238 lines
30 KiB
XML
238 lines
30 KiB
XML
<document id="4615D9E6B5A3BDE0F610C1FC1FB97859" ID-CLB-Dataset="8377" ID-DOI="10.5281/zenodo.4523442" ID-GBIF-Dataset="0d0d29eb-b376-4f5d-934d-dd987ff0cbff" ID-ISSN="0253-116X" ID-Zenodo-Dep="4523442" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612222933374" checkinUser="felipe" docAuthor="Assing, V." docDate="2007" docId="191487A5FFF4FFAEFF2D51D13C51FEEE" docLanguage="en" docName="LinzerbiolBeitr.39.1.0015-0021.pdf" docOrigin="Linzer biologische Beiträge 39 (1)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Pseudobium pakistanicum Assing 2007, nov.sp." docType="treatment" docVersion="6" lastPageNumber="21" masterDocId="E52DFFDDFFF0FFA8FFBA50483850FFE8" masterDocTitle="On the genus Pseudobium MULSANT & REY II. A new species from Pakistan and additional records (Insecta: Coleoptera: Staphylinidae: Paederinae)" masterLastPageNumber="21" masterPageNumber="15" pageNumber="19" updateTime="1698926238243" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="9FB8E4153FF0BBEB9A4BFD7E6FEA44CD">
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<mods:title id="136ED162CE7EF2E660FD2E66393D476C">On the genus Pseudobium MULSANT & REY II. A new species from Pakistan and additional records (Insecta: Coleoptera: Staphylinidae: Paederinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="1B6E8ABF608D4402E4E74B15649CCE6C" type="personal">
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<mods:namePart id="367A1A5511D1309CDA0FAA6F000628EC">Assing, V.</mods:namePart>
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<mods:typeOfResource id="4369ED9C043B2C975F6AC5BA0141857D">text</mods:typeOfResource>
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<mods:title id="4A5C1FFF78BF94B584D0977558D27FC0">Linzer biologische Beiträge</mods:title>
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<mods:part id="A0C333C205BBF8D928E14B17B65F87F9">
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<mods:date id="CD95D167BCF3744BBFDE06FF825C4CF1">2007</mods:date>
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<mods:detail id="DAD72491923D75177295061B37BA94E4" type="pubDate">
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<mods:number id="EBF85802BE16BD1D2D25CCD1503757D2">2007-07-23</mods:number>
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<mods:detail id="C0EAD4861F366481238F42923041F6C2" type="volume">
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<mods:number id="04E43B8D4E4BFBA07495C1F7C9682F85">39</mods:number>
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<mods:detail id="7AF0C7C42DE53590D39DEBCC04F97530" type="issue">
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<mods:number id="917C042229EBD490CD7C73B8C5DE69BE">1</mods:number>
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<mods:start id="05173EC07D855AFE11D12AD29FD2FD71">15</mods:start>
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<mods:end id="B04FD9A1059F460FAEDC88BA0D709556">21</mods:end>
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<mods:classification id="EEEBFD043234502A71F07C1C3CBDDF83">journal article</mods:classification>
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<mods:identifier id="11E1E0F23D4F461ACDC683887ADE6A7B" type="DOI">10.5281/zenodo.4523442</mods:identifier>
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<mods:identifier id="67A805F67061DDFCF8782A9150545A3A" type="ISSN">0253-116X</mods:identifier>
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<mods:identifier id="5B67AD98B8022818C0E0F3ADBB493A24" type="Zenodo-Dep">4523442</mods:identifier>
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</mods:mods>
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<treatment id="191487A5FFF4FFAEFF2D51D13C51FEEE" ID-DOI="http://doi.org/10.5281/zenodo.4523482" ID-GBIF-Taxon="177311772" ID-Zenodo-Dep="4523482" LSID="urn:lsid:plazi:treatment:191487A5FFF4FFAEFF2D51D13C51FEEE" httpUri="http://treatment.plazi.org/id/191487A5FFF4FFAEFF2D51D13C51FEEE" lastPageId="6" lastPageNumber="21" pageId="4" pageNumber="19">
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<subSubSection id="D9A76538FFF4FFACFF2D51D13AEDFE47" box="[151,701,409,431]" pageId="4" pageNumber="19" type="nomenclature">
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<paragraph id="910236B3FFF4FFACFF2D51D13AEDFE47" blockId="4.[151,701,409,431]" box="[151,701,409,431]" pageId="4" pageNumber="19">
