE1" box="[189,371,1294,1316]" country="Vietnam" name="Kon Tum" pageId="1" pageNumber="290">Kon Tum Province</collectingRegion>
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,
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<collectingMunicipality id="716F571ECD23E81AFEF9916BFDA8FAE1" box="[380,544,1294,1316]" pageId="1" pageNumber="290">Dak Glei District</collectingMunicipality>
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,
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<location id="946B9BBFCD23E81AFDAC916BFD78FAE1" LSID="urn:lsid:plazi:treatment:191D7C72CD23E81FFF7D90F4FF23FD10:946B9BBFCD23E81AFDAC916BFD78FAE1" box="[553,752,1294,1316]" county="Male" latitude="15.3" longLatPrecision="1285" longitude="8.016666" municipality="Dak Glei District" name="Ngoc Linh Mountain" pageId="1" pageNumber="290" stateProvince="Kon Tum">Ngoc Linh Mountain</location>
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(15°00'–
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<geoCoordinate id="F480ABA3CD23E81AFF40914EFE9CFA84" box="[197,276,1323,1345]" degrees="15" direction="north" minutes="18" orientation="latitude" pageId="1" pageNumber="290" precision="925" value="15.3">15°18'N</geoCoordinate>
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, 107°41'–
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<geoCoordinate id="F480ABA3CD23E81AFEF3914EFE4CFA84" box="[374,452,1323,1345]" degrees="08" direction="east" minutes="01" orientation="longitude" pageId="1" pageNumber="290" precision="925" value="8.016666">08°01'E</geoCoordinate>
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), primary forest,
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<elevation id="1A992A57CD23E81AFDEE914EFF17FA98" metricMagnitude="3" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" pageId="1" pageNumber="290" unit="m" value="1850.0" valueMax="1900.0" valueMin="1800.0">
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<quantity id="564C6081CD23E81AFDEE914EFD78FA85" box="[619,752,1323,1344]" metricMagnitude="3" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" pageId="1" pageNumber="290" unit="m" value="1850.0" valueMax="1900.0" valueMin="1800.0">1800–1900 m</quantity>
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a.s.l.
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</elevation>
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, coll.
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<collectorName id="3C41A8B2CD23E81AFF59912DFE01FA98" box="[220,393,1351,1373]" pageId="1" pageNumber="290">Nguyen Duc Anh</collectorName>
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,
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<date id="E50AEBA4CD23E81AFE11912DFD87FA98" box="[404,527,1351,1373]" pageId="1" pageNumber="290" value="2006-03-11">
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||
<collectingDate id="F54E124CCD23E81AFE11912DFD87FA98" box="[404,527,1351,1373]" pageId="1" pageNumber="290" value="2006-03-11">11 Mar.2006</collectingDate>
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</date>
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–
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<date id="E50AEBA4CD23E81AFDAC912DFD10FA98" box="[553,664,1351,1373]" pageId="1" pageNumber="290" value="2006-04-09">
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<collectingDate id="F54E124CCD23E81AFDAC912DFD10FA98" box="[553,664,1351,1373]" pageId="1" pageNumber="290" value="2006-04-09">9 Apr.2006</collectingDate>
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</date>
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.
