treatments-xml/data/19/15/5B/19155BD5B564958FCF00A8FC0B1AA234.xml

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<mods:title id="ECA83D096F1E11DF5B79ABFB56734886">Three new species of woodlizards (Hoplocercinae, Enyalioides) from northwestern South America</mods:title>
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<mods:namePart id="74AB5F417E1D7099177FA9758CAB335F">Venegas, Pablo J.</mods:namePart>
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<treatment id="19155BD5B564958FCF00A8FC0B1AA234" ID-GBIF-Taxon="152059152" LSID="urn:lsid:zoobank.org:act:4AE55600-2B8F-446B-B702-B55BDC3FF1EF" httpUri="http://treatment.plazi.org/id/19155BD5B564958FCF00A8FC0B1AA234" lastPageId="4" lastPageNumber="111" pageId="2" pageNumber="109">
<subSubSection id="4476D1819C598542FB637F44915282C1" pageId="2" pageNumber="109" type="multiple">
<paragraph id="B15F4FF33A1F7DEF8334CC0CBB7CF753" pageId="2" pageNumber="109">Taxon classification Animalia Squamata Hoplocercidae</paragraph>
</subSubSection>
<subSubSection id="5772AEF2FC0692307DD0EBF8925B30CA" pageId="2" pageNumber="109" type="nomenclature">
<paragraph id="A05FBD7B57DD6C4CFAE6D4614EE64A27" pageId="2" pageNumber="109">
<taxonomicName id="4BEE6F97BEB596471A3ABC71C2633683" ID-CoL="6FH4C" LSID="http://zoobank.org/4AE55600-2B8F-446B-B702-B55BDC3FF1EF" class="Squamata" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides altotambo" pageId="2" pageNumber="109" phylum="Chordata" rank="species" species="altotambo">Enyalioides altotambo</taxonomicName>
<taxonomicNameLabel id="C9095C6D39B86AE81A1215C527473FAF" pageId="2" pageNumber="109">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="B8E13FAB32CE456FE1FD4563BB46C661" pageId="2" pageNumber="109" type="reference_group">
<paragraph id="256EC9A052D5EF412D49594E21546079" pageId="2" pageNumber="109">
<taxonomicName id="4DDA430693FAB4498143D3250E8C1D62" LSID="http://zoobank.org/4AE55600-2B8F-446B-B702-B55BDC3FF1EF" class="Squamata" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides altotambo" pageId="2" pageNumber="109" phylum="Chordata" rank="species" species="altotambo">Enyalioides altotambo</taxonomicName>
Proposed standard English name: Alto Tambo woodlizards
</paragraph>
<paragraph id="EA8B01894E803F72F2B98014D125AF4B" pageId="2" pageNumber="109">
<taxonomicName id="FD94B12009865C1F0E2DF35EB341DC50" LSID="http://zoobank.org/4AE55600-2B8F-446B-B702-B55BDC3FF1EF" class="Squamata" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides altotambo" pageId="2" pageNumber="109" phylum="Chordata" rank="species" species="altotambo">Enyalioides altotambo</taxonomicName>
Proposed standard Spanish name: lagartijas de palo de Alto Tambo
</paragraph>
<paragraph id="7BF0844BF710E6ADB4CD349552FA7839" pageId="2" pageNumber="109">
<taxonomicName id="188DB6229E5FAE82852764045FBCDEBF" class="Squamata" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides oshaughnessyi" pageId="2" pageNumber="109" phylum="Chordata" rank="species" species="oshaughnessyi">Enyalioides oshaughnessyi</taxonomicName>
(part) 
<bibRefCitation id="C73287B94BF36794A0CBA5A5D87C4459" author="Torres-Carvajal, O" journalOrPublisher="Zootaxa" pageId="16" pageNumber="123" pagination="1 - 44" title="A systematic revision of Neotropical lizards in the clade Hoplocercinae (Squamata: Iguania)." volume="2752" year="2011">Torres-Carvajal et al. 2011</bibRefCitation>
: 23.