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<heading id="CA4A81DFFFF4FFACFF2D51D13AEDFE47" box="[151,701,409,431]" fontSize="8" level="2" pageId="4" pageNumber="19" reason="2">
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<emphasis id="A3C9EAA1FFF4FFACFF2D51D13AEDFE47" bold="true" box="[151,701,409,431]" pageId="4" pageNumber="19">
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<taxonomicName id="56BD4D30FFF4FFACFF2D51D139FCFE47" ID-CoL="78BXW" authority="Assing, 2007" authorityName="Assing" authorityYear="2007" box="[151,428,409,431]" class="Insecta" family="Staphylinidae" genus="Pseudobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="pakistanicum" status="sp. nov.">
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<emphasis id="A3C9EAA1FFF4FFACFF2D51D139FCFE47" bold="true" box="[151,428,409,431]" italics="true" pageId="4" pageNumber="19">Pseudobium pakistanicum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="B8FA57DAFFF4FFACFE0851D139AEFE47" box="[434,510,409,431]" pageId="4" pageNumber="19" rank="species">nov.sp.</taxonomicNameLabel>
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(
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<figureCitation id="09862A36FFF4FFACFDB151D13A33FE47" box="[523,611,409,431]" captionStart="Figs 1-7" captionStartId="5.[151,191,1520,1540]" captionTargetBox="[155,1011,465,1506]" captionTargetId="figure-157@5.[151,1058,569,1045]" captionTargetPageId="5" captionText="Figs 1-7: Pseudobium pakistanicum nov.sp. (holotype). (1) forebody; (2) head; (3) antenna; (4) male sternite VIII; (5-6) aedeagus in lateral and in ventral view; (7) apical part of aedeagus in ventral view. Scale bars: 1: 1.0 mm; 2-4: 0.2 mm; 5-7: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4523446" httpUri="https://zenodo.org/record/4523446/files/figure.png" pageId="4" pageNumber="19">Figs 1-7</figureCitation>
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, Map 3)
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="D9A76538FFF4FFAEFF2D51F73C51FEEE" lastPageId="6" lastPageNumber="21" pageId="4" pageNumber="19" type="materials_examined">
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<paragraph id="910236B3FFF4FFACFF2D51F73A14FDE9" blockId="4.[151,1059,446,513]" pageId="4" pageNumber="19">
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<materialsCitation id="21D53CEEFFF4FFACFF2D51F73B47FE02" ID-GBIF-Occurrence="3033346306" collectingDate="2005-09-02" collectingDateMax="2005-09-12" collectingDateMin="2005-09-02" collectorName="V. Gurko" country="Pakistan" county="Tanai village" elevation="2000" latitude="32.2" location="Northwest Frontier Province" longLatPrecision="1212" longitude="69.683334" municipality="South Waziristan" pageId="4" pageNumber="19" specimenCount="1" typeStatus="holotype">
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<typeStatus id="4E068811FFF4FFACFF2D51F738BCFE3B" box="[151,236,447,467]" pageId="4" pageNumber="19" type="holotype">Holotype</typeStatus>
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Ƌ:
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<collectingCountry id="E9AA7623FFF4FFACFEAF51F73935FE3B" box="[277,357,447,467]" name="Pakistan" pageId="4" pageNumber="19">Pakistan</collectingCountry>
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:
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<location id="94626068FFF4FFACFECB51F73A2DFE3B" LSID="urn:lsid:plazi:treatment:191487A5FFF4FFAEFF2D51D13C51FEEE:94626068FFF4FFACFECB51F73A2DFE3B" box="[369,637,447,467]" country="Pakistan" county="Tanai village" latitude="32.2" longLatPrecision="1212" longitude="69.683334" municipality="South Waziristan" name="Northwest Frontier Province" pageId="4" pageNumber="19">Northwest Frontier Province</location>
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:
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<collectingMunicipality id="7166ACC9FFF4FFACFD3251F73B78FE3B" box="[648,808,447,467]" pageId="4" pageNumber="19">South Waziristan</collectingMunicipality>
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agency near
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<collectingCounty id="78634E3FFFF4FFACFC1351F73C73FE3B" box="[937,1059,447,467]" pageId="4" pageNumber="19">Tanai village</collectingCounty>
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[ca.