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</materialsCitation>
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</paragraph>
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<paragraph id="910BCD64CD23E81AFFF49100FE39F981" blockId="1.[113,752,1264,1604]" pageId="1" pageNumber="290">
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<materialsCitation id="21DCC739CD23E81AFFF49100FD64FABF" box="[113,748,1379,1402]" collectingDate="2006-03-11" collectingDateMax="2006-04-09" collectingDateMin="2006-03-11" collectorName="Nguyen Duc Anh" county="Male" elevation="1850" latitude="15.3" location="Ngoc Linh Mountain" longLatPrecision="1285" longitude="8.016666" municipality="Dak Glei District" pageId="1" pageNumber="290" specimenCount="1" specimenCount-female="1" stateProvince="Kon Tum" typeStatus="paratype">
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<typeStatus id="4E0F73C6CD23E81AFFF49100FF59FABF" box="[113,209,1381,1402]" pageId="1" pageNumber="290" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="87B206EDCD23E81AFF5D9101FEA2FABF" box="[216,298,1380,1402]" pageId="1" pageNumber="290" type="female">1 female</specimenCount>
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(
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<emphasis id="A3C01176CD23E81AFEB09106FE39FABC" bold="true" box="[309,433,1379,1401]" pageId="1" pageNumber="290">IEBR-100P)</emphasis>
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, collected together with holotype
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</materialsCitation>
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;
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<materialsCitation id="21DCC739CD23E81AFFF491E4FF71FA28" collectingDate="2004-03-25" collectingDateMax="2004-04-11" collectingDateMin="2004-03-25" collectorName="Nguyen Duc Anh" elevation="1700" latitude="15.3" location="Ngoc Linh Mountain" longLatPrecision="1285" longitude="8.016666" municipality="Dak Glei District" pageId="1" pageNumber="290" specimenCount="6" specimenCount-female="2" specimenCount-male="4" stateProvince="Kon Tum" typeStatus="paratype">
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<specimenCount id="87B206EDCD23E81AFFF491E4FF4AFA52" box="[113,194,1409,1431]" pageId="1" pageNumber="290" type="male">4 males</specimenCount>
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,
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<specimenCount id="87B206EDCD23E81AFF4A91E4FEB9FA52" box="[207,305,1409,1431]" pageId="1" pageNumber="290" type="female">2 females</specimenCount>
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(
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<emphasis id="A3C01176CD23E81AFEC691E4FE23FA53" bold="true" box="[323,427,1409,1430]" pageId="1" pageNumber="290">IEBR-101</emphasis>
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),
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<collectingRegion id="53700386CD23E81AFE4691E7FD0FFA52" box="[451,647,1409,1431]" country="Vietnam" name="Kon Tum" pageId="1" pageNumber="290">Kon Tum Province</collectingRegion>
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,
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<collectingMunicipality id="716F571ECD23E81AFD1191E4FF34FA71" pageId="1" pageNumber="290">Dak Glei District</collectingMunicipality>
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,
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<location id="946B9BBFCD23E81AFF4191FBFE05FA71" LSID="urn:lsid:plazi:treatment:191D7C72CD23E81FFF7D90F4FF23FD10:946B9BBFCD23E81AFF4191FBFE05FA71" box="[196,397,1438,1460]" latitude="15.3" longLatPrecision="1285" longitude="8.016666" municipality="Dak Glei District" name="Ngoc Linh Mountain" pageId="1" pageNumber="290" stateProvince="Kon Tum">Ngoc Linh Mountain</location>
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(15°00'–
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<geoCoordinate id="F480ABA3CD23E81AFE6291FBFDBEFA71" box="[487,566,1438,1460]" degrees="15" direction="north" minutes="18" orientation="latitude" pageId="1" pageNumber="290" precision="925" value="15.3">15°18'N</geoCoordinate>
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, 107°41'–
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<geoCoordinate id="F480ABA3CD23E81AFD1291FBFD6DFA71" box="[663,741,1438,1460]" degrees="08" direction="east" minutes="01" orientation="longitude" pageId="1" pageNumber="290" precision="925" value="8.016666">08°01'E</geoCoordinate>
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), secondary forest,
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||
<elevation id="1A992A57CD23E81AFE9291DEFE18FA14" box="[279,400,1467,1489]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="290" unit="m" value="1700.0">
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<quantity id="564C6081CD23E81AFE9291DEFED6FA15" box="[279,350,1467,1488]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="290" unit="m" value="1700.0">1700 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
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, coll.