</paragraph>
</subSubSection>
<subSubSection id="FB3CE96FF942F65AED00193418431B67" pageId="2" pageNumber="109" type="type material">
<paragraph id="8D176CF9897106C00AD58491106BE735" pageId="2" pageNumber="109">Type material.</paragraph>
<paragraph id="E5F6F69FDB72441723B3EB7A9307577E" pageId="2" pageNumber="109">
Holotype. QCAZ 8073 (Fig. 1), an adult male from Alto Tambo, 5 km on road to Placer, Bosque Integral Otokiki, 
<geoCoordinate id="10302DE37C8C8BD65F3E14EB7C7B3BC1" direction="north" orientation="latitude" precision="1" value="0.906">0.90600°N</geoCoordinate>
; - 
<geoCoordinate id="85FDDF3D91A6DF93C9C02C5B1DA1B36D" direction="west" orientation="longitude" precision="1" value="-78.606">78.60600°W</geoCoordinate>
(DD), 620 m, Provincia Esmeraldas, Ecuador, collected on 2 May 2010 by I.G. Tapia, D. Almeida-Reinoso, J.M. Guayasamin and L.A. Coloma.
</paragraph>
<caption id="857029B2DA79692DAB8BF63EE8D0B203" pageId="2" pageNumber="109">
<paragraph id="80A1D6EF7F5E87485DA5C13D68BC8FED" pageId="2" pageNumber="109">
Figure 1. Holotype (QCAZ 8073, adult male, SVL = 119 mm) of 
<taxonomicName id="962E5C78722AC0D5982C08B2E12CD0C8" class="Squamata" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides altotambo" pageId="2" pageNumber="109" phylum="Chordata" rank="species" species="altotambo">Enyalioides altotambo</taxonomicName>
in dorsal (top) and ventral (bottom) views. Photographs by Luis A. Coloma.
</paragraph>
</caption>
<paragraph id="9BF6978D052E0CF35641614F6197C837" pageId="2" pageNumber="109">
Paratype. ECUADOR: Provincia Esmeraldas: QCAZ 6671, adult female, Alto Tambo, Balthazar river, 
<geoCoordinate id="4B75BF5AE8FE17B38FA6FD67AC63A813" direction="north" orientation="latitude" precision="1" value="0.9">0.90000°N</geoCoordinate>
; - 
<geoCoordinate id="CC538749670161CDE686B5AFFEDB04BA" direction="west" orientation="longitude" precision="1" value="-78.61667">78.61667°W</geoCoordinate>
, 645 m, collected on 5 November 2005 by F. Ayala-Varela and I.G. Tapia.
</paragraph>
</subSubSection>
<subSubSection id="E0DA04DA0DCA2A3A22A0FDA94D1B0472" pageId="2" pageNumber="109" type="diagnosis">
<paragraph id="97DC33B8A041766A0673D49487DF64A2" pageId="2" pageNumber="109">Diagnosis.</paragraph>
<paragraph id="0DBC42855F1719E92FCCE927BEF95F1D" pageId="2" pageNumber="109">
<taxonomicName id="5EF4087D0A21AE2B7A3427602DDD24D9" class="Squamata" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides altotambo" pageId="2" pageNumber="109" phylum="Chordata" rank="species" species="altotambo">Enyalioides altotambo</taxonomicName>
differs from other species of 
<taxonomicName id="180069626BE7AA830675C4E6DFA53E3D" class="Squamata" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides" pageId="2" pageNumber="109" phylum="Chordata" rank="genus">Enyalioides</taxonomicName>
, except for 
<taxonomicName id="18AA59D16761BFA14F8D96E226D039B3" class="Squamata" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides oshaughnessyi" pageId="2" pageNumber="109" phylum="Chordata" rank="species" species="oshaughnessyi">Enyalioides oshaughnessyi</taxonomicName>
, in having dorsal scales that are both smooth and homogeneous in size. It can be distinguished from 
<taxonomicName id="258E14DA006C4503B7A51DDB095AB37D" class="Squamata" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides oshaughnessyi" pageId="2" pageNumber="109" phylum="Chordata" rank="species" species="oshaughnessyi">Enyalioides oshaughnessyi</taxonomicName>
(character states in parentheses) by the following characters: iris brown in both sexes (iris bright red in both sexes); scales on lateral edge of skull roof just posterior to superciliaries strongly projected (moderately projected); adults of both sexes with light green spots on dorsum (if present, spots turquoise or blue); adult males with a black medial patch on gular region not extending dorsally to form an antehumeral bar (black patch under gular fold extending dorsally to form a short antehumeral bar); scales on flanks almost homogenous in size (flank scales heterogeneous in size, with a few enlarged, circular, keeled scales); pale postympanic stripe on lateral aspect of neck in both sexes (pale postympanic spot in both sexes), posterior surface of thighs without enlarged scales (scattered enlarged scales), tail length/total length 0.57-0.60 (0.59-0.62).
</paragraph>
</subSubSection>
<subSubSection id="B6BB49BA6E0A465C48B082E5AB328C16" pageId="2" pageNumber="109" type="description of holotype">
<paragraph id="0DACF64D446EE2CAFBB524A59975002A" pageId="2" pageNumber="109">Description of holotype.</paragraph>
<paragraph id="D5BFDDEA6AB13DAD3727A4FB3EC26F1F" pageId="2" pageNumber="109">
Male (Fig. 1); SVL = 119 mm; TL = 160 mm; maximum head width = 21.9 mm; head length = 29.8 mm; head height = 20.3 mm; dorsal head scales keeled or multicarinate, projected dorsally; parietal eye present; eight scales immediately posterior to superciliaries conical, dorsolaterally projected, and conspicuously larger than adjacent scales; temporal scales small, pyramidal, low; one large conical pretympanic scale; superciliaries 17; canthals five; postrostrals three; supralabials 13 if counted to a point below middle of eye; rostral divided into three small scales, similar in size to adjacent supralabials; one longitudinal row of lorilabials between suboculars and supralabials at level of middle of eye, longitudinal rows of lorilabials anterior to this point two; loreal region with small, keeled, and juxtaposed scales; nasal at level of supralabials 
<normalizedToken id="5CB7D8AB9454680DEF4E48B8532B8F93" originalValue="V–VI">V-VI</normalizedToken>
; infralabials 11 if counted to a point below middle of eye; mental (1.68 mm wide 
<normalizedToken id="8BCC31E48D503A9337149413D78ED7AD" originalValue="×">x</normalizedToken>
1.98 mm high) slightly wider and 1.5 times higher than adjacent infralabials; postmentals three; gulars ventrally projected and separated from each other by skin covered with tiny granular scales; gular fold complete midventrally, extending dorsally and posteriorly to form an antehumeral fold; neck with some oblique folds, and a dorsolateral row of enlarged scales; distal aspect of oblique fold immediately anterior to antehumeral fold with approximately six enlarged scales similar in size to gulars, but more than three or four times the size of adjacent fold scales.
</paragraph>
<paragraph id="B5F51A131448925241A5CFF1D858DF68" pageId="2" pageNumber="109">Vertebral crest strongly projected and decreasing in size posteriorly, with vertebrals on neck at least four times higher than those between hind limbs; crest bifurcates at a point approximately 10 mm posterior to the cloaca, and extends onto tail about 1/3 its length; body flanks between fore and hind limbs with slight dorsolateral fold; scales on dorsolateral fold slightly larger than adjacent scales; dorsal and flank scales small, smooth, imbricate, more or less homogeneous in size; ventral scales imbricate, keeled, rectangular or rhomboid, with a posterolateral mucron; ventrals more than twice the size (area) of dorsals.</paragraph>
<paragraph id="138FD0CAFD44766B5D2EE16E98A80435" pageId="2" pageNumber="109">Limb scales keeled dorsally and keeled or feebly keeled ventrally; scales on dorsal and posterior aspects of thighs heterogeneous in size, with most scales less than half the size of those scales on anterior and ventral aspects, separated from each other by skin covered with tiny granular scales; subdigitals on finger IV 25; subdigitals on toe IV 29; femoral pores on each side one; tail laterally compressed and gradually tapering posteriorly; caudal scales smooth at the base of tail, becoming keeled and imbricate towards tip, gradually increasing in size posteriorly on lateral and dorsal aspects of each caudal segment; caudals larger ventrally than dorsally; individual caudal segments three scales long ventrally and seven scales long dorsally.</paragraph>
</subSubSection>
<subSubSection id="5FD8421FA2D83DDDC0A646114D30FF25" pageId="2" pageNumber="109" type="color in life of holotype">
<paragraph id="BAA89B96E7D22DA14897885F7AD63F94" pageId="2" pageNumber="109">Color in life of holotype</paragraph>
<paragraph id="7EA3AE610A086AB91422274575A40C7A" pageId="2" pageNumber="109">(Fig. 1). Head light green with a few black and dark brown scales; superciliaries, canthals and labials yellow; bluish cream blotch, wider than high, behind tympanum; pretympanic scales bluish cream; dorsal body background light green with a fine dark brown reticulation and scattered bluish cream scales; vertebrals yellowish green; tail green with incomplete dark brown rings; black irregular marks on limbs, covering most of hands and feet; chin white; gular region bluish cream anterolaterally grading into yellowish green and then bluish green posteriorly, with a posteromedial black patch; ventral aspect of body, limbs and tail dirty cream; flank color pattern extending onto ventrolateral aspect of body; iris brown with golden ring around pupil.</paragraph>
</subSubSection>
<subSubSection id="8C1FE56B0DB8D23E832652EF390F85E1" lastPageId="3" lastPageNumber="110" pageId="2" pageNumber="109" type="variation">
<paragraph id="67C6183903E127774A9BCCF7059E2AEA" pageId="2" pageNumber="109">Variation.</paragraph>
<paragraph id="81ED17A1D0894513942068DB0910316A" lastPageId="3" lastPageNumber="110" pageId="2" pageNumber="109">
Variation in meristic and morphometric characters of 
<taxonomicName id="3B98114F55FD3E675DBBE9DA95F73C1C" class="Squamata" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides altotambo" pageId="3" pageNumber="110" phylum="Chordata" rank="species" species="altotambo">
<pageBreakToken id="FB24D76E39E92D0E77573F00F0228C98" pageId="3" pageNumber="110" start="start">Enyalioides</pageBreakToken>
altotambo
</taxonomicName>
are presented in Table 2. The single female paratype (QCAZ 6671; Fig. 2) is similar in lepidosis and color patterns to the holotype. It differs from the holotype in lacking a black gular patch, and in having a longer pale postympanic stripe, a yellow chin, and a yellow gular region. Furthermore, the scales on the lateral edge of the skull roof and those forming the vertebral crest are more projected in the female (Fig. 2); however, this variation could be ontogenetic rather than sexual because the female is larger (SVL = 134 mm) than the male (SVL = 119).
</paragraph>
<caption id="4AF31D5C429660A7837CE5FA76D6657B" pageId="3" pageNumber="110">
<paragraph id="329A96DF46C7B2BD7D0038BDFA417914" pageId="3" pageNumber="110">
Figure 2. Paratype (QCAZ 6671, adult female, SVL = 132 mm) of 
<taxonomicName id="62D648B83EBD37958F7AAFE92BB50336" class="Squamata" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides altotambo" pageId="3" pageNumber="110" phylum="Chordata" rank="species" species="altotambo">Enyalioides altotambo</taxonomicName>
. Photograph by Luis A. Coloma.
</paragraph>
</caption>
<caption id="84380A7DB0D46ABE57C654BEAE24CAEE" ID-Table-UUID="84380A7DB0D46ABE57C654BEAE24CAEE" httpUri="http://table.plazi.org/id/84380A7DB0D46ABE57C654BEAE24CAEE" pageId="3" pageNumber="110">
<paragraph id="361DB21270250FA3B44B695C05B9EAAE" pageId="3" pageNumber="110">
Table 2. Summary of morphological characters and measurements (mm) of 
<taxonomicName id="E5DCA82745061B2BE618254B18511938" class="Squamata" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides altotambo" pageId="3" pageNumber="110" phylum="Chordata" rank="species" species="altotambo">Enyalioides altotambo</taxonomicName>
, 
<taxonomicName id="4A5C3BCB90914436625B571B694C4452" class="Squamata" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides anisolepis" pageId="3" pageNumber="110" phylum="Chordata" rank="species" species="anisolepis">Enyalioides anisolepis</taxonomicName>
and 
<taxonomicName id="8A54B3565D216FC3F12DA52100F50590" class="Squamata" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides sophiarothschildae" pageId="3" pageNumber="110" phylum="Chordata" rank="species" species="sophiarothschildae">Enyalioides sophiarothschildae</taxonomicName>
. Range (first line) and mean 
<normalizedToken id="898FF1D2E0E23FE855A91FEEBE7D1A55" originalValue="±">+/-</normalizedToken>
standard deviation (second line) are given for quantitative characters, except when there was no variation.
</paragraph>
</caption>
<paragraph id="37A487736DD57AAAF576AD3BF7991925" pageId="3" pageNumber="110">
<table id="ECCC789E44E5B205A04740E3B7BB502A" pageId="3" pageNumber="110">
<tr id="6A3C28F4D0E3AADF6A7F6B6A4C5E3698" pageId="3" pageNumber="110">
<th id="051B58CD556A7A0CE1B8A66EA8DE41B0" colspan="1" pageId="3" pageNumber="110" rowspan="1">Character</th>
<th id="346F9E4E9C0898D02E03774CA7F640CE" colspan="1" pageId="3" pageNumber="110" rowspan="1">
<taxonomicName id="8237AA307AF1C47F60A5D4DDAD2BCD0B" class="Squamata" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides altotambo" pageId="3" pageNumber="110" phylum="Chordata" rank="species" species="altotambo">Enyalioides altotambo</taxonomicName>
N = 2
</th>
<th id="963F29829AFF374E9AF2A10D3D594612" colspan="1" pageId="3" pageNumber="110" rowspan="1">
<taxonomicName id="806F7FEB9DA78446404D20DA4978F584" class="Squamata" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides anisolepis" pageId="3" pageNumber="110" phylum="Chordata" rank="species" species="anisolepis">Enyalioides anisolepis</taxonomicName>
N = 15
</th>
<th id="29CAC1D8B921D1636ED71AA30FD96791" colspan="1" pageId="3" pageNumber="110" rowspan="1">
<taxonomicName id="195711C2DB3AF10CCA866C2E825443E7" class="Squamata" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides sophiarothschildae" pageId="3" pageNumber="110" phylum="Chordata" rank="species" species="sophiarothschildae">Enyalioides sophiarothschildae</taxonomicName>
N = 3
</th>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection id="E4EB800C3E8701C6E44918D718D56340" pageId="3" pageNumber="110" type="distribution">
<paragraph id="ABB64E5D1136FC62D3473F06456DF59D" pageId="3" pageNumber="110">Distribution.</paragraph>
<paragraph id="43F85792C63A609902617BBBDC1ACA24" pageId="3" pageNumber="110">
<taxonomicName id="D3E7AC24DF7F0B3FA338E471CBEBA7A8" class="Squamata" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides altotambo" pageId="3" pageNumber="110" phylum="Chordata" rank="species" species="altotambo">Enyalioides altotambo</taxonomicName>
is only known from two adjacent localities at 620-645 m in the Chocoan rainforests of northwestern Ecuador (Fig. 3). Female paratype QCAZ 6671 was found at 5:30 pm with its head facing up on a tree trunk.
</paragraph>
<caption id="FA74CAB678BB9273CBF806780FEADA66" pageId="3" pageNumber="110">
<paragraph id="A90E0C8A76D9021AEB305BB448EDD370" pageId="3" pageNumber="110">
Figure 3. Distribution of 
<taxonomicName id="EBC1F58D7AB4E410FBB33E21D4AEB2DD" class="Squamata" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides altotambo" pageId="3" pageNumber="110" phylum="Chordata" rank="species" species="altotambo">Enyalioides altotambo</taxonomicName>
(circles), 
<taxonomicName id="BF13A260EC41787CDD2AA6DD5C804346" class="Squamata" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides anisolepis" pageId="3" pageNumber="110" phylum="Chordata" rank="species" species="anisolepis">Enyalioides anisolepis</taxonomicName>
(triangles) and 
<taxonomicName id="04C2F5AAF84D6A43F91904E623551F6C" class="Squamata" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides sophiarothschildae" pageId="3" pageNumber="110" phylum="Chordata" rank="species" species="sophiarothschildae">Enyalioides sophiarothschildae</taxonomicName>
(squares).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="71EDAB3EC03805E74041BFC5E6DCE718" pageId="3" pageNumber="110" type="etymology">
<paragraph id="C56990A944A5F9CB5257334C0834082C" pageId="3" pageNumber="110">Etymology.</paragraph>
<paragraph id="5437A9583222DD0E3A649DE10735282F" pageId="3" pageNumber="110">
The specific epithet is a noun in apposition and refers to Alto Tambo, Provincia Esmeraldas, Ecuador, a village on the Ibarra-San Lorenzo road where 
<taxonomicName id="04EDEE1031651F008EF3CAAE4D99E2F6" class="Squamata" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides altotambo" pageId="3" pageNumber="110" phylum="Chordata" rank="species" species="altotambo">Enyalioides altotambo</taxonomicName>
was discovered.
</paragraph>
</subSubSection>
<subSubSection id="73A1B65D4299F1963B72B46677363584" lastPageId="4" lastPageNumber="111" pageId="3" pageNumber="110" type="remarks">
<paragraph id="9192BD4A21B7FF8698EA72C3189EFA7B" pageId="3" pageNumber="110">Remarks.</paragraph>
<paragraph id="4DB2CCFE9DD35E2CFE3DBE36DBD4157C" lastPageId="4" lastPageNumber="111" pageId="3" pageNumber="110">
Although previously referred to 
<taxonomicName id="425314874F2F875C4B61065F11229A76" class="Squamata" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides oshaughnessyi" pageId="4" pageNumber="111" phylum="Chordata" rank="species" species="oshaughnessyi">
<pageBreakToken id="9E6E2060DD646F00DCB6B654F5F1FA4A" pageId="4" pageNumber="111" start="start">Enyalioides</pageBreakToken>
oshaughnessyi
</taxonomicName>
, the possibility that the specimens here named 
<taxonomicName id="99164EEE398C3B47C87C778750482EBB" class="Squamata" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides altotambo" pageId="4" pageNumber="111" phylum="Chordata" rank="species" species="altotambo">Enyalioides altotambo</taxonomicName>
represented a distinct species was recognized in previous studies. In a phylogenetic analysis of hoplocercine lizards, 
<bibRefCitation id="7F702112759B097DC07A794C5ABE319D" author="Torres-Carvajal, O" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="16" pageNumber="123" pagination="31 - 43" title="Phylogeny of hoplocercine lizards (Squamata: Iguania) with estimates of relative divergence times." url="10.1016/j.ympev.2008.10.002" volume="50" year="2009">Torres-Carvajal and de Queiroz (2009)</bibRefCitation>
found &quot; 
<taxonomicName id="688E66E9FA43AA01D71F782DE3B6BB0A" class="Squamata" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides oshaughnessyi" pageId="4" pageNumber="111" phylum="Chordata" rank="species" species="oshaughnessyi">Enyalioides oshaughnessyi</taxonomicName>
&quot; to be paraphyletic relative to 
<taxonomicName id="36437B5E784F790A1F00082FEF89FB87" class="Squamata" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides touzeti" pageId="4" pageNumber="111" phylum="Chordata" rank="species" species="touzeti">Enyalioides touzeti</taxonomicName>
based on three samples of &quot; 
<taxonomicName id="FA6EFF330607F547940D51B0FC2EA9EB" class="Squamata" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides oshaughnessyi" pageId="4" pageNumber="111" phylum="Chordata" rank="species" species="oshaughnessyi">Enyalioides oshaughnessyi</taxonomicName>
&quot;. One of them corresponded to the paratype of 
<taxonomicName id="3D6B340BDCDA849434AF777F55EA69E2" class="Squamata" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides altotambo" pageId="4" pageNumber="111" phylum="Chordata" rank="species" species="altotambo">Enyalioides altotambo</taxonomicName>
, and was sister to a clade containing the sister taxa 
<taxonomicName id="8DD38F6D55860DBBB689561A5A07BB55" class="Squamata" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides touzeti" pageId="4" pageNumber="111" phylum="Chordata" rank="species" species="touzeti">Enyalioides touzeti</taxonomicName>
and 
<taxonomicName id="DE25B406B2480E62973379B256F85AF0" class="Squamata" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides oshaughnessyi" pageId="4" pageNumber="111" phylum="Chordata" rank="species" species="oshaughnessyi">Enyalioides oshaughnessyi</taxonomicName>
. 
<bibRefCitation id="9D01B99A38EEB6DF25FE055D62D2F31C" author="Torres-Carvajal, O" journalOrPublisher="Zootaxa" pageId="16" pageNumber="123" pagination="1 - 44" title="A systematic revision of Neotropical lizards in the clade Hoplocercinae (Squamata: Iguania)." volume="2752" year="2011">Torres-Carvajal et al. (2011)</bibRefCitation>
noted that the color of the iris in live specimens of &quot; 
<taxonomicName id="F6691F3796A5FA31E3247EA6A2A65873" class="Squamata" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides oshaughnessyi" pageId="4" pageNumber="111" phylum="Chordata" rank="species" species="oshaughnessyi">Enyalioides oshaughnessyi</taxonomicName>
&quot; from Alto Tambo was not bright red as in live specimens of &quot; 
<taxonomicName id="54E88200EBF9ABDBBE87F10CFA2F43BF" class="Squamata" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides oshaughnessyi" pageId="4" pageNumber="111" phylum="Chordata" rank="species" species="oshaughnessyi">Enyalioides oshaughnessyi</taxonomicName>
&quot; from other localities and suggested that the two forms represented separate species. Nonetheless, these authors found no other differences between the two potential species and refrained from associating the name 
<taxonomicName id="F4C79D2DA9EAFF425226EC49E81C9A28" class="Squamata" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides oshaughnessyi" pageId="4" pageNumber="111" phylum="Chordata" rank="species" species="oshaughnessyi">Enyalioides oshaughnessyi</taxonomicName>
with one versus the other because the type locality data of 
<taxonomicName id="E642ED5A78C9B71E48649D3BF29C1DC4" class="Squamata" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides oshaughnessyi" pageId="4" pageNumber="111" phylum="Chordata" rank="species" species="oshaughnessyi">Enyalioides oshaughnessyi</taxonomicName>
is vague (
<normalizedToken id="58DCFA3ED74F26442AFC71162097CDDF" originalValue="“Ecuador”">&quot;Ecuador&quot;</normalizedToken>
), and the color of the iris was not recorded in its original description (
<bibRefCitation id="2F2FF188C26AF90E46EA404F84AD2E18" author="Boulenger, GA" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="15" pageNumber="122" pagination="246 - 247" title="Description of a new species of Enyalius in the Brussels Museum." url="10.1111/j.1096-3642.1881.tb01283.x" volume="1881" year="1881">Boulenger 1881</bibRefCitation>
). Here we recognize known populations other than that at Alto Tambo as 
<taxonomicName id="3299F12866CE2AF05531F9CBCA9696EF" class="Squamata" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides oshaughnessyi" pageId="4" pageNumber="111" phylum="Chordata" rank="species" species="oshaughnessyi">Enyalioides oshaughnessyi</taxonomicName>
based on the enlarged, circular and keeled scales scattered on the flanks of 
<taxonomicName id="96D1B46C193C42D078C93ADCA5893AC5" class="Squamata" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides oshaughnessyi" pageId="4" pageNumber="111" phylum="Chordata" rank="species" species="oshaughnessyi">Enyalioides oshaughnessyi</taxonomicName>
(absent in 
<taxonomicName id="96D145AD9324970659038839B0B13FC7" class="Squamata" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides altotambo" pageId="4" pageNumber="111" phylum="Chordata" rank="species" species="altotambo">Enyalioides altotambo</taxonomicName>
), as described and illustrated in its original description (Fig. 4; 
<bibRefCitation id="F6C05FB42800A99F610A39233BC62710" author="Boulenger, GA" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="15" pageNumber="122" pagination="246 - 247" title="Description of a new species of Enyalius in the Brussels Museum." url="10.1111/j.1096-3642.1881.tb01283.x" volume="1881" year="1881">Boulenger 1881</bibRefCitation>
).
</paragraph>
<caption id="383427A660D78CC51F06691D543FA8BE" pageId="4" pageNumber="111">
<paragraph id="B9A610FBFA62E81C402F3DBB76457CBF" pageId="4" pageNumber="111">
Figure 4. Holotype of 
<taxonomicName id="4CEDB826AB247A2F526F4AC7CF4E83D0" class="Squamata" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides oshaughnessyi" pageId="4" pageNumber="111" phylum="Chordata" rank="species" species="oshaughnessyi">Enyalioides oshaughnessyi</taxonomicName>
(MRHN [Museum Royal 
<normalizedToken id="56C3CB2E4D76C06F48CD469287FCEC02" originalValue="d’Histoire">d'Histoire</normalizedToken>
Naturelle, Belgium] 2009, adult male). Illustration taken from original description (
<bibRefCitation id="AAF6F047BD6B8B3E9160BDBB8042BB58" author="Boulenger, GA" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="15" pageNumber="122" pagination="246 - 247" title="Description of a new species of Enyalius in the Brussels Museum." url="10.1111/j.1096-3642.1881.tb01283.x" volume="1881" year="1881">Boulenger 1881</bibRefCitation>
).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>