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<geoCoordinate id="F4895074FFF4FFACFF7B519E395AFE02" box="[193,266,470,490]" degrees="32" direction="north" minutes="12" orientation="latitude" pageId="4" pageNumber="19" precision="925" value="32.2">32°12N</geoCoordinate>
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,
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<geoCoordinate id="F4895074FFF4FFACFEAD519E390DFE02" box="[279,349,470,490]" degrees="69" direction="east" minutes="41" orientation="longitude" pageId="4" pageNumber="19" precision="925" value="69.683334">69°41E</geoCoordinate>
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] /
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<date id="E5031073FFF4FFACFEC0519E39ACFE02" box="[378,508,470,490]" pageId="4" pageNumber="19" value="2005-09-02" valueMax="2005-09-12" valueMin="2005-09-02">
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<collectingDate id="F547E99BFFF4FFACFEC0519E39ACFE02" box="[378,508,470,490]" pageId="4" pageNumber="19" value="2005-09-02" valueMax="2005-09-12" valueMin="2005-09-02">2.-12.IX.2005</collectingDate>
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</date>
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,
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<quantity id="56459B56FFF4FFACFDB3519E3AD8FE02" box="[521,648,470,490]" metricMagnitude="3" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="4" pageNumber="19" unit="m" value="2000.0" valueMax="2500.0" valueMin="1500.0">
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<elevation id="1A90D180FFF4FFACFDB3519E3AD8FE02" box="[521,648,470,490]" metricMagnitude="3" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="4" pageNumber="19" unit="m" value="2000.0" valueMax="2500.0" valueMin="1500.0">1500-2500 m</elevation>
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</quantity>
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, leg.
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<collectorName id="3C485365FFF4FFACFD04519E3B47FE02" box="[702,791,470,490]" pageId="4" pageNumber="19">V. Gurko</collectorName>
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</materialsCitation>
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/
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<typeStatus id="4E068811FFF4FFACFC95519E3BDEFE02" box="[815,910,470,490]" pageId="4" pageNumber="19" type="holotype">Holotypus</typeStatus>
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Ƌ
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<taxonomicName id="56BD4D30FFF4FFACFC08519D3940FDE8" authority="Assing, 2007" authorityName="Assing" authorityYear="2007" class="Insecta" family="Staphylinidae" genus="Pseudobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="pakistanicum" status="sp. nov.">
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<emphasis id="A3C9EAA1FFF4FFACFC08519D3940FDE8" italics="true" pageId="4" pageNumber="19">Pseudobium pakistanicum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="B8FA57DAFFF4FFACFEAC51A53912FDE9" box="[278,322,493,513]" pageId="4" pageNumber="19" rank="species">sp.n.</taxonomicNameLabel>
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det. V. Assing 2007 (cAss).
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</paragraph>
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<paragraph id="910236B3FFF4FFACFF2D52473A5AFD90" blockId="4.[151,1060,526,632]" pageId="4" pageNumber="19">
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D e s c r i p t i o n: Measurements (in mm) and ratios (
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<typeStatus id="4E068811FFF4FFACFD5952463B12FDCC" box="[739,834,526,548]" pageId="4" pageNumber="19" type="holotype">holotype</typeStatus>
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): AL: 1.36; HL: 0.57; HW: 0.44; PW: 0.41; PL: 0.65; EL: 0.65; EW: 0.59; AW: 0.51; TiL: 0.45; TaL: 0.32; ML: 0.47; TL: 4.4; HL/HW: 1.31; PW/HW: 0.93; PW/PL: 0.63; EL/PL: 1.00; EW/PW: 1.44; AW/EW: 0.87; TiL/TaL: 1.43.
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</paragraph>
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<paragraph id="910236B3FFF4FFACFF2D52CE39F8FCE4" blockId="4.[151,1059,646,780]" pageId="4" pageNumber="19">
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Relatively small species (see measurements), of similar size as
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<taxonomicName id="56BD4D30FFF4FFACFCFF52CF3B81FD74" authorityName="JARRIGE" authorityYear="1949" box="[837,977,647,668]" class="Insecta" family="Staphylinidae" genus="Pseudobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="cypriacum">
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<emphasis id="A3C9EAA1FFF4FFACFCFF52CF3B81FD74" box="[837,977,647,668]" italics="true" pageId="4" pageNumber="19">P. cypriacum</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="56BD4D30FFF4FFACFBB652CF3956FD5F" class="Insecta" family="Staphylinidae" genus="Pseudobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="hellenicum">
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<emphasis id="A3C9EAA1FFF4FFACFBB652CF3956FD5F" italics="true" pageId="4" pageNumber="19">P. hellenicum</emphasis>
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</taxonomicName>
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. Coloration similar to that of
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<taxonomicName id="56BD4D30FFF4FFACFD8F52EB3A93FD5F" box="[565,707,674,696]" class="Insecta" family="Staphylinidae" genus="Pseudobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="anatolicum">
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<emphasis id="A3C9EAA1FFF4FFACFD8F52EB3A93FD5F" box="[565,707,674,696]" italics="true" pageId="4" pageNumber="19">P. anatolicum</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="56BD4D30FFF4FFACFD4F52EB3BD1FD5F" box="[757,897,674,696]" class="Insecta" family="Staphylinidae" genus="Pseudobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="hellenicum">
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<emphasis id="A3C9EAA1FFF4FFACFD4F52EB3BD1FD5F" box="[757,897,674,696]" italics="true" pageId="4" pageNumber="19">P. hellenicum</emphasis>
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</taxonomicName>
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: body blackish, with the posterior margin of the elytra yellowish; legs reddish yellow to yellowish brown, with the meso- and metafemora slightly darker; antennae brown, with the basal two antennomeres reddish.