|
||
<collectorName id="3C41A8B2CD23E81AFE4D91DEFDE6FA14" box="[456,622,1467,1489]" pageId="1" pageNumber="290">Nguyen Duc Anh</collectorName>
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,
|
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<date id="E50AEBA4CD23E81AFDF291DEFD67FA15" box="[631,751,1467,1489]" pageId="1" pageNumber="290" value="2004-03-25">
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||
<collectingDate id="F54E124CCD23E81AFDF291DEFD67FA15" box="[631,751,1467,1489]" pageId="1" pageNumber="290" value="2004-03-25">25 Mar.2004</collectingDate>
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||
</date>
|
||
–
|
||
<date id="E50AEBA4CD23E81AFF0791BDFF71FA28" box="[130,249,1496,1517]" pageId="1" pageNumber="290" value="2004-04-11">
|
||
<collectingDate id="F54E124CCD23E81AFF0791BDFF71FA28" box="[130,249,1496,1517]" pageId="1" pageNumber="290" value="2004-04-11">11 Apr.2004</collectingDate>
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||
</date>
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||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="21DCC739CD23E81AFE8491BDFE25F981" collectingDate="2006-03-21" collectingDateMax="2006-04-09" collectingDateMin="2006-03-21" collectorName="Nguyen Duc Anh" elevation="2000" latitude="15.3" location="Ngoc Linh Mountain" longLatPrecision="1285" longitude="8.016666" municipality="Dak Glei District" pageId="1" pageNumber="290" specimenCount="6" specimenCount-female="2" specimenCount-male="4" stateProvince="Kon Tum" typeStatus="paratype">
|
||
<specimenCount id="87B206EDCD23E81AFE8491BDFEC4FA28" box="[257,332,1495,1517]" pageId="1" pageNumber="290" type="male">4 males</specimenCount>
|
||
,
|
||
<specimenCount id="87B206EDCD23E81AFED091BDFE38FA28" box="[341,432,1495,1517]" pageId="1" pageNumber="290" type="female">2 females</specimenCount>
|
||
(
|
||
<emphasis id="A3C01176CD23E81AFE3891B2FDABFA29" bold="true" box="[445,547,1495,1516]" pageId="1" pageNumber="290">IEBR-102</emphasis>
|
||
),
|
||
<collectingRegion id="53700386CD23E81AFDB391BDFD65FA28" box="[566,749,1495,1517]" country="Vietnam" name="Kon Tum" pageId="1" pageNumber="290">Kon Tum Province</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="716F571ECD23E81AFFF49191FE9FF9CF" box="[113,279,1524,1546]" pageId="1" pageNumber="290">Dak Glei District</collectingMunicipality>
|
||
,
|
||
<location id="946B9BBFCD23E81AFEA59190FE60F9CF" LSID="urn:lsid:plazi:treatment:191D7C72CD23E81FFF7D90F4FF23FD10:946B9BBFCD23E81AFEA59190FE60F9CF" box="[288,488,1524,1546]" latitude="15.3" longLatPrecision="1285" longitude="8.016666" municipality="Dak Glei District" name="Ngoc Linh Mountain" pageId="1" pageNumber="290" stateProvince="Kon Tum">Ngoc Linh Mountain</location>
|
||
(15°00'–
|
||
<geoCoordinate id="F480ABA3CD23E81AFDC79191FD19F9CF" box="[578,657,1524,1546]" degrees="15" direction="north" minutes="18" orientation="latitude" pageId="1" pageNumber="290" precision="925" value="15.3">15°18'N</geoCoordinate>
|
||
, 107°41'–
|
||
<geoCoordinate id="F480ABA3CD23E81AFFF49274FF49F9E2" box="[113,193,1553,1575]" degrees="08" direction="east" minutes="01" orientation="longitude" pageId="1" pageNumber="290" precision="925" value="8.016666">08°01'E</geoCoordinate>
|
||
), primary forest,
|
||
<elevation id="1A992A57CD23E81AFEF49274FDB9F9E2" box="[369,561,1553,1575]" metricMagnitude="3" metricUnit="m" metricValue="2.0" metricValueMax="2.1" metricValueMin="1.9" pageId="1" pageNumber="290" unit="m" value="2000.0" valueMax="2100.0" valueMin="1900.0">
|
||
<quantity id="564C6081CD23E81AFEF49274FE72F9E2" box="[369,506,1553,1575]" metricMagnitude="3" metricUnit="m" metricValue="2.0" metricValueMax="2.1" metricValueMin="1.9" pageId="1" pageNumber="290" unit="m" value="2000.0" valueMax="2100.0" valueMin="1900.0">1900–2100 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
, coll.
|
||
<collectorName id="3C41A8B2CD23E81AFDF79277FF15F981" pageId="1" pageNumber="290">Nguyen Duc Anh</collectorName>
|
||
,
|
||
<date id="E50AEBA4CD23E81AFF22924BFEACF981" box="[167,292,1582,1604]" pageId="1" pageNumber="290" value="2006-03-21">
|
||
<collectingDate id="F54E124CCD23E81AFF22924BFEACF981" box="[167,292,1582,1604]" pageId="1" pageNumber="290" value="2006-03-21">21 Mar.2006</collectingDate>
|
||
</date>
|
||
–
|
||
<date id="E50AEBA4CD23E81AFEBB924BFE25F981" box="[318,429,1582,1604]" pageId="1" pageNumber="290" value="2006-04-09">
|
||
<collectingDate id="F54E124CCD23E81AFEBB924BFE25F981" box="[318,429,1582,1604]" pageId="1" pageNumber="290" value="2006-04-09">9 Apr.2006</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D9AE9EEFCD23E81AFFF49209FE58F8C1" pageId="1" pageNumber="290" type="diagnosis">
|
||
<paragraph id="910BCD64CD23E81AFFF49209FE58F8C1" blockId="1.[113,752,1644,1796]" pageId="1" pageNumber="290">
|
||
<emphasis id="A3C01176CD23E81AFFF49209FF6DF941" bold="true" box="[113,229,1644,1668]" italics="true" pageId="1" pageNumber="290">Diagnosis.</emphasis>
|
||
— The species is very similar to
|
||
<taxonomicName id="56B4B6E7CD23E81AFDD59208FD78F941" baseAuthorityName="Attems" baseAuthorityYear="1937" box="[592,752,1645,1668]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="290" phylum="Arthropoda" rank="species" species="hilaris">
|
||
<emphasis id="A3C01176CD23E81AFDD59208FD78F941" box="[592,752,1645,1668]" italics="true" pageId="1" pageNumber="290">Tylopus hilaris</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="56B4B6E7CD23E81AFF2392E8FEB3F961" authorityName="Golovatch" authorityYear="1984" box="[166,315,1677,1700]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="290" phylum="Arthropoda" rank="species" species="hilaroides">
|
||
<emphasis id="A3C01176CD23E81AFF2392E8FEB3F961" box="[166,315,1677,1700]" italics="true" pageId="1" pageNumber="290">T. hilaroides</emphasis>
|
||
</taxonomicName>
|
||
in general appearance and gonopod conformation, but it can be distinguished from them in having the gonopod process
|
||
<emphasis id="A3C01176CD23E81AFED992A9FEE3F921" bold="true" box="[348,363,1740,1764]" pageId="1" pageNumber="290">h</emphasis>
|
||
with two branches (one spiniform, and the other a serrated lamina).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D9AE9EEFCD23E81AFFF4934EFD5CF841" pageId="1" pageNumber="290" type="etymology">
|
||
<paragraph id="910BCD64CD23E81AFFF4934EFD5CF841" blockId="1.[113,751,1835,1925]" pageId="1" pageNumber="290">
|
||
<emphasis id="A3C01176CD23E81AFFF4934EFF64F881" bold="true" box="[113,236,1835,1860]" italics="true" pageId="1" pageNumber="290">Etymology.</emphasis>
|
||
— “
|
||
<taxonomicName id="56B4B6E7CD23E81AFEA49348FE12F881" authorityName="Nguyen" authorityYear="2012" box="[289,410,1837,1860]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="290" phylum="Arthropoda" rank="species" species="phanluongi">
|
||
<emphasis id="A3C01176CD23E81AFEA49348FE12F881" box="[289,410,1837,1860]" italics="true" pageId="1" pageNumber="290">phanluongi</emphasis>
|
||
</taxonomicName>
|
||
”, named in honour of Dr. Phan Luong of the Vietnamese-Russian Tropical Center for his major contribution to the ecological surveys in
|
||
<collectingCountry id="E9A38DF4CD23E81AFDF79309FD58F841" box="[626,720,1900,1924]" name="Vietnam" pageId="1" pageNumber="290">Vietnam</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D9AE9EEFCD23E81FFFF493C9FD45FDB0" lastPageId="4" lastPageNumber="293" pageId="1" pageNumber="290" type="description">
|
||
<paragraph id="910BCD64CD23E81AFFF493C9FD2AF7E1" blockId="1.[113,752,1964,2084]" pageId="1" pageNumber="290">
|
||
<emphasis id="A3C01176CD23E81AFFF493C9FF7FF801" bold="true" box="[113,247,1964,1988]" italics="true" pageId="1" pageNumber="290">Description.</emphasis>
|
||
— Size: Body length 37–[40]–
|
||
<quantity id="564C6081CD23E81AFDCA93C9FD13F801" box="[591,667,1964,1988]" metricMagnitude="-2" metricUnit="m" metricValue="4.1" pageId="1" pageNumber="290" unit="mm" value="41.0">41 mm</quantity>
|
||
(male),
|
||
<quantity id="564C6081CD23E81AFFF493A9FF6DF821" box="[113,229,1996,2020]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="4.2" metricValueMin="3.8" pageId="1" pageNumber="290" unit="mm" value="40.0" valueMax="42.0" valueMin="38.0">38–42 mm</quantity>
|
||
(female), width of midbody prozona 3.1–[3.3]–
|
||
<quantity id="564C6081CD23E81AFFF49389FF40F7C1" box="[113,200,2028,2052]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="1" pageNumber="290" unit="mm" value="3.4">3.4 mm</quantity>
|
||
(male) and
|
||
<quantity id="564C6081CD23E81AFED39389FE55F7C1" box="[342,477,2028,2052]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" metricValueMax="3.6" metricValueMin="3.2" pageId="1" pageNumber="290" unit="mm" value="3.4" valueMax="3.6" valueMin="3.2">3.2–3.6 mm</quantity>
|
||
(female), and metazona 4.1–[4.3]–
|
||
<quantity id="564C6081CD23E81AFF5B9C69FEBAF7E1" box="[222,306,2060,2084]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="1" pageNumber="290" unit="mm" value="4.4">4.4 mm</quantity>
|
||
(male) and
|
||
<quantity id="564C6081CD23E81AFE329C69FDB0F7E1" box="[439,568,2060,2084]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" metricValueMax="4.6" metricValueMin="4.2" pageId="1" pageNumber="290" unit="mm" value="4.4" valueMax="4.6" valueMin="4.2">4.2–4.6 mm</quantity>
|
||
(female).
|
||
</paragraph>
|
||
<paragraph id="910BCD64CD23E81AFCA990F6FBD4FAEE" blockId="1.[812,1452,1171,2091]" pageId="1" pageNumber="290">Colouration: Body generally castaneous brown, but margin of posterior part of prozona, anterior area of metazona, metatergal transverse sulcus, distal part of antennomere 6, and whole antennomere 7 blackish brown. Legs, paraterga and sterna brownish yellow.</paragraph>
|
||
<paragraph id="910BCD64CD23E81AFCA99156FB7AFA0E" blockId="1.[812,1452,1171,2091]" pageId="1" pageNumber="290">
|
||
Head: Slightly broader than collum. Region between antennal sockets with sparse setae, labrum more setose. Frons slightly convex, divided into two parts by a distinct epicranial suture. Antenna long, claviform, reaching body ring 4 laterally. Antennomere 3=4=5>6=2>1&
|
||
<quantity id="564C6081CD23E81AFBF091D6FB28FA0E" box="[1141,1184,1459,1483]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="1" pageNumber="290" unit="in" value="7.0">7 in</quantity>
|
||
length.
|
||
</paragraph>
|
||
<paragraph id="910BCD64CD23E81AFCA991B6FAC8F9AE" blockId="1.[812,1452,1171,2091]" pageId="1" pageNumber="290">Collum: Slightly narrower than body ring 2; surface shining, finely rugose in anterior half and strongly rugose in posterior half, without granules or tubercles, with a row of 4+4 setae near anterior margin. Transverse sulcus vestigial, partly obliterated. Paratergum minute, sub-semicircular.</paragraph>
|
||
<paragraph id="910BCD64CD23E81AFCA99216FC3BF84E" blockId="1.[812,1452,1171,2091]" pageId="1" pageNumber="290">
|
||
Body ring 3<2=4<
|
||
<quantity id="564C6081CD23E81AFC7C9216FBACF94E" box="[1017,1060,1651,1675]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="1" pageNumber="290" unit="in" value="5.0">5 in</quantity>
|
||
width, body parallel-sided on rings 5–16, thereafter gradually tapering towards telson. Surface of prozona shining, smooth. Metatergal surface shining, slightly rugulose on anterior half and considerably rugose on posterior half. Metatergal transverse sulcus present on body rings 5–19, finely striate, reaching base of paraterga (
|
||
<figureCitation id="098FD1E1CD23E81AFCB19356FC03F88E" box="[820,907,1843,1867]" captionStart="Fig" captionStartId="2.[137,172,2059,2081]" captionTargetBox="[139,1480,356,1904]" captionTargetId="figure-8@2.[107,1493,337,1932]" captionTargetPageId="2" captionText="Fig. 1. Tylopus phanluongi, new species, holotype: body ring 10, dorsal view (A); hypoproct, ventral view (B); sternite 5, posterioventral view (C); leg 9 (D); right gonopod, mesal view (E)and lateral view (F). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5347273" httpUri="https://zenodo.org/record/5347273/files/figure.png" pageId="1" pageNumber="290">Fig. 1A</figureCitation>
|
||
). Metaterga of body rings 2–16 with rows of 2+2 and of 3+3 setae in front of and behind transverse sulcus, respectively.
|
||
</paragraph>
|
||
<paragraph id="910BCD64CD23E81AFCA993F6FADAF7CE" blockId="1.[812,1452,1171,2091]" pageId="1" pageNumber="290">Metaterga: Body rings 17–18 with three rows of 2+2, 1+1 and 4+4 setae; second and third rows located behind transverse sulcus. Metatergum 19 with rows of 2+2 and 5+5 setae in front of and behind transverse sulcus, respectively.</paragraph>
|
||
<paragraph id="910BCD64CD23E818FCA99C76FEB7FEF0" blockId="1.[812,1452,1171,2091]" lastBlockId="3.[113,752,189,469]" lastPageId="3" lastPageNumber="292" pageId="1" pageNumber="290">Pleura: With dense microgranules. Pleurosternal keels small, beak-like or spiniform on body rings 2–5, absent on subsequent body rings. Axial line evident only in anterior half of metaterga. Waist between pro- and metazona narrow, deep, finely striate.</paragraph>
|
||
<caption id="C5CB9DECCD20E819FF0C9C6EFB81F7F8" ID-DOI="http://doi.org/10.5281/zenodo.5347273" ID-Zenodo-Dep="5347273" httpUri="https://zenodo.org/record/5347273/files/figure.png" pageId="2" pageNumber="291" startId="2.[137,172,2059,2081]" targetBox="[139,1480,356,1904]" targetPageId="2">
|
||
<paragraph id="910BCD64CD20E819FF0C9C6EFB81F7F8" blockId="2.[137,1474,2059,2110]" pageId="2" pageNumber="291">
|
||
Fig. 1.
|
||
<taxonomicName id="56B4B6E7CD20E819FF4A9C69FE05F7E5" authority="Nguyen, 2012" authorityName="Nguyen" authorityYear="2012" box="[207,397,2060,2080]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="291" phylum="Arthropoda" rank="species" species="phanluongi" status="sp. nov.">
|
||
<emphasis id="A3C01176CD20E819FF4A9C69FE05F7E5" box="[207,397,2060,2080]" italics="true" pageId="2" pageNumber="291">Tylopus phanluongi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="B8F3AC0DCD20E819FE1E9C69FD9AF7E4" box="[411,530,2059,2081]" pageId="2" pageNumber="291" rank="species">new species</taxonomicNameLabel>
|
||
, holotype: body ring 10, dorsal view (A); hypoproct, ventral view (B); sternite 5, posterioventral view (C); leg 9 (D); right gonopod, mesal view (E)and lateral view (F). Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="910BCD64CD21E818FFF49558FAC0FF30" blockId="3.[113,752,189,469]" lastBlockId="3.[812,1452,189,469]" pageId="3" pageNumber="292">
|
||
Paraterga: Well developed, slightly lower than metatergal surface, beak-like on body rings 5–14, more protruding on body rings 15–19. Paraterga surpassing posterior contour of metaterga. Calluses small, with one or two setiferous incisions on lateral side of poreless and pore-bearing paraterga, respectively. Ozopores located on lateral side of paraterga of body rings 5, 7, 9–10, 12–13, 15–19 (
|
||
<figureCitation id="098FD1E1CD21E818FB6094B8FAB3FF30" box="[1253,1339,221,245]" captionStart="Fig" captionStartId="2.[137,172,2059,2081]" captionTargetBox="[139,1480,356,1904]" captionTargetId="figure-8@2.[107,1493,337,1932]" captionTargetPageId="2" captionText="Fig. 1. Tylopus phanluongi, new species, holotype: body ring 10, dorsal view (A); hypoproct, ventral view (B); sternite 5, posterioventral view (C); leg 9 (D); right gonopod, mesal view (E)and lateral view (F). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5347273" httpUri="https://zenodo.org/record/5347273/files/figure.png" pageId="3" pageNumber="292">Fig. 1A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="910BCD64CD21E818FCA99498FCD2FEB0" blockId="3.[812,1452,189,469]" pageId="3" pageNumber="292">
|
||
Telson: Epiproct short, broadly truncated, with two small terminal tubercles. Tip with four spinnerets. Hypoproct trapeziform, with two distolateral setiferous knobs (
|
||
<figureCitation id="098FD1E1CD21E818FAFA9558FCC5FEB0" captionStart="Fig" captionStartId="2.[137,172,2059,2081]" captionTargetBox="[139,1480,356,1904]" captionTargetId="figure-8@2.[107,1493,337,1932]" captionTargetPageId="2" captionText="Fig. 1. Tylopus phanluongi, new species, holotype: body ring 10, dorsal view (A); hypoproct, ventral view (B); sternite 5, posterioventral view (C); leg 9 (D); right gonopod, mesal view (E)and lateral view (F). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5347273" httpUri="https://zenodo.org/record/5347273/files/figure.png" pageId="3" pageNumber="292">Fig. 1B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="910BCD64CD21E818FCA99518FAC3FE10" blockId="3.[812,1452,189,469]" pageId="3" pageNumber="292">
|
||
Sterna densely setose, without any modification; transverse and longitudinal sutures distinct. Male sternum 5 with a setiferous bifid lamina between coxae 4 (
|
||
<figureCitation id="098FD1E1CD21E818FB6C95D8FAB6FE10" box="[1257,1342,445,469]" captionStart="Fig" captionStartId="2.[137,172,2059,2081]" captionTargetBox="[139,1480,356,1904]" captionTargetId="figure-8@2.[107,1493,337,1932]" captionTargetPageId="2" captionText="Fig. 1. Tylopus phanluongi, new species, holotype: body ring 10, dorsal view (A); hypoproct, ventral view (B); sternite 5, posterioventral view (C); leg 9 (D); right gonopod, mesal view (E)and lateral view (F). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5347273" httpUri="https://zenodo.org/record/5347273/files/figure.png" pageId="3" pageNumber="292">Fig. 1C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="C5CB9DECCD21E818FFF49C49FA89F787" ID-DOI="http://doi.org/10.5281/zenodo.5347275" ID-Zenodo-Dep="5347275" box="[113,1281,2092,2114]" httpUri="https://zenodo.org/record/5347275/files/figure.png" pageId="3" pageNumber="292" startId="3.[113,148,2092,2114]" targetBox="[113,1452,549,2054]" targetPageId="3">
|
||
<paragraph id="910BCD64CD21E818FFF49C49FA89F787" blockId="3.[113,1281,2092,2114]" box="[113,1281,2092,2114]" pageId="3" pageNumber="292">
|
||
Fig. 2.
|
||
<taxonomicName id="56B4B6E7CD21E818FF329C48FEFEF784" authority="Nguyen, 2012" authorityName="Nguyen" authorityYear="2012" box="[183,374,2093,2113]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="292" phylum="Arthropoda" rank="species" species="phanluongi" status="sp. nov.">
|
||
<emphasis id="A3C01176CD21E818FF329C48FEFEF784" box="[183,374,2093,2113]" italics="true" pageId="3" pageNumber="292">Tylopus phanluongi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="B8F3AC0DCD21E818FE069C49FE72F787" box="[387,506,2092,2114]" pageId="3" pageNumber="292" rank="species">new species</taxonomicNameLabel>
|
||
, holotype: right gonopod, mesal view (A); postfemoral region, mesal view (B).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="910BCD64CD26E81FFF0D94DBFF7BFEF0" blockId="4.[136,775,189,629]" pageId="4" pageNumber="293">
|
||
Legs: Slender, about 1.8–2 times as long as midbody height, without tarsal brushes. Prefemora swollen dorsally. Ventral part of prefemora and femora with dense microtubercles (
|
||
<figureCitation id="098FD1E1CD26E81FFF159578FF6DFEF0" box="[144,229,285,309]" captionStart="Fig" captionStartId="2.[137,172,2059,2081]" captionTargetBox="[139,1480,356,1904]" captionTargetId="figure-8@2.[107,1493,337,1932]" captionTargetPageId="2" captionText="Fig. 1. Tylopus phanluongi, new species, holotype: body ring 10, dorsal view (A); hypoproct, ventral view (B); sternite 5, posterioventral view (C); leg 9 (D); right gonopod, mesal view (E)and lateral view (F). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5347273" httpUri="https://zenodo.org/record/5347273/files/figure.png" pageId="4" pageNumber="293">Fig. 1D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="910BCD64CD26E81FFF0D9558FD45FDB0" blockId="4.[136,775,189,629]" pageId="4" pageNumber="293">
|
||
Gonopods (
|
||
<figureCitation id="098FD1E1CD26E81FFE8D9558FEF4FE90" box="[264,380,317,341]" captionStart="Fig" captionStartId="2.[137,172,2059,2081]" captionTargetBox="[139,1480,356,1904]" captionTargetId="figure-8@2.[107,1493,337,1932]" captionTargetPageId="2" captionText="Fig. 1. Tylopus phanluongi, new species, holotype: body ring 10, dorsal view (A); hypoproct, ventral view (B); sternite 5, posterioventral view (C); leg 9 (D); right gonopod, mesal view (E)and lateral view (F). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5347273" httpUri="https://zenodo.org/record/5347273/files/figure.png" pageId="4" pageNumber="293">Figs. 1E,F</figureCitation>
|
||
,
|
||
<figureCitation id="098FD1E1CD26E81FFE0E9558FE41FE90" box="[395,457,317,341]" captionStart="Fig" captionStartId="3.[113,148,2092,2114]" captionTargetBox="[113,1452,549,2054]" captionTargetId="figure-80@3.[113,1453,548,2054]" captionTargetPageId="3" captionText="Fig. 2. Tylopus phanluongi, new species, holotype: right gonopod, mesal view (A); postfemoral region, mesal view (B)." figureDoi="http://doi.org/10.5281/zenodo.5347275" httpUri="https://zenodo.org/record/5347275/files/figure.png" pageId="4" pageNumber="293">2A,B</figureCitation>
|
||
): Coxite cylindrical, slightly shorter than femorite; distoventral part sparsely setose. Prefemorite densely setose, set off from femorite by an oblique sulcus on lateral side. Femorite stout, expanded distally. Postfemoral region demarcated from femorite by a distinct sulcus laterally. Lamina
|
||
<emphasis id="A3C01176CD26E81FFE0A95B8FE1EFE30" bold="true" box="[399,406,477,501]" pageId="4" pageNumber="293">l</emphasis>
|
||
present, somewhat overlapped by a well-developed, serrated process
|
||
<emphasis id="A3C01176CD26E81FFD839598FD90FDD0" bold="true" box="[518,536,509,533]" pageId="4" pageNumber="293">z.</emphasis>
|
||
Process
|
||
<emphasis id="A3C01176CD26E81FFDFE9598FD02FDD0" bold="true" box="[635,650,509,533]" pageId="4" pageNumber="293">h</emphasis>
|
||
short, with 2 branches, one longer and spiniform, the other shorter and slightly serrated. Lobes
|
||
<emphasis id="A3C01176CD26E81FFEFA9658FE1DFD90" bold="true" box="[383,405,573,597]" pageId="4" pageNumber="293">m</emphasis>
|
||
and
|
||
<emphasis id="A3C01176CD26E81FFE409658FE5CFD90" bold="true" box="[453,468,573,597]" pageId="4" pageNumber="293">n</emphasis>
|
||
absent. Solenophore strongly coiled, completely sheathing flagelliform solenomere.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D9AE9EEFCD26E81FFF0D96F9FF23FD10" pageId="4" pageNumber="293" type="distribution">
|
||
<paragraph id="910BCD64CD26E81FFF0D96F9FF23FD10" blockId="4.[136,775,668,725]" pageId="4" pageNumber="293">
|
||
<emphasis id="A3C01176CD26E81FFF0D96F9FE9DFD70" bold="true" box="[136,277,668,693]" italics="true" pageId="4" pageNumber="293">Distribution.</emphasis>
|
||
— Known only from the
|
||
<typeStatus id="4E0F73C6CD26E81FFDC596FBFDE6FD73" box="[576,622,670,694]" pageId="4" pageNumber="293">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="098FD1E1CD26E81FFD5E96F8FF2BFD10" captionStart="Fig" captionStartId="21.[113,147,1802,1824]" captionTargetBox="[113,753,1083,1781]" captionTargetId="figure-736@21.[113,753,1083,1779]" captionTargetPageId="21" captionText="Fig. 18.Distribution map ofthe millipede genus Tylopus Jeekel, 1968 in Vietnam. 1: Sa Pa, Lao Cai Prov. (T. crassipes, T. procurvus, T. nodulipes, T. sigma, T. sapaensis, T. maculatus, T. magnicus); 2: Van Ban, Lao Cai Prov. (T. crassipes, T. nodulipes); 3: Xuan Son NP (T. strongylosomoides, T. golovatchi); 4: Tam Dao NP (T. tamdaoensis, T. roseiparaterga, T. strongylosomoides); 5: Cuc Phuong NP (T. granulatus, T. hilaroides, T. topali); 6: Pu Mat NP (T. nodulipes); 7: Huong Son, Ha Tinh prov. (T. nodulipes); 8: Bach Ma–Ba Na NP (T. hilaris, T. multilatus); 9: Ngoc Linh Mts., Kon Tum prov. (T. hilaris, T. phanluongi); 10: Bi Doup–Nui Ba NP (T. spinisterna)." figureDoi="http://doi.org/10.5281/zenodo.5347309" httpUri="https://zenodo.org/record/5347309/files/figure.png" pageId="4" pageNumber="293">Fig. 18</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |