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<document id="78936B4747714FF3322AC8AD8F60A8E3" ID-CLB-Dataset="22378" ID-DOI="10.5852/ejt.2018.437" ID-GBIF-Dataset="29c5b07d-8712-4d58-9666-850f5761b080" ID-Zenodo-Dep="3787887" ID-ZooBank="AF27DA05-746B-459D-AC8C-4F059EAA4146" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588707111488" checkinUser="valdenar" docAuthor="Nunes, Luis Gabriel de O. A., Nunes, Rafael V. &amp; Vaz-De, Fernando Z." docDate="2018" docId="191B87A8FF8E3C26FE67FE6BFD52D837" docLanguage="en" docName="ejt-437_nunes_nunes_vaz-de-mello.pdf.imf" docOrigin="European Journal of Taxonomy 437" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Canthon (Goniocanthon) bicolor Castelnau 1840" docType="treatment" docVersion="6" lastPageNumber="24" masterDocId="E522FFD0FF9E3C3FFF97FFF9FFEDD95B" masterDocTitle="Taxonomic revision of the South American subgenus Canthon (Goniocanthon) Pereira &amp; Martínez, 1956 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini)" masterLastPageNumber="31" masterPageNumber="1" pageNumber="17" updateTime="1698838631990" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="78C7330009DAC8F194FFACDEFD7B218C">Taxonomic revision of the South American subgenus Canthon (Goniocanthon) Pereira &amp; Martínez, 1956 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini)</mods:title>
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<mods:namePart id="E2C163EB36B5DE72B6DCB9B936CDEF9C">Nunes, Luis Gabriel de O. A.</mods:namePart>
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<mods:namePart id="6323C9CCA67CC785E9D7BB60383FC0CB">Nunes, Rafael V.</mods:namePart>
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<mods:affiliation id="CDAFCD9D37A199B52C922EFF320E3F67">Universidade Federal de Mato Grosso, Instituto de Biociências, Programa de Pós Graduação em Ecologia e Conservação da Biodiversidade, Av. Fernando Correa da Costa, 2367, Boa Esperança, Cuiabá-MT, 78060 - 900, Brazil. Universidade Federal de Mato Grosso, Instituto de Biociências, Departamento de Biologia e Zoologia, Av. Fernando Correa da Costa, 2367, Boa Esperança, Cuiabá-MT, 78060 - 900, Brazil.</mods:affiliation>
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<paragraph id="910D36BEFF8E3C2FFE67FE6BFBA9D8F6" blockId="16.[496,1092,402,465]" box="[496,1092,402,429]" pageId="16" pageNumber="17">
<heading id="CA4581D2FF8E3C2FFE67FE6BFBA9D8F6" box="[496,1092,402,429]" centered="true" fontSize="11" level="2" pageId="16" pageNumber="17" reason="2">
<taxonomicName id="56B24D3DFF8E3C2FFE67FE6BFBA9D8F6" ID-CoL="68XHJ" authority="Castelnau, 1840" authorityName="Castelnau" authorityYear="1840" box="[496,1092,402,429]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="bicolor" subGenus="Goniocanthon">
<emphasis id="A3C6EAACFF8E3C2FFE67FE6BFC92D8F6" bold="true" box="[496,895,402,429]" italics="true" pageId="16" pageNumber="17">Canthon (Goniocanthon) bicolor</emphasis>
<bibRefCitation id="F5234B4FFF8E3C2FFC10FE6AFBA9D8F6" author="Castelnau Comte" box="[903,1092,403,429]" pageId="16" pageNumber="17" refId="ref16713" refString="Castelnau Comte de. 1840. Histoire naturelle des insectes coleopteres. Tome Deuxieme. Societe bibliophile, Paris. https: // doi. org / 10.5962 / bhl. title. 36538" type="book" year="1840">Castelnau, 1840</bibRefCitation>
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<paragraph id="910D36BEFF8E3C2FFD70FE4EFCA0D88A" blockId="16.[496,1092,402,465]" box="[743,845,439,465]" pageId="16" pageNumber="17">
<figureCitation id="09892A3BFF8E3C2FFD70FE4EFCDFD88A" box="[743,818,439,465]" captionStart="Fig" captionStartId="19.[189,232,1894,1920]" captionTargetBox="[202,1379,264,1882]" captionTargetId="figure@19.[189,1399,263,1882]" captionTargetPageId="19" captionText="Fig. 4. Canthon (G.) bicolor Castelnau, 1840. A. Habitus, dorsal view. B. Ventral surface of meso- and metafemora and metasternal anterior lobe not convex. C. Dorsal view showing convex pygidium. D. Male protibial spur. E. Female protibial spur. F. Paramera lateral view. G. Paramera dorsal view. Scale bar: A = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3787895" httpUri="https://zenodo.org/record/3787895/files/figure.png" pageId="16" pageNumber="17">Figs 4</figureCitation>
, 
<figureCitation id="09892A3BFF8E3C2FFCA9FE4EFCA0D88A" box="[830,845,439,465]" captionStart="Fig" captionStartId="23.[189,232,1861,1887]" captionTargetBox="[191,1287,1049,1818]" captionTargetId="figure@23.[189,1399,1049,1820]" captionTargetPageId="23" captionText="Fig. 8. Distribution map for subspecies of Goniocanthon: C. (G.) fulgidus martinezi subsp. nov. (green triangle); C. (G.) fulgidus pereirai subsp. nov. (red triangle); C. (G.) fulgidus fulgidus Redtenbacher, 1868 (yellow triangle); C. (G.) smaragdulus subviridis Schmidt, 1922 stat. rev. (green circle); C. (G.) smaragdulus smaragdulus (Fabricius, 1781) (blue circle) and C. bicolor Castelnau, 1840 (brown square)." figureDoi="http://doi.org/10.5281/zenodo.3787905" httpUri="https://zenodo.org/record/3787905/files/figure.png" pageId="16" pageNumber="17">8</figureCitation>
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<treatmentCitation id="101310AFFF8E3C2FFF2AFE00FD9EDB48" author="Castelnau Comte" box="[189,627,505,531]" page="69" pageId="16" pageNumber="17" year="1840">
<taxonomicName id="56B24D3DFF8E3C2FFF2AFE00FD9EDB48" ID-CoL="68XHJ" authority="Castelnau, 1840: 69" authorityName="Castelnau" authorityPageNumber="69" authorityYear="1840" box="[189,627,505,531]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="A3C6EAACFF8E3C2FFF2AFE00FE93DB48" box="[189,382,505,531]" italics="true" pageId="16" pageNumber="17">Canthon bicolor</emphasis>
<bibRefCitation id="F5234B4FFF8E3C2FFE12FE00FD9EDB48" author="Castelnau Comte" box="[389,627,505,531]" pageId="16" pageNumber="17" refId="ref16713" refString="Castelnau Comte de. 1840. Histoire naturelle des insectes coleopteres. Tome Deuxieme. Societe bibliophile, Paris. https: // doi. org / 10.5962 / bhl. title. 36538" type="book" year="1840">Castelnau, 1840: 69</bibRefCitation>
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.
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<paragraph id="910D36BEFF8E3C2FFF2AFDC2FBC9DB99" blockId="16.[189,1399,571,706]" pageId="16" pageNumber="17">
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<taxonomicName id="56B24D3DFF8E3C2FFF2AFDC2FE93DB0E" ID-CoL="68XHJ" authorityName="Castelnau" authorityYear="1840" box="[189,382,571,597]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="A3C6EAACFF8E3C2FFF2AFDC2FE93DB0E" box="[189,382,571,597]" italics="true" pageId="16" pageNumber="17">Canthon bicolor</emphasis>
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– 
<treatmentCitation id="101310AFFF8E3C2FFE0EFDC2FDB6DB0E" author="Harold E." box="[409,603,571,597]" page="14" pageId="16" pageNumber="17" year="1868">
<bibRefCitation id="F5234B4FFF8E3C2FFE0EFDC2FDB6DB0E" author="Harold E." box="[409,603,571,597]" pageId="16" pageNumber="17" pagination="1 - 144" refId="ref17251" refString="Harold E. 1868. Monographie der Gattung Canthon. Berliner entomologische Zeitschrift 12: 1 - 144." type="journal article" year="1868">Harold 1868: 14</bibRefCitation>
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, 71 (key). — 
<treatmentCitation id="101310AFFF8E3C2FFD6DFDC2FB9ADB0E" author="Gemminger M. &amp; Harold E." box="[762,1143,571,597]" page="990" pageId="16" pageNumber="17" year="1869">
<bibRefCitation id="F5234B4FFF8E3C2FFD6DFDC2FB9ADB0E" author="Gemminger M. &amp; Harold E." box="[762,1143,571,597]" pageId="16" pageNumber="17" refId="ref17044" refString="Gemminger M. &amp; Harold E. 1869. Catalogus coleopterorum hucusque descriptorum synonymicus et systematicus. Tome IV: Scarabaeidae. Sumptu E. H. Gummi, Monaco. https: // doi. org / 10.5962 / bhl. title. 9089" type="book" year="1869">Gemminger &amp; Harold 1869: 990</bibRefCitation>
</treatmentCitation>
(catalogue). — 
<treatmentCitation id="101310AFFF8E3C2FFAA3FDC2FEBDDB22" author="Gillet J. J. E." page="28" pageId="16" pageNumber="17" year="1911">
<bibRefCitation id="F5234B4FFF8E3C2FFAA3FDC2FEBDDB22" author="Gillet J. J. E." pageId="16" pageNumber="17" refId="ref17089" refString="Gillet J. J. E. 1911. Coleopterorum catalogus. Pars 38: Scarabaeidae: Coprinae. I. W. Junk, Berlin." type="book" year="1911">Gillet 1911: 28</bibRefCitation>
</treatmentCitation>
(catalogue). — 
<treatmentCitation id="101310AFFF8E3C2FFD9AFDA6FD0DDB22" author="Schmidt A." box="[525,736,607,633]" page="64" pageId="16" pageNumber="17" year="1922">
<bibRefCitation id="F5234B4FFF8E3C2FFD9AFDA6FD0DDB22" author="Schmidt A." box="[525,736,607,633]" pageId="16" pageNumber="17" pagination="61 - 103" refId="ref18074" refString="Schmidt A. 1922. 1. Bestimmungstabelle der mir bekannten Canthon- Arten. 2. Verbreitungsgebiete der Canthon - Arten. 3. Neubeschreibungen von Canthon, Saprosites, Mendidius, Euparia und Ataenius. Archiv fur Naturgeschichte 88 (3): 61 - 103." type="journal article" year="1922">Schmidt 1922: 64</bibRefCitation>
</treatmentCitation>
, 73 (key). — 
<treatmentCitation id="101310AFFF8E3C2FFC17FDA6FB86DB22" author="Balthasar V." box="[896,1131,607,633]" page="189" pageId="16" pageNumber="17" year="1939">
<bibRefCitation id="F5234B4FFF8E3C2FFC17FDA6FB86DB22" author="Balthasar V." box="[896,1131,607,633]" pageId="16" pageNumber="17" pagination="179 - 238" refId="ref16169" refString="Balthasar V. 1939. Eine Vorstudie zur Monographie der Gattung Canthon Hffsg. Folia Zoologica et Hydrobiologica 9 (2): 179 - 238." type="journal article" year="1939">Balthasar 1939: 189</bibRefCitation>
</treatmentCitation>
(key). — 
<treatmentCitation id="101310AFFF8E3C2FFB75FDA6FE8CDBC6" author="Blackwelder R." page="198" pageId="16" pageNumber="17" year="1944">
<bibRefCitation id="F5234B4FFF8E3C2FFB75FDA6FE8CDBC6" author="Blackwelder R." pageId="16" pageNumber="17" pagination="189 - 341" refId="ref16238" refString="Blackwelder R. 1944. Checklist of the coleopterous insects of Mexico, Central America the West Indies and South America. Part 2. Bulletin of the United States National Museum 185: 189 - 341." type="journal article" year="1944">Blackwelder 1944: 198</bibRefCitation>
</treatmentCitation>
(catalogue). — 
<treatmentCitation id="101310AFFF8E3C2FFDB5FD7AFC90DBC6" author="Vulcano M. A. &amp; Pereira F. S." box="[546,893,643,669]" page="604" pageId="16" pageNumber="17" year="1964">
<bibRefCitation id="F5234B4FFF8E3C2FFDB5FD7AFC90DBC6" author="Vulcano M. A. &amp; Pereira F. S." box="[546,893,643,669]" pageId="16" pageNumber="17" pagination="570 - 685" refId="ref18506" refString="Vulcano M. A. &amp; Pereira F. S. 1964. Catalogue of the Canthonini (Col. Scarab.) inhabiting the Western Hemisphere. Entomologische Arbeiten aus dem Museum G. Frey 15 (2): 570 - 685." type="journal article" year="1964">Vulcano &amp; Pereira 1964: 604</bibRefCitation>
</treatmentCitation>
(catalogue). — 
<treatmentCitation id="101310AFFF8E3C2FFBA9FD7AFEE7DB9A" author="Halffter G. &amp; Martinez A." page="39" pageId="16" pageNumber="17" year="1977">
<bibRefCitation id="F5234B4FFF8E3C2FFBA9FD7AFEE7DB9A" author="Halffter G. &amp; Martinez A." pageId="16" pageNumber="17" pagination="29 - 107" refId="ref17216" refString="Halffter G. &amp; Martinez A. 1977. Revision monografica de los Canthonina americanos, IV parte. Clave para generos y subgeneros. Folia Entomologica Mexicana. 38: 29 - 107." type="journal article" year="1977">Halffter &amp; Martínez 1977: 39</bibRefCitation>
</treatmentCitation>
, 70 (taxonomic remarks). — 
<treatmentCitation id="101310AFFF8E3C2FFDF4FD5EFC6FDB9A" author="Vaz-de-Mello F. Z." box="[611,898,679,705]" page="191" pageId="16" pageNumber="17" year="2000">
<bibRefCitation id="F5234B4FFF8E3C2FFDF4FD5EFC6FDB9A" author="Vaz-de-Mello F. Z." box="[611,898,679,705]" pageId="16" pageNumber="17" pagination="183 - 195" refId="ref18376" refString="Vaz-de-Mello F. Z. 2000. Estado atual de conhecimentos dos Scarabaeidae s. str. (Coleoptera: Scarabaeoidea) do Brasil. In: Martin-Piera F., Morrone J. J. &amp; Melic A. (eds) Hacya un proyecto CYTED para el inventario y estimacion de la diversidad entomologica en Iberoamerica: PRIBES- 2000: Vol. 1: 183 - 195. Monografias Tercer Milenio y Sociedad Entomologica Aragonesa, Zaragoza." type="journal article" year="2000">Vaz-de-Mello 2000: 191</bibRefCitation>
</treatmentCitation>
(species list).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="D9A86535FF8E3C2FFF2AFD10FA8ADD76" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="910D36BEFF8E3C2FFF2AFD10FEDBDA58" blockId="16.[189,310,745,771]" box="[189,310,745,771]" pageId="16" pageNumber="17">
<heading id="CA4581D2FF8E3C2FFF2AFD10FEDBDA58" bold="true" box="[189,310,745,771]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="A3C6EAACFF8E3C2FFF2AFD10FEDBDA58" bold="true" box="[189,310,745,771]" pageId="16" pageNumber="17">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="910D36BEFF8E3C2FFF2AFCE1FA8ADD76" blockId="16.[189,1399,792,1070]" pageId="16" pageNumber="17">
In the subgenus, 
<taxonomicName id="56B24D3DFF8E3C2FFEE9FCE1FD07DA69" authority="Castelnau, 1840" authorityName="Castelnau" authorityYear="1840" box="[382,746,792,818]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="bicolor" subGenus="Goniocanthon">
<emphasis id="A3C6EAACFF8E3C2FFEE9FCE1FDC7DA69" box="[382,554,792,818]" italics="true" pageId="16" pageNumber="17">C. (G.) bicolor</emphasis>
<bibRefCitation id="F5234B4FFF8E3C2FFDB8FCE1FD07DA69" author="Castelnau Comte" box="[559,746,792,818]" pageId="16" pageNumber="17" refId="ref16713" refString="Castelnau Comte de. 1840. Histoire naturelle des insectes coleopteres. Tome Deuxieme. Societe bibliophile, Paris. https: // doi. org / 10.5962 / bhl. title. 36538" type="book" year="1840">Castelnau, 1840</bibRefCitation>
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is separated by the following combination of characters: ventral surface of meso- and metafemora smooth (
<figureCitation id="09892A3BFF8E3C2FFC9BFCC5FC84DA0D" box="[780,873,828,854]" captionStart="Fig" captionStartId="19.[189,232,1894,1920]" captionTargetBox="[202,1379,264,1882]" captionTargetId="figure@19.[189,1399,263,1882]" captionTargetPageId="19" captionText="Fig. 4. Canthon (G.) bicolor Castelnau, 1840. A. Habitus, dorsal view. B. Ventral surface of meso- and metafemora and metasternal anterior lobe not convex. C. Dorsal view showing convex pygidium. D. Male protibial spur. E. Female protibial spur. F. Paramera lateral view. G. Paramera dorsal view. Scale bar: A = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3787895" httpUri="https://zenodo.org/record/3787895/files/figure.png" pageId="16" pageNumber="17">Fig. 4B</figureCitation>
); dorsal surface of pronotum and elytral disc with different colours, pronotum with light green metallic sheen and elytra with dark green metallic sheen (
<figureCitation id="09892A3BFF8E3C2FFE84FC7DFE99DAC5" box="[275,372,900,926]" captionStart="Fig" captionStartId="19.[189,232,1894,1920]" captionTargetBox="[202,1379,264,1882]" captionTargetId="figure@19.[189,1399,263,1882]" captionTargetPageId="19" captionText="Fig. 4. Canthon (G.) bicolor Castelnau, 1840. A. Habitus, dorsal view. B. Ventral surface of meso- and metafemora and metasternal anterior lobe not convex. C. Dorsal view showing convex pygidium. D. Male protibial spur. E. Female protibial spur. F. Paramera lateral view. G. Paramera dorsal view. Scale bar: A = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3787895" httpUri="https://zenodo.org/record/3787895/files/figure.png" pageId="16" pageNumber="17">Fig. 4A</figureCitation>
). Metasternal anterior lobe flat (
<figureCitation id="09892A3BFF8E3C2FFD6DFC7DFCB4DAC5" box="[762,857,900,926]" captionStart="Fig" captionStartId="19.[189,232,1894,1920]" captionTargetBox="[202,1379,264,1882]" captionTargetId="figure@19.[189,1399,263,1882]" captionTargetPageId="19" captionText="Fig. 4. Canthon (G.) bicolor Castelnau, 1840. A. Habitus, dorsal view. B. Ventral surface of meso- and metafemora and metasternal anterior lobe not convex. C. Dorsal view showing convex pygidium. D. Male protibial spur. E. Female protibial spur. F. Paramera lateral view. G. Paramera dorsal view. Scale bar: A = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3787895" httpUri="https://zenodo.org/record/3787895/files/figure.png" pageId="16" pageNumber="17">Fig. 4B</figureCitation>
). Profemur without teeth on anterior margin. Paramera, in lateral view (
<figureCitation id="09892A3BFF8E3C2FFD94FC5EFD8FDA99" box="[515,610,935,962]" captionStart="Fig" captionStartId="19.[189,232,1894,1920]" captionTargetBox="[202,1379,264,1882]" captionTargetId="figure@19.[189,1399,263,1882]" captionTargetPageId="19" captionText="Fig. 4. Canthon (G.) bicolor Castelnau, 1840. A. Habitus, dorsal view. B. Ventral surface of meso- and metafemora and metasternal anterior lobe not convex. C. Dorsal view showing convex pygidium. D. Male protibial spur. E. Female protibial spur. F. Paramera lateral view. G. Paramera dorsal view. Scale bar: A = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3787895" httpUri="https://zenodo.org/record/3787895/files/figure.png" pageId="16" pageNumber="17">Fig. 4F</figureCitation>
): flattened; apex semicircular; upper angle rounded; lower angle triangular. In dorsal view (
<figureCitation id="09892A3BFF8E3C2FFE62FC32FDB8DABE" box="[501,597,971,997]" captionStart="Fig" captionStartId="19.[189,232,1894,1920]" captionTargetBox="[202,1379,264,1882]" captionTargetId="figure@19.[189,1399,263,1882]" captionTargetPageId="19" captionText="Fig. 4. Canthon (G.) bicolor Castelnau, 1840. A. Habitus, dorsal view. B. Ventral surface of meso- and metafemora and metasternal anterior lobe not convex. C. Dorsal view showing convex pygidium. D. Male protibial spur. E. Female protibial spur. F. Paramera lateral view. G. Paramera dorsal view. Scale bar: A = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3787895" httpUri="https://zenodo.org/record/3787895/files/figure.png" pageId="16" pageNumber="17">Fig. 4G</figureCitation>
). Internal sac (
<figureCitation id="09892A3BFF8E3C2FFC95FC32FCA1DABE" box="[770,844,971,997]" captionStart="Fig" captionStartId="24.[189,232,1964,1990]" captionTargetBox="[281,1315,437,1914]" captionTargetId="figure@24.[264,1324,425,1923]" captionTargetPageId="24" captionText="Fig. 9. Sclerites of the internal sac of Canthon (G.) bicolor Castelnau, 1840. Abbreviations: see Material and methods. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3787907" httpUri="https://zenodo.org/record/3787907/files/figure.png" pageId="16" pageNumber="17">Fig. 9</figureCitation>
): FLP not completely involving A forming two duct with spiniform apex. SA lateral to FLP + A complex absent. MP distal to A, divided in MP 1 and MP 2, MP 1 semicircular, MP 2 fishhook-shaped with two hook; FLP + A complex involved by MP 1.
</paragraph>
</subSubSection>
<subSubSection id="D9A86535FF8E3C2DFF2AFBACFC1BDB5E" lastPageId="18" lastPageNumber="19" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="910D36BEFF8E3C2FFF2AFBACFE47DD34" blockId="16.[189,426,1109,1135]" box="[189,426,1109,1135]" pageId="16" pageNumber="17">
<heading id="CA4581D2FF8E3C2FFF2AFBACFE47DD34" bold="true" box="[189,426,1109,1135]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="A3C6EAACFF8E3C2FFF2AFBACFE47DD34" bold="true" box="[189,426,1109,1135]" pageId="16" pageNumber="17">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="21DA3CE3FF8E3C2FFF7DFB7CFD2ADC09" ID-GBIF-Occurrence="2610406305" pageId="16" pageNumber="17" typeStatus="lectotype">
<paragraph id="910D36BEFF8E3C2FFF7DFB7CFC1ADDC4" blockId="16.[189,1399,1156,1363]" box="[234,1015,1156,1183]" pageId="16" pageNumber="17">
<emphasis id="A3C6EAACFF8E3C2FFF7DFB7CFE8BDDC4" bold="true" box="[234,358,1157,1183]" pageId="16" pageNumber="17">
<typeStatus id="4E09881CFF8E3C2FFF7DFB7CFE8BDDC4" box="[234,358,1157,1183]" pageId="16" pageNumber="17" type="lectotype">Lectotype</typeStatus>
</emphasis>
(for 
<taxonomicName id="56B24D3DFF8E3C2FFE36FB7DFCC7DDC4" authority="Castelnau, 1840" authorityName="Castelnau" authorityYear="1840" box="[417,810,1156,1183]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="A3C6EAACFF8E3C2FFE36FB7DFD8FDDC5" box="[417,610,1156,1182]" italics="true" pageId="16" pageNumber="17">Canthon bicolor</emphasis>
<bibRefCitation id="F5234B4FFF8E3C2FFDFEFB7DFCC7DDC4" author="Castelnau Comte" box="[617,810,1156,1183]" pageId="16" pageNumber="17" refId="ref16713" refString="Castelnau Comte de. 1840. Histoire naturelle des insectes coleopteres. Tome Deuxieme. Societe bibliophile, Paris. https: // doi. org / 10.5962 / bhl. title. 36538" type="book" year="1840">Castelnau, 1840</bibRefCitation>
</taxonomicName>
; here designated)
</paragraph>
<paragraph id="910D36BEFF8E3C2FFF2AFB51FD2ADC09" blockId="16.[189,1399,1156,1363]" pageId="16" pageNumber="17">
<collectingCountry id="E9A5762EFF8E3C2FFF2AFB51FE5DDD99" box="[189,432,1192,1218]" name="French Guiana" pageId="16" pageNumber="17">FRENCH GUIANA</collectingCountry>
: 
<specimenCount id="87B4FD37FF8E3C2FFE2BFB50FE3FDD98" box="[444,466,1193,1219]" pageId="16" pageNumber="17" type="female">♀</specimenCount>
, 
<taxonomicName id="56B24D3DFF8E3C2FFE76FB51FCADDD98" authority="Bicolor DeLaporte" authorityName="Bicolor DeLaporte" box="[481,832,1192,1219]" class="Insecta" family="Scarabaeidae" genus="Coprobius" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="A3C6EAACFF8E3C2FFE76FB51FD56DD99" box="[481,699,1192,1218]" italics="true" pageId="16" pageNumber="17">Coprobius Bicolor</emphasis>
DeLaporte
</taxonomicName>
anim. art. p. 69 
<taxonomicName id="56B24D3DFF8E3C2FFB95FB51FB8ADD99" authorityName="sensu Halffter &amp; Martinez" authorityYear="1977" box="[1026,1127,1192,1218]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="A3C6EAACFF8E3C2FFB95FB51FB8ADD99" box="[1026,1127,1192,1218]" italics="true" pageId="16" pageNumber="17">Canthon</emphasis>
</taxonomicName>
nº 6 Cayennae Maroni D’DeLaporte (rectangular green label, border black) / 
<emphasis id="A3C6EAACFF8E3C2FFCC6FB34FC8BDDBC" box="[849,870,1229,1255]" italics="true" pageId="16" pageNumber="17">G</emphasis>
(small red label) / A. Gory coll. purchased 
<date id="E50C107EFF8E3C2FFF2AFB09FED1DC50" box="[189,316,1264,1291]" pageId="16" pageNumber="17" value="1849" valueMax="1850">1849-1850</date>
From H.L. 
<collectionCode id="F7A3AE7BFF8E3C2FFE2EFB09FDFDDC51" box="[441,528,1264,1290]" pageId="16" pageNumber="17">GORY</collectionCode>
(white label) standing over: 
<taxonomicName id="56B24D3DFF8E3C2FFCF6FB09FAD3DC51" authority="DeLaporte, Cayennae" authorityName="DeLaporte, Cayennae" box="[865,1342,1264,1291]" class="Insecta" family="Scarabaeidae" genus="Coprobius" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="A3C6EAACFF8E3C2FFCF6FB09FBDADC51" box="[865,1079,1264,1290]" italics="true" pageId="16" pageNumber="17">Coprobius bicolor</emphasis>
DeLaporte, Cayennae
</taxonomicName>
OX. 
<collectionCode id="F7A3AE7BFF8E3C2FFF2AFAECFF18DC74" box="[189,245,1301,1327]" country="0" httpUri="http://grbio.org/cool/xxq3-2kv6" name="University of Northern Iowa" pageId="16" pageNumber="17">UNI</collectionCode>
. MUS. NAT. 
<collectionCode id="F7A3AE7BFF8E3C2FFE0CFAEDFE33DC75" box="[411,478,1300,1326]" pageId="16" pageNumber="17">HIST</collectionCode>
. (
<collectionCode id="F7A3AE7BFF8E3C2FFE64FAEDFD8BDC75" box="[499,614,1300,1326]" pageId="16" pageNumber="17">OUMNH</collectionCode>
) (white label) / 
<typeStatus id="4E09881CFF8E3C2FFCBDFAEDFC3CDC75" box="[810,977,1300,1326]" pageId="16" pageNumber="17" type="lectotype">LECTOTYPE</typeStatus>
<specimenCount id="87B4FD37FF8E3C2FFC4BFAECFC1FDC74" box="[988,1010,1301,1327]" pageId="16" pageNumber="17" type="female">♀</specimenCount>
<taxonomicName id="56B24D3DFF8E3C2FFC6CFAEDFAEBDC75" authority="Cast." authorityName="Castelnau" authorityYear="1840" box="[1019,1286,1300,1326]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="A3C6EAACFF8E3C2FFC6CFAEDFAEBDC75" box="[1019,1286,1300,1326]" italics="true" pageId="16" pageNumber="17">Canthon bicolor Cast.</emphasis>
</taxonomicName>
des. F.Z. Vaz-de-Mello, 2013” (red label) (
<collectionCode id="F7A3AE7BFF8E3C2FFDD2FAC1FD56DC09" box="[581,699,1336,1362]" pageId="16" pageNumber="17">OUMNH</collectionCode>
).
</paragraph>
</materialsCitation>
<paragraph id="910D36BEFF8E3C2FFF7DFA83FD75DCCE" blockId="16.[189,1399,1402,1679]" box="[234,664,1402,1429]" pageId="16" pageNumber="17">
<emphasis id="A3C6EAACFF8E3C2FFF7DFA83FE22DCCF" bold="true" box="[234,463,1402,1428]" pageId="16" pageNumber="17">Non-type material</emphasis>
(
<specimenCount id="87B4FD37FF8E3C2FFE48FA83FD7DDCCE" box="[479,656,1402,1429]" pageId="16" pageNumber="17" type="generic">248 specimens</specimenCount>
)
</paragraph>
<paragraph id="910D36BEFF8E3C2FFF2AFA67FD8CDFD4" blockId="16.[189,1399,1402,1679]" pageId="16" pageNumber="17">
<collectingCountry id="E9A5762EFF8E3C2FFF2AFA67FE87DCE3" box="[189,362,1438,1464]" name="Venezuela" pageId="16" pageNumber="17">VENEZUELA</collectingCountry>
– 
<emphasis id="A3C6EAACFF8E3C2FFE13FA64FE0CDCE3" bold="true" box="[388,481,1437,1464]" pageId="16" pageNumber="17">
<collectingRegion id="5376F85CFF8E3C2FFE13FA64FE0CDCE3" box="[388,481,1437,1464]" country="Venezuela" name="Bolivar" pageId="16" pageNumber="17">Bolívar</collectingRegion>
</emphasis>
: 
<specimenCount id="87B4FD37FF8E3C2FFE78FA67FDDDDCE3" box="[495,560,1438,1464]" pageId="16" pageNumber="17" type="female">3 ♀♀</specimenCount>
,Altiplanicie de Nuria (“El Hormiguero, Meseta de Nuria”), 
<quantity id="564A9B5BFF8E3C2FFB7DFA67FADEDCE3" box="[1258,1331,1438,1465]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="16" pageNumber="17" unit="m" value="500.0">500 m</quantity>
a.s.l., Expediction Instituto Zoologia Agricola de la Faculd de Agronomia, 
<date id="E50C107EFF8E3C2FFB92FA3BFB3ADC87" box="[1029,1239,1474,1500]" pageId="16" pageNumber="17" value="1974-12-13" valueMax="1974-12-17" valueMin="1974-12-13">13–17 Dec. 1974</date>
, Universidad Central de Venezuela leg. (
<collectionCode id="F7A3AE7BFF8E3C2FFD97FA1FFDB0DF5B" box="[512,605,1510,1536]" pageId="16" pageNumber="17">CMNC</collectionCode>
); 
<specimenCount id="87B4FD37FF8E3C2FFDE4FA1FFD55DF5B" box="[627,696,1510,1536]" pageId="16" pageNumber="17" type="female">2 ♀♀</specimenCount>
, Sinfontes, 
<quantity id="564A9B5BFF8E3C2FFCD0FA1FFC7EDF5B" box="[839,915,1510,1536]" metricMagnitude="4" metricUnit="m" metricValue="2.2" pageId="16" pageNumber="17" unit="km" value="22.0">22 km</quantity>
S of El Dorado, flight interception trap, 
<date id="E50C107EFF8E3C2FFF2AF9F0FE59DF7F" box="[189,436,1545,1572]" pageId="16" pageNumber="17" value="1987-06-25" valueMax="1987-07-12" valueMin="1987-06-25">25 Jun.–12 Jul. 1987</date>
, S. and J. Peck leg. (
<collectionCode id="F7A3AE7BFF8E3C2FFD3FF9F3FCE8DF7F" box="[680,773,1546,1572]" pageId="16" pageNumber="17">CMNC</collectionCode>
); 
<specimenCount id="87B4FD37FF8E3C2FFC8DF9F3FCABDF7F" box="[794,838,1546,1572]" pageId="16" pageNumber="17" type="female">1 ♀</specimenCount>
, Sinfontes, 
<quantity id="564A9B5BFF8E3C2FFC44F9F3FBF0DF78" box="[979,1053,1545,1572]" metricMagnitude="4" metricUnit="m" metricValue="3.3" pageId="16" pageNumber="17" unit="km" value="33.0">33 km</quantity>
S of El Dorado, 
<quantity id="564A9B5BFF8E3C2FFB71F9F3FADCDF7F" box="[1254,1329,1546,1572]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="16" pageNumber="17" unit="m" value="220.0">220 m</quantity>
a.s.l., 
<date id="E50C107EFF8E3C2FFF2AF9D7FE83DF1C" box="[189,366,1581,1608]" pageId="16" pageNumber="17" value="1986-08-02" valueMax="1986-08-07" valueMin="1986-08-02">2–7 Aug. 1986</date>
, B. Gill leg. (
<collectionCode id="F7A3AE7BFF8E3C2FFD98F9D7FD81DF13" box="[527,620,1582,1608]" pageId="16" pageNumber="17">CMNC</collectionCode>
); 
<specimenCount id="87B4FD37FF8E3C2FFD16F9D7FD43DF13" box="[641,686,1582,1608]" pageId="16" pageNumber="17" type="female">1 ♀</specimenCount>
, Sinfontes, El Bochinche, Reserva Florestal Imataca, 
<quantity id="564A9B5BFF8E3C2FFABCF9D7FA9ADF13" box="[1323,1399,1582,1608]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="16" pageNumber="17" unit="m" value="200.0">200 m</quantity>
a.s.l., Expediction Instituto Zoologia Agricola de la Faculd de Agronomia, 
<date id="E50C107EFF8E3C2FFBB6F9A8FB31DF30" box="[1057,1244,1617,1644]" pageId="16" pageNumber="17" value="1974-12-06" valueMax="1974-12-13" valueMin="1974-12-06">6–13 Dec. 1974</date>
, Universidad Central de Venezuela leg. (
<collectionCode id="F7A3AE7BFF8E3C2FFE6FF98CFDB8DFD4" box="[504,597,1653,1679]" pageId="16" pageNumber="17">CMNC</collectionCode>
).
</paragraph>
<paragraph id="910D36BEFF8E3C2FFF2AF94EFEDDDE42" blockId="16.[189,1399,1717,1817]" pageId="16" pageNumber="17">
<collectingCountry id="E9A5762EFF8E3C2FFF2AF94EFED3DF8A" box="[189,318,1719,1745]" name="Guyana" pageId="16" pageNumber="17">GUYANA</collectingCountry>
: 2 ♁♁, 
<specimenCount id="87B4FD37FF8E3C2FFE09F94EFE26DF89" box="[414,459,1719,1746]" pageId="16" pageNumber="17" type="female">1 ♀</specimenCount>
, no locality, 
<date id="E50C107EFF8E3C2FFDFFF941FD08DF8A" box="[616,741,1719,1746]" pageId="16" pageNumber="17" value="1987-03">Mar. 1987</date>
, [local collector leg.] (
<collectionCode id="F7A3AE7BFF8E3C2FFC63F94EFB87DF8A" box="[1012,1130,1719,1745]" pageId="16" pageNumber="17">OUMNH</collectionCode>
); 1 unsexed specimen, Essequibo River, Monkey Jump, 
<date id="E50C107EFF8E3C2FFDDAF922FD19DFAE" box="[589,756,1755,1781]" pageId="16" pageNumber="17" value="1929-08-19">19 Aug. 1929</date>
, Oxford University Expedition to British Guiana leg. (
<collectionCode id="F7A3AE7BFF8E3C2FFF52F906FEC9DE42" box="[197,292,1791,1817]" pageId="16" pageNumber="17">BMNH</collectionCode>
).
</paragraph>
<paragraph id="910D36BEFF8E3C2FFF2AF8B8FA9BDEB0" blockId="16.[189,1399,1855,2027]" pageId="16" pageNumber="17">
<collectingCountry id="E9A5762EFF8E3C2FFF2AF8B8FEA4DE00" box="[189,329,1857,1883]" name="Suriname" pageId="16" pageNumber="17">SURINAM</collectingCountry>
: 3 unsexed specimens, no more data (
<collectionCode id="F7A3AE7BFF8E3C2FFC9DF8B8FC84DE00" box="[778,873,1857,1883]" pageId="16" pageNumber="17">BMNH</collectionCode>
); 1 ♁, no more data (
<collectionCode id="F7A3AE7BFF8E3C2FFBF1F8B8FB54DE00" box="[1126,1209,1857,1883]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="16" pageNumber="17" type="Museum">MZSP</collectionCode>
); 1 ♁, A.J. Buis and 
<collectionCode id="F7A3AE7BFF8E3C2FFF79F89FFEEEDEDB" box="[238,259,1894,1920]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="16" pageNumber="17" type="Museum">V</collectionCode>
. Balthasar leg. (
<collectionCode id="F7A3AE7BFF8E3C2FFE57F89CFDF7DE24" box="[448,538,1893,1919]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="16" pageNumber="17">NMPC</collectionCode>
); 
<specimenCount id="87B4FD37FF8E3C2FFDB9F89CFDB7DEDB" box="[558,602,1893,1920]" pageId="16" pageNumber="17" type="female">1 ♀</specimenCount>
, no locality or date, 
<collectionCode id="F7A3AE7BFF8E3C2FFCDAF89FFC8FDEDB" box="[845,866,1894,1920]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="16" pageNumber="17" type="Museum">V</collectionCode>
. Balthasar leg. (
<collectionCode id="F7A3AE7BFF8E3C2FFB89F89CFB95DE24" box="[1054,1144,1893,1919]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="16" pageNumber="17">NMPC</collectionCode>
); 
<specimenCount id="87B4FD37FF8E3C2FFB1AF89CFB54DEDB" box="[1165,1209,1893,1920]" pageId="16" pageNumber="17" type="female">1 ♀</specimenCount>
, no locality and date, Dr. Will leg. (
<collectionCode id="F7A3AE7BFF8E3C2FFE0AF870FE15DEF8" box="[413,504,1929,1955]" pageId="16" pageNumber="17">ZMHB</collectionCode>
). – 
<emphasis id="A3C6EAACFF8E3C2FFDB6F870FD54DEF8" bold="true" box="[545,697,1929,1955]" pageId="16" pageNumber="17">
<collectingRegion id="5376F85CFF8E3C2FFDB6F870FD54DEF8" box="[545,697,1929,1955]" country="Suriname" name="Paramaribo" pageId="16" pageNumber="17">Paramaribo</collectingRegion>
</emphasis>
: 2 ♁♁, 
<specimenCount id="87B4FD37FF8E3C2FFC8EF870FCA8DEF8" box="[793,837,1929,1955]" pageId="16" pageNumber="17" type="female">1 ♀</specimenCount>
, no locality or date, S. 
<collectionCode id="F7A3AE7BFF8E3C2FFBDDF870FB8FDEF8" box="[1098,1122,1929,1955]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="16" pageNumber="17" type="Museum">V</collectionCode>
. Heller leg. (
<collectionCode id="F7A3AE7BFF8E3C2FFB6EF870FAB9DEF8" box="[1273,1364,1929,1955]" pageId="16" pageNumber="17">ZMHB</collectionCode>
). – 
<emphasis id="A3C6EAACFF8E3C2FFF2AF854FE8EDE9C" bold="true" box="[189,355,1965,1991]" pageId="16" pageNumber="17">
<collectingRegion id="5376F85CFF8E3C2FFF2AF854FE8EDE9C" box="[189,355,1965,1991]" country="Suriname" name="Commewijne" pageId="16" pageNumber="17">Commewijne</collectingRegion>
</emphasis>
: 1 ♁, 
<specimenCount id="87B4FD37FF8E3C2FFE27F854FE1EDE9C" box="[432,499,1965,1991]" pageId="16" pageNumber="17" type="female">2 ♀♀</specimenCount>
, Akintosoela, 
<geoCoordinate id="F4865079FF8E3C2FFD0AF852FCDCDE9C" box="[669,817,1963,1991]" degrees="05" direction="north" minutes="16" orientation="latitude" pageId="16" pageNumber="17" precision="15" seconds="17" value="5.271389">05º16′17″ N</geoCoordinate>
, 
<geoCoordinate id="F4865079FF8E3C2FFCA9F852FC38DE9C" box="[830,981,1963,1991]" degrees="54" direction="west" minutes="55" orientation="longitude" pageId="16" pageNumber="17" precision="15" seconds="15" value="-54.920834">54º55′15″ W</geoCoordinate>
, 
<quantity id="564A9B5BFF8E3C2FFC75F854FBF3DE9C" box="[994,1054,1965,1992]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="16" pageNumber="17" unit="m" value="50.0">50 m</quantity>
a.s.l., dung trap, 
<date id="E50C107EFF8E3C2FFB7BF854FA9CDE9C" box="[1260,1393,1965,1991]" pageId="16" pageNumber="17" value="1999-07-02">2 Jul. 1999</date>
, Z. Falin leg. (
<collectionCode id="F7A3AE7BFF8E3C2FFEF0F828FE29DEB0" box="[359,452,2001,2027]" pageId="16" pageNumber="17">CMNC</collectionCode>
); 
<specimenCount id="87B4FD37FF8E3C2FFE4BF828FDE1DEB0" box="[476,524,2001,2027]" pageId="16" pageNumber="17" type="female">1 ♀</specimenCount>
, Akintosoela, 
<geoCoordinate id="F4865079FF8E3C2FFD2AF836FCB9DEB0" box="[701,852,1999,2027]" degrees="05" direction="north" minutes="16" orientation="latitude" pageId="16" pageNumber="17" precision="15" seconds="17" value="5.271389">05º16′17″ N</geoCoordinate>
, 
<geoCoordinate id="F4865079FF8E3C2FFCF3F836FC13DEB0" box="[868,1022,1999,2027]" degrees="54" direction="west" minutes="55" orientation="longitude" pageId="16" pageNumber="17" precision="15" seconds="15" value="-54.920834">54º55′15″ W</geoCoordinate>
, 
<quantity id="564A9B5BFF8E3C2FFB99F828FBA0DEB0" box="[1038,1101,2001,2027]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="16" pageNumber="17" unit="m" value="40.0">40 m</quantity>
a.s.l., flight interception
</paragraph>
<paragraph id="910D36BEFF8F3C2EFF2AFEF5FCC8D811" blockId="17.[189,1398,265,330]" pageId="17" pageNumber="18">
trap, 
<date id="E50C107EFF8F3C2EFF6EFEF2FE90D87E" box="[249,381,267,293]" pageId="17" pageNumber="18" value="1999-07-03">3 Jul. 1999</date>
, Z. Falin leg. (
<collectionCode id="F7A3AE7BFF8F3C2EFDBFFEF2FD68D87E" box="[552,645,267,293]" pageId="17" pageNumber="18">CMNC</collectionCode>
). – 
<emphasis id="A3C6EAACFF8F3C2EFD39FEF2FCD5D87E" bold="true" box="[686,824,267,293]" pageId="17" pageNumber="18">
<collectingRegion id="5376F85CFF8F3C2EFD39FEF2FCD5D87E" box="[686,824,267,293]" country="Suriname" name="Marowijne" pageId="17" pageNumber="18">Marowijne</collectingRegion>
</emphasis>
: 
<specimenCount id="87B4FD37FF8F3C2EFCDFFEF2FC99D87D" box="[840,884,267,294]" pageId="17" pageNumber="18" type="female">1 ♀</specimenCount>
, Palumeu, 
<geoCoordinate id="F4865079FF8F3C2EFC6FFEF0FB66D87D" box="[1016,1163,265,294]" degrees="03" direction="north" minutes="20" orientation="latitude" pageId="17" pageNumber="18" precision="15" seconds="56" value="3.3488889">03º20′56″ N</geoCoordinate>
, 
<geoCoordinate id="F4865079FF8F3C2EFB01FEF0FAC0D87D" box="[1174,1325,265,294]" degrees="55" direction="west" minutes="26" orientation="longitude" pageId="17" pageNumber="18" precision="15" seconds="18" value="-55.438335">55º26′18″ W</geoCoordinate>
, flight interception trap, 
<date id="E50C107EFF8F3C2EFE1AFEC9FDFCD811" box="[397,529,303,330]" pageId="17" pageNumber="18" value="1999-07-09">9 Jul. 1999</date>
, Z. Falin leg. (
<collectionCode id="F7A3AE7BFF8F3C2EFD2BFEC9FCF4D811" box="[700,793,304,330]" pageId="17" pageNumber="18">CMNC</collectionCode>
).
</paragraph>
<paragraph id="910D36BEFF8F3C2EFF2AFE8BFE13DEB0" blockId="17.[189,1399,370,2027]" pageId="17" pageNumber="18">
<collectingCountry id="E9A5762EFF8F3C2EFF2AFE8BFE5ED8D7" box="[189,435,370,396]" name="French Guiana" pageId="17" pageNumber="18">FRENCH GUIANA</collectingCountry>
: 1 unsexed specimen, no locality, 1981, G. Tavakilian leg. (
<collectionCode id="F7A3AE7BFF8F3C2EFB12FE8BFB08D8D7" box="[1157,1253,370,396]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="17" pageNumber="18">MNHN</collectionCode>
); 
<specimenCount id="87B4FD37FF8F3C2EFB6BFE8BFAC6D8D6" box="[1276,1323,370,397]" pageId="17" pageNumber="18" type="female">1 ♀</specimenCount>
, “D-5 4k SE Tngmd Jct”, 
<date id="E50C107EFF8F3C2EFE3AFE6FFDBED8EA" box="[429,595,406,433]" pageId="17" pageNumber="18" value="1995-08-24">24 Aug. 1995</date>
, J.E. Wappes leg. (
<collectionCode id="F7A3AE7BFF8F3C2EFCADFE6FFC7AD8EB" box="[826,919,406,432]" pageId="17" pageNumber="18">CMNC</collectionCode>
); 
<specimenCount id="87B4FD37FF8F3C2EFC39FE6FFC31D8EA" box="[942,988,406,433]" pageId="17" pageNumber="18" type="female">1 ♀</specimenCount>
, PK 45, Piste del Beligan, 
<date id="E50C107EFF8F3C2EFAB0FE6EFF17D88E" pageId="17" pageNumber="18" value="2003-03-07">7 Mar. 2003</date>
, no collector (
<collectionCode id="F7A3AE7BFF8F3C2EFE31FE42FDEDD88E" box="[422,512,443,469]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="17" pageNumber="18">NMPC</collectionCode>
); 1 ♁, Cayenne, 1915, no collector (
<collectionCode id="F7A3AE7BFF8F3C2EFC24FE42FBEBD88E" box="[947,1030,443,469]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="17" pageNumber="18" type="Museum">MZSP</collectionCode>
); 1 ♁, Cayenne (“ 
<quantity id="564A9B5BFF8F3C2EFB4BFE43FAC9D88F" box="[1244,1316,442,468]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="18" unit="km" value="20.0">20km</quantity>
SW”), 
<geoCoordinate id="F4865079FF8F3C2EFF2AFE24FEBED8A2" box="[189,339,477,505]" degrees="04" direction="north" minutes="48" orientation="latitude" pageId="17" pageNumber="18" precision="15" seconds="18" value="4.8050003">04º48′18″ N</geoCoordinate>
, 
<geoCoordinate id="F4865079FF8F3C2EFEF5FE24FE16D8A2" box="[354,507,477,505]" degrees="52" direction="west" minutes="28" orientation="longitude" pageId="17" pageNumber="18" precision="15" seconds="41" value="-52.478058">52º28′41″ W</geoCoordinate>
, 
<quantity id="564A9B5BFF8F3C2EFD9DFE26FDA4D8A2" box="[522,585,479,505]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="17" pageNumber="18" unit="m" value="30.0">30 m</quantity>
a.s.l., flight interception trap, 
<date id="E50C107EFF8F3C2EFC2EFE26FB5BD8A2" box="[953,1206,479,505]" pageId="17" pageNumber="18" value="1997-05-29" valueMax="1997-06-09" valueMin="1997-05-29">29 May–9 Jun. 1997</date>
, J. Ashe and 
<collectionCode id="F7A3AE7BFF8F3C2EFACBFE26FA9ED8A2" box="[1372,1395,479,505]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="17" pageNumber="18">R</collectionCode>
. Brooks leg. (
<collectionCode id="F7A3AE7BFF8F3C2EFEC4FDFAFE5DDB46" box="[339,432,515,541]" pageId="17" pageNumber="18">CMNC</collectionCode>
); 1 ♁, 
<specimenCount id="87B4FD37FF8F3C2EFE6AFDFAFDC5DB45" box="[509,552,515,542]" pageId="17" pageNumber="18" type="female">1 ♀</specimenCount>
, Cayenne, Camopi, Oyapock river, Ilet Massikiri, [dung], 
<date id="E50C107EFF8F3C2EFB45FDFAFA9CDB46" box="[1234,1393,515,542]" pageId="17" pageNumber="18" value="1969-11-17">17 Nov. 1969</date>
, G. Halffter leg. (
<collectionCode id="F7A3AE7BFF8F3C2EFE17FDDEFE30DB1A" box="[384,477,551,577]" pageId="17" pageNumber="18">CMNC</collectionCode>
); 7 ♁♁, 
<specimenCount id="87B4FD37FF8F3C2EFDD5FDDEFD79DB19" box="[578,660,551,578]" pageId="17" pageNumber="18" type="female">12 ♀♀</specimenCount>
, Cayenne, Montagne des Chevaux, 
<geoCoordinate id="F4865079FF8F3C2EFBABFDDCFB22DB19" box="[1084,1231,549,578]" degrees="04" direction="north" minutes="44" orientation="latitude" pageId="17" pageNumber="18" precision="15" seconds="56" value="4.748889">04º44′56″ N</geoCoordinate>
, 
<geoCoordinate id="F4865079FF8F3C2EFB4CFDDCFA9FDB19" box="[1243,1394,549,578]" degrees="52" direction="west" minutes="26" orientation="longitude" pageId="17" pageNumber="18" precision="15" seconds="28" value="-52.441113">52º26′28″ W</geoCoordinate>
, 
<quantity id="564A9B5BFF8F3C2EFF2AFDB2FF11DB3D" box="[189,252,587,614]" metricMagnitude="1" metricUnit="m" metricValue="7.5" pageId="17" pageNumber="18" unit="m" value="75.0">75 m</quantity>
a.s.l., 
<date id="E50C107EFF8F3C2EFED9FDB2FE02DB3D" box="[334,495,587,614]" pageId="17" pageNumber="18" value="2011-09-25">25 Sep. 2011</date>
, 
<collectionCode id="F7A3AE7BFF8F3C2EFE68FDB5FDA1DB3D" box="[511,588,588,614]" pageId="17" pageNumber="18">SEAG</collectionCode>
leg. (
<collectionCode id="F7A3AE7BFF8F3C2EFD04FDB5FD06DB3D" box="[659,747,588,614]" pageId="17" pageNumber="18">CEMT</collectionCode>
); 4 ♁♁, 
<specimenCount id="87B4FD37FF8F3C2EFCCEFDB5FC65DB3D" box="[857,904,588,614]" pageId="17" pageNumber="18" type="female">1 ♀</specimenCount>
, same data as for preceding, but 
<date id="E50C107EFF8F3C2EFAB7FDB5FF15DBD1" pageId="17" pageNumber="18" value="2013-01-27">27 Jan. 2013</date>
(
<collectionCode id="F7A3AE7BFF8F3C2EFE90FD89FEB2DBD1" box="[263,351,624,650]" pageId="17" pageNumber="18">CEMT</collectionCode>
); 4 ♁♁, 
<specimenCount id="87B4FD37FF8F3C2EFE54FD89FDEBDBD1" box="[451,518,624,650]" pageId="17" pageNumber="18" type="female">7 ♀♀</specimenCount>
, Cayenne, Kourou, flight interception trap, 
<date id="E50C107EFF8F3C2EFB92FD89FB9ADBD1" box="[1029,1143,624,650]" pageId="17" pageNumber="18" value="2013-01">Jan. 2013</date>
, 
<collectionCode id="F7A3AE7BFF8F3C2EFB14FD89FB3DDBD1" box="[1155,1232,624,650]" pageId="17" pageNumber="18">SEAG</collectionCode>
leg. (
<collectionCode id="F7A3AE7BFF8F3C2EFA85FD89FA87DBD1" box="[1298,1386,624,650]" pageId="17" pageNumber="18">CEMT</collectionCode>
); 3 ♁♁, 
<specimenCount id="87B4FD37FF8F3C2EFE84FD6DFEAFDBF5" box="[275,322,660,686]" pageId="17" pageNumber="18" type="female">1 ♀</specimenCount>
, Cayenne, La Chaumière, 
<date id="E50C107EFF8F3C2EFD12FD6DFCEDDBF5" box="[645,768,660,686]" pageId="17" pageNumber="18" value="1978-04">Apr. 1978</date>
, P. Arnaud leg. (
<collectionCode id="F7A3AE7BFF8F3C2EFC5BFD6DFBC4DBF5" box="[972,1065,660,686]" pageId="17" pageNumber="18">CMNC</collectionCode>
); 
<specimenCount id="87B4FD37FF8F3C2EFBD7FD6DFB68DBF5" box="[1088,1157,660,686]" pageId="17" pageNumber="18" type="female">2 ♀♀</specimenCount>
, Cayenne, [Réserve naturelle nationale des] Nouragues, 
<geoCoordinate id="F4865079FF8F3C2EFDF0FD4FFD3FDB88" box="[615,722,694,723]" degrees="04" direction="north" minutes="05" orientation="latitude" pageId="17" pageNumber="18" precision="925" value="4.0833335">04º05′ N</geoCoordinate>
, 
<geoCoordinate id="F4865079FF8F3C2EFD48FD4FFCA0DB88" box="[735,845,694,723]" degrees="52" direction="west" minutes="40" orientation="longitude" pageId="17" pageNumber="18" precision="925" value="-52.666668">52º40′ W</geoCoordinate>
, 
<quantity id="564A9B5BFF8F3C2EFCCEFD41FC49DB88" box="[857,932,696,723]" metricMagnitude="2" metricUnit="m" metricValue="1.55" pageId="17" pageNumber="18" unit="m" value="155.0">155 m</quantity>
a.s.l., 
<date id="E50C107EFF8F3C2EFC64FD40FB82DB89" box="[1011,1135,696,723]" pageId="17" pageNumber="18" value="1997-03">Mar. 1997</date>
, F. Feer leg. (
<collectionCode id="F7A3AE7BFF8F3C2EFA85FD41FA87DB89" box="[1298,1386,696,722]" pageId="17" pageNumber="18">CEMT</collectionCode>
); 3 ♁♁, 
<specimenCount id="87B4FD37FF8F3C2EFE80FD24FEAADBAC" box="[279,327,733,759]" pageId="17" pageNumber="18" type="female">1 ♀</specimenCount>
, same data as for preceding, but 
<date id="E50C107EFF8F3C2EFD7DFD24FC8BDBAC" box="[746,870,733,759]" pageId="17" pageNumber="18" value="2003-05">May 2003</date>
(
<collectionCode id="F7A3AE7BFF8F3C2EFCEDFD24FC3FDBAC" box="[890,978,733,759]" pageId="17" pageNumber="18">CEMT</collectionCode>
); 5 ♁♁, 
<specimenCount id="87B4FD37FF8F3C2EFBD2FD24FB61DBAC" box="[1093,1164,733,759]" pageId="17" pageNumber="18" type="female">4 ♀♀</specimenCount>
, Cayenne, Paracou Field Station, 
<geoCoordinate id="F4865079FF8F3C2EFEF0FD06FE3EDA40" box="[359,467,767,795]" degrees="05" direction="north" minutes="02" orientation="latitude" pageId="17" pageNumber="18" precision="925" value="5.0333333">05º02′ N</geoCoordinate>
, 
<geoCoordinate id="F4865079FF8F3C2EFE75FD06FDBCDA40" box="[482,593,767,795]" degrees="53" direction="west" minutes="00" orientation="longitude" pageId="17" pageNumber="18" precision="925" value="-53.0">53º00′ W</geoCoordinate>
, 
<quantity id="564A9B5BFF8F3C2EFDC8FCF9FD70DA40" box="[607,669,768,795]" metricMagnitude="1" metricUnit="m" metricValue="5.5" pageId="17" pageNumber="18" unit="m" value="55.0">55 m</quantity>
a.s.l., 
<date id="E50C107EFF8F3C2EFD78FCF8FC8ADA40" box="[751,871,769,795]" pageId="17" pageNumber="18" value="2003-10">Oct. 2003</date>
, F. Feer leg. (
<collectionCode id="F7A3AE7BFF8F3C2EFB87FCF8FB85DA40" box="[1040,1128,769,795]" pageId="17" pageNumber="18">CEMT</collectionCode>
); 1 ♁, 
<specimenCount id="87B4FD37FF8F3C2EFB29FCF8FB00DA40" box="[1214,1261,769,795]" pageId="17" pageNumber="18" type="female">1 ♀</specimenCount>
, 2 unsexed specimens, Cayenne, Roura (“ 
<quantity id="564A9B5BFF8F3C2EFDBCFCDCFD94DA64" box="[555,633,805,831]" metricMagnitude="3" metricUnit="m" metricValue="8.4" pageId="17" pageNumber="18" unit="km" value="8.4">8.4km</quantity>
SSE”), 
<geoCoordinate id="F4865079FF8F3C2EFD75FCDAFC97DA64" box="[738,890,803,831]" degrees="04" direction="north" minutes="40" orientation="latitude" pageId="17" pageNumber="18" precision="15" seconds="41" value="4.6780553">04º40′41″ N</geoCoordinate>
, 
<geoCoordinate id="F4865079FF8F3C2EFC1DFCDAFBCBDA64" box="[906,1062,803,831]" degrees="52" direction="west" minutes="13" orientation="longitude" pageId="17" pageNumber="18" precision="15" seconds="25" value="-52.22361">52º13′25″ W</geoCoordinate>
, flight interception trap, 
<date id="E50C107EFF8F3C2EFACEFCDCFE73DA38" pageId="17" pageNumber="18" value="1997-05-29" valueMax="1997-06-10" valueMin="1997-05-29">29 May–10 Jun. 1997</date>
, J. Ashe and 
<collectionCode id="F7A3AE7BFF8F3C2EFDD7FCB3FDBADA3F" box="[576,599,842,868]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="17" pageNumber="18">R</collectionCode>
. Brooks leg. (
<collectionCode id="F7A3AE7BFF8F3C2EFD69FCB0FCB6DA38" box="[766,859,841,867]" pageId="17" pageNumber="18">CMNC</collectionCode>
); 
<specimenCount id="87B4FD37FF8F3C2EFCE6FCB0FC5BDA3F" box="[881,950,841,868]" pageId="17" pageNumber="18" type="female">2 ♀♀</specimenCount>
, Cayenne, Roura, Montagne de Kaw, 
<date id="E50C107EFF8F3C2EFF2AFC94FEC6DADC" box="[189,299,877,903]" pageId="17" pageNumber="18" value="1992-07">Jul. 1992</date>
, A. Chaminade leg. (
<collectionCode id="F7A3AE7BFF8F3C2EFDB5FC94FD97DADC" box="[546,634,877,903]" pageId="17" pageNumber="18">CEMT</collectionCode>
); 3 ♁♁, 
<specimenCount id="87B4FD37FF8F3C2EFD77FC97FCCEDAD3" box="[736,803,878,904]" pageId="17" pageNumber="18" type="female">7 ♀♀</specimenCount>
, Cayenne, Saint-Georges-de-l’Oyapock, 
<geoCoordinate id="F4865079FF8F3C2EFA9FFC95FA9FDAD3" box="[1288,1394,876,904]" degrees="03" direction="north" minutes="54" orientation="latitude" pageId="17" pageNumber="18" precision="925" value="3.9">03º54′ N</geoCoordinate>
, 
<geoCoordinate id="F4865079FF8F3C2EFF2AFC69FEC0DAF7" box="[189,301,912,940]" degrees="51" direction="west" minutes="48" orientation="longitude" pageId="17" pageNumber="18" precision="925" value="-51.8">51º48′ W</geoCoordinate>
, 
<quantity id="564A9B5BFF8F3C2EFEACFC68FE97DAF7" box="[315,378,913,940]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="17" pageNumber="18" unit="m" value="35.0">35 m</quantity>
a.s.l., 
<date id="E50C107EFF8F3C2EFE5CFC6BFDA4DAF7" box="[459,585,914,940]" pageId="17" pageNumber="18" value="2014-05">May 2014</date>
, F. Feer leg. (
<collectionCode id="F7A3AE7BFF8F3C2EFD65FC6BFCA7DAF7" box="[754,842,914,940]" pageId="17" pageNumber="18">CEMT</collectionCode>
); 4 unsexed specimens, Nouveau Chantier, no date or collector (
<collectionCode id="F7A3AE7BFF8F3C2EFE1BFC4FFE05DA8B" box="[396,488,950,976]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="17" pageNumber="18">MNHN</collectionCode>
– Le Moult collection); 1 ♁, Nouveau Chantier, no date or collector (
<collectionCode id="F7A3AE7BFF8F3C2EFAB7FC4FFA9ADA8B" box="[1312,1399,950,976]" pageId="17" pageNumber="18">ZMHB</collectionCode>
– Le Moult collection); 
<specimenCount id="87B4FD37FF8F3C2EFE73FC23FD82DAAF" box="[484,623,986,1012]" pageId="17" pageNumber="18" type="generic">1 specimen</specimenCount>
, Nouveau Chantier, Bas-Maroni, no date or collector (
<collectionCode id="F7A3AE7BFF8F3C2EFA9EFC23FA84DAAF" box="[1289,1385,986,1012]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="17" pageNumber="18">MNHN</collectionCode>
); 6 ♁♁, 
<specimenCount id="87B4FD37FF8F3C2EFE87FC07FE8EDD42" box="[272,355,1022,1049]" pageId="17" pageNumber="18" type="female">12 ♀♀</specimenCount>
, 3 unsexed specimens, 
<taxonomicName id="56B24D3DFF8F3C2EFDEBFC07FD22DD43" box="[636,719,1022,1048]" class="Reptilia" family="Colubridae" genus="Regina" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="genus">Régina</taxonomicName>
(“S of 
<taxonomicName id="56B24D3DFF8F3C2EFCBFFC07FC6FDD42" box="[808,898,1022,1049]" class="Reptilia" family="Colubridae" genus="Regina" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="genus">Régina</taxonomicName>
”), 
<date id="E50C107EFF8F3C2EFC35FC07FBA8DD43" box="[930,1093,1022,1049]" pageId="17" pageNumber="18" value="2006-12-30">30 Dec. 2006</date>
, M. Snižek leg. (
<collectionCode id="F7A3AE7BFF8F3C2EFA87FC07FA87DD43" box="[1296,1386,1022,1048]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="17" pageNumber="18">NMPC</collectionCode>
); 4 ♁♁, 
<specimenCount id="87B4FD37FF8F3C2EFE84FBDAFEB5DD66" box="[275,344,1059,1085]" pageId="17" pageNumber="18" type="female">3 ♀♀</specimenCount>
, Roura, Fourgassie, 
<date id="E50C107EFF8F3C2EFDC5FBDBFCCEDD67" box="[594,803,1058,1085]" pageId="17" pageNumber="18" value="2006-12-15" valueMax="2006-12-30" valueMin="2006-12-15">15–30 Dec. 2006</date>
, M. Snižek leg. (
<collectionCode id="F7A3AE7BFF8F3C2EFC63FBDAFBA3DD66" box="[1012,1102,1059,1085]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="17" pageNumber="18">NMPC</collectionCode>
); 
<specimenCount id="87B4FD37FF8F3C2EFBF3FBDAFB7EDD66" box="[1124,1171,1059,1085]" pageId="17" pageNumber="18" type="female">1 ♀</specimenCount>
, Saint-Laurent-du- Maroni, no date and collector (
<collectionCode id="F7A3AE7BFF8F3C2EFDADFBBEFD7CDD3A" box="[570,657,1095,1121]" pageId="17" pageNumber="18">ZMHB</collectionCode>
– Le Moult collection); 1 unsexed specimen, Saint-Laurentdu-Maroni (“Maroni”), no date or collector (
<collectionCode id="F7A3AE7BFF8F3C2EFD4EFB92FCD4DDDE" box="[729,825,1131,1157]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="17" pageNumber="18">MNHN</collectionCode>
); 
<specimenCount id="87B4FD37FF8F3C2EFCC7FB92FC92DDDD" box="[848,895,1131,1158]" pageId="17" pageNumber="18" type="female">1 ♀</specimenCount>
, Saint-Laurent-du-Maroni (“Maroni”), no date or collector (
<collectionCode id="F7A3AE7BFF8F3C2EFE07FB76FE06DDF2" box="[400,491,1167,1193]" pageId="17" pageNumber="18">ZMHB</collectionCode>
); 1 ♁, 
<specimenCount id="87B4FD37FF8F3C2EFDA8FB76FD69DDF1" box="[575,644,1167,1194]" pageId="17" pageNumber="18" type="female">6 ♀♀</specimenCount>
, Saint-Laurent-du-Maroni (“Bas-Maroni”), no date or collector (
<collectionCode id="F7A3AE7BFF8F3C2EFF52FB4DFEE4DD95" box="[197,265,1204,1230]" pageId="17" pageNumber="18">ISNB</collectionCode>
– Le Moult collection); 1 unsexed specimen, Saint-Laurent-du-Maroni, “Jun”, no collector (
<collectionCode id="F7A3AE7BFF8F3C2EFF52FB21FECDDDA9" box="[197,288,1240,1266]" pageId="17" pageNumber="18">ZMHB</collectionCode>
); 1 ♁, Saint-Laurent-du-Maroni (“Maroni”), 1915, no collector (
<collectionCode id="F7A3AE7BFF8F3C2EFB8AFB21FB97DDA9" box="[1053,1146,1240,1266]" pageId="17" pageNumber="18">CMNC</collectionCode>
); 
<specimenCount id="87B4FD37FF8F3C2EFB18FB21FB51DDA9" box="[1167,1212,1240,1266]" pageId="17" pageNumber="18" type="female">1 ♀</specimenCount>
, Saint-Laurentdu-Maroni (“Maroni”), 
<date id="E50C107EFF8F3C2EFE43FB05FDA9DC4C" box="[468,580,1276,1303]" pageId="17" pageNumber="18" value="1925-09">Set. 1925</date>
, no collector (
<collectionCode id="F7A3AE7BFF8F3C2EFD7DFB05FCD0DC4D" box="[746,829,1276,1302]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="17" pageNumber="18" type="Museum">MZSP</collectionCode>
); 1 ♁, 
<specimenCount id="87B4FD37FF8F3C2EFC1CFB05FC5ADC4D" box="[907,951,1276,1302]" pageId="17" pageNumber="18" type="female">1 ♀</specimenCount>
, Saint-Laurent-du-Maroni, 
<date id="E50C107EFF8F3C2EFB6DFB05FA9CDC4D" box="[1274,1393,1276,1302]" pageId="17" pageNumber="18" value="1976-04">Apr. 1976</date>
, F. Chalumeau leg. (
<collectionCode id="F7A3AE7BFF8F3C2EFE33FAD9FE11DC61" box="[420,508,1312,1338]" pageId="17" pageNumber="18">CEMT</collectionCode>
); 2 ♁♁, 
<specimenCount id="87B4FD37FF8F3C2EFDF4FAD9FD7DDC60" box="[611,656,1312,1339]" pageId="17" pageNumber="18" type="female">1 ♀</specimenCount>
, Saint-Laurent-du-Maroni, 
<date id="E50C107EFF8F3C2EFC42FAD9FBA3DC61" box="[981,1102,1312,1338]" pageId="17" pageNumber="18" value="1976-04">Apr. 1976</date>
, P. Arnaud leg. (
<collectionCode id="F7A3AE7BFF8F3C2EFA85FAD9FA87DC61" box="[1298,1386,1312,1338]" pageId="17" pageNumber="18">CEMT</collectionCode>
); 1 ♁, Saint-Laurent-du-Maroni, Bélvédère de Saül (“Saül”), 
<date id="E50C107EFF8F3C2EFCEEFABDFC0ADC04" box="[889,999,1348,1375]" pageId="17" pageNumber="18" value="1979-07">Jul. 1979</date>
, G. Nazareth leg. (
<collectionCode id="F7A3AE7BFF8F3C2EFB55FABCFAF7DC04" box="[1218,1306,1349,1375]" pageId="17" pageNumber="18">CEMT</collectionCode>
); 6 ♁♁, 
<specimenCount id="87B4FD37FF8F3C2EFF2AFA90FEE9DCD8" box="[189,260,1385,1411]" pageId="17" pageNumber="18" type="female">3 ♀♀</specimenCount>
, Saint-Laurent-du-Maroni, Bélvédère de Saül, 
<geoCoordinate id="F4865079FF8F3C2EFCD4FA9EFC37DCD8" box="[835,986,1383,1411]" degrees="03" direction="north" minutes="37" orientation="latitude" pageId="17" pageNumber="18" precision="15" seconds="22" value="3.622778">03º37′22″ N</geoCoordinate>
, 
<geoCoordinate id="F4865079FF8F3C2EFC7DFA9EFB68DCD8" box="[1002,1157,1383,1411]" degrees="53" direction="west" minutes="12" orientation="longitude" pageId="17" pageNumber="18" precision="15" seconds="57" value="-53.215836">53º12′57″ W</geoCoordinate>
, 
<quantity id="564A9B5BFF8F3C2EFB02FA90FB0EDCD8" box="[1173,1251,1385,1411]" metricMagnitude="2" metricUnit="m" metricValue="3.26" pageId="17" pageNumber="18" unit="m" value="326.0">326 m</quantity>
a.s.l., flight interception trap, 
<date id="E50C107EFF8F3C2EFE1CFA74FDCADCFC" box="[395,551,1421,1447]" pageId="17" pageNumber="18" value="2011-05-13">13 May 2011</date>
, 
<collectionCode id="F7A3AE7BFF8F3C2EFDA4FA74FD6DDCFC" box="[563,640,1421,1447]" pageId="17" pageNumber="18">SEAG</collectionCode>
leg. (
<collectionCode id="F7A3AE7BFF8F3C2EFD57FA74FCF5DCFC" box="[704,792,1421,1447]" pageId="17" pageNumber="18">CEMT</collectionCode>
); 7 ♁♁, 
<specimenCount id="87B4FD37FF8F3C2EFCEDFA74FC51DCFC" box="[890,956,1421,1447]" pageId="17" pageNumber="18" type="female">3 ♀♀</specimenCount>
, Saint-Laurent-du-Maroni, Bélvédère de Saül, 
<geoCoordinate id="F4865079FF8F3C2EFEBFFA56FE50DC97" box="[296,445,1455,1484]" degrees="03" direction="north" minutes="37" orientation="latitude" pageId="17" pageNumber="18" precision="15" seconds="22" value="3.622778">03º37′22″ N</geoCoordinate>
, 
<geoCoordinate id="F4865079FF8F3C2EFE5CFA56FD89DC97" box="[459,612,1455,1484]" degrees="53" direction="west" minutes="12" orientation="longitude" pageId="17" pageNumber="18" precision="15" seconds="57" value="-53.215836">53º12′57″ W</geoCoordinate>
, 
<quantity id="564A9B5BFF8F3C2EFDE4FA48FD52DC97" box="[627,703,1457,1484]" metricMagnitude="2" metricUnit="m" metricValue="3.26" pageId="17" pageNumber="18" unit="m" value="326.0">326 m</quantity>
a.s.l., 
<date id="E50C107EFF8F3C2EFC86FA48FC4ADC90" box="[785,935,1457,1483]" pageId="17" pageNumber="18" value="2011-07-21">21 Jul. 2011</date>
, 
<collectionCode id="F7A3AE7BFF8F3C2EFC21FA48FBEEDC90" box="[950,1027,1457,1483]" pageId="17" pageNumber="18">SEAG</collectionCode>
leg. (
<collectionCode id="F7A3AE7BFF8F3C2EFBDEFA48FB4CDC90" box="[1097,1185,1457,1483]" pageId="17" pageNumber="18">CEMT</collectionCode>
); 14 ♁♁, 
<specimenCount id="87B4FD37FF8F3C2EFA8BFA48FA9DDC97" box="[1308,1392,1457,1484]" pageId="17" pageNumber="18" type="female">18 ♀♀</specimenCount>
, 2 unsexed specimens, Saint-Laurent-du-Maroni (“ 
<quantity id="564A9B5BFF8F3C2EFC99FA2CFCB8DCB4" box="[782,853,1493,1519]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="18" unit="km" value="20.0">20km</quantity>
E of St. Laurent du Maroni”), [route] Crique Naï, 
<date id="E50C107EFF8F3C2EFF6EFA03FE4DDF48" box="[249,416,1529,1556]" pageId="17" pageNumber="18" value="2006-12-10">10 Dec. 2006</date>
, M. Snižek leg. (
<collectionCode id="F7A3AE7BFF8F3C2EFDE6FA03FD26DF4F" box="[625,715,1530,1556]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="17" pageNumber="18">NMPC</collectionCode>
); 1 ♁, 
<specimenCount id="87B4FD37FF8F3C2EFCB4FA03FCBFDF4F" box="[803,850,1530,1556]" pageId="17" pageNumber="18" type="female">1 ♀</specimenCount>
, Saint-Laurent-du-Maroni, Maripasoula, Saül (“ 
<quantity id="564A9B5BFF8F3C2EFF47F9E7FEE5DF63" box="[208,264,1566,1592]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="17" pageNumber="18" unit="km" value="7.0">7km</quantity>
N”), Les Eaux Claires, 
<geoCoordinate id="F4865079FF8F3C2EFD8DF9E5FD46DF63" box="[538,683,1564,1592]" degrees="03" direction="north" minutes="39" orientation="latitude" pageId="17" pageNumber="18" precision="15" seconds="46" value="3.662778">03º39′46″ N</geoCoordinate>
, 
<geoCoordinate id="F4865079FF8F3C2EFD21F9E5FCA6DF63" box="[694,843,1564,1592]" degrees="53" direction="west" minutes="13" orientation="longitude" pageId="17" pageNumber="18" precision="15" seconds="19" value="-53.221947">53º13′19″ W</geoCoordinate>
, human faeces baited trap, 
<date id="E50C107EFF8F3C2EFB15F9E7FA9CDF63" box="[1154,1393,1566,1592]" pageId="17" pageNumber="18" value="1997-05-31" valueMax="1997-06-03" valueMin="1997-05-31">31 May–3 Jun. 1997</date>
, J. Ashe and 
<collectionCode id="F7A3AE7BFF8F3C2EFEDDF9BAFE8CDF06" box="[330,353,1603,1629]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="17" pageNumber="18">R</collectionCode>
. Brooks leg. (
<collectionCode id="F7A3AE7BFF8F3C2EFD95F9BBFDB2DF07" box="[514,607,1602,1628]" pageId="17" pageNumber="18">CMNC</collectionCode>
); 1 ♁, Saint-Laurent-du-Maroni, Maripasoula, Saül (“ 
<quantity id="564A9B5BFF8F3C2EFB44F9BBFAE6DF07" box="[1235,1291,1602,1628]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="17" pageNumber="18" unit="km" value="7.0">7km</quantity>
N”), Les Eaux Claires, 
<geoCoordinate id="F4865079FF8F3C2EFEF4F99DFE1BDFDA" box="[355,502,1636,1665]" degrees="03" direction="north" minutes="39" orientation="latitude" pageId="17" pageNumber="18" precision="15" seconds="46" value="3.662778">03º39′46″ N</geoCoordinate>
, 
<geoCoordinate id="F4865079FF8F3C2EFD94F99DFD74DFDA" box="[515,665,1636,1665]" degrees="53" direction="west" minutes="13" orientation="longitude" pageId="17" pageNumber="18" precision="15" seconds="19" value="-53.221947">53º13′19″ W</geoCoordinate>
, flight interception trap, 
<date id="E50C107EFF8F3C2EFC2CF99FFB89DFDB" box="[955,1124,1638,1665]" pageId="17" pageNumber="18" value="1997-06-01" valueMax="1997-06-08" valueMin="1997-06-01">1–8 Jun. 1997</date>
, J. Ashe and 
<collectionCode id="F7A3AE7BFF8F3C2EFB68F99EFAFBDFDA" box="[1279,1302,1639,1665]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="17" pageNumber="18">R</collectionCode>
. Brooks leg. (
<collectionCode id="F7A3AE7BFF8F3C2EFF60F972FEB9DFFE" box="[247,340,1675,1701]" pageId="17" pageNumber="18">CMNC</collectionCode>
); 2 ♁♁, Saint-Laurent-du-Maroni, Maripasoula, Saül (“ 
<quantity id="564A9B5BFF8F3C2EFC4CF973FBFEDFFF" box="[987,1043,1674,1700]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="17" pageNumber="18" unit="km" value="7.0">7km</quantity>
N”), Les Eaux Claires, Mount La Fumée, 
<geoCoordinate id="F4865079FF8F3C2EFED1F954FE31DF92" box="[326,476,1709,1737]" degrees="03" direction="north" minutes="39" orientation="latitude" pageId="17" pageNumber="18" precision="15" seconds="46" value="3.662778">03º39′46″ N</geoCoordinate>
, 
<geoCoordinate id="F4865079FF8F3C2EFE7DF954FD6EDF92" box="[490,643,1709,1737]" degrees="53" direction="west" minutes="13" orientation="longitude" pageId="17" pageNumber="18" precision="15" seconds="19" value="-53.221947">53º13′19″ W</geoCoordinate>
, flight interception trap, 
<date id="E50C107EFF8F3C2EFC3AF956FBB7DF92" box="[941,1114,1711,1737]" pageId="17" pageNumber="18" value="1997-06-01" valueMax="1997-06-08" valueMin="1997-06-01">1–8 Jun. 1997</date>
, J. Ashe and 
<collectionCode id="F7A3AE7BFF8F3C2EFB6AF956FAF9DF92" box="[1277,1300,1711,1737]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="17" pageNumber="18">R</collectionCode>
. Brooks leg. (
<collectionCode id="F7A3AE7BFF8F3C2EFF6EF92AFEBBDFB6" box="[249,342,1747,1773]" pageId="17" pageNumber="18">CMNC</collectionCode>
); 
<specimenCount id="87B4FD37FF8F3C2EFEFCF92AFE75DFB5" box="[363,408,1747,1774]" pageId="17" pageNumber="18" type="female">1 ♀</specimenCount>
, Saint-Laurent-du-Maroni, Maripasoula, Saül (“ 
<quantity id="564A9B5BFF8F3C2EFC45F92AFBE7DFB6" box="[978,1034,1747,1773]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="17" pageNumber="18" unit="km" value="7.0">7km</quantity>
N”), Les Eaux Claires, Mount La Fumée, 
<geoCoordinate id="F4865079FF8F3C2EFED1F90CFE36DE49" box="[326,475,1781,1810]" degrees="03" direction="north" minutes="39" orientation="latitude" pageId="17" pageNumber="18" precision="15" seconds="46" value="3.662778">03º39′46″ N</geoCoordinate>
, 
<geoCoordinate id="F4865079FF8F3C2EFE7EF90CFD6FDE49" box="[489,642,1781,1810]" degrees="53" direction="west" minutes="13" orientation="longitude" pageId="17" pageNumber="18" precision="15" seconds="19" value="-53.221947">53º13′19″ W</geoCoordinate>
, human faeces trap, 
<date id="E50C107EFF8F3C2EFCEDF90EFBCBDE4A" box="[890,1062,1783,1810]" pageId="17" pageNumber="18" value="1997-06-01" valueMax="1997-06-08" valueMin="1997-06-01">1–8 Jun. 1997</date>
, J. Ashe and 
<collectionCode id="F7A3AE7BFF8F3C2EFB5EF901FB0DDE49" box="[1225,1248,1784,1810]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="17" pageNumber="18">R</collectionCode>
. Brooks leg. (
<collectionCode id="F7A3AE7BFF8F3C2EFF52F8E5FECFDE6D" box="[197,290,1820,1846]" pageId="17" pageNumber="18">CMNC</collectionCode>
); 
<specimenCount id="87B4FD37FF8F3C2EFEADF8E5FE6CDE6D" box="[314,385,1820,1846]" pageId="17" pageNumber="18" type="female">3 ♀♀</specimenCount>
, Saint-Laurent-du-Maroni, Maripasoula, Saül (“ 
<quantity id="564A9B5BFF8F3C2EFC53F8E2FC10DE6E" box="[964,1021,1819,1845]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="17" pageNumber="18" unit="km" value="7.0">7km</quantity>
N”), Les Eaux Claires, Mount La Fumée, 
<geoCoordinate id="F4865079FF8F3C2EFED2F8C7FE34DE01" box="[325,473,1854,1882]" degrees="03" direction="north" minutes="39" orientation="latitude" pageId="17" pageNumber="18" precision="15" seconds="46" value="3.662778">03º39′46″ N</geoCoordinate>
, 
<geoCoordinate id="F4865079FF8F3C2EFE70F8C7FD92DE01" box="[487,639,1854,1882]" degrees="53" direction="west" minutes="13" orientation="longitude" pageId="17" pageNumber="18" precision="15" seconds="19" value="-53.221947">53º13′19″ W</geoCoordinate>
, 
<quantity id="564A9B5BFF8F3C2EFD1BF8B9FD35DE01" box="[652,728,1856,1882]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="17" pageNumber="18" unit="m" value="300.0">300 m</quantity>
a.s.l., human faeces trap, 
<date id="E50C107EFF8F3C2EFB87F8B9FB56DE01" box="[1040,1211,1856,1882]" pageId="17" pageNumber="18" value="1997-06-04" valueMax="1997-06-08" valueMin="1997-06-04">4–8 Jun. 1997</date>
, J. Ashe and 
<collectionCode id="F7A3AE7BFF8F3C2EFACBF8B9FA9EDE01" box="[1372,1395,1856,1882]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="17" pageNumber="18">R</collectionCode>
. Brooks leg. (
<collectionCode id="F7A3AE7BFF8F3C2EFEC5F89DFE42DE25" box="[338,431,1892,1918]" pageId="17" pageNumber="18">CMNC</collectionCode>
); 2 ♁♁, Saint-Laurent-du-Maroni, Maripasoula, Saül, Mount Galbao, 
<geoCoordinate id="F4865079FF8F3C2EFB77F89BFA9FDE24" box="[1248,1394,1890,1919]" degrees="03" direction="north" minutes="37" orientation="latitude" pageId="17" pageNumber="18" precision="15" seconds="18" value="3.621667">03º37′18″ N</geoCoordinate>
, 
<geoCoordinate id="F4865079FF8F3C2EFF2AF87FFEB9DEF8" box="[189,340,1926,1955]" degrees="53" direction="west" minutes="16" orientation="longitude" pageId="17" pageNumber="18" precision="15" seconds="42" value="-53.27833">53º16′42″ W</geoCoordinate>
, 
<quantity id="564A9B5BFF8F3C2EFEF7F871FE46DEF8" box="[352,427,1928,1955]" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="17" pageNumber="18" unit="m" value="740.0">740 m</quantity>
a.s.l., flight interception trap, 
<date id="E50C107EFF8F3C2EFC99F871FC5ADEF9" box="[782,951,1928,1955]" pageId="17" pageNumber="18" value="1997-06-05" valueMax="1997-06-07" valueMin="1997-06-05">5–7 Jun. 1997</date>
, J. Ashe and 
<collectionCode id="F7A3AE7BFF8F3C2EFBC5F870FB84DEF8" box="[1106,1129,1929,1955]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="17" pageNumber="18">R</collectionCode>
. Brooks leg. (
<collectionCode id="F7A3AE7BFF8F3C2EFA9AF871FA87DEF9" box="[1293,1386,1928,1954]" pageId="17" pageNumber="18">CMNC</collectionCode>
); 5 ♁♁, 
<specimenCount id="87B4FD37FF8F3C2EFE84F854FEB5DE9C" box="[275,344,1965,1991]" pageId="17" pageNumber="18" type="female">2 ♀♀</specimenCount>
, Sinnamary, Saint-Eugène, Barrage de Petit Saut, 
<geoCoordinate id="F4865079FF8F3C2EFC2FF852FBC9DE9C" box="[952,1060,1963,1991]" degrees="04" direction="north" minutes="51" orientation="latitude" pageId="17" pageNumber="18" precision="925" value="4.85">04º51′ N</geoCoordinate>
, 
<geoCoordinate id="F4865079FF8F3C2EFBA5F852FB4FDE9C" box="[1074,1186,1963,1991]" degrees="53" direction="west" minutes="04" orientation="longitude" pageId="17" pageNumber="18" precision="925" value="-53.066666">53º04′ W</geoCoordinate>
, 
<quantity id="564A9B5BFF8F3C2EFB27F854FB03DE9C" box="[1200,1262,1965,1991]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="17" pageNumber="18" unit="m" value="90.0">90 m</quantity>
a.s.l., 
<date id="E50C107EFF8F3C2EFAD7F854FF17DEB0" pageId="17" pageNumber="18" value="1998-12">Dec. 1998</date>
, F. Feer leg. (
<collectionCode id="F7A3AE7BFF8F3C2EFE0EF828FE1CDEB0" box="[409,497,2001,2027]" pageId="17" pageNumber="18">CEMT</collectionCode>
).
</paragraph>
<paragraph id="910D36BEFF8C3C2DFF2AFEF5FC1BDB5E" blockId="18.[189,1399,267,517]" pageId="18" pageNumber="19">
<collectingCountry id="E9A5762EFF8C3C2DFF2AFEF5FEC1D87D" box="[189,300,268,294]" name="Brazil" pageId="18" pageNumber="19">BRAZIL</collectingCountry>
: 1 unsexed specimen, no more data (
<collectionCode id="F7A3AE7BFF8C3C2DFD42FEF2FCD8D87E" box="[725,821,267,293]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="18" pageNumber="19">MNHN</collectionCode>
); 
<specimenCount id="87B4FD37FF8C3C2DFCDFFEF2FC9ED87D" box="[840,883,267,294]" pageId="18" pageNumber="19" type="female">1 ♀</specimenCount>
, illegible information (collector?) (
<collectionCode id="F7A3AE7BFF8C3C2DFA98FEF2FA87D87E" box="[1295,1386,267,293]" pageId="18" pageNumber="19">ZMHB</collectionCode>
). – 
<emphasis id="A3C6EAACFF8C3C2DFF45FEC9FEC0D810" bold="true" box="[210,301,304,331]" pageId="18" pageNumber="19">
<collectingRegion id="5376F85CFF8C3C2DFF45FEC9FEC0D810" box="[210,301,304,331]" country="Brazil" name="Amapa" pageId="18" pageNumber="19">Amapá</collectingRegion>
</emphasis>
: 1 ♁, 
<specimenCount id="87B4FD37FF8C3C2DFEE4FEC8FE73D810" box="[371,414,305,331]" pageId="18" pageNumber="19" type="female">1 ♀</specimenCount>
, Serra do Navio, 
<date id="E50C107EFF8C3C2DFDF0FEC8FD35D810" box="[615,728,304,331]" pageId="18" pageNumber="19" value="1957-01">Jan. 1957</date>
, Pereira &amp; Martínez leg. (
<collectionCode id="F7A3AE7BFF8C3C2DFB95FEC8FBB2D810" box="[1026,1119,305,331]" pageId="18" pageNumber="19">CMNC</collectionCode>
); 
<specimenCount id="87B4FD37FF8C3C2DFBE5FEC8FB59D810" box="[1138,1204,305,331]" pageId="18" pageNumber="19" type="female">4 ♀♀</specimenCount>
, Serra do Navio, 
<date id="E50C107EFF8C3C2DFF2AFEAFFED8D82A" box="[189,309,342,369]" pageId="18" pageNumber="19" value="1957-09">Sep. 1957</date>
, Pereira &amp; Martínez leg. (
<collectionCode id="F7A3AE7BFF8C3C2DFDFFFEAFFD28D82B" box="[616,709,342,368]" pageId="18" pageNumber="19">CMNC</collectionCode>
); 1 ♁, 2 unsexed specimens, Serra do Navio, 
<geoCoordinate id="F4865079FF8C3C2DFB49FEADFA9FD82B" box="[1246,1394,340,368]" degrees="00" direction="north" minutes="53" orientation="latitude" pageId="18" pageNumber="19" precision="15" seconds="06" value="0.885">00º53′06″ N</geoCoordinate>
, 
<geoCoordinate id="F4865079FF8C3C2DFF2AFE80FEB4D8CD" box="[189,345,377,406]" degrees="51" direction="west" minutes="52" orientation="longitude" pageId="18" pageNumber="19" precision="15" seconds="53" value="-51.881386">51º52′53″ W</geoCoordinate>
, 
<date id="E50C107EFF8C3C2DFEFEFE82FE0AD8CE" box="[361,487,379,405]" pageId="18" pageNumber="19" value="2000-09">Sep. 2000</date>
, 
<collectionCode id="F7A3AE7BFF8C3C2DFE6FFE85FDE2D8CD" box="[504,527,380,406]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="18" pageNumber="19">R</collectionCode>
. Ribon leg. (
<collectionCode id="F7A3AE7BFF8C3C2DFD24FE82FCE6D8CE" box="[691,779,379,405]" pageId="18" pageNumber="19">CEMT</collectionCode>
). – 
<emphasis id="A3C6EAACFF8C3C2DFCA8FE83FC96D8CE" bold="true" box="[831,891,378,405]" pageId="18" pageNumber="19">
<collectingRegion id="5376F85CFF8C3C2DFCA8FE83FC96D8CE" box="[831,891,378,405]" country="Brazil" name="Para" pageId="18" pageNumber="19">Pará</collectingRegion>
</emphasis>
: 6 unsexed specimens, Almeirim, Monte Dourado, Pacanari, 0º81′ S, 
<geoCoordinate id="F4865079FF8C3C2DFD85FE67FD6CD8E0" box="[530,641,414,443]" degrees="52" direction="west" minutes="56" orientation="longitude" pageId="18" pageNumber="19" precision="925" value="-52.933334">52º56′ W</geoCoordinate>
, 
<date id="E50C107EFF8C3C2DFD18FE58FCE3D8E1" box="[655,782,416,443]" pageId="18" pageNumber="19" value="2006-03">Mar. 2006</date>
, 
<collectionCode id="F7A3AE7BFF8C3C2DFC8CFE58FCC3D8E0" box="[795,814,417,443]" name="Tavera, Department of Geology and Geophysics" pageId="18" pageNumber="19">T</collectionCode>
. Gardner and M. Hernández leg. (
<collectionCode id="F7A3AE7BFF8C3C2DFB5CFE59FAC7D8E1" box="[1227,1322,416,442]" pageId="18" pageNumber="19">BMNH</collectionCode>
); 1 ♁, 
<specimenCount id="87B4FD37FF8C3C2DFF2AFE3FFF04D8BB" box="[189,233,454,480]" pageId="18" pageNumber="19" type="female">1 ♀</specimenCount>
, Almeirim, Monte Dourado, 
<geoCoordinate id="F4865079FF8C3C2DFDA9FE3DFD02D8BB" box="[574,751,452,480]" degrees="00" direction="south" minutes="50" orientation="latitude" pageId="18" pageNumber="19" precision="1" seconds="08.89" value="-0.83580273">00º50′08.89″ S</geoCoordinate>
, 
<geoCoordinate id="F4865079FF8C3C2DFD6CFE3DFC7CD8BB" box="[763,913,452,480]" degrees="53" direction="west" minutes="02" orientation="longitude" pageId="18" pageNumber="19" precision="15" seconds="34" value="-53.042778">53º02′34″ W</geoCoordinate>
, 
<date id="E50C107EFF8C3C2DFC0AFE3FFBF5D8BB" box="[925,1048,454,480]" pageId="18" pageNumber="19" value="2009-03">Mar. 2009</date>
, Schiffler 
<emphasis id="A3C6EAACFF8C3C2DFB18FE3FFB3DD8BB" box="[1167,1232,454,480]" italics="true" pageId="18" pageNumber="19">et. al.</emphasis>
leg. (
<collectionCode id="F7A3AE7BFF8C3C2DFA85FE3FFA87D8BB" box="[1298,1386,454,480]" pageId="18" pageNumber="19">CEMT</collectionCode>
); 3 unsexed specimens, Santarém, 
<date id="E50C107EFF8C3C2DFDD6FE12FD51DB5E" box="[577,700,491,517]" pageId="18" pageNumber="19" value="1990-03">Mar. 1990</date>
, O. Schmitt leg. (
<collectionCode id="F7A3AE7BFF8C3C2DFC1EFE12FC07DB5E" box="[905,1002,491,517]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="18" pageNumber="19">MNHN</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="D9A86535FF8C3C2DFF2AFDD7FE0EDA55" pageId="18" pageNumber="19" type="discussion">
<paragraph id="910D36BEFF8C3C2DFF2AFDD7FD7FDB13" blockId="18.[189,658,558,584]" box="[189,658,558,584]" pageId="18" pageNumber="19">
<heading id="CA4581D2FF8C3C2DFF2AFDD7FD7FDB13" bold="true" box="[189,658,558,584]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="A3C6EAACFF8C3C2DFF2AFDD7FD7FDB13" bold="true" box="[189,658,558,584]" pageId="18" pageNumber="19">
Remarks on the 
<typeStatus id="4E09881CFF8C3C2DFE1CFDD7FE16DB13" box="[395,507,558,584]" pageId="18" pageNumber="19" type="lectotype">lectotype</typeStatus>
designation
</emphasis>
</heading>
</paragraph>
<paragraph id="910D36BEFF8C3C2DFF2AFDA6FE0EDA55" blockId="18.[189,1399,607,782]" pageId="18" pageNumber="19">
The specimen designated herein as 
<typeStatus id="4E09881CFF8C3C2DFDF3FDA6FD3DDB22" box="[612,720,607,633]" pageId="18" pageNumber="19" type="lectotype">lectotype</typeStatus>
has a label with Pierre François Marie Auguste Dejean’s handwriting indicating that François Louis Nompar de Caumont Laporte, the Comte of Castelnau, is the author of 
<taxonomicName id="56B24D3DFF8C3C2DFEA4FD50FE65DB98" authorityName="Castelnau" authorityYear="1840" box="[307,392,681,707]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="A3C6EAACFF8C3C2DFEA4FD50FE65DB98" box="[307,392,681,707]" italics="true" pageId="18" pageNumber="19">bicolor</emphasis>
</taxonomicName>
and the collector of that specimen. We are aware that Castelnau sold or donated his collections, but there is no evidence to state that all 
<typeStatus id="4E09881CFF8C3C2DFC9AFD36FC99DBB2" box="[781,884,719,745]" pageId="18" pageNumber="19" type="syntype">syntypes</typeStatus>
of Castelnau were sold to a single collection or went to a single place.
</paragraph>
</subSubSection>
<subSubSection id="D9A86535FF8C3C28FF2AFCCEFBFCD8A4" lastPageId="23" lastPageNumber="24" pageId="18" pageNumber="19" type="description">
<paragraph id="910D36BEFF8C3C2DFF2AFCCEFE86DA0A" blockId="18.[189,363,823,849]" box="[189,363,823,849]" pageId="18" pageNumber="19">
<heading id="CA4581D2FF8C3C2DFF2AFCCEFE86DA0A" bold="true" box="[189,363,823,849]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="A3C6EAACFF8C3C2DFF2AFCCEFE86DA0A" bold="true" box="[189,363,823,849]" pageId="18" pageNumber="19">Redescription</emphasis>
</heading>
</paragraph>
<paragraph id="910D36BEFF8C3C2DFF2AFC91FC8ADAFC" blockId="18.[189,1399,872,936]" pageId="18" pageNumber="19">
MALE. Dorsal surface pronotum and elytra disc with different colours, pronotum with light green metallic sheen and elytra with dark green metallic sheen (
<figureCitation id="09892A3BFF8C3C2DFD6CFC74FCB7DAFC" box="[763,858,909,935]" captionStart="Fig" captionStartId="19.[189,232,1894,1920]" captionTargetBox="[202,1379,264,1882]" captionTargetId="figure@19.[189,1399,263,1882]" captionTargetPageId="19" captionText="Fig. 4. Canthon (G.) bicolor Castelnau, 1840. A. Habitus, dorsal view. B. Ventral surface of meso- and metafemora and metasternal anterior lobe not convex. C. Dorsal view showing convex pygidium. D. Male protibial spur. E. Female protibial spur. F. Paramera lateral view. G. Paramera dorsal view. Scale bar: A = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3787895" httpUri="https://zenodo.org/record/3787895/files/figure.png" pageId="18" pageNumber="19">Fig. 4A</figureCitation>
).
</paragraph>
<paragraph id="910D36BEFF8C3C2DFF2AFC28FD90DD6D" blockId="18.[189,1399,977,1078]" pageId="18" pageNumber="19">HEAD. Frontal surface smooth, lacking horns or tubercles. Clypeal teeth with rounded apex, with thin setae on apex and separated by a U-shaped emargination. Clypeo-genal junction slightly emarginated, rounded. Clypeo-genal suture present.</paragraph>
<paragraph id="910D36BEFF8C3C2DFF2AFBA6FE3ADD98" blockId="18.[189,1399,1119,1220]" pageId="18" pageNumber="19">PRONOTUM. Disc simply convex, smooth, lacking microscultpure (120× magnification). Anterior and posterior angles acute. Lateral borders effaced. Posterior border with lateral semicircular emarginations near to posterior angles.</paragraph>
<paragraph id="910D36BEFF8C3C2DFF2AFB14FE74DC77" blockId="18.[189,1399,1260,1324]" pageId="18" pageNumber="19">HYPOMERON. Lacking transverse keel, surface black with chagrined microsculpture. Sparse setae present at anterior portion.</paragraph>
<paragraph id="910D36BEFF8C3C2DFF2AFAAFFCD1DC34" blockId="18.[189,828,1365,1392]" box="[189,828,1365,1392]" pageId="18" pageNumber="19">
METASTERNUM. Metasternal anterior lobe flat (
<figureCitation id="09892A3BFF8C3C2DFD45FAACFCC2DC34" box="[722,815,1365,1391]" captionStart="Fig" captionStartId="19.[189,232,1894,1920]" captionTargetBox="[202,1379,264,1882]" captionTargetId="figure@19.[189,1399,263,1882]" captionTargetPageId="19" captionText="Fig. 4. Canthon (G.) bicolor Castelnau, 1840. A. Habitus, dorsal view. B. Ventral surface of meso- and metafemora and metasternal anterior lobe not convex. C. Dorsal view showing convex pygidium. D. Male protibial spur. E. Female protibial spur. F. Paramera lateral view. G. Paramera dorsal view. Scale bar: A = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3787895" httpUri="https://zenodo.org/record/3787895/files/figure.png" pageId="18" pageNumber="19">Fig. 4B</figureCitation>
).
</paragraph>
<paragraph id="910D36BEFF8C3C2DFF2AFA60FE84DC83" blockId="18.[189,1399,1431,1496]" pageId="18" pageNumber="19">
ELYTRA. Striae weakly impressed, more distinct near elytral basis, 8 
<superScript id="66C79BF6FF8C3C2DFC42FA6EFC0FDCFE" attach="left" box="[981,994,1431,1445]" fontSize="6" pageId="18" pageNumber="19">th</superScript>
stria carinate along its basal third. Interstriae flat.
</paragraph>
<paragraph id="910D36BEFF8C3C2DFF2AF9FBFDEFDFEB" blockId="18.[189,1399,1537,1713]" pageId="18" pageNumber="19">
LEGS. Protibiae three-toothed, dorsal face with two tufts of setae present on each tibial teeth. Protibial spur truncated, larger than tall and apically emarginated, forming two spines (
<figureCitation id="09892A3BFF8C3C2DFBDBF9DEFB46DF1A" box="[1100,1195,1575,1601]" captionStart="Fig" captionStartId="19.[189,232,1894,1920]" captionTargetBox="[202,1379,264,1882]" captionTargetId="figure@19.[189,1399,263,1882]" captionTargetPageId="19" captionText="Fig. 4. Canthon (G.) bicolor Castelnau, 1840. A. Habitus, dorsal view. B. Ventral surface of meso- and metafemora and metasternal anterior lobe not convex. C. Dorsal view showing convex pygidium. D. Male protibial spur. E. Female protibial spur. F. Paramera lateral view. G. Paramera dorsal view. Scale bar: A = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3787895" httpUri="https://zenodo.org/record/3787895/files/figure.png" pageId="18" pageNumber="19">Fig. 4D</figureCitation>
). External border of protibia serrated. Profemur ventral face brown, glabrous, without a distal tooth on its anterior border. Ventral face of meso- and metatibiae rugose on posterior half (
<figureCitation id="09892A3BFF8C3C2DFC34F988FBECDFD0" box="[931,1025,1649,1675]" captionStart="Fig" captionStartId="19.[189,232,1894,1920]" captionTargetBox="[202,1379,264,1882]" captionTargetId="figure@19.[189,1399,263,1882]" captionTargetPageId="19" captionText="Fig. 4. Canthon (G.) bicolor Castelnau, 1840. A. Habitus, dorsal view. B. Ventral surface of meso- and metafemora and metasternal anterior lobe not convex. C. Dorsal view showing convex pygidium. D. Male protibial spur. E. Female protibial spur. F. Paramera lateral view. G. Paramera dorsal view. Scale bar: A = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3787895" httpUri="https://zenodo.org/record/3787895/files/figure.png" pageId="18" pageNumber="19">Fig. 4B</figureCitation>
). Meta-trochanter having a hair tuft near femur articulation.
</paragraph>
<paragraph id="910D36BEFF8C3C2DFF2AF923FBD8DFAF" blockId="18.[189,1077,1754,1781]" box="[189,1077,1754,1781]" pageId="18" pageNumber="19">ABDOMEN. Ventrites glabrous, lacking punctures and not narrowed medially.</paragraph>
<paragraph id="910D36BEFF8C3C2DFF2AF8E7FE23DE06" blockId="18.[189,1398,1821,1885]" pageId="18" pageNumber="19">PYGIDIUM. Convex. Basal line delimiting pygidium from pro-pygidium. Metallic sheen pattern similar to that of pronotal disc.</paragraph>
<paragraph id="910D36BEFF8C3C2DFF2AF87EFDC9DEB0" blockId="18.[189,1399,1926,2027]" pageId="18" pageNumber="19">
PARAMERA. In lateral view (
<figureCitation id="09892A3BFF8C3C2DFD9DF87FFD85DEFB" box="[522,616,1926,1952]" captionStart="Fig" captionStartId="19.[189,232,1894,1920]" captionTargetBox="[202,1379,264,1882]" captionTargetId="figure@19.[189,1399,263,1882]" captionTargetPageId="19" captionText="Fig. 4. Canthon (G.) bicolor Castelnau, 1840. A. Habitus, dorsal view. B. Ventral surface of meso- and metafemora and metasternal anterior lobe not convex. C. Dorsal view showing convex pygidium. D. Male protibial spur. E. Female protibial spur. F. Paramera lateral view. G. Paramera dorsal view. Scale bar: A = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3787895" httpUri="https://zenodo.org/record/3787895/files/figure.png" pageId="18" pageNumber="19">Fig. 4F</figureCitation>
): flattened; apex semicircular; upper angle rounded; lower angle triangular. In dorsal view (
<figureCitation id="09892A3BFF8C3C2DFE69F852FD8DDE9D" box="[510,608,1963,1990]" captionStart="Fig" captionStartId="19.[189,232,1894,1920]" captionTargetBox="[202,1379,264,1882]" captionTargetId="figure@19.[189,1399,263,1882]" captionTargetPageId="19" captionText="Fig. 4. Canthon (G.) bicolor Castelnau, 1840. A. Habitus, dorsal view. B. Ventral surface of meso- and metafemora and metasternal anterior lobe not convex. C. Dorsal view showing convex pygidium. D. Male protibial spur. E. Female protibial spur. F. Paramera lateral view. G. Paramera dorsal view. Scale bar: A = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3787895" httpUri="https://zenodo.org/record/3787895/files/figure.png" pageId="18" pageNumber="19">Fig. 4G</figureCitation>
): membranous portion wider at central portion and narrower near apex and basis. Apex rounded.
</paragraph>
<caption id="C5CD6636FF8D3C2CFF2AF89FFE26DEB1" ID-DOI="http://doi.org/10.5281/zenodo.3787895" ID-Zenodo-Dep="3787895" httpUri="https://zenodo.org/record/3787895/files/figure.png" pageId="19" pageNumber="20" startId="19.[189,232,1894,1920]" targetBox="[202,1379,264,1882]" targetPageId="19">
<paragraph id="910D36BEFF8D3C2CFF2AF89FFE26DEB1" blockId="19.[189,1399,1894,2027]" pageId="19" pageNumber="20">
<emphasis id="A3C6EAACFF8D3C2CFF2AF89FFEE3DEDB" bold="true" box="[189,270,1894,1920]" pageId="19" pageNumber="20">Fig. 4.</emphasis>
<taxonomicName id="56B24D3DFF8D3C2CFE8EF89FFD0ADEDB" authority="Castelnau, 1840" authorityName="Castelnau" authorityYear="1840" box="[281,743,1894,1921]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="bicolor" subGenus="Goniocanthon">
<emphasis id="A3C6EAACFF8D3C2CFE8EF89FFDF4DEDB" box="[281,537,1894,1921]" italics="true" pageId="19" pageNumber="20">Canthon (G.) bicolor</emphasis>
<bibRefCitation id="F5234B4FFF8D3C2CFDB4F89FFD0ADEDB" author="Castelnau Comte" box="[547,743,1894,1920]" pageId="19" pageNumber="20" refId="ref16713" refString="Castelnau Comte de. 1840. Histoire naturelle des insectes coleopteres. Tome Deuxieme. Societe bibliophile, Paris. https: // doi. org / 10.5962 / bhl. title. 36538" type="book" year="1840">Castelnau, 1840</bibRefCitation>
</taxonomicName>
. 
<emphasis id="A3C6EAACFF8D3C2CFD61F89FFCE6DEDB" bold="true" box="[758,779,1894,1920]" pageId="19" pageNumber="20">A</emphasis>
. Habitus, dorsal view. 
<emphasis id="A3C6EAACFF8D3C2CFBB0F89EFBD6DEDA" bold="true" box="[1063,1083,1895,1921]" pageId="19" pageNumber="20">B</emphasis>
. Ventral surface of meso- and metafemora and metasternal anterior lobe not convex. 
<emphasis id="A3C6EAACFF8D3C2CFCEFF870FC60DEF8" bold="true" box="[888,909,1929,1955]" pageId="19" pageNumber="20">C</emphasis>
. Dorsal view showing convex pygidium. 
<emphasis id="A3C6EAACFF8D3C2CFF2AF854FF3FDE9C" bold="true" box="[189,210,1965,1991]" pageId="19" pageNumber="20">D</emphasis>
. Male protibial spur. 
<emphasis id="A3C6EAACFF8D3C2CFE4EF854FE00DE9C" bold="true" box="[473,493,1965,1991]" pageId="19" pageNumber="20">E</emphasis>
. Female protibial spur. 
<emphasis id="A3C6EAACFF8D3C2CFC99F854FCCDDE9C" bold="true" box="[782,800,1965,1991]" pageId="19" pageNumber="20">F</emphasis>
. Paramera lateral view. 
<emphasis id="A3C6EAACFF8D3C2CFBD1F854FBB0DE9C" bold="true" box="[1094,1117,1965,1991]" pageId="19" pageNumber="20">G</emphasis>
. Paramera dorsal view. Scale bar: A = 5.0 mm.
</paragraph>
</caption>
<caption id="C5CD6636FF8A3C2BFF2AF855FC66DEB1" ID-DOI="http://doi.org/10.5281/zenodo.3787897" ID-Zenodo-Dep="3787897" httpUri="https://zenodo.org/record/3787897/files/figure.png" pageId="20" pageNumber="21" startId="20.[189,232,1964,1990]" targetBox="[227,1373,428,1901]" targetPageId="20">
<paragraph id="910D36BEFF8A3C2BFF2AF855FC66DEB1" blockId="20.[189,1399,1964,2026]" pageId="20" pageNumber="21">
<emphasis id="A3C6EAACFF8A3C2BFF2AF855FEE0DE9C" bold="true" box="[189,269,1964,1991]" pageId="20" pageNumber="21">Fig. 5.</emphasis>
Sclerites of the internal sac of 
<taxonomicName id="56B24D3DFF8A3C2BFD18F855FAEADE9D" authority="Schmidt, 1922" authorityName="Schmidt" authorityYear="1922" box="[655,1287,1964,1991]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="subSpecies" species="smaragdulus" status="stat. nov." subGenus="Goniocanthon" subSpecies="subviridis">
<emphasis id="A3C6EAACFF8A3C2BFD18F855FBA3DE9D" box="[655,1102,1964,1991]" italics="true" pageId="20" pageNumber="21">Canthon (G.) smaragdulus subviridis</emphasis>
<bibRefCitation id="F5234B4FFF8A3C2BFBCEF855FAEADE9D" author="Schmidt A." box="[1113,1287,1964,1990]" pageId="20" pageNumber="21" pagination="61 - 103" refId="ref18074" refString="Schmidt A. 1922. 1. Bestimmungstabelle der mir bekannten Canthon- Arten. 2. Verbreitungsgebiete der Canthon - Arten. 3. Neubeschreibungen von Canthon, Saprosites, Mendidius, Euparia und Ataenius. Archiv fur Naturgeschichte 88 (3): 61 - 103." type="journal article" year="1922">Schmidt, 1922</bibRefCitation>
</taxonomicName>
<taxonomicNameLabel id="B8F557D7FF8A3C2BFA86F854FA9ADE9C" box="[1297,1399,1965,1991]" pageId="20" pageNumber="21">stat. rev.</taxonomicNameLabel>
Abbreviations: see Material and methods. Scale bar: 1.0 mm.
</paragraph>
</caption>
<caption id="C5CD6636FF8B3C2AFF2AF855FC24DEB1" ID-DOI="http://doi.org/10.5281/zenodo.3787899" ID-Zenodo-Dep="3787899" httpUri="https://zenodo.org/record/3787899/files/figure.png" pageId="21" pageNumber="22" startId="21.[189,232,1964,1990]" targetBox="[218,1370,265,1931]" targetPageId="21">
<paragraph id="910D36BEFF8B3C2AFF2AF855FC24DEB1" blockId="21.[189,1399,1964,2026]" pageId="21" pageNumber="22">
<emphasis id="A3C6EAACFF8B3C2AFF2AF855FEE4DE9D" bold="true" box="[189,265,1964,1990]" pageId="21" pageNumber="22">Fig. 6.</emphasis>
<taxonomicName id="56B24D3DFF8B3C2AFE98F855FC5ADE9D" authority="smaragdulus (Fabricius, 1781)" authorityName="smaragdulus (Fabricius" authorityYear="1781" baseAuthorityName="Fabricius" baseAuthorityYear="1781" box="[271,951,1964,1991]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="subSpecies" species="smaragdulus" subGenus="Goniocanthon" subSpecies="smaragdulus">
<emphasis id="A3C6EAACFF8B3C2AFE98F855FD0BDE9D" box="[271,742,1964,1991]" italics="true" pageId="21" pageNumber="22">Canthon (G.) smaragdulus smaragdulus</emphasis>
(
<bibRefCitation id="F5234B4FFF8B3C2AFD64F855FC43DE9D" author="Fabricius J. C." box="[755,942,1964,1990]" pageId="21" pageNumber="22" refId="ref16842" refString="Fabricius J. C. 1781. Species insectorum exhibentes eorum differentias specificas, synonyma auctorum, loca Natalia, metamorphosin adiectis observationibus, descriptionibus. Tom I. Carol Ernest, Hamburg and Kiel. https: // doi. org / 10.5962 / bhl. title. 36509" type="book" year="1781">Fabricius, 1781</bibRefCitation>
)
</taxonomicName>
, position of the sclerites of the internal sac. Abbreviations: see Material and methods. Scale bars: 0.5 mm.
</paragraph>
</caption>
<caption id="C5CD6636FF883C29FF2AF854FD44DEB0" ID-DOI="http://doi.org/10.5281/zenodo.3787903" ID-Zenodo-Dep="3787903" httpUri="https://zenodo.org/record/3787903/files/figure.png" pageId="22" pageNumber="23" startId="22.[189,232,1965,1991]" targetBox="[290,1323,479,1903]" targetPageId="22">
<paragraph id="910D36BEFF883C29FF2AF854FD44DEB0" blockId="22.[189,1398,1965,2027]" pageId="22" pageNumber="23">
<emphasis id="A3C6EAACFF883C29FF2AF854FEE1DE9C" bold="true" box="[189,268,1965,1991]" pageId="22" pageNumber="23">Fig. 7.</emphasis>
Sclerites of the internal sac of 
<taxonomicName id="56B24D3DFF883C29FD12F854FBEEDE9C" authority="Nunes &amp; Nunes &amp; Vaz-De, 2018" authorityName="Nunes &amp; Nunes &amp; Vaz-De" authorityYear="2018" box="[645,1027,1965,1992]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="subSpecies" species="fulgidus" status="subsp. nov." subGenus="Goniocanthon" subSpecies="martinezi">
<emphasis id="A3C6EAACFF883C29FD12F854FBEEDE9C" box="[645,1027,1965,1992]" italics="true" pageId="22" pageNumber="23">Canthon (G.) fulgidus martinezi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B8F557D7FF883C29FB9BF854FB7CDE93" box="[1036,1169,1965,1992]" pageId="22" pageNumber="23" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
Abbreviations: 
<emphasis id="A3C6EAACFF883C29FAC6F854FA9BDE9C" bold="true" box="[1361,1398,1965,1991]" pageId="22" pageNumber="23">see</emphasis>
Material and methods. Scale bar: 1.0 mm.
</paragraph>
</caption>
<paragraph id="910D36BEFF893C28FF2AFEF5FC83D837" blockId="23.[189,1399,267,364]" pageId="23" pageNumber="24">
INTERNAL SAC (
<figureCitation id="09892A3BFF893C28FEF2FEF2FE41D87E" box="[357,428,267,294]" captionStart="Fig" captionStartId="24.[189,232,1964,1990]" captionTargetBox="[281,1315,437,1914]" captionTargetId="figure@24.[264,1324,425,1923]" captionTargetPageId="24" captionText="Fig. 9. Sclerites of the internal sac of Canthon (G.) bicolor Castelnau, 1840. Abbreviations: see Material and methods. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3787907" httpUri="https://zenodo.org/record/3787907/files/figure.png" pageId="23" pageNumber="24">Fig. 9</figureCitation>
). FLP not completely involving A forming two duct with spiniform apex. MP distal to A, divided in MP 1 and MP 2, MP 1 semicircular, MP 1 fishhook-shaped with two hook; FLP + A complex involved by MP 2. SRP with lateral prolongation.
</paragraph>
<paragraph id="910D36BEFF893C28FF2AFE6AFE03D8F6" blockId="23.[189,494,403,429]" box="[189,494,403,429]" pageId="23" pageNumber="24">
<heading id="CA4581D2FF893C28FF2AFE6AFE03D8F6" bold="true" box="[189,494,403,429]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
<emphasis id="A3C6EAACFF893C28FF2AFE6AFE03D8F6" bold="true" box="[189,494,403,429]" pageId="23" pageNumber="24">Morphological variation</emphasis>
</heading>
</paragraph>
<paragraph id="910D36BEFF893C28FF2AFE3BFBFCD8A4" blockId="23.[189,1398,449,511]" pageId="23" pageNumber="24">
BL: 
<quantity id="564A9B5BFF893C28FF78FE3BFE82D887" box="[239,367,450,476]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="23" pageNumber="24" unit="mm" value="11.5" valueMax="13.0" valueMin="10.0">10–13 mm</quantity>
, PW: 
<quantity id="564A9B5BFF893C28FE38FE38FDCAD887" box="[431,551,449,476]" metricMagnitude="-3" metricUnit="m" metricValue="7.75" metricValueMax="8.5" metricValueMin="7.0" pageId="23" pageNumber="24" unit="mm" value="7.75" valueMax="8.5" valueMin="7.0">7–8.5 mm</quantity>
. Female protibial spur as wide as long, with apical U-shaped emargination (
<figureCitation id="09892A3BFF893C28FF51FE1CFECFD8A4" box="[198,290,485,511]" captionStart="Fig" captionStartId="19.[189,232,1894,1920]" captionTargetBox="[202,1379,264,1882]" captionTargetId="figure@19.[189,1399,263,1882]" captionTargetPageId="19" captionText="Fig. 4. Canthon (G.) bicolor Castelnau, 1840. A. Habitus, dorsal view. B. Ventral surface of meso- and metafemora and metasternal anterior lobe not convex. C. Dorsal view showing convex pygidium. D. Male protibial spur. E. Female protibial spur. F. Paramera lateral view. G. Paramera dorsal view. Scale bar: A = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3787895" httpUri="https://zenodo.org/record/3787895/files/figure.png" pageId="23" pageNumber="24">Fig. 4E</figureCitation>
). Pygidium convexity less protuberant in females than in males.
</paragraph>
</subSubSection>
<subSubSection id="D9A86535FF893C26FF2AFDDFFD52D837" lastPageId="25" lastPageNumber="26" pageId="23" pageNumber="24" type="distribution">
<paragraph id="910D36BEFF893C28FF2AFDDFFEBBDB1B" blockId="23.[189,342,550,576]" box="[189,342,550,576]" pageId="23" pageNumber="24">
<heading id="CA4581D2FF893C28FF2AFDDFFEBBDB1B" bold="true" box="[189,342,550,576]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
<emphasis id="A3C6EAACFF893C28FF2AFDDFFEBBDB1B" bold="true" box="[189,342,550,576]" pageId="23" pageNumber="24">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="910D36BEFF893C28FF2AFDACFEF0DBC9" blockId="23.[189,1399,597,658]" pageId="23" pageNumber="24">
North of the Amazon forest, in 
<collectingCountry id="E9A5762EFF893C28FDB2FDACFD4CDB34" box="[549,673,597,623]" name="Venezuela" pageId="23" pageNumber="24">Venezuela</collectingCountry>
, 
<collectingCountry id="E9A5762EFF893C28FD3CFDACFCE4DB34" box="[683,777,597,623]" name="Guyana" pageId="23" pageNumber="24">Guyana</collectingCountry>
, 
<collectingCountry id="E9A5762EFF893C28FC83FDACFC97DB34" box="[788,890,597,623]" name="Suriname" pageId="23" pageNumber="24">Surinam</collectingCountry>
, French Guiana and 
<collectingCountry id="E9A5762EFF893C28FBFFFDACFB5DDB34" box="[1128,1200,597,623]" name="Brazil" pageId="23" pageNumber="24">Brazil</collectingCountry>
(state of 
<collectingRegion id="5376F85CFF893C28FA8FFDACFA82DB34" box="[1304,1391,597,623]" country="Brazil" name="Amapa" pageId="23" pageNumber="24">Amapá</collectingRegion>
) (
<figureCitation id="09892A3BFF893C28FF51FD81FEE0DBC9" box="[198,269,632,658]" captionStart="Fig" captionStartId="23.[189,232,1861,1887]" captionTargetBox="[191,1287,1049,1818]" captionTargetId="figure@23.[189,1399,1049,1820]" captionTargetPageId="23" captionText="Fig. 8. Distribution map for subspecies of Goniocanthon: C. (G.) fulgidus martinezi subsp. nov. (green triangle); C. (G.) fulgidus pereirai subsp. nov. (red triangle); C. (G.) fulgidus fulgidus Redtenbacher, 1868 (yellow triangle); C. (G.) smaragdulus subviridis Schmidt, 1922 stat. rev. (green circle); C. (G.) smaragdulus smaragdulus (Fabricius, 1781) (blue circle) and C. bicolor Castelnau, 1840 (brown square)." figureDoi="http://doi.org/10.5281/zenodo.3787905" httpUri="https://zenodo.org/record/3787905/files/figure.png" pageId="23" pageNumber="24">Fig. 8</figureCitation>
).
</paragraph>
<paragraph id="910D36BEFF893C28FF2AFD40FE5ADB8F" blockId="23.[189,439,697,724]" box="[189,439,697,724]" pageId="23" pageNumber="24">
<heading id="CA4581D2FF893C28FF2AFD40FE5ADB8F" bold="true" box="[189,439,697,724]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
<emphasis id="A3C6EAACFF893C28FF2AFD40FE5ADB8F" bold="true" box="[189,439,697,724]" pageId="23" pageNumber="24">Taxonomic remarks</emphasis>
</heading>
</paragraph>
<paragraph id="910D36BEFF893C26FF2AFD11FD52D837" blockId="23.[189,1399,744,982]" lastBlockId="25.[189,1398,267,364]" lastPageId="25" lastPageNumber="26" pageId="23" pageNumber="24">
This is the first time in the taxonomic history of 
<taxonomicName id="56B24D3DFF893C28FC8AFD11FC6FDA59" authorityName="sensu Halffter &amp; Martinez" authorityYear="1977" box="[797,898,744,770]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis id="A3C6EAACFF893C28FC8AFD11FC6FDA59" box="[797,898,744,770]" italics="true" pageId="23" pageNumber="24">Canthon</emphasis>
</taxonomicName>
that 
<taxonomicName id="56B24D3DFF893C28FC52FD10FBACDA59" authorityName="Castelnau" authorityYear="1840" box="[965,1089,744,771]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="A3C6EAACFF893C28FC52FD10FBACDA59" box="[965,1089,744,771]" italics="true" pageId="23" pageNumber="24">C. bicolor</emphasis>
</taxonomicName>
is considered part of the subgenus 
<emphasis id="A3C6EAACFF893C28FEA4FCF2FE0FDA7E" box="[307,482,779,805]" italics="true" pageId="23" pageNumber="24">
<taxonomicName id="56B24D3DFF893C28FEA4FCF2FE32DA7E" authorityName="Pereira &amp; Martinez" authorityYear="1956" box="[307,479,779,805]" class="Insecta" family="Scarabaeidae" genus="Goniocanthon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">Goniocanthon</taxonomicName>
.
</emphasis>
The following character combination supports this decision: metafemora not claviform, hypomeron lacking transverse keel and pygidium strongly convex, an exclusive character of 
<taxonomicName id="56B24D3DFF893C28FF2AFCABFE88DA37" authorityName="Pereira &amp; Martinez" authorityYear="1956" box="[189,357,850,876]" class="Insecta" family="Scarabaeidae" genus="Goniocanthon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis id="A3C6EAACFF893C28FF2AFCABFE88DA37" box="[189,357,850,876]" italics="true" pageId="23" pageNumber="24">Goniocanthon</emphasis>
</taxonomicName>
among the genus. The latter character was already cited by some authors for 
<taxonomicName id="56B24D3DFF893C28FB69FCABFA9ADA37" authorityName="Castelnau" authorityYear="1840" box="[1278,1399,850,876]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="A3C6EAACFF893C28FB69FCABFA9ADA37" box="[1278,1399,850,876]" italics="true" pageId="23" pageNumber="24">C. bicolor</emphasis>
</taxonomicName>
(e.g., 
<bibRefCitation id="F5234B4FFF893C28FE93FC8CFE73DAD4" author="Harold E." box="[260,414,885,911]" pageId="23" pageNumber="24" pagination="1 - 144" refId="ref17251" refString="Harold E. 1868. Monographie der Gattung Canthon. Berliner entomologische Zeitschrift 12: 1 - 144." type="journal article" year="1868">Harold 1868</bibRefCitation>
; 
<bibRefCitation id="F5234B4FFF893C28FE39FC8CFDB4DAD4" author="Schmidt A." box="[430,601,885,911]" pageId="23" pageNumber="24" pagination="61 - 103" refId="ref18074" refString="Schmidt A. 1922. 1. Bestimmungstabelle der mir bekannten Canthon- Arten. 2. Verbreitungsgebiete der Canthon - Arten. 3. Neubeschreibungen von Canthon, Saprosites, Mendidius, Euparia und Ataenius. Archiv fur Naturgeschichte 88 (3): 61 - 103." type="journal article" year="1922">Schmidt 1922</bibRefCitation>
; 
<bibRefCitation id="F5234B4FFF893C28FDFEFC8CFCCFDAD4" author="Balthasar V." box="[617,802,885,911]" pageId="23" pageNumber="24" pagination="179 - 238" refId="ref16169" refString="Balthasar V. 1939. Eine Vorstudie zur Monographie der Gattung Canthon Hffsg. Folia Zoologica et Hydrobiologica 9 (2): 179 - 238." type="journal article" year="1939">Balthasar 1939</bibRefCitation>
). In a preliminary phylogenetic analysis, 
<bibRefCitation id="F5234B4FFF893C28FA8AFC8CFEB8DAE9" author="Medina C. A. &amp; Scholtz C. H. &amp; Gill B. D." pageId="23" pageNumber="24" pagination="23 - 68" refId="ref17576" refString="Medina C. A., Scholtz C. H. &amp; Gill B. D. 2003. Morphological variation and systematics of Canthon Hoffmansegg 1817, and related genera of New World Canthonini dung beetles (Coleoptera, Scarabaeinae). Deutsche entomologische Zeitschrift 50 (1): 23 - 68." type="journal article" year="2003">
Medina 
<emphasis id="A3C6EAACFF893C28FF2AFC60FF11DAE9" box="[189,252,920,946]" italics="true" pageId="23" pageNumber="24">et al.</emphasis>
(2003)
</bibRefCitation>
place this species close to the subgenera 
<taxonomicName id="56B24D3DFF893C28FCCEFC61FBECDAE9" authorityName="Pereira &amp; Martinez" authorityYear="1956" box="[857,1025,920,946]" class="Insecta" family="Scarabaeidae" genus="Goniocanthon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis id="A3C6EAACFF893C28FCCEFC61FBECDAE9" box="[857,1025,920,946]" italics="true" pageId="23" pageNumber="24">Goniocanthon</emphasis>
</taxonomicName>
and 
<taxonomicName id="56B24D3DFF893C28FBD5FC61FB30DAE9" authorityName="Pereira" authorityYear="1953" box="[1090,1245,920,946]" class="Insecta" family="Scarabaeidae" genus="Peltecanthon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis id="A3C6EAACFF893C28FBD5FC61FB30DAE9" box="[1090,1245,920,946]" italics="true" pageId="23" pageNumber="24">Peltecanthon</emphasis>
</taxonomicName>
, mainly due to male internal sac sclerites. The main morphological difference of 
<taxonomicName id="56B24D3DFF893C28FC64FC45FB81DA8E" authorityName="Castelnau" authorityYear="1840" box="[1011,1132,955,982]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="A3C6EAACFF893C28FC64FC45FB81DA8E" box="[1011,1132,955,982]" italics="true" pageId="23" pageNumber="24">C. bicolor</emphasis>
</taxonomicName>
among 
<taxonomicName id="56B24D3DFF893C28FB58FC42FA9ADA8E" authorityName="Pereira &amp; Martinez" authorityYear="1956" box="[1231,1399,955,981]" class="Insecta" family="Scarabaeidae" genus="Goniocanthon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis id="A3C6EAACFF893C28FB58FC42FA9ADA8E" box="[1231,1399,955,981]" italics="true" pageId="23" pageNumber="24">Goniocanthon</emphasis>
</taxonomicName>
is the flat metasternum anterior lobe and the MP sclerite in the internal sac, which is divided in two structures (MP1 and MP2, 
<figureCitation id="09892A3BFF873C26FD92FED7FDA3D812" box="[517,590,302,329]" captionStart="Fig" captionStartId="24.[189,232,1964,1990]" captionTargetBox="[281,1315,437,1914]" captionTargetId="figure@24.[264,1324,425,1923]" captionTargetPageId="24" captionText="Fig. 9. Sclerites of the internal sac of Canthon (G.) bicolor Castelnau, 1840. Abbreviations: see Material and methods. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3787907" httpUri="https://zenodo.org/record/3787907/files/figure.png" pageId="25" pageNumber="26">Fig. 9</figureCitation>
). The relationship with 
<taxonomicName id="56B24D3DFF873C26FCE6FED6FBE1D812" authorityName="Pereira" authorityYear="1953" box="[881,1036,303,329]" class="Insecta" family="Scarabaeidae" genus="Peltecanthon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="A3C6EAACFF873C26FCE6FED6FBE1D812" box="[881,1036,303,329]" italics="true" pageId="25" pageNumber="26">Peltecanthon</emphasis>
</taxonomicName>
should be investigated with a more comprehensive phylogenetic analysis.
</paragraph>
</subSubSection>
</treatment>
</document>