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</paragraph>
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<paragraph id="910236B3FFF4FFACFF2D53523A90FC48" blockId="4.[151,1059,794,928]" pageId="4" pageNumber="19">
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Head distinctly oblong (
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<figureCitation id="09862A36FFF4FFACFE3F53523991FCD8" box="[389,449,794,816]" captionStart="Figs 1-7" captionStartId="5.[151,191,1520,1540]" captionTargetBox="[155,1011,465,1506]" captionTargetId="figure-157@5.[151,1058,569,1045]" captionTargetPageId="5" captionText="Figs 1-7: Pseudobium pakistanicum nov.sp. (holotype). (1) forebody; (2) head; (3) antenna; (4) male sternite VIII; (5-6) aedeagus in lateral and in ventral view; (7) apical part of aedeagus in ventral view. Scale bars: 1: 1.0 mm; 2-4: 0.2 mm; 5-7: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4523446" httpUri="https://zenodo.org/record/4523446/files/figure.png" pageId="4" pageNumber="19">Fig. 2</figureCitation>
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), of quadrangular shape, lateral margins subparallel, posterior angles moderately marked; puncturation rather coarse and well-delimited, interstices in central dorsal area 1-2 times as wide as diameter of punctures; microsculpture absent; eyes large and projecting from lateral outline of head (
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<figureCitation id="09862A36FFF4FFACFD3F53263A92FC6C" box="[645,706,878,900]" captionStart="Figs 1-7" captionStartId="5.[151,191,1520,1540]" captionTargetBox="[155,1011,465,1506]" captionTargetId="figure-157@5.[151,1058,569,1045]" captionTargetPageId="5" captionText="Figs 1-7: Pseudobium pakistanicum nov.sp. (holotype). (1) forebody; (2) head; (3) antenna; (4) male sternite VIII; (5-6) aedeagus in lateral and in ventral view; (7) apical part of aedeagus in ventral view. Scale bars: 1: 1.0 mm; 2-4: 0.2 mm; 5-7: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4523446" httpUri="https://zenodo.org/record/4523446/files/figure.png" pageId="4" pageNumber="19">Fig. 2</figureCitation>
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), slightly more than half the length of postocular region in dorsal view; antenna as in
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<figureCitation id="09862A36FFF4FFACFD3B53C23AEDFC48" box="[641,701,906,928]" captionStart="Figs 1-7" captionStartId="5.[151,191,1520,1540]" captionTargetBox="[155,1011,465,1506]" captionTargetId="figure-157@5.[151,1058,569,1045]" captionTargetPageId="5" captionText="Figs 1-7: Pseudobium pakistanicum nov.sp. (holotype). (1) forebody; (2) head; (3) antenna; (4) male sternite VIII; (5-6) aedeagus in lateral and in ventral view; (7) apical part of aedeagus in ventral view. Scale bars: 1: 1.0 mm; 2-4: 0.2 mm; 5-7: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4523446" httpUri="https://zenodo.org/record/4523446/files/figure.png" pageId="4" pageNumber="19">Fig. 3</figureCitation>
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.
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</paragraph>
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<paragraph id="910236B3FFF4FFACFF2D53E739BDFC08" blockId="4.[151,1059,942,992]" pageId="4" pageNumber="19">
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Pronotum narrower than head and subparallel (
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<figureCitation id="09862A36FFF4FFACFD3853E63A90FC2C" box="[642,704,942,964]" captionStart="Figs 1-7" captionStartId="5.[151,191,1520,1540]" captionTargetBox="[155,1011,465,1506]" captionTargetId="figure-157@5.[151,1058,569,1045]" captionTargetPageId="5" captionText="Figs 1-7: Pseudobium pakistanicum nov.sp. (holotype). (1) forebody; (2) head; (3) antenna; (4) male sternite VIII; (5-6) aedeagus in lateral and in ventral view; (7) apical part of aedeagus in ventral view. Scale bars: 1: 1.0 mm; 2-4: 0.2 mm; 5-7: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4523446" httpUri="https://zenodo.org/record/4523446/files/figure.png" pageId="4" pageNumber="19">Fig. 1</figureCitation>
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); dorsal rows composed of 15-20 punctures; microsculpture absent.
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</paragraph>
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<paragraph id="910236B3FFF4FFACFF2D53A63B39FBD4" blockId="4.[151,1059,1006,1084]" pageId="4" pageNumber="19">Elytra distinctly wider than and at suture as long as pronotum (see ratios EW/PW and EL/PL); punctures of similar size as those of pronotum and arranged in distinct rows, these rows becoming obsolete posteriorly. Hind wings fully developed.</paragraph>
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<paragraph id="910236B3FFF4FFACFF2D540239C1FB5C" blockId="4.[151,1059,1098,1204]" pageId="4" pageNumber="19">Abdomen narrower than elytra, widest at segment VII; tergites III-V with shallow anterior impression; puncturation of anterior impressions of tergites III-V moderately coarse, on remainder of tergal surfaces fine; microsculpture shallow; posterior margin of tergite VII with palisade fringe.</paragraph>
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<paragraph id="910236B3FFF4FFACFF2D54893B84FB1C" blockId="4.[151,1059,1217,1268]" pageId="4" pageNumber="19">
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Ƌ: sternite VII unmodified; posterior margin of sternite VIII deeply excised, this excision anteriorly concave, not acute (
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<figureCitation id="09862A36FFF4FFACFE01549639A7FB1C" box="[443,503,1246,1268]" captionStart="Figs 1-7" captionStartId="5.[151,191,1520,1540]" captionTargetBox="[155,1011,465,1506]" captionTargetId="figure-157@5.[151,1058,569,1045]" captionTargetPageId="5" captionText="Figs 1-7: Pseudobium pakistanicum nov.sp. (holotype). (1) forebody; (2) head; (3) antenna; (4) male sternite VIII; (5-6) aedeagus in lateral and in ventral view; (7) apical part of aedeagus in ventral view. Scale bars: 1: 1.0 mm; 2-4: 0.2 mm; 5-7: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4523446" httpUri="https://zenodo.org/record/4523446/files/figure.png" pageId="4" pageNumber="19">Fig. 4</figureCitation>
|
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); aedeagus of highly distinctive shape (
|
|
<figureCitation id="09862A36FFF4FFACFCC954963B97FB1C" box="[883,967,1246,1268]" captionStart="Figs 1-7" captionStartId="5.[151,191,1520,1540]" captionTargetBox="[155,1011,465,1506]" captionTargetId="figure-157@5.[151,1058,569,1045]" captionTargetPageId="5" captionText="Figs 1-7: Pseudobium pakistanicum nov.sp. (holotype). (1) forebody; (2) head; (3) antenna; (4) male sternite VIII; (5-6) aedeagus in lateral and in ventral view; (7) apical part of aedeagus in ventral view. Scale bars: 1: 1.0 mm; 2-4: 0.2 mm; 5-7: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4523446" httpUri="https://zenodo.org/record/4523446/files/figure.png" pageId="4" pageNumber="19">Figs 5-7</figureCitation>
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).
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</paragraph>
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<paragraph id="910236B3FFF4FFACFF2D554B39FEFADC" blockId="4.[151,1059,1282,1332]" pageId="4" pageNumber="19">E t y m o l o g y: The name (Lat., adj.) is derived from the name of the country where the species was discovered.</paragraph>
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<paragraph id="910236B3FFF4FFADFF2D550A3A47FED4" blockId="4.[151,1059,1346,1480]" lastBlockId="5.[151,1059,210,316]" lastPageId="5" lastPageNumber="20" pageId="4" pageNumber="19">
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C o m p a r a t i v e n o t e s:
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<taxonomicName id="56BD4D30FFF4FFACFE4F550A3B51FABF" authorityName="Assing" authorityYear="2007" box="[501,769,1346,1367]" class="Insecta" family="Staphylinidae" genus="Pseudobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="pakistanicum">
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<emphasis id="A3C9EAA1FFF4FFACFE4F550A3B51FABF" box="[501,769,1346,1367]" italics="true" pageId="4" pageNumber="19">Pseudobium pakistanicum</emphasis>
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</taxonomicName>
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|
is distinguished from all its congeners by the distinctive morphology of the aedeagus, from most species also by the the shape of the posterior excision of the male sternite VIII (in Western Palaearctic congeners mostly acutely angled anteriorly). From geographically close
|
|
<taxonomicName id="56BD4D30FFF4FFACFCEA55DE3B9DFA43" authorityName="MULSANT & REY II. A" authorityYear="1878" box="[848,973,1430,1451]" class="Insecta" family="Staphylinidae" genus="Pseudobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="genus">
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<emphasis id="A3C9EAA1FFF4FFACFCEA55DE3B9DFA43" box="[848,973,1430,1451]" italics="true" pageId="4" pageNumber="19">Pseudobium</emphasis>
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</taxonomicName>
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|
species, some of whose primary sexual characters are unknown, it is separated as follows: * from
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<emphasis id="A3C9EAA1FFF5FFADFEB9509B39D5FF00" box="[259,389,211,232]" italics="true" pageId="5" pageNumber="20">P. asiaticum</emphasis>
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|
<bibRefCitation id="F52C4B42FFF5FFADFE2A509A3A62FF00" author="GUSAROV V" box="[400,562,210,232]" pageId="5" pageNumber="20" pagination="431 - 446" refId="ref3155" refString="GUSAROV V. I. (1994): New and little-known Palaearctic Paederinae (Coleoptera: Staphylinidae). - Annales de la Societe Entomologique de France (N. S.) 30: 431 - 446." type="journal article" year="1994">GUSAROV 1994</bibRefCitation>
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|
(Middle Asia) by the darker coloration of the forebody (
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<emphasis id="A3C9EAA1FFF5FFADFE8950A739E3FEEC" box="[307,435,239,260]" italics="true" pageId="5" pageNumber="20">P. asiaticum</emphasis>
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|
: head reddish brown, pronotum and abdomen brown, elytra reddish brown with three indistinct dark brown spots), and the different shape of the aedeagus (see
|
|
<bibRefCitation id="F52C4B42FFF5FFADFEE3516E3A42FED4" author="GUSAROV V" box="[345,530,294,316]" pageId="5" pageNumber="20" pagination="431 - 446" refId="ref3155" refString="GUSAROV V. I. (1994): New and little-known Palaearctic Paederinae (Coleoptera: Staphylinidae). - Annales de la Societe Entomologique de France (N. S.) 30: 431 - 446." type="journal article" year="1994">GUSAROV (1994))</bibRefCitation>
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;
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</paragraph>
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<footnote id="F2A62ABDFFF4FFACFF2D559F3BD0F9C0" pageId="4" pageNumber="19">
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<paragraph id="910236B3FFF4FFACFF2D559F3BD0F9C0" blockId="4.[151,1058,1495,1576]" pageId="4" pageNumber="19">
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|
* from
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<emphasis id="A3C9EAA1FFF4FFACFEBE559039C5FA05" box="[260,405,1496,1517]" italics="true" pageId="4" pageNumber="19">P. laeviventre</emphasis>
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|
CHAMPION 1922 (Afghanistan, India),
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|
<emphasis id="A3C9EAA1FFF4FFACFC8655903BE0FA04" box="[828,944,1495,1517]" italics="true" pageId="4" pageNumber="19">P. assimile</emphasis>
|
|
CAMERON 1924, and
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|
<emphasis id="A3C9EAA1FFF4FFACFE8E55BD3994F9E2" box="[308,452,1525,1546]" italics="true" pageId="4" pageNumber="19">P. semiflavum</emphasis>
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CAMERON 1924 (India) by much darker coloration at least of the head and pronotum (in the other species yellowish to reddish);
|
|
</paragraph>
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|
</footnote>
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|
<paragraph id="910236B3FFF5FFADFF2D51023A09FE94" blockId="5.[151,1059,330,380]" pageId="5" pageNumber="20">
|
|
* from
|
|
<taxonomicName id="56BD4D30FFF5FFADFEBA51033939FEB7" box="[256,361,330,352]" class="Insecta" family="Staphylinidae" genus="Pseudobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="richteri">
|
|
<emphasis id="A3C9EAA1FFF5FFADFEBA51033939FEB7" box="[256,361,330,352]" italics="true" pageId="5" pageNumber="20">P. richteri</emphasis>
|
|
</taxonomicName>
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|
and
|
|
<taxonomicName id="56BD4D30FFF5FFADFE2751033A07FE88" authority="(Iran)" baseAuthorityName="Iran" box="[413,599,330,352]" class="Insecta" family="Staphylinidae" genus="Pseudobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="iranicum">
|
|
<emphasis id="A3C9EAA1FFF5FFADFE2751033A45FE88" box="[413,533,331,352]" italics="true" pageId="5" pageNumber="20">P. iranicum</emphasis>
|
|
(
|
|
<collectingCountry id="E9AA7623FFF5FFADFD9F51023A00FE88" box="[549,592,330,352]" name="Iran" pageId="5" pageNumber="20">Iran</collectingCountry>
|
|
)
|
|
</taxonomicName>
|
|
by distinctly smaller size, darker coloration, and the different shape of the aedeagus.
|
|
</paragraph>
|
|
<paragraph id="910236B3FFF5FFADFF2D51C33BABFE54" blockId="5.[151,1059,394,444]" pageId="5" pageNumber="20">
|
|
For illustrations of the male primary sexual characters of other Palaearctic
|
|
<taxonomicName id="56BD4D30FFF5FFADFC1051C23C73FE77" authorityName="MULSANT & REY II. A" authorityYear="1878" box="[938,1059,394,415]" class="Insecta" family="Staphylinidae" genus="Pseudobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="19" phylum="Arthropoda" rank="genus">
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|
<emphasis id="A3C9EAA1FFF5FFADFC1051C23C73FE77" box="[938,1059,394,415]" italics="true" pageId="5" pageNumber="20">Pseudobium</emphasis>
|
|
</taxonomicName>
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|
species see
|
|
<bibRefCitation id="F52C4B42FFF5FFADFEBD51EE39FAFE54" author="COIFFAIT H" box="[263,426,422,444]" pageId="5" pageNumber="20" pagination="1 - 440" refId="ref3047" refString="COIFFAIT H. (1982): Coleopteres Staphylinidae de la region palearctique occidentale. IV. Sous famille Paederinae. Tribu Paederini 1 (Paederi, Lathrobii). - Supplement a la Nouvelle Revue d'Entomologie 12 (4): 1 - 440." type="journal article" year="1982">COIFFAIT (1982)</bibRefCitation>
|
|
,
|
|
<bibRefCitation id="F52C4B42FFF5FFADFE0951EF3A11FE54" author="BOHAC J" box="[435,577,422,444]" pageId="5" pageNumber="20" pagination="434 - 443" refId="ref3012" refString="BOHAC J. (1988): Descriptions of new species of the subfamily Paederinae (Coleoptera, Staphylinidae) from the Palaearctic region. - Acta Entomologica Bohemoslovaca 85: 434 - 443." type="journal article" year="1988">BOHÁČ (1988)</bibRefCitation>
|
|
,
|
|
<bibRefCitation id="F52C4B42FFF5FFADFDF151EE3ABDFE54" author="GUSAROV V" box="[587,749,422,444]" pageId="5" pageNumber="20" pagination="65 - 74" refId="ref3128" refString="GUSAROV V. I. (1993): New and little-known Palaearctic Staphylinidae (Coleoptera). - Zoosystematica Rossica 1: 65 - 74." type="journal article" year="1993">GUSAROV (1993</bibRefCitation>
|
|
,
|
|
<bibRefCitation id="F52C4B42FFF5FFADFD4D51EE3B7BFE54" author="GUSAROV V" box="[759,811,422,444]" pageId="5" pageNumber="20" pagination="431 - 446" refId="ref3155" refString="GUSAROV V. I. (1994): New and little-known Palaearctic Paederinae (Coleoptera: Staphylinidae). - Annales de la Societe Entomologique de France (N. S.) 30: 431 - 446." type="journal article" year="1994">1994</bibRefCitation>
|
|
), and
|
|
<bibRefCitation id="F52C4B42FFF5FFADFCDE51EE3BA7FE54" author="ASSING V" box="[868,1015,422,444]" pageId="5" pageNumber="20" pagination="385 - 391" refId="ref2973" refString="ASSING V. (2006): New species and records of Pseudobium MULSANT & REY (Insecta: Coleoptera: Staphylinidae: Paederinae). - Linzer biologische Beitrage 38 (1): 385 - 391." type="journal article" year="2006">ASSING (2006)</bibRefCitation>
|
|
.
|
|
</paragraph>
|
|
<caption id="C5C2663BFFF5FFADFF2D55B83A87F9D9" ID-DOI="http://doi.org/10.5281/zenodo.4523446" ID-Zenodo-Dep="4523446" httpUri="https://zenodo.org/record/4523446/files/figure.png" pageId="5" pageNumber="20" startId="5.[151,191,1520,1540]" targetBox="[155,1011,465,1506]" targetPageId="5">
|
|
<paragraph id="910236B3FFF5FFADFF2D55B83A87F9D9" blockId="5.[151,1059,1518,1585]" pageId="5" pageNumber="20">
|
|
<emphasis id="A3C9EAA1FFF5FFADFF2D55B838B7F9EC" bold="true" box="[151,231,1520,1540]" pageId="5" pageNumber="20">Figs 1-7</emphasis>
|
|
:
|
|
<taxonomicName id="56BD4D30FFF5FFADFF4D55A639B9F9EB" authority="Assing, 2007" authorityName="Assing" authorityYear="2007" box="[247,489,1518,1539]" class="Insecta" family="Staphylinidae" genus="Pseudobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="pakistanicum" status="sp. nov.">
|
|
<emphasis id="A3C9EAA1FFF5FFADFF4D55A639B9F9EB" box="[247,489,1518,1539]" italics="true" pageId="5" pageNumber="20">Pseudobium pakistanicum</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="B8FA57DAFFF5FFADFE4855B83A65F9EC" box="[498,565,1520,1540]" pageId="5" pageNumber="20" rank="species">nov.sp.</taxonomicNameLabel>
|
|
(holotype). (
|
|
<emphasis id="A3C9EAA1FFF5FFADFD0855B83AEDF9EC" bold="true" box="[690,701,1520,1540]" pageId="5" pageNumber="20">1</emphasis>
|
|
) forebody; (
|
|
<emphasis id="A3C9EAA1FFF5FFADFC8255B83B13F9EC" bold="true" box="[824,835,1520,1540]" pageId="5" pageNumber="20">2</emphasis>
|
|
) head; (
|
|
<emphasis id="A3C9EAA1FFF5FFADFC2C55B83BF1F9EC" bold="true" box="[918,929,1520,1540]" pageId="5" pageNumber="20">3</emphasis>
|
|
) antenna; (
|
|
<emphasis id="A3C9EAA1FFF5FFADFBAA55B83C4BF9EC" bold="true" box="[1040,1051,1520,1540]" pageId="5" pageNumber="20">4</emphasis>
|
|
) male sternite VIII; (
|
|
<emphasis id="A3C9EAA1FFF5FFADFEEC564F3925F9F3" bold="true" box="[342,373,1543,1563]" pageId="5" pageNumber="20">5-6</emphasis>
|
|
) aedeagus in lateral and in ventral view; (
|
|
<emphasis id="A3C9EAA1FFF5FFADFCB5564F3B4AF9F3" bold="true" box="[783,794,1543,1563]" pageId="5" pageNumber="20">7</emphasis>
|
|
) apical part of aedeagus in ventral view. Scale bars: 1: 1.0 mm; 2-4: 0.2 mm; 5-7: 0.1 mm.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="910236B3FFF6FFAEFF2D509C3C51FEEE" blockId="6.[151,1058,211,262]" pageId="6" pageNumber="21">
|
|
D i s t r i b u t i o n a n d b i o n o m i c s: The
|
|
<typeStatus id="4E068811FFF6FFAEFD76509C3AA0FF01" box="[716,752,212,233]" pageId="6" pageNumber="21">type</typeStatus>
|
|
locality is situated in northwestern
|
|
<collectingCountry id="E9AA7623FFF6FFAEFF5250B8396AFEEE" box="[232,314,240,262]" name="Pakistan" pageId="6" pageNumber="21">Pakistan</collectingCountry>
|
|
at an altitude of
|
|
<quantity id="56459B56FFF6FFAEFE6650B83A32FEEE" box="[476,610,240,262]" metricMagnitude="3" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="6" pageNumber="21" unit="m" value="2000.0" valueMax="2500.0" valueMin="1500.0">1500-2500 m</quantity>
|
|
. The
|
|
<typeStatus id="4E068811FFF6FFAEFD2C50B83ABBFEEE" box="[662,747,240,262]" pageId="6" pageNumber="21" type="holotype">holotype</typeStatus>
|
|
was collected in September.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |