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<document id="98964518CEAA9F1DB5B7F787530C1293" ID-CLB-Dataset="21417" ID-DOI="10.5852/ejt.2020.689" ID-GBIF-Dataset="74c501e2-be96-45ef-9f59-d3ea64b028c4" ID-Zenodo-Dep="3961738" ID-ZooBank="4E8B68C9-4F9F-4244-9A2C-192046753782" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1595860179353" checkinUser="valdenar" docAuthor="Bruno, Maria Cristina, Cottarelli, Vezio, Marrone, Federico, Grasso, Rosario, Stefani, Erika, Vecchioni, Luca &amp; Spena, Maria Teresa" docDate="2020" docId="183E87BFCD6BFFB6FE2E71D3FA99C039" docLanguage="en" docName="ejt-689_bruno_cottarelli_marrone.pdf" docOrigin="European Journal of Taxonomy 689" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Cottarellicaris sanctiangeli Bruno &amp; Cottarelli &amp; Marrone &amp; Grasso &amp; Stefani &amp; Vecchioni &amp; Spena 2020, sp. nov." docType="treatment" docUuid="BFD63C80-116A-49C7-B10D-547D0146CB5A" docUuidSource="ZooBank" docVersion="7" lastPageNumber="19" masterDocId="E407FFC7CD63FFA4FFFD723BFF9DC50A" masterDocTitle="Morphological and molecular characterization of three new Parastenocarididae (Copepoda: Harpacticoida) from caves in Southern Italy" masterLastPageNumber="46" masterPageNumber="1" pageNumber="9" updateTime="1698845518282" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="793560AD04B701B970F51818AD3330A1">Morphological and molecular characterization of three new Parastenocarididae (Copepoda: Harpacticoida) from caves in Southern Italy</mods:title>
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<mods:namePart id="C7AE00CF036B6D670D3616DB009EC2E6">Bruno, Maria Cristina</mods:namePart>
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<mods:affiliation id="1790CF748AA54E48272E83718565DFAD">Research and Innovation Centre, Fondazione Edmund Mach, Via E. Mach 1, 38010 San Michele allAdige (TN), Italy. Department for Innovation in Biological, Agro-food and Forest Systems, Tuscia University, Largo dellUniversità snc, 01100 Viterbo, Italy. Department of Biological, Chemical and Pharmaceutical Sciences and Technologies, University of Palermo, Via Archirafi 18, 90123, Palermo, Italy. Department of Biological, Geological and Environmental Sciences, Catania University, Via Androne 81, 95124 Catania, Italy.</mods:affiliation>
<mods:nameIdentifier id="7F8D982427CB63502E958F8084274C17" type="email">cristina.bruno@fmach.it</mods:nameIdentifier>
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<mods:namePart id="687620E2CEA8DDFCD4F2EFD685D12282">Cottarelli, Vezio</mods:namePart>
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<mods:affiliation id="546A70DC7B41DC9216D4526B8F7A5B35">Department for Innovation in Biological, Agro-food and Forest Systems, Tuscia University, Largo dellUniversità snc, 01100 Viterbo, Italy.</mods:affiliation>
<mods:nameIdentifier id="C96EEF143890A52C81D4DDDECE80BCE9" type="email">cottarel@unitus.it</mods:nameIdentifier>
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<mods:namePart id="593195627E59C7E6F7546C9EEA0B9F44">Marrone, Federico</mods:namePart>
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<mods:affiliation id="995662BD8355D6559383826277512363">Department of Biological, Chemical and Pharmaceutical Sciences and Technologies, University of Palermo, Via Archirafi 18, 90123, Palermo, Italy.</mods:affiliation>
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<mods:namePart id="BCF8B3AE2A47896C14F65E81ED68E1ED">Grasso, Rosario</mods:namePart>
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<mods:affiliation id="2DE348F3CFD293AA5D12CBB2E77BEEC1">Department of Biological, Geological and Environmental Sciences, Catania University, Via Androne 81, 95124 Catania, Italy.</mods:affiliation>
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<mods:namePart id="EE854B5296E789D420FD27DC958BADA0">Stefani, Erika</mods:namePart>
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<mods:affiliation id="51F079B6F080CFB4D918320D95961587">Research and Innovation Centre, Fondazione Edmund Mach, Via E. Mach 1, 38010 San Michele allAdige (TN), Italy. Department for Innovation in Biological, Agro-food and Forest Systems, Tuscia University, Largo dellUniversità snc, 01100 Viterbo, Italy. Department of Biological, Chemical and Pharmaceutical Sciences and Technologies, University of Palermo, Via Archirafi 18, 90123, Palermo, Italy. Department of Biological, Geological and Environmental Sciences, Catania University, Via Androne 81, 95124 Catania, Italy.</mods:affiliation>
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<mods:namePart id="0B6370D1BDBCA13320B86336500BBC63">Vecchioni, Luca</mods:namePart>
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<mods:affiliation id="C339C66CB66F51F69951BBD433279713">Department of Biological, Chemical and Pharmaceutical Sciences and Technologies, University of Palermo, Via Archirafi 18, 90123, Palermo, Italy.</mods:affiliation>
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<mods:namePart id="29AEDC8AE8CF53355CF45A9A1B6EB55E">Spena, Maria Teresa</mods:namePart>
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<mods:date id="08FF214B5F39C1EF5F5CD56670903C84">2020</mods:date>
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<treatment id="183E87BFCD6BFFB6FE2E71D3FA99C039" ID-DOI="http://doi.org/10.5281/zenodo.4327933" ID-GBIF-Taxon="166003066" ID-Zenodo-Dep="4327933" LSID="urn:lsid:zoobank.org:act:BFD63C80-116A-49C7-B10D-547D0146CB5A" httpUri="http://treatment.plazi.org/id/183E87BFCD6BFFB6FE2E71D3FA99C039" lastPageId="18" lastPageNumber="19" pageId="8" pageNumber="9">
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<heading id="CB6081C5CD6BFFACFE2E71D3FBFCC109" box="[467,1121,1000,1027]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="9" reason="2">
<taxonomicName id="57974D2ACD6BFFACFE2E71D3FC62C108" authority="Bruno &amp; Cottarelli" authorityName="Bruno &amp; Cottarelli &amp; Marrone &amp; Grasso &amp; Stefani &amp; Vecchioni &amp; Spena" authorityYear="2020" box="[467,1023,1000,1027]" class="Maxillopoda" family="Parastenocarididae" genus="Cottarellicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="sanctiangeli" status="sp. nov.">
<emphasis id="A2E3EABBCD6BFFACFE2E71D3FC87C109" bold="true" box="[467,794,1000,1027]" italics="true" pageId="8" pageNumber="9">Cottarellicaris sanctiangeli</emphasis>
Bruno &amp; Cottarelli
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<figureCitation id="08AC2A2CCD6BFFACFD5D7614FC95C143" box="[672,776,1071,1097]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStart-6="Fig" captionStartId-0="10.[189,232,1966,1992]" captionStartId-1="11.[189,232,1825,1851]" captionStartId-2="12.[189,232,1895,1921]" captionStartId-3="14.[189,232,1862,1888]" captionStartId-4="15.[189,232,1825,1851]" captionStartId-5="17.[189,232,1859,1885]" captionStartId-6="18.[188,231,1033,1059]" captionTargetBox-0="[222,1398,508,1920]" captionTargetBox-1="[195,1384,351,1760]" captionTargetBox-2="[211,1394,915,1855]" captionTargetBox-3="[223,1358,296,1822]" captionTargetBox-4="[200,1378,266,1782]" captionTargetBox-5="[214,1388,351,1791]" captionTargetBox-6="[342,1246,265,991]" captionTargetId-0="figure@10.[189,1398,496,1928]" captionTargetId-1="figure@11.[189,1398,340,1777]" captionTargetId-2="figure@12.[189,1398,904,1877]" captionTargetId-3="figure@14.[189,1398,294,1829]" captionTargetId-4="figure@15.[189,1398,237,1792]" captionTargetId-5="figure@17.[189,1398,338,1818]" captionTargetId-6="figure@18.[342,1246,265,991]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionTargetPageId-3="14" captionTargetPageId-4="15" captionTargetPageId-5="17" captionTargetPageId-6="18" captionText-0="Fig. 2. Cottarellicaris sanctiangeli Bruno &amp; Cottarelli sp. nov. from Grotta superiore di SantAngelo (NHMUK). a. ♂, habitus. b. ♀, habitus. Scale bar: 50 µm." captionText-1="Fig. 3. Cottarellicaris sanctiangeli Bruno &amp; Cottarelli sp. nov., ♂ (NHMUK). a. Anal somite, anal operculum and caudal rami, dorsal view. b. Anal somite, anal operculum and caudal rami, dorsal view (variability). c. Anal somite, anal operculum and caudal rami, dorsal view. d. Anal somite, anal operculum and caudal rami, lateral view (variability). e. Anal somite, anal operculum and caudal rami, lateral view (variability). a: Grotta superiore di SantAngelo; b, d: Grotta del Banco di ferro; c, e: Grotta Vucco Ucciardo. Scale bar: 50 µm." captionText-2="Fig. 4. Cottarellicaris sanctiangeli Bruno &amp; Cottarelli sp. nov., ♂ (NHMUK). a. Rostrum and A1, ventral view. b. Rostrum and A1, ventral view. c. A1, disarticulated (antennular segments marked with Roman numerals). d. A2. e. Mdb. f. Mx1. g. Mx2. h. Mxp. a, eg: Grotta Vucco Ucciardo; bd, h: Grotta superiore di SantAngelo. Scale bar: 50 µm." captionText-3="Fig. 5. Cottarellicaris sanctiangeli Bruno &amp; Cottarelli sp. nov., ♂ (NHMUK). a. P1, coxa, basis and enp, inner view. b. P1, basis and exp, outer view. c. P2, posterior view. d. P3, anterior view. e. P3, posterior view (variability). f. P4, anterior view. g. P4, intercoxal sclerite, coxa, basis and enp, outer view. h. P4, posterior view. a, c, e, g: Grotta Vucco Ucciardo; b, h: Grotta del Banco di ferro; d, f: Grotta superiore di SantAngelo. Scale bar: 50 µm." captionText-4="Fig. 6. Cottarellicaris sanctiangeli Bruno &amp; Cottarelli sp. nov. (NHMUK). a. ♂, P5, P6, first and second urosomites, ventral view. b. ♂, P5. c. ♂, P5 and first urosomite, ventral view. d. ♂, P5. e. ♀, first urosomite, P5, P6, genital double-somite and genital field, ventral view. f. ♀, anal somite, anal operculum and caudal rami, dorsal view. g. ♀, anal somite, anal operculum and caudal rami, lateral view. ab, e, g: Grotta Vucco Ucciardo; c: Grotta del Banco di ferro; d, f: Grotta superiore di SantAngelo. Scale bar: 50 µm." captionText-5="Fig. 7. Cottarellicaris sanctiangeli Bruno &amp; Cottarelli sp. nov., ♀ (NHMUK). a. Anal somite, anal operculum and caudal rami, dorsal view (variability). b. Anal somite, anal operculum and caudal rami, lateral view (variability). c. Rostrum, A1. d. P1, anterior view. e. P2, inner view. f. P3, posterior view. g. Coxa, basis, enp P3, inner view. h. P4, posterior view. i. Coxa, basis, enp P4, inner view (variability). a, ci: Grotta Grotta Vucco Ucciardo; b: Grotta del Banco di ferro. Scale bar: 50 µm." captionText-6="Fig. 8. Cottarellicaris sanctiangeli Bruno &amp; Cottarelli sp. nov. (CIME). a. ♂, anal somite, anal operculum and caudal rami, ventro-lateral view. b. ♀, anal somite, anal operculum and caudal rami, lateral view. c. ♀, P5, P6, genital double-somite and genital field, ventral view (outermost seta, which is folded behind the P5 arrowed). d. ♀, enp P4. Grotta superiore di SantAngelo. Scale bar: 50 µm." figureDoi-0="http://doi.org/10.5281/zenodo.3961742" figureDoi-1="http://doi.org/10.5281/zenodo.3961744" figureDoi-2="http://doi.org/10.5281/zenodo.3961746" figureDoi-3="http://doi.org/10.5281/zenodo.3961748" figureDoi-4="http://doi.org/10.5281/zenodo.3961750" figureDoi-5="http://doi.org/10.5281/zenodo.3961752" figureDoi-6="http://doi.org/10.5281/zenodo.3961754" httpUri-0="https://zenodo.org/record/3961742/files/figure.png" httpUri-1="https://zenodo.org/record/3961744/files/figure.png" httpUri-2="https://zenodo.org/record/3961746/files/figure.png" httpUri-3="https://zenodo.org/record/3961748/files/figure.png" httpUri-4="https://zenodo.org/record/3961750/files/figure.png" httpUri-5="https://zenodo.org/record/3961752/files/figure.png" httpUri-6="https://zenodo.org/record/3961754/files/figure.png" pageId="8" pageNumber="9">
Figs 2
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8
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;
<tableCitation id="DD150312CD6BFFACFCE97614FC09C143" box="[788,916,1071,1097]" captionStart-0="Table 1" captionStart-1="Table 2" captionStart-2="Table 3" captionStartId-0="3.[189,258,268,294]" captionStartId-1="6.[189,258,268,294]" captionStartId-2="33.[189,258,268,294]" captionTargetPageId-0="3" captionText-0="Table 1. Estimated distances between caves and from the seashore for each cave." captionText-1="Table 2. List of species used for phylogenetic analysis, collection site, number of specimens for which DNA was successfully extracted and amplified for the two investigated fragments (COI and 18S), number of haplotypes recorded for each species/fragment, and GenBank accession numbers." captionText-2="Table 3. Pairwise distances (p-distance model) among COI sequences between the analysed species of Parastenocarididae." httpUri-0="http://table.plazi.org/id/C4E86621CD60FFA7FF407337FBE8C42C" pageId="8" pageNumber="9" tableUuid-0="C4E86621CD60FFA7FF407337FBE8C42C">Tables 13</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="D88D6522CD6BFFACFF40764BFE7CC029" pageId="8" pageNumber="9" type="diagnosis">
<paragraph id="902836A9CD6BFFACFF40764BFEABC180" blockId="8.[189,310,1136,1162]" box="[189,310,1136,1162]" pageId="8" pageNumber="9">
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<emphasis id="A2E3EABBCD6BFFACFF40764BFEABC180" bold="true" box="[189,310,1136,1162]" pageId="8" pageNumber="9">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="902836A9CD6BFFACFF4076A4FE7CC029" blockId="8.[189,1399,1183,1315]" pageId="8" pageNumber="9">
<taxonomicName id="57974D2ACD6BFFACFF4076A4FD7BC1B3" authority="Bruno &amp; Cottarelli" authorityName="Bruno &amp; Cottarelli &amp; Marrone &amp; Grasso &amp; Stefani &amp; Vecchioni &amp; Spena" authorityYear="2020" box="[189,742,1183,1209]" class="Maxillopoda" family="Parastenocarididae" genus="Cottarellicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="sanctiangeli" status="sp. nov.">
<emphasis id="A2E3EABBCD6BFFACFF4076A4FE62C1B3" box="[189,511,1183,1209]" italics="true" pageId="8" pageNumber="9">Cottarellicaris sanctiangeli</emphasis>
Bruno &amp; Cottarelli
</taxonomicName>
<taxonomicNameLabel id="B9D057C0CD6BFFACFD1376A4FCD4C1B3" box="[750,841,1183,1209]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
is characterized by the peculiar morphology of the male P4 enp, with apically curved inner tip and, mainly, with a thin proximal outgrowth as long as the apical (crenulate) lamellar outgrowth. Females are characterized by the P3 enp being shorter than the corresponding exp-1.
</paragraph>
</subSubSection>
<subSubSection id="D88D6522CD6BFFACFF407771FB8EC0D3" pageId="8" pageNumber="9" type="etymology">
<paragraph id="902836A9CD6BFFACFF407771FED9C06E" blockId="8.[189,324,1354,1380]" box="[189,324,1354,1380]" pageId="8" pageNumber="9">
<heading id="CB6081C5CD6BFFACFF407771FED9C06E" bold="true" box="[189,324,1354,1380]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="A2E3EABBCD6BFFACFF407771FED9C06E" bold="true" box="[189,324,1354,1380]" pageId="8" pageNumber="9">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="902836A9CD6BFFACFF407743FB8EC0D3" blockId="8.[189,1399,1400,1497]" pageId="8" pageNumber="9">
The species epithet is the masculine genitive of the Latin words
<emphasis id="A2E3EABBCD6BFFACFC567742FB9EC098" box="[939,1027,1401,1426]" italics="true" pageId="8" pageNumber="9">sanctus</emphasis>
and
<emphasis id="A2E3EABBCD6BFFACFBB67743FB3BC098" box="[1099,1190,1400,1426]" italics="true" pageId="8" pageNumber="9">angelus</emphasis>
, meaning Saint and angel, respectively, and refer to the locus typicus of this
<taxonomicName id="57974D2ACD6BFFACFC5977A7FBCDC0BC" authorityName="Schminke" authorityYear="2013" box="[932,1104,1436,1462]" class="Maxillopoda" family="Parastenocarididae" genus="Cottarellicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="A2E3EABBCD6BFFACFC5977A7FBCDC0BC" box="[932,1104,1436,1462]" italics="true" pageId="8" pageNumber="9">Cottarellicaris</emphasis>
</taxonomicName>
, i.e., a cave dedicated to Saint Michael the Archangel, whose cult is widespread in Southern
<collectingCountry id="E8807639CD6BFFACFC2A7784FB92C0D3" box="[983,1039,1471,1497]" name="Italy" pageId="8" pageNumber="9">Italy</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="D88D6522CD6BFFADFF40743BFADBC12A" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="902836A9CD6BFFACFF40743BFE37C310" blockId="8.[189,426,1536,1562]" box="[189,426,1536,1562]" pageId="8" pageNumber="9">
<heading id="CB6081C5CD6BFFACFF40743BFE37C310" bold="true" box="[189,426,1536,1562]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="A2E3EABBCD6BFFACFF40743BFE37C310" bold="true" box="[189,426,1536,1562]" pageId="8" pageNumber="9">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="902836A9CD6BFFACFF177414FEC1C343" blockId="8.[189,1399,1583,1750]" box="[234,348,1583,1609]" pageId="8" pageNumber="9">
<emphasis id="A2E3EABBCD6BFFACFF177414FEC1C343" bold="true" box="[234,348,1583,1609]" pageId="8" pageNumber="9">
<typeStatus id="4F2C880BCD6BFFACFF177414FEC1C343" box="[234,348,1583,1609]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="902836A9CD6BFFACFF407469FE7CC3DC" blockId="8.[189,1399,1583,1750]" pageId="8" pageNumber="9">
<materialsCitation id="20FF3CF4CD6BFFACFF407469FE7CC3DC" ID-GBIF-Occurrence="2818212324" collectingDate="2017-05-09" collectionCode="R, T" collectorName="R. Grasso &amp; M. T. Spena" country="Italy" county="Grotta Superiore di Sant'Angelo" latitude="39.788956" location="Cassano allo Ionio" longLatPrecision="1" longitude="16.30711" municipality="Complesso Grotte di Sant'Angelo" pageId="8" pageNumber="9" specimenCode="NHMUK 2020.4" specimenCount="1" stateProvince="Cosenza Province" typeStatus="holotype">
<collectingCountry id="E8807639CD6BFFACFF407469FE92C366" box="[189,271,1618,1644]" name="Italy" pageId="8" pageNumber="9">ITALY</collectingCountry>
<specimenCount id="8691FD20CD6BFFACFEDE7468FEA6C367" box="[291,315,1619,1645]" pageId="8" pageNumber="9" type="male"></specimenCount>
;
<collectingRegion id="5253F84BCD6BFFACFEBB7469FD84C366" box="[326,537,1618,1644]" pageId="8" pageNumber="9">Cosenza Province</collectingRegion>
,
<location id="95486072CD6BFFACFDDE7469FD61C366" LSID="urn:lsid:plazi:treatment:183E87BFCD6BFFB6FE2E71D3FA99C039:95486072CD6BFFACFDDE7469FD61C366" box="[547,764,1618,1644]" country="Italy" county="Grotta Superiore di Sant'Angelo" latitude="39.788956" longLatPrecision="1" longitude="16.30711" municipality="Complesso Grotte di Sant'Angelo" name="Cassano allo Ionio" pageId="8" pageNumber="9" stateProvince="Cosenza Province">Cassano allo Ionio</location>
,
<location id="95486072CD6BFFACFCF87469FB12C366" LSID="urn:lsid:plazi:treatment:183E87BFCD6BFFB6FE2E71D3FA99C039:95486072CD6BFFACFCF87469FB12C366" box="[773,1167,1618,1644]" country="Italy" county="Grotta Superiore di Sant'Angelo" latitude="39.788956" longLatPrecision="1" longitude="16.30711" municipality="Complesso Grotte di Sant'Angelo" name="Complesso Grotte di Sant'Angelo" pageId="8" pageNumber="9" stateProvince="Cosenza Province">Complesso Grotte di SantAngelo</location>
,
<location id="95486072CD6BFFACFB647469FEC9C385" LSID="urn:lsid:plazi:treatment:183E87BFCD6BFFB6FE2E71D3FA99C039:95486072CD6BFFACFB647469FEC9C385" country="Italy" county="Grotta Superiore di Sant'Angelo" latitude="39.788956" longLatPrecision="1" longitude="16.30711" municipality="Complesso Grotte di Sant'Angelo" name="Grotta Superiore di Sant'Angelo" pageId="8" pageNumber="9" stateProvince="Cosenza Province">Grotta Superiore di SantAngelo</location>
, pool 2; approximate coordinates
<geoCoordinate id="F5A3506ECD6BFFACFD1C7448FC04C39A" box="[737,921,1651,1680]" degrees="39" direction="north" minutes="47" orientation="latitude" pageId="8" pageNumber="9" precision="1" seconds="20.24" value="39.788956">39°4720.24ʺ N</geoCoordinate>
,
<geoCoordinate id="F5A3506ECD6BFFACFC587448FBC7C39A" box="[933,1114,1651,1680]" degrees="16" direction="east" minutes="18" orientation="longitude" pageId="8" pageNumber="9" precision="1" seconds="25.60" value="16.30711">16°1825.60ʺ E</geoCoordinate>
;
<date id="E4291069CD6BFFACFB9A744DFB6AC39A" box="[1127,1271,1654,1680]" pageId="8" pageNumber="9" value="2017-05-09">
<collectingDate id="F46DE981CD6BFFACFB9A744DFB6AC39A" box="[1127,1271,1654,1680]" pageId="8" pageNumber="9" value="2017-05-09">9 May 2017</collectingDate>
</date>
;
<collectorName id="3D62537FCD6BFFACFAFE744DFAEBC39A" box="[1283,1398,1654,1680]" pageId="8" pageNumber="9">R. Grasso</collectorName>
and
<collectorName id="3D62537FCD6BFFACFF1074A2FEE9C3B9" box="[237,372,1689,1715]" pageId="8" pageNumber="9">M.T. Spena</collectorName>
leg.; dissected and mounted on one slide labelled “
<taxonomicName id="57974D2ACD6BFFACFC3C74A2FA9CC3B9" authorityName="Bruno &amp; Cottarelli &amp; Marrone &amp; Grasso &amp; Stefani &amp; Vecchioni &amp; Spena" authorityYear="2020" box="[961,1281,1689,1715]" class="Maxillopoda" family="Parastenocarididae" genus="Cottarellicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="sanctiangeli">
<emphasis id="A2E3EABBCD6BFFACFC3C74A2FA9CC3B9" box="[961,1281,1689,1715]" italics="true" pageId="8" pageNumber="9">Cottarellicaris sanctiangeli</emphasis>
</taxonomicName>
holotype: male”;
<specimenCode id="C0319ED2CD6BFFACFEEE7487FE43C3DC" box="[275,478,1724,1750]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="8" pageNumber="9" type="Museum">NHMUK 2020.4</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="902836A9CD6BFFACFF1774C5FEFAC212" blockId="8.[189,1399,1790,2027]" box="[234,359,1790,1816]" pageId="8" pageNumber="9">
<emphasis id="A2E3EABBCD6BFFACFF1774C5FEFAC212" bold="true" box="[234,359,1790,1816]" pageId="8" pageNumber="9">
<typeStatus id="4F2C880BCD6BFFACFF1774C5FEFAC212" box="[234,359,1790,1816]" pageId="8" pageNumber="9" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="902836A9CD6BFFADFF40751AFDE7C4DE" blockId="8.[189,1399,1790,2027]" lastBlockId="9.[189,1399,267,469]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<materialsCitation id="20FF3CF4CD6BFFACFF40751AFBA0C254" ID-GBIF-Occurrence="2818212314" collectingDate="2017-05-09" collectionCode="NHMUK" collectorName="Grasso &amp; Spena" country="Italy" county="Grotta Superiore di Sant'Angelo" latitude="39.788956" location="ITALY" longLatPrecision="1" longitude="16.30711" municipality="Complesso Grotte di Sant'Angelo" pageId="8" pageNumber="9" specimenCode="NHMUK 2020.5, 2020.6" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<collectingCountry id="E8807639CD6BFFACFF40751AFE92C231" box="[189,271,1825,1851]" name="Italy" pageId="8" pageNumber="9">ITALY</collectingCountry>
<specimenCount id="8691FD20CD6BFFACFED5751AFEF0C231" box="[296,365,1825,1851]" pageId="8" pageNumber="9" type="male">2 ♂♂</specimenCount>
; same collection data as for holotype; each dissected and mounted on one slide labelled “
<taxonomicName id="57974D2ACD6BFFACFF37757FFD8CC254" authorityName="Bruno &amp; Cottarelli &amp; Marrone &amp; Grasso &amp; Stefani &amp; Vecchioni &amp; Spena" authorityYear="2020" box="[202,529,1860,1886]" class="Maxillopoda" family="Parastenocarididae" genus="Cottarellicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="sanctiangeli">
<emphasis id="A2E3EABBCD6BFFACFF37757FFD8CC254" box="[202,529,1860,1886]" italics="true" pageId="8" pageNumber="9">Cottarellicaris sanctiangeli</emphasis>
</taxonomicName>
paratype: male”;
<specimenCode id="C0319ED2CD6BFFACFD0C757FFC20C255" box="[753,957,1860,1887]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="8" pageNumber="9" type="Museum">NHMUK 2020.5</specimenCode>
to
<specimenCode id="C0319ED2CD6BFFACFC11757FFBA0C254" box="[1004,1085,1860,1886]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="8" pageNumber="9" type="Museum">2020.6</specimenCode>
</materialsCitation>
<materialsCitation id="20FF3CF4CD6BFFACFBA2757FFCCBC2AE" ID-GBIF-Occurrence="2818212320" collectingDate="2017-05-09" collectionCode="NHMUK" collectorName="Grasso &amp; Spena" country="Italy" county="Grotta Superiore di Sant'Angelo" latitude="39.788956" location="Cosenza Province" longLatPrecision="1" longitude="16.30711" municipality="Complesso Grotte di Sant'Angelo" pageId="8" pageNumber="9" specimenCode="NHMUK 2020.7, 2020.8" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="8691FD20CD6BFFACFBA2757FFB35C254" box="[1119,1192,1860,1886]" pageId="8" pageNumber="9" type="male">2 ♂♂</specimenCount>
; same collection data as for holotype but pool 4; dissected and mounted each on one slide labelled “
<taxonomicName id="57974D2ACD6BFFACFB36755CFED1C2AE" authorityName="Bruno &amp; Cottarelli &amp; Marrone &amp; Grasso &amp; Stefani &amp; Vecchioni &amp; Spena" authorityYear="2020" class="Maxillopoda" family="Parastenocarididae" genus="Cottarellicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="sanctiangeli">
<emphasis id="A2E3EABBCD6BFFACFB36755CFED1C2AE" italics="true" pageId="8" pageNumber="9">Cottarellicaris sanctiangeli</emphasis>
</taxonomicName>
paratype: male”;
<specimenCode id="C0319ED2CD6BFFACFDE675B0FD7FC2AE" box="[539,738,1930,1957]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="8" pageNumber="9" type="Museum">NHMUK 2020.7</specimenCode>
to
<specimenCode id="C0319ED2CD6BFFACFCF875B1FCCBC2AE" box="[773,854,1930,1956]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="8" pageNumber="9" type="Museum">2020.8</specimenCode>
</materialsCitation>
<materialsCitation id="20FF3CF4CD6BFFACFC9075B1FE1AC2E1" ID-GBIF-Occurrence="2818212316" collectingDate="2017-05-09" collectionCode="NHMUK" collectorName="Grasso &amp; Spena" country="Italy" county="Grotta Superiore di Sant'Angelo" latitude="39.788956" location="Cosenza Province" longLatPrecision="1" longitude="16.30711" municipality="Complesso Grotte di Sant'Angelo" pageId="8" pageNumber="9" specimenCode="NHMUK 2020.9" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="8691FD20CD6BFFACFC9075B1FC06C2AF" box="[877,923,1930,1957]" pageId="8" pageNumber="9" type="male">1 ♂</specimenCount>
; same collection data as for holotype but pool 4; undissected and mounted on one slide labelled “
<taxonomicName id="57974D2ACD6BFFACFC9E7596FB3AC2CD" authorityName="Bruno &amp; Cottarelli &amp; Marrone &amp; Grasso &amp; Stefani &amp; Vecchioni &amp; Spena" authorityYear="2020" box="[867,1191,1965,1991]" class="Maxillopoda" family="Parastenocarididae" genus="Cottarellicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="sanctiangeli">
<emphasis id="A2E3EABBCD6BFFACFC9E7596FB3AC2CD" box="[867,1191,1965,1991]" italics="true" pageId="8" pageNumber="9">Cottarellicaris sanctiangeli</emphasis>
</taxonomicName>
paratype: male”;
<specimenCode id="C0319ED2CD6BFFACFF4075EAFE1AC2E1" box="[189,391,2001,2027]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="8" pageNumber="9" type="Museum">NHMUK 2020.9</specimenCode>
</materialsCitation>
<materialsCitation id="20FF3CF4CD6BFFADFE5B75EAFB74C42C" ID-GBIF-Occurrence="2818212301" collectingDate="2017-05-09" collectionCode="NHMUK" collectorName="Grasso &amp; Spena" country="Italy" county="Grotta Superiore di Sant'Angelo" lastPageId="9" lastPageNumber="10" latitude="39.788956" location="Cosenza Province" longLatPrecision="1" longitude="16.30711" municipality="Complesso Grotte di Sant'Angelo" pageId="8" pageNumber="9" specimenCode="NHMUK 2020.10, 2020.12" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
<specimenCount id="8691FD20CD6BFFACFE5B75EAFE70C2E1" box="[422,493,2001,2027]" pageId="8" pageNumber="9" type="female">3 ♀♀</specimenCount>
; same collection data as for holotype; each dissected and mounted on one slide labelled “
<taxonomicName id="57974D2ACD6AFFADFE967330FD31C42F" authorityName="Bruno &amp; Cottarelli &amp; Marrone &amp; Grasso &amp; Stefani &amp; Vecchioni &amp; Spena" authorityYear="2020" box="[363,684,267,293]" class="Maxillopoda" family="Parastenocarididae" genus="Cottarellicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="sanctiangeli">
<emphasis id="A2E3EABBCD6AFFADFE967330FD31C42F" box="[363,684,267,293]" italics="true" pageId="9" pageNumber="10">Cottarellicaris sanctiangeli</emphasis>
</taxonomicName>
paratype: female”;
<specimenCode id="C0319ED2CD6AFFADFC6F7337FBFBC42C" box="[914,1126,268,294]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="9" pageNumber="10" type="Museum">NHMUK 2020.10</specimenCode>
to
<specimenCode id="C0319ED2CD6AFFADFB777337FB74C42C" box="[1162,1257,268,294]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="9" pageNumber="10" type="Museum">2020.12</specimenCode>
</materialsCitation>
<materialsCitation id="20FF3CF4CD6AFFADFAFC7337FE66C461" ID-GBIF-Occurrence="2818212317" collectingDate="2017-05-09" collectionCode="NHMUK" collectorName="Grasso &amp; Spena" country="Italy" county="Grotta Superiore di Sant'Angelo" latitude="39.788956" location="Cosenza Province" longLatPrecision="1" longitude="16.30711" municipality="Complesso Grotte di Sant'Angelo" pageId="9" pageNumber="10" specimenCode="NHMUK 2020.13" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="8691FD20CD6AFFADFAFC7337FAB0C42C" box="[1281,1325,268,294]" pageId="9" pageNumber="10" type="female">1 ♀</specimenCount>
; same collection data as for holotype; mounted on one slide labelled “
<taxonomicName id="57974D2ACD6AFFADFC467315FB62C442" authorityName="Bruno &amp; Cottarelli &amp; Marrone &amp; Grasso &amp; Stefani &amp; Vecchioni &amp; Spena" authorityYear="2020" box="[955,1279,302,328]" class="Maxillopoda" family="Parastenocarididae" genus="Cottarellicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="sanctiangeli">
<emphasis id="A2E3EABBCD6AFFADFC467315FB62C442" box="[955,1279,302,328]" italics="true" pageId="9" pageNumber="10">Cottarellicaris sanctiangeli</emphasis>
</taxonomicName>
paratype: female”;
<specimenCode id="C0319ED2CD6AFFADFED57369FE66C461" box="[296,507,337,364]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="9" pageNumber="10" type="Museum">NHMUK 2020.13</specimenCode>
</materialsCitation>
<materialsCitation id="20FF3CF4CD6AFFADFDEC736AFB46C484" ID-GBIF-Occurrence="2818212321" collectingDate="2017-05-09" collectionCode="NHMUK" collectorName="Grasso &amp; Spena" country="Italy" county="Grotta Superiore di Sant'Angelo" latitude="39.788956" location="Cosenza Province" longLatPrecision="1" longitude="16.30711" municipality="Complesso Grotte di Sant'Angelo" pageId="9" pageNumber="10" specimenCode="NHMUK 2020.14" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="8691FD20CD6AFFADFDEC736AFDA1C466" box="[529,572,337,364]" pageId="9" pageNumber="10" type="female">1 ♀</specimenCount>
; same collection data as for holotype but pool 4; dissected and mounted on one slide labelled “
<taxonomicName id="57974D2ACD6AFFADFE33734FFC8EC484" authorityName="Bruno &amp; Cottarelli &amp; Marrone &amp; Grasso &amp; Stefani &amp; Vecchioni &amp; Spena" authorityYear="2020" box="[462,787,372,398]" class="Maxillopoda" family="Parastenocarididae" genus="Cottarellicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="sanctiangeli">
<emphasis id="A2E3EABBCD6AFFADFE33734FFC8EC484" box="[462,787,372,398]" italics="true" pageId="9" pageNumber="10">Cottarellicaris sanctiangeli</emphasis>
</taxonomicName>
paratype: female”;
<specimenCode id="C0319ED2CD6AFFADFBFE734EFB46C484" box="[1027,1243,372,399]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="9" pageNumber="10" type="Museum">NHMUK 2020.14</specimenCode>
</materialsCitation>
<materialsCitation id="20FF3CF4CD6AFFADFB04734FFDEEC4DF" ID-GBIF-Occurrence="2818212310" collectingDate="2017-05-09" collectionCode="NHMUK" collectorName="Grasso &amp; Spena" country="Italy" county="Grotta Superiore di Sant'Angelo" latitude="39.788956" location="Cosenza Province" longLatPrecision="1" longitude="16.30711" municipality="Complesso Grotte di Sant'Angelo" pageId="9" pageNumber="10" specimenCode="NHMUK 2020.15" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="8691FD20CD6AFFADFB04734FFAB4C485" box="[1273,1321,372,399]" pageId="9" pageNumber="10" type="female">1 ♀</specimenCount>
; same collection data as for holotype but pool 4; mounted on one slide labelled “
<taxonomicName id="57974D2ACD6AFFADFBCE73ACFAEBC4BB" authorityName="Bruno &amp; Cottarelli &amp; Marrone &amp; Grasso &amp; Stefani &amp; Vecchioni &amp; Spena" authorityYear="2020" box="[1075,1398,407,433]" class="Maxillopoda" family="Parastenocarididae" genus="Cottarellicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="sanctiangeli">
<emphasis id="A2E3EABBCD6AFFADFBCE73ACFAEBC4BB" box="[1075,1398,407,433]" italics="true" pageId="9" pageNumber="10">Cottarellicaris sanctiangeli</emphasis>
</taxonomicName>
paratype: female”;
<specimenCode id="C0319ED2CD6AFFADFE637381FDEEC4DF" box="[414,627,442,469]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="9" pageNumber="10" type="Museum">NHMUK 2020.15</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="902836A9CD6AFFADFF1773C0FE3BC71F" blockId="9.[189,1399,507,1056]" box="[234,422,507,533]" pageId="9" pageNumber="10">
<emphasis id="A2E3EABBCD6AFFADFF1773C0FE3BC71F" bold="true" box="[234,422,507,533]" pageId="9" pageNumber="10">Other material</emphasis>
</paragraph>
<paragraph id="902836A9CD6AFFADFF407025FADBC12A" blockId="9.[189,1399,507,1056]" pageId="9" pageNumber="10">
<materialsCitation id="20FF3CF4CD6AFFADFF407025FC27C777" ID-GBIF-Occurrence="2818212325" collectingDate="2013-04-25" collectionCode="R, T" collectorName="R. Grasso &amp; M. T. Spena" country="Italy" county="Vucco Ucciardo" latitude="39.78432" location="Cassano allo Ionio" longLatPrecision="1" longitude="16.320799" municipality="Ionio" pageId="9" pageNumber="10" specimenCode="NHMUK 2020.16" specimenCount="1" stateProvince="Cosenza Province">
<collectingCountry id="E8807639CD6AFFADFF407025FE92C732" box="[189,271,542,568]" name="Italy" pageId="9" pageNumber="10">ITALY</collectingCountry>
<specimenCount id="8691FD20CD6AFFADFEDA7025FEC9C732" box="[295,340,542,568]" pageId="9" pageNumber="10">1 ♂</specimenCount>
;
<collectingRegion id="5253F84BCD6AFFADFE9C7025FDAAC732" box="[353,567,542,568]" pageId="9" pageNumber="10">Cosenza Province</collectingRegion>
,
<location id="95486072CD6AFFADFDBF7025FC82C732" LSID="urn:lsid:plazi:treatment:183E87BFCD6BFFB6FE2E71D3FA99C039:95486072CD6AFFADFDBF7025FC82C732" box="[578,799,542,568]" country="Italy" county="Vucco Ucciardo" latitude="39.78432" longLatPrecision="1" longitude="16.320799" municipality="Ionio" name="Cassano allo Ionio" pageId="9" pageNumber="10" stateProvince="Cosenza Province">Cassano allo Ionio</location>
,
<location id="95486072CD6AFFADFCD47025FC77C732" LSID="urn:lsid:plazi:treatment:183E87BFCD6BFFB6FE2E71D3FA99C039:95486072CD6AFFADFCD47025FC77C732" box="[809,1002,542,568]" country="Italy" county="Vucco Ucciardo" latitude="39.78432" longLatPrecision="1" longitude="16.320799" municipality="Ionio" name="Vucco Ucciardo" pageId="9" pageNumber="10" stateProvince="Cosenza Province">Vucco Ucciardo</location>
, pool 1; approximate coordinates;
<geoCoordinate id="F5A3506ECD6AFFADFF407004FEF7C751" box="[189,362,575,603]" degrees="39" direction="north" minutes="47" orientation="latitude" pageId="9" pageNumber="10" precision="1" seconds="3.56" value="39.78432">39°473.56ʺ N</geoCoordinate>
,
<geoCoordinate id="F5A3506ECD6AFFADFE857004FDACC751" box="[376,561,575,603]" degrees="16" direction="east" minutes="19" orientation="longitude" pageId="9" pageNumber="10" precision="1" seconds="14.87" value="16.320799">16°1914.87″ E</geoCoordinate>
;
<date id="E4291069CD6AFFADFDBD707BFD7CC751" box="[576,737,576,603]" pageId="9" pageNumber="10" value="2013-04-25">
<collectingDate id="F46DE981CD6AFFADFDBD707BFD7CC751" box="[576,737,576,603]" pageId="9" pageNumber="10" value="2013-04-25">25 Apr. 2013</collectingDate>
</date>
;
<collectorName id="3D62537FCD6AFFADFD12707AFCF8C751" box="[751,869,577,603]" pageId="9" pageNumber="10">R. Grasso</collectorName>
and
<collectorName id="3D62537FCD6AFFADFC5E707AFBB0C751" box="[931,1069,577,603]" pageId="9" pageNumber="10">M.T. Spena</collectorName>
leg.; mounted on one slide labelled “
<taxonomicName id="57974D2ACD6AFFADFECC7058FDEBC777" authorityName="Bruno &amp; Cottarelli &amp; Marrone &amp; Grasso &amp; Stefani &amp; Vecchioni &amp; Spena" authorityYear="2020" box="[305,630,611,637]" class="Maxillopoda" family="Parastenocarididae" genus="Cottarellicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="sanctiangeli">
<emphasis id="A2E3EABBCD6AFFADFECC7058FDEBC777" box="[305,630,611,637]" italics="true" pageId="9" pageNumber="10">Cottarellicaris sanctiangeli</emphasis>
</taxonomicName>
: male”;
<specimenCode id="C0319ED2CD6AFFADFD1F705FFC27C777" box="[738,954,611,638]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="9" pageNumber="10" type="Museum">NHMUK 2020.16</specimenCode>
</materialsCitation>
<materialsCitation id="20FF3CF4CD6AFFADFC27705FFDB8C7C9" ID-GBIF-Occurrence="2818212302" collectingDate="2013-04-25" collectionCode="NHMUK" collectorName="Grasso &amp; Spena" country="Italy" county="Vucco Ucciardo" latitude="39.78432" location="Cosenza Province" longLatPrecision="1" longitude="16.320799" municipality="Ionio" pageId="9" pageNumber="10" specimenCode="NHMUK 2020.17, 2020.20" specimenCount="4" specimenCount-male="4" stateProvince="Cosenza Province">
<specimenCount id="8691FD20CD6AFFADFC27705FFBBCC774" box="[986,1057,612,638]" pageId="9" pageNumber="10" type="male">4 ♂♂</specimenCount>
; same collection data as for preceding; each dissected and mounted on one slide labelled “
<taxonomicName id="57974D2ACD6AFFADFC3670BDFA8EC7AA" authorityName="Bruno &amp; Cottarelli &amp; Marrone &amp; Grasso &amp; Stefani &amp; Vecchioni &amp; Spena" authorityYear="2020" box="[971,1299,646,672]" class="Maxillopoda" family="Parastenocarididae" genus="Cottarellicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="sanctiangeli">
<emphasis id="A2E3EABBCD6AFFADFC3670BDFA8EC7AA" box="[971,1299,646,672]" italics="true" pageId="9" pageNumber="10">Cottarellicaris sanctiangeli</emphasis>
</taxonomicName>
: male”;
<specimenCode id="C0319ED2CD6AFFADFF407091FE0AC7C9" box="[189,407,681,708]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="9" pageNumber="10" type="Museum">NHMUK 2020.17</specimenCode>
to
<specimenCode id="C0319ED2CD6AFFADFE3B7092FDB8C7C9" box="[454,549,681,707]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="9" pageNumber="10" type="Museum">2020.20</specimenCode>
</materialsCitation>
<materialsCitation id="20FF3CF4CD6AFFADFDB57092FC55C7EC" ID-GBIF-Occurrence="2818212306" collectingDate="2013-04-25" collectionCode="NHMUK" collectorName="Grasso &amp; Spena" country="Italy" county="Vucco Ucciardo" latitude="39.78432" location="Cosenza Province" longLatPrecision="1" longitude="16.320799" municipality="Ionio" pageId="9" pageNumber="10" specimenCode="NHMUK 2020.21" specimenCount="1" specimenCount-female="1" stateProvince="Cosenza Province">
<specimenCount id="8691FD20CD6AFFADFDB57092FDE7C7CE" box="[584,634,681,708]" pageId="9" pageNumber="10" type="female">1 ♀</specimenCount>
; same collection data as for preceding; mounted on one slide labelled “
<taxonomicName id="57974D2ACD6AFFADFECD70F7FDEEC7EC" authorityName="Bruno &amp; Cottarelli &amp; Marrone &amp; Grasso &amp; Stefani &amp; Vecchioni &amp; Spena" authorityYear="2020" box="[304,627,716,742]" class="Maxillopoda" family="Parastenocarididae" genus="Cottarellicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="sanctiangeli">
<emphasis id="A2E3EABBCD6AFFADFECD70F7FDEEC7EC" box="[304,627,716,742]" italics="true" pageId="9" pageNumber="10">Cottarellicaris sanctiangeli</emphasis>
</taxonomicName>
: female”;
<specimenCode id="C0319ED2CD6AFFADFD0F70F6FC55C7EC" box="[754,968,716,743]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="9" pageNumber="10" type="Museum">NHMUK 2020.21</specimenCode>
</materialsCitation>
<materialsCitation id="20FF3CF4CD6AFFADFC1970F7FD82C627" ID-GBIF-Occurrence="2818212319" collectingDate="2013-04-25" collectionCode="NHMUK" collectorName="Grasso &amp; Spena" country="Italy" county="Vucco Ucciardo" latitude="39.78432" location="Cosenza Province" longLatPrecision="1" longitude="16.320799" municipality="Ionio" pageId="9" pageNumber="10" specimenCode="NHMUK 2020.22, 2020.25" specimenCount="4" specimenCount-female="4" stateProvince="Cosenza Province">
<specimenCount id="8691FD20CD6AFFADFC1970F7FBB5C7ED" box="[996,1064,716,743]" pageId="9" pageNumber="10" type="female">4 ♀♀</specimenCount>
; same collection data as for preceding; each dissected and mounted on one slide labelled “
<taxonomicName id="57974D2ACD6AFFADFC4570D4FB63C603" authorityName="Bruno &amp; Cottarelli &amp; Marrone &amp; Grasso &amp; Stefani &amp; Vecchioni &amp; Spena" authorityYear="2020" box="[952,1278,751,777]" class="Maxillopoda" family="Parastenocarididae" genus="Cottarellicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="sanctiangeli">
<emphasis id="A2E3EABBCD6AFFADFC4570D4FB63C603" box="[952,1278,751,777]" italics="true" pageId="9" pageNumber="10">Cottarellicaris sanctiangeli</emphasis>
</taxonomicName>
: female”;
<specimenCode id="C0319ED2CD6AFFADFF407129FE08C626" box="[189,405,786,812]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="9" pageNumber="10" type="Museum">NHMUK 2020.22</specimenCode>
to
<specimenCode id="C0319ED2CD6AFFADFE3D7129FD82C627" box="[448,543,786,813]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="9" pageNumber="10" type="Museum">2020.25</specimenCode>
</materialsCitation>
<materialsCitation id="20FF3CF4CD6AFFADFDC07129FD87C69E" ID-GBIF-Occurrence="2818212322" collectingDate="2015-05-27" collectionCode="T" collectorName="Grasso &amp; M. T. Spena" country="Italy" county="San Lorenzo Bellizzi" latitude="39.873993" location="Grotta del Banco di ferro" longLatPrecision="1" longitude="16.34093" municipality="San Lorenzo Bellizzi" pageId="9" pageNumber="10" specimenCode="NHMUK 2020.26, 2020.27" specimenCount="2" specimenCount-male="2" stateProvince="Cosenza Province">
<specimenCount id="8691FD20CD6AFFADFDC07129FD19C626" box="[573,644,786,812]" pageId="9" pageNumber="10" type="male">2 ♂♂</specimenCount>
;
<collectingRegion id="5253F84BCD6AFFADFD697129FCF1C626" box="[660,876,786,812]" pageId="9" pageNumber="10">Cosenza Province</collectingRegion>
,
<collectingCounty id="79494E25CD6AFFADFC877129FBE7C626" box="[890,1146,786,812]" pageId="9" pageNumber="10">San Lorenzo Bellizzi</collectingCounty>
,
<location id="95486072CD6AFFADFB757129FF64C645" LSID="urn:lsid:plazi:treatment:183E87BFCD6BFFB6FE2E71D3FA99C039:95486072CD6AFFADFB757129FF64C645" country="Italy" county="San Lorenzo Bellizzi" latitude="39.873993" longLatPrecision="1" longitude="16.34093" municipality="San Lorenzo Bellizzi" name="Grotta del Banco di ferro" pageId="9" pageNumber="10" stateProvince="Cosenza Province">Grotta del Banco di ferro</location>
, pool 3; approximate coordinates:
<geoCoordinate id="F5A3506ECD6AFFADFD5F7108FCC2C645" box="[674,863,819,847]" degrees="39" direction="north" minutes="52" orientation="latitude" pageId="9" pageNumber="10" precision="1" seconds="26.38" value="39.873993">39°5226.38ʺ N</geoCoordinate>
,
<geoCoordinate id="F5A3506ECD6AFFADFC937108FBB5C645" box="[878,1064,819,847]" degrees="16" direction="east" minutes="20" orientation="longitude" pageId="9" pageNumber="10" precision="1" seconds="27.35" value="16.34093">16°2027.35ʺ E</geoCoordinate>
;
<date id="E4291069CD6AFFADFBC5710EFB7DC645" box="[1080,1248,820,847]" pageId="9" pageNumber="10" value="2015-05-27">
<collectingDate id="F46DE981CD6AFFADFBC5710EFB7DC645" box="[1080,1248,820,847]" pageId="9" pageNumber="10" value="2015-05-27">27 May 2015</collectingDate>
</date>
;
<collectorName id="3D62537FCD6AFFADFB0D710EFADCC645" box="[1264,1345,821,847]" pageId="9" pageNumber="10">Grasso</collectorName>
and
<collectorName id="3D62537FCD6AFFADFF407163FED9C678" box="[189,324,856,882]" pageId="9" pageNumber="10">M.T. Spena</collectorName>
leg.; each dissected and mounted on one slide labelled “
<taxonomicName id="57974D2ACD6AFFADFC2A716CFA84C67B" authorityName="Bruno &amp; Cottarelli &amp; Marrone &amp; Grasso &amp; Stefani &amp; Vecchioni &amp; Spena" authorityYear="2020" box="[983,1305,855,881]" class="Maxillopoda" family="Parastenocarididae" genus="Cottarellicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="sanctiangeli">
<emphasis id="A2E3EABBCD6AFFADFC2A716CFA84C67B" box="[983,1305,855,881]" italics="true" pageId="9" pageNumber="10">Cottarellicaris sanctiangeli</emphasis>
</taxonomicName>
: male”;
<specimenCode id="C0319ED2CD6AFFADFF407140FE0EC69E" box="[189,403,890,917]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="9" pageNumber="10" type="Museum">NHMUK 2020.26</specimenCode>
to
<specimenCode id="C0319ED2CD6AFFADFE467141FD87C69E" box="[443,538,890,916]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="9" pageNumber="10" type="Museum">2020.27</specimenCode>
</materialsCitation>
<materialsCitation id="20FF3CF4CD6AFFADFDC87141FB54C6BD" ID-GBIF-Occurrence="2818212312" collectingDate="2015-05-27" collectionCode="NHMUK" collectorName="Grasso &amp; Spena" country="Italy" county="Grotta del Banco" latitude="39.873993" location="Grotta del Banco di ferro" longLatPrecision="1" longitude="16.34093" municipality="San Lorenzo Bellizzi" pageId="9" pageNumber="10" specimenCode="NHMUK 2020.28, 2020.29" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Cosenza Province">
<specimenCount id="8691FD20CD6AFFADFDC87141FDFEC69F" box="[565,611,890,917]" pageId="9" pageNumber="10" type="male">1 ♂</specimenCount>
,
<specimenCount id="8691FD20CD6AFFADFD8F7141FD02C69F" box="[626,671,890,917]" pageId="9" pageNumber="10" type="female">1 ♀</specimenCount>
; same collection data as for preceding; mounted on one slide labelled “
<taxonomicName id="57974D2ACD6AFFADFECD71A6FDE9C6BD" authorityName="Bruno &amp; Cottarelli &amp; Marrone &amp; Grasso &amp; Stefani &amp; Vecchioni &amp; Spena" authorityYear="2020" box="[304,628,925,951]" class="Maxillopoda" family="Parastenocarididae" genus="Cottarellicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="sanctiangeli">
<emphasis id="A2E3EABBCD6AFFADFECD71A6FDE9C6BD" box="[304,628,925,951]" italics="true" pageId="9" pageNumber="10">Cottarellicaris sanctiangeli</emphasis>
</taxonomicName>
: male and female”;
<specimenCode id="C0319ED2CD6AFFADFC9471A5FBDDC6BD" box="[873,1088,925,952]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="9" pageNumber="10" type="Museum">NHMUK 2020.28</specimenCode>
to
<specimenCode id="C0319ED2CD6AFFADFB9771A6FB54C6BD" box="[1130,1225,925,951]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="9" pageNumber="10" type="Museum">2020.29</specimenCode>
</materialsCitation>
<materialsCitation id="20FF3CF4CD6AFFADFB1B71A6FCA6C6F7" ID-GBIF-Occurrence="2818212305" collectingDate="2015-05-27" collectionCode="NHMUK" collectorName="Grasso &amp; Spena" country="Italy" county="Grotta del Banco" latitude="39.873993" location="Cosenza Province" longLatPrecision="1" longitude="16.34093" municipality="San Lorenzo Bellizzi" pageId="9" pageNumber="10" specimenCode="NHMUK 2020.30, 2020.31" specimenCount="2" specimenCount-female="2" stateProvince="Cosenza Province">
<specimenCount id="8691FD20CD6AFFADFB1B71A6FAB6C6B2" box="[1254,1323,925,952]" pageId="9" pageNumber="10" type="female">2 ♀♀</specimenCount>
, same collection data as for preceding; each dissected and mounted on one slide labelled “
<taxonomicName id="57974D2ACD6AFFADFB3671FBFED1C6F7" authorityName="Bruno &amp; Cottarelli &amp; Marrone &amp; Grasso &amp; Stefani &amp; Vecchioni &amp; Spena" authorityYear="2020" class="Maxillopoda" family="Parastenocarididae" genus="Cottarellicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="sanctiangeli">
<emphasis id="A2E3EABBCD6AFFADFB3671FBFED1C6F7" italics="true" pageId="9" pageNumber="10">Cottarellicaris sanctiangeli</emphasis>
</taxonomicName>
: female”;
<specimenCode id="C0319ED2CD6AFFADFE2F71D8FD31C6F7" box="[466,684,995,1021]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="9" pageNumber="10" type="Museum">NHMUK 2020.30</specimenCode>
to
<specimenCode id="C0319ED2CD6AFFADFD2171D8FCA6C6F7" box="[732,827,995,1021]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="9" pageNumber="10" type="Museum">2020.31</specimenCode>
</materialsCitation>
<materialsCitation id="20FF3CF4CD6AFFADFCA371D8FAA2C12A" ID-GBIF-Occurrence="2818212308" collectingDate="2017-05-10" collectionCode="NHMUK" collectorName="Grasso &amp; Spena" country="Italy" county="Grotta del Banco" latitude="39.873993" location="Cosenza Province" longLatPrecision="1" longitude="16.34093" municipality="San Lorenzo Bellizzi" pageId="9" pageNumber="10" specimenCode="NHMUK 2020.32" specimenCount="1" specimenCount-female="1" stateProvince="Cosenza Province">
<specimenCount id="8691FD20CD6AFFADFCA371D8FC12C6F7" box="[862,911,995,1021]" pageId="9" pageNumber="10" type="female">1 ♀</specimenCount>
, same collection data as for preceding;
<date id="E4291069CD6AFFADFF40763DFEC0C12A" box="[189,349,1030,1056]" pageId="9" pageNumber="10" value="2017-05-10">
<collectingDate id="F46DE981CD6AFFADFF40763DFEC0C12A" box="[189,349,1030,1056]" pageId="9" pageNumber="10" value="2017-05-10">10 May 2017</collectingDate>
</date>
; mounted on slide labelled “
<taxonomicName id="57974D2ACD6AFFADFD56763DFC70C12A" authorityName="Bruno &amp; Cottarelli &amp; Marrone &amp; Grasso &amp; Stefani &amp; Vecchioni &amp; Spena" authorityYear="2020" box="[683,1005,1030,1056]" class="Maxillopoda" family="Parastenocarididae" genus="Cottarellicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="sanctiangeli">
<emphasis id="A2E3EABBCD6AFFADFD56763DFC70C12A" box="[683,1005,1030,1056]" italics="true" pageId="9" pageNumber="10">Cottarellicaris sanctiangeli</emphasis>
</taxonomicName>
: female”;
<specimenCode id="C0319ED2CD6AFFADFB97763DFAA2C12A" box="[1130,1343,1030,1056]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="9" pageNumber="10" type="Museum">NHMUK 2020.32</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D88D6522CD6AFFB6FF40767CFA99C039" lastPageId="18" lastPageNumber="19" pageId="9" pageNumber="10" type="description">
<paragraph id="902836A9CD6AFFADFF40767CFED3C16B" blockId="9.[189,334,1095,1121]" box="[189,334,1095,1121]" pageId="9" pageNumber="10">
<heading id="CB6081C5CD6AFFADFF40767CFED3C16B" bold="true" box="[189,334,1095,1121]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
<emphasis id="A2E3EABBCD6AFFADFF40767CFED3C16B" bold="true" box="[189,334,1095,1121]" pageId="9" pageNumber="10">Description</emphasis>
</heading>
</paragraph>
<paragraph id="902836A9CD6AFFADFF17764EFEE8C185" blockId="9.[189,1399,1141,1516]" box="[234,373,1141,1167]" pageId="9" pageNumber="10">
<emphasis id="A2E3EABBCD6AFFADFF17764EFEE8C185" bold="true" box="[234,373,1141,1167]" pageId="9" pageNumber="10">Adult male</emphasis>
</paragraph>
<paragraph id="902836A9CD6AFFADFF4076A3FC37C0E6" blockId="9.[189,1399,1141,1516]" pageId="9" pageNumber="10">
BODY. Unpigmented, nauplius eye absent. Total body length, measured from tip of rostrum to posterior margin of caudal rami (excluding caudal setae) from
<quantity id="576F9B4CCD6AFFADFCBC7680FC6AC1DF" box="[833,1015,1211,1238]" metricMagnitude="-1" metricUnit="m" metricValue="3.1399999999999997" metricValueMax="3.3299999999999996" metricValueMin="2.95" pageId="9" pageNumber="10" unit="mm" value="314.0" valueMax="333.0" valueMin="295.0">295 to 333 µm</quantity>
, mean 311 μm (n = 6). Habitus (
<figureCitation id="08AC2A2CCD6AFFADFF3B76E5FE87C1F2" box="[198,282,1246,1272]" captionStart="Fig" captionStartId="10.[189,232,1966,1992]" captionTargetBox="[222,1398,508,1920]" captionTargetId="figure@10.[189,1398,496,1928]" captionTargetPageId="10" captionText="Fig. 2. Cottarellicaris sanctiangeli Bruno &amp; Cottarelli sp. nov. from Grotta superiore di SantAngelo (NHMUK). a. ♂, habitus. b. ♀, habitus. Scale bar: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3961742" httpUri="https://zenodo.org/record/3961742/files/figure.png" pageId="9" pageNumber="10">Fig. 2a</figureCitation>
) cylindrical and slender, without any demarcation between prosome and urosome; prosome to urosome ratio: 0.90. Free pedigerous somites without any lateral or dorsal expansions, all connected by well-developed arthrodial membranes.Integument weakly sclerotized, without cuticular pits, ornamented with sensilla on all somites except preanal one. Cuticular windows on urosomites and cephalothorax not observed. Cephalothorax representing about 20 % of total body length. Anal somite (
<figureCitation id="08AC2A2CCD6AFFADFB6E7752FA9DC08E" box="[1171,1280,1385,1412]" captionStart="Fig" captionStartId="11.[189,232,1825,1851]" captionTargetBox="[195,1384,351,1760]" captionTargetId="figure@11.[189,1398,340,1777]" captionTargetPageId="11" captionText="Fig. 3. Cottarellicaris sanctiangeli Bruno &amp; Cottarelli sp. nov., ♂ (NHMUK). a. Anal somite, anal operculum and caudal rami, dorsal view. b. Anal somite, anal operculum and caudal rami, dorsal view (variability). c. Anal somite, anal operculum and caudal rami, dorsal view. d. Anal somite, anal operculum and caudal rami, lateral view (variability). e. Anal somite, anal operculum and caudal rami, lateral view (variability). a: Grotta superiore di SantAngelo; b, d: Grotta del Banco di ferro; c, e: Grotta Vucco Ucciardo. Scale bar: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3961744" httpUri="https://zenodo.org/record/3961744/files/figure.png" pageId="9" pageNumber="10">Fig. 3a, c</figureCitation>
) with pair of large dorsal sensilla at base of anal operculum, pair of cuticular lateral pores (one pore on each side) on proximal margin. Anal operculum (
<figureCitation id="08AC2A2CCD6AFFADFD787794FD6BC0C3" box="[645,758,1455,1481]" captionStart="Fig" captionStartId="11.[189,232,1825,1851]" captionTargetBox="[195,1384,351,1760]" captionTargetId="figure@11.[189,1398,340,1777]" captionTargetPageId="11" captionText="Fig. 3. Cottarellicaris sanctiangeli Bruno &amp; Cottarelli sp. nov., ♂ (NHMUK). a. Anal somite, anal operculum and caudal rami, dorsal view. b. Anal somite, anal operculum and caudal rami, dorsal view (variability). c. Anal somite, anal operculum and caudal rami, dorsal view. d. Anal somite, anal operculum and caudal rami, lateral view (variability). e. Anal somite, anal operculum and caudal rami, lateral view (variability). a: Grotta superiore di SantAngelo; b, d: Grotta del Banco di ferro; c, e: Grotta Vucco Ucciardo. Scale bar: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3961744" httpUri="https://zenodo.org/record/3961744/files/figure.png" pageId="9" pageNumber="10">Fig. 3a, c</figureCitation>
) well-developed, ornamented with row of spinules on outer surface, with straight distal margin. Anal sinus wide open.
</paragraph>
<paragraph id="902836A9CD6AFFADFF407428FD1DC3F4" blockId="9.[189,1399,1555,1790]" pageId="9" pageNumber="10">
CAUDAL RAMI (
<figureCitation id="08AC2A2CCD6AFFADFE907428FE47C327" box="[365,474,1555,1581]" captionStart="Fig" captionStartId="11.[189,232,1825,1851]" captionTargetBox="[195,1384,351,1760]" captionTargetId="figure@11.[189,1398,340,1777]" captionTargetPageId="11" captionText="Fig. 3. Cottarellicaris sanctiangeli Bruno &amp; Cottarelli sp. nov., ♂ (NHMUK). a. Anal somite, anal operculum and caudal rami, dorsal view. b. Anal somite, anal operculum and caudal rami, dorsal view (variability). c. Anal somite, anal operculum and caudal rami, dorsal view. d. Anal somite, anal operculum and caudal rami, lateral view (variability). e. Anal somite, anal operculum and caudal rami, lateral view (variability). a: Grotta superiore di SantAngelo; b, d: Grotta del Banco di ferro; c, e: Grotta Vucco Ucciardo. Scale bar: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3961744" httpUri="https://zenodo.org/record/3961744/files/figure.png" pageId="9" pageNumber="10">Fig. 3a, c</figureCitation>
). Shorter than anal somite, approximately cylindrical, slightly divergent; length to width ratio: 2.7. Anterolateral accessory seta (I) and anterolateral seta (II) subequal in length, both setae inserted together distally at 5/6 length of caudal ramus, smooth. Posterolateral seta (III) missing. Outer terminal seta (IV) long and pinnate (length seta/length caudal ramus: 1.5), inserted subterminally and projecting outwards; inner terminal seta (V) without fracture plane. Terminal accessory seta (VI) short (length seta/length caudal ramus: 0.7) and smooth. Dorsal seta (VII) articulate, inserted distally at 5/6 length of the caudal ramus, smooth.
</paragraph>
<paragraph id="902836A9CD6AFFADFF40751EFD40C268" blockId="9.[189,1398,1829,1891]" pageId="9" pageNumber="10">
ROSTRUM (
<figureCitation id="08AC2A2CCD6AFFADFEC3751EFE1EC24A" box="[318,387,1829,1856]" captionStart="Fig" captionStartId="12.[189,232,1895,1921]" captionTargetBox="[211,1394,915,1855]" captionTargetId="figure@12.[189,1398,904,1877]" captionTargetPageId="12" captionText="Fig. 4. Cottarellicaris sanctiangeli Bruno &amp; Cottarelli sp. nov., ♂ (NHMUK). a. Rostrum and A1, ventral view. b. Rostrum and A1, ventral view. c. A1, disarticulated (antennular segments marked with Roman numerals). d. A2. e. Mdb. f. Mx1. g. Mx2. h. Mxp. a, eg: Grotta Vucco Ucciardo; bd, h: Grotta superiore di SantAngelo. Scale bar: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3961746" httpUri="https://zenodo.org/record/3961746/files/figure.png" pageId="9" pageNumber="10">Fig. 4</figureCitation>
ab). Small, not demarcated at base, almost reaching distal margin of first antennulary segment, ornamented with two dorsal sensilla.
</paragraph>
<paragraph id="902836A9CD6AFFAEFF4075B1FAEBC49A" blockId="9.[189,1399,1930,2027]" lastBlockId="10.[189,1399,267,400]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
A1 (
<figureCitation id="08AC2A2CCD6AFFADFF0975B1FEA1C2AE" box="[244,316,1930,1957]" captionStart="Fig" captionStartId="12.[189,232,1895,1921]" captionTargetBox="[211,1394,915,1855]" captionTargetId="figure@12.[189,1398,904,1877]" captionTargetPageId="12" captionText="Fig. 4. Cottarellicaris sanctiangeli Bruno &amp; Cottarelli sp. nov., ♂ (NHMUK). a. Rostrum and A1, ventral view. b. Rostrum and A1, ventral view. c. A1, disarticulated (antennular segments marked with Roman numerals). d. A2. e. Mdb. f. Mx1. g. Mx2. h. Mxp. a, eg: Grotta Vucco Ucciardo; bd, h: Grotta superiore di SantAngelo. Scale bar: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3961746" httpUri="https://zenodo.org/record/3961746/files/figure.png" pageId="9" pageNumber="10">Fig. 4</figureCitation>
ac). Prehensile, eight-segmented pocket-knife
<typeStatus id="4F2C880BCD6AFFADFC8C75B0FC39C2AF" box="[881,932,1931,1957]" pageId="9" pageNumber="10">type</typeStatus>
<emphasis id="A2E3EABBCD6AFFADFC5075B0FC73C2AE" box="[941,1006,1931,1956]" italics="true" pageId="9" pageNumber="10">sensu</emphasis>
<bibRefCitation id="F4064B58CD6AFFADFC0575B1FB54C2AE" author="Schminke H. K." box="[1016,1225,1930,1956]" pageId="9" pageNumber="10" pagination="343 - 367" refId="ref25363" refString="Schminke H. K. 2010. High-level phylogenetic relationships within Parastenocarididae (Copepoda, Harpacticoida). Crustaceana 83: 343 - 367. https: // doi. org / 10.1163 / 001121610 X 12627655658168" type="journal article" year="2010">Schminke (2010)</bibRefCitation>
. First segment short with transversal row of spinules; second segment longest, with six setae, longest seta unipinnate; third segment with four distal bare setae; fourth segment reduced to a small sclerite with two short setae; fifth segment enlarged with inner round expansion carrying one short and two longer subequal setae, and distal tubercle with one seta and one large aesthetasc, reaching past end of eighth segment; sixth segment bare, partially fused to previous one; seventh segment bare, distal anterior corner protruding as curved apophysis ending in bilobate tip; eighth segment with seven setae and apical acrothek represented two
</paragraph>
<caption id="C4E86621CD69FFAEFF407595FCF7C2E6" ID-DOI="http://doi.org/10.5281/zenodo.3961742" ID-Zenodo-Dep="3961742" httpUri="https://zenodo.org/record/3961742/files/figure.png" pageId="10" pageNumber="11" startId="10.[189,232,1966,1992]" targetBox="[222,1398,508,1920]" targetPageId="10">
<paragraph id="902836A9CD69FFAEFF407595FCF7C2E6" blockId="10.[189,1398,1966,2028]" pageId="10" pageNumber="11">
<emphasis id="A2E3EABBCD69FFAEFF407595FE90C2C2" bold="true" box="[189,269,1966,1992]" pageId="10" pageNumber="11">Fig. 2.</emphasis>
<taxonomicName id="57974D2ACD69FFAEFEEB7595FCDBC2C2" authority="Bruno &amp; Cottarelli" authorityName="Bruno &amp; Cottarelli &amp; Marrone &amp; Grasso &amp; Stefani &amp; Vecchioni &amp; Spena" authorityYear="2020" box="[278,838,1966,1992]" class="Maxillopoda" family="Parastenocarididae" genus="Cottarellicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="sanctiangeli" status="sp. nov.">
<emphasis id="A2E3EABBCD69FFAEFEEB7595FDC7C2C2" box="[278,602,1966,1992]" italics="true" pageId="10" pageNumber="11">Cottarellicaris sanctiangeli</emphasis>
Bruno &amp; Cottarelli
</taxonomicName>
<taxonomicNameLabel id="B9D057C0CD69FFAEFCB27595FC31C2C2" box="[847,940,1966,1992]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
from Grotta superiore di SantAngelo (NHMUK).
<emphasis id="A2E3EABBCD69FFAEFEB375EAFEC0C2E1" bold="true" box="[334,349,2001,2027]" pageId="10" pageNumber="11">a</emphasis>
. ♂, habitus.
<emphasis id="A2E3EABBCD69FFAEFE0E75EAFD9EC2E1" bold="true" box="[499,515,2001,2027]" pageId="10" pageNumber="11">b</emphasis>
. ♀, habitus. Scale bar: 50 µm.
</paragraph>
</caption>
<caption id="C4E86621CD68FFAFFF40751AFDC4C2E1" ID-DOI="http://doi.org/10.5281/zenodo.3961744" ID-Zenodo-Dep="3961744" httpUri="https://zenodo.org/record/3961744/files/figure.png" pageId="11" pageNumber="12" startId="11.[189,232,1825,1851]" targetBox="[195,1384,351,1760]" targetPageId="11">
<paragraph id="902836A9CD68FFAFFF40751AFDC4C2E1" blockId="11.[189,1399,1825,2027]" pageId="11" pageNumber="12">
<emphasis id="A2E3EABBCD68FFAFFF40751AFE93C231" bold="true" box="[189,270,1825,1851]" pageId="11" pageNumber="12">Fig. 3.</emphasis>
<taxonomicName id="57974D2ACD68FFAFFEE4751AFCCDC231" authority="Bruno &amp; Cottarelli" authorityName="Bruno &amp; Cottarelli &amp; Marrone &amp; Grasso &amp; Stefani &amp; Vecchioni &amp; Spena" authorityYear="2020" box="[281,848,1825,1851]" class="Maxillopoda" family="Parastenocarididae" genus="Cottarellicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="sanctiangeli" status="sp. nov.">
<emphasis id="A2E3EABBCD68FFAFFEE4751AFDC2C231" box="[281,607,1825,1851]" italics="true" pageId="11" pageNumber="12">Cottarellicaris sanctiangeli</emphasis>
Bruno &amp; Cottarelli
</taxonomicName>
<taxonomicNameLabel id="B9D057C0CD68FFAFFCA6751AFC21C231" box="[859,956,1825,1851]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
, ♂ (NHMUK).
<emphasis id="A2E3EABBCD68FFAFFB7C751AFB0DC231" bold="true" box="[1153,1168,1825,1851]" pageId="11" pageNumber="12">a</emphasis>
. Anal somite, anal operculum and caudal rami, dorsal view.
<emphasis id="A2E3EABBCD68FFAFFD3C757FFD4CC254" bold="true" box="[705,721,1860,1886]" pageId="11" pageNumber="12">b</emphasis>
. Anal somite, anal operculum and caudal rami, dorsal view (variability).
<emphasis id="A2E3EABBCD68FFAFFE61755CFE34C28B" bold="true" box="[412,425,1895,1921]" pageId="11" pageNumber="12">c</emphasis>
. Anal somite, anal operculum and caudal rami, dorsal view.
<emphasis id="A2E3EABBCD68FFAFFB78755CFB08C28B" bold="true" box="[1157,1173,1895,1921]" pageId="11" pageNumber="12">d</emphasis>
. Anal somite, anal operculum and caudal rami, lateral view (variability).
<emphasis id="A2E3EABBCD68FFAFFCC675B0FCD5C2AF" bold="true" box="[827,840,1931,1957]" pageId="11" pageNumber="12">e</emphasis>
. Anal somite, anal operculum and caudal rami, lateral view (variability). a: Grotta superiore di SantAngelo; b, d: Grotta del Banco di ferro; c, e: Grotta Vucco Ucciardo. Scale bar: 50 µm.
</paragraph>
</caption>
<paragraph id="902836A9CD6FFFA8FF407337FC1AC443" blockId="12.[189,1398,267,329]" pageId="12" pageNumber="13">setae and long, slender aesthetasc. Armature formula: 1-[0], 2-[1 unipinnate + 5 bare], 3-[4 bare], 4-[2 bare], 5-[4 bare + ae], 6-[0], 7-[0], 8-[7 bare + (2 bare + ae)].</paragraph>
<paragraph id="902836A9CD6FFFA8FF40734BFC44C4FE" blockId="12.[189,1398,368,500]" pageId="12" pageNumber="13">
A2 (
<figureCitation id="08AC2A2CCD6FFFA8FF0C734BFEDAC481" box="[241,327,368,395]" captionStart="Fig" captionStartId="12.[189,232,1895,1921]" captionTargetBox="[211,1394,915,1855]" captionTargetId="figure@12.[189,1398,904,1877]" captionTargetPageId="12" captionText="Fig. 4. Cottarellicaris sanctiangeli Bruno &amp; Cottarelli sp. nov., ♂ (NHMUK). a. Rostrum and A1, ventral view. b. Rostrum and A1, ventral view. c. A1, disarticulated (antennular segments marked with Roman numerals). d. A2. e. Mdb. f. Mx1. g. Mx2. h. Mxp. a, eg: Grotta Vucco Ucciardo; bd, h: Grotta superiore di SantAngelo. Scale bar: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3961746" httpUri="https://zenodo.org/record/3961746/files/figure.png" pageId="12" pageNumber="13">Fig. 4d</figureCitation>
). Coxa unarmed; allobasis with transverse row of spinules on inner margin. Exp represented by small segment merged with allobasis, with pinnate apical seta. Enp bearing two short subdistal unipinnate spines and five longer distal elements: two of them geniculated, one transformed with furcate tip, all elements unipinnate, with long spinules near their insertions.
</paragraph>
<paragraph id="902836A9CD6FFFA8FF407027FC03C752" blockId="12.[189,1399,539,601]" pageId="12" pageNumber="13">
MDB (
<figureCitation id="08AC2A2CCD6FFFA8FEF77020FEFEC73F" box="[266,355,539,566]" captionStart="Fig" captionStartId="12.[189,232,1895,1921]" captionTargetBox="[211,1394,915,1855]" captionTargetId="figure@12.[189,1398,904,1877]" captionTargetPageId="12" captionText="Fig. 4. Cottarellicaris sanctiangeli Bruno &amp; Cottarelli sp. nov., ♂ (NHMUK). a. Rostrum and A1, ventral view. b. Rostrum and A1, ventral view. c. A1, disarticulated (antennular segments marked with Roman numerals). d. A2. e. Mdb. f. Mx1. g. Mx2. h. Mxp. a, eg: Grotta Vucco Ucciardo; bd, h: Grotta superiore di SantAngelo. Scale bar: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3961746" httpUri="https://zenodo.org/record/3961746/files/figure.png" pageId="12" pageNumber="13">Fig. 4e</figureCitation>
). Coxal gnathobase bare, cutting edge with apical teeth and short bipinnate seta. Onesegmented palp, with two smooth, apical setae of equal length.
</paragraph>
<paragraph id="902836A9CD6FFFA8FF4070BBFD52C7EB" blockId="12.[189,1398,640,737]" pageId="12" pageNumber="13">
<collectingCountry id="E8807639CD6FFFA8FF4070BBFF70C790" box="[189,237,640,666]" name="Mexico" pageId="12" pageNumber="13">MX</collectingCountry>
1 (
<figureCitation id="08AC2A2CCD6FFFA8FEFB70BBFEC5C791" box="[262,344,640,667]" captionStart="Fig" captionStartId="12.[189,232,1895,1921]" captionTargetBox="[211,1394,915,1855]" captionTargetId="figure@12.[189,1398,904,1877]" captionTargetPageId="12" captionText="Fig. 4. Cottarellicaris sanctiangeli Bruno &amp; Cottarelli sp. nov., ♂ (NHMUK). a. Rostrum and A1, ventral view. b. Rostrum and A1, ventral view. c. A1, disarticulated (antennular segments marked with Roman numerals). d. A2. e. Mdb. f. Mx1. g. Mx2. h. Mxp. a, eg: Grotta Vucco Ucciardo; bd, h: Grotta superiore di SantAngelo. Scale bar: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3961746" httpUri="https://zenodo.org/record/3961746/files/figure.png" pageId="12" pageNumber="13">Fig. 4f</figureCitation>
). Praecoxal arthrite with three apical curved robust spines apically denticled, one subdistal curved seta. Coxal endite long, with one apical seta. Basis cylindrical, with two distal bare setae. Enp and exp absent (fused to basis without trace).
</paragraph>
<paragraph id="902836A9CD6FFFA8FF407133FAEAC64F" blockId="12.[189,1399,776,837]" pageId="12" pageNumber="13">
<collectingCountry id="E8807639CD6FFFA8FF407133FF74C628" box="[189,233,776,802]" name="Mexico" pageId="12" pageNumber="13">MX</collectingCountry>
2 (
<figureCitation id="08AC2A2CCD6FFFA8FEF77133FEF9C628" box="[266,356,776,802]" captionStart="Fig" captionStartId="12.[189,232,1895,1921]" captionTargetBox="[211,1394,915,1855]" captionTargetId="figure@12.[189,1398,904,1877]" captionTargetPageId="12" captionText="Fig. 4. Cottarellicaris sanctiangeli Bruno &amp; Cottarelli sp. nov., ♂ (NHMUK). a. Rostrum and A1, ventral view. b. Rostrum and A1, ventral view. c. A1, disarticulated (antennular segments marked with Roman numerals). d. A2. e. Mdb. f. Mx1. g. Mx2. h. Mxp. a, eg: Grotta Vucco Ucciardo; bd, h: Grotta superiore di SantAngelo. Scale bar: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3961746" httpUri="https://zenodo.org/record/3961746/files/figure.png" pageId="12" pageNumber="13">
Fig.
<quantity id="576F9B4CCD6FFFA8FEBE7133FEF9C628" box="[323,356,776,802]" metricMagnitude="-3" metricUnit="kg" metricValue="4.0" pageId="12" pageNumber="13" unit="g" value="4.0">4g</quantity>
</figureCitation>
). Basis with two endites, proximal endite short, with one thin, bare seta; distal endite cylindrical, longer, armed apically with two subequal thin bare setae and one transformed, leaf-like
</paragraph>
<caption id="C4E86621CD6FFFA8FF40755CFD5FC2E1" ID-DOI="http://doi.org/10.5281/zenodo.3961746" ID-Zenodo-Dep="3961746" httpUri="https://zenodo.org/record/3961746/files/figure.png" pageId="12" pageNumber="13" startId="12.[189,232,1895,1921]" targetBox="[211,1394,915,1855]" targetPageId="12">
<paragraph id="902836A9CD6FFFA8FF40755CFD5FC2E1" blockId="12.[189,1398,1894,2027]" pageId="12" pageNumber="13">
<emphasis id="A2E3EABBCD6FFFA8FF40755CFE92C28B" bold="true" box="[189,271,1895,1921]" pageId="12" pageNumber="13">Fig. 4.</emphasis>
<taxonomicName id="57974D2ACD6FFFA8FEE7755DFCCFC28B" authority="Bruno &amp; Cottarelli" authorityName="Bruno &amp; Cottarelli &amp; Marrone &amp; Grasso &amp; Stefani &amp; Vecchioni &amp; Spena" authorityYear="2020" box="[282,850,1894,1921]" class="Maxillopoda" family="Parastenocarididae" genus="Cottarellicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="sanctiangeli" status="sp. nov.">
<emphasis id="A2E3EABBCD6FFFA8FEE7755DFDFDC28A" box="[282,608,1894,1920]" italics="true" pageId="12" pageNumber="13">Cottarellicaris sanctiangeli</emphasis>
Bruno &amp; Cottarelli
</taxonomicName>
<taxonomicNameLabel id="B9D057C0CD6FFFA8FCA0755CFC22C28B" box="[861,959,1895,1921]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
, ♂ (NHMUK).
<emphasis id="A2E3EABBCD6FFFA8FB78755CFB09C28B" bold="true" box="[1157,1172,1895,1921]" pageId="12" pageNumber="13">a</emphasis>
. Rostrum and A1, ventral view.
<emphasis id="A2E3EABBCD6FFFA8FEA075B1FEF0C2AE" bold="true" box="[349,365,1930,1956]" pageId="12" pageNumber="13">b</emphasis>
. Rostrum and A1, ventral view.
<emphasis id="A2E3EABBCD6FFFA8FD1175B1FD64C2AE" bold="true" box="[748,761,1930,1956]" pageId="12" pageNumber="13">c</emphasis>
. A1, disarticulated (antennular segments marked with Roman numerals).
<emphasis id="A2E3EABBCD6FFFA8FE667596FE36C2CD" bold="true" box="[411,427,1965,1991]" pageId="12" pageNumber="13">d</emphasis>
. A2.
<emphasis id="A2E3EABBCD6FFFA8FE157596FE68C2CD" bold="true" box="[488,501,1965,1991]" pageId="12" pageNumber="13">e</emphasis>
. Mdb.
<emphasis id="A2E3EABBCD6FFFA8FDBA7596FDCCC2CD" bold="true" box="[583,593,1965,1991]" pageId="12" pageNumber="13">f</emphasis>
. Mx1.
<emphasis id="A2E3EABBCD6FFFA8FD5E7596FD2FC2CD" bold="true" box="[675,690,1965,1991]" pageId="12" pageNumber="13">g</emphasis>
. Mx2.
<emphasis id="A2E3EABBCD6FFFA8FCF97596FC89C2CD" bold="true" box="[772,788,1965,1991]" pageId="12" pageNumber="13">h</emphasis>
. Mxp. a, eg: Grotta Vucco Ucciardo; bd, h: Grotta superiore di SantAngelo. Scale bar: 50 µm.
</paragraph>
</caption>
<paragraph id="902836A9CD6EFFA9FF407330FDFCC443" blockId="13.[189,1399,267,329]" pageId="13" pageNumber="14">pinnate seta; proximal endopodal segment drawn into apical unipinnate claw; distal endopodal segment with two long setae of equal length.</paragraph>
<paragraph id="902836A9CD6EFFA9FF40734AFC8EC4A4" blockId="13.[189,1398,368,430]" pageId="13" pageNumber="14">
MXP (
<figureCitation id="08AC2A2CCD6EFFA9FEF9734BFEC6C481" box="[260,347,368,395]" captionStart="Fig" captionStartId="12.[189,232,1895,1921]" captionTargetBox="[211,1394,915,1855]" captionTargetId="figure@12.[189,1398,904,1877]" captionTargetPageId="12" captionText="Fig. 4. Cottarellicaris sanctiangeli Bruno &amp; Cottarelli sp. nov., ♂ (NHMUK). a. Rostrum and A1, ventral view. b. Rostrum and A1, ventral view. c. A1, disarticulated (antennular segments marked with Roman numerals). d. A2. e. Mdb. f. Mx1. g. Mx2. h. Mxp. a, eg: Grotta Vucco Ucciardo; bd, h: Grotta superiore di SantAngelo. Scale bar: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3961746" httpUri="https://zenodo.org/record/3961746/files/figure.png" pageId="13" pageNumber="14">Fig. 4h</figureCitation>
). Subchelate, composed of small and unarmed syncoxa, basis slim and elongate, unarmed, 1-segmented enp fused to the claw-like apical seta.
</paragraph>
<paragraph id="902836A9CD6EFFA9FF4073EDFB5CC7ED" blockId="13.[189,1399,469,744]" pageId="13" pageNumber="14">
P1 (
<figureCitation id="08AC2A2CCD6EFFA9FF1073EEFEADC4FA" box="[237,304,469,496]" captionStart="Fig" captionStartId="14.[189,232,1862,1888]" captionTargetBox="[223,1358,296,1822]" captionTargetId="figure@14.[189,1398,294,1829]" captionTargetPageId="14" captionText="Fig. 5. Cottarellicaris sanctiangeli Bruno &amp; Cottarelli sp. nov., ♂ (NHMUK). a. P1, coxa, basis and enp, inner view. b. P1, basis and exp, outer view. c. P2, posterior view. d. P3, anterior view. e. P3, posterior view (variability). f. P4, anterior view. g. P4, intercoxal sclerite, coxa, basis and enp, outer view. h. P4, posterior view. a, c, e, g: Grotta Vucco Ucciardo; b, h: Grotta del Banco di ferro; d, f: Grotta superiore di SantAngelo. Scale bar: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3961748" httpUri="https://zenodo.org/record/3961748/files/figure.png" pageId="13" pageNumber="14">Fig. 5</figureCitation>
ab). With smooth and small intercoxal sclerite; coxa bare. Basis large, armed with small seta and lamellar hook with rounded tip on the inner margin near the enp insertion. Exp three-segmented, slightly shorter than endopod; exp-1 with thin pinnate seta on outer distal corner; exp-2 shortest and unarmed, ornamented with row of spinules along the subdistal and distal margins; exp-3 with two geniculate unipinnate apical setae and two subapical unipinnate setae. Enp two-segmented; enp-1 as long as first two segments of corresponding exp, with two transversal rows of few spinules on outer margin. Enp-2 thinner and shorter than enp-1, with two spinules at ¼ of inner margin and spinules along distal margin, with long, geniculate unipinnate seta and shorter unipinnate seta apically.
</paragraph>
<paragraph id="902836A9CD6EFFA9FF407134FDFEC12A" blockId="13.[189,1399,783,1057]" pageId="13" pageNumber="14">
P2 (
<figureCitation id="08AC2A2CCD6EFFA9FF137134FEDEC620" box="[238,323,783,810]" captionStart="Fig" captionStartId="14.[189,232,1862,1888]" captionTargetBox="[223,1358,296,1822]" captionTargetId="figure@14.[189,1398,294,1829]" captionTargetPageId="14" captionText="Fig. 5. Cottarellicaris sanctiangeli Bruno &amp; Cottarelli sp. nov., ♂ (NHMUK). a. P1, coxa, basis and enp, inner view. b. P1, basis and exp, outer view. c. P2, posterior view. d. P3, anterior view. e. P3, posterior view (variability). f. P4, anterior view. g. P4, intercoxal sclerite, coxa, basis and enp, outer view. h. P4, posterior view. a, c, e, g: Grotta Vucco Ucciardo; b, h: Grotta del Banco di ferro; d, f: Grotta superiore di SantAngelo. Scale bar: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3961748" httpUri="https://zenodo.org/record/3961748/files/figure.png" pageId="13" pageNumber="14">Fig. 5c</figureCitation>
). With smooth intercoxal sclerite, twice as wide as long, with concave distal margin. Coxa bare. Basis unarmed, with row of five spinules on outer margin. Exp three-segmented, exp-1 longest, with row of two spinules at ¼ of outer margin, longitudinal row of three spinules proximal to strong distolateral unipinnate spine. Second and third segments of same length; exp-2 unarmed, with row of spinules on distolateral corner, exp-3 armed with subapical outer unipinnate spine, apical bipinnate seta and outer unipinnate spine, ornamented with subapical spinule and inner hyaline frill. Enp onesegmented, about half length of exp-1, represented by cylindrical segment, with apical seta about as long as segment and three short spinules.
</paragraph>
<paragraph id="902836A9CD6EFFA9FF407673FDD3C03C" blockId="13.[189,1399,1096,1335]" pageId="13" pageNumber="14">
P3 (
<figureCitation id="08AC2A2CCD6EFFA9FF137673FED8C169" box="[238,325,1096,1123]" captionStart="Fig" captionStartId="14.[189,232,1862,1888]" captionTargetBox="[223,1358,296,1822]" captionTargetId="figure@14.[189,1398,294,1829]" captionTargetPageId="14" captionText="Fig. 5. Cottarellicaris sanctiangeli Bruno &amp; Cottarelli sp. nov., ♂ (NHMUK). a. P1, coxa, basis and enp, inner view. b. P1, basis and exp, outer view. c. P2, posterior view. d. P3, anterior view. e. P3, posterior view (variability). f. P4, anterior view. g. P4, intercoxal sclerite, coxa, basis and enp, outer view. h. P4, posterior view. a, c, e, g: Grotta Vucco Ucciardo; b, h: Grotta del Banco di ferro; d, f: Grotta superiore di SantAngelo. Scale bar: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3961748" httpUri="https://zenodo.org/record/3961748/files/figure.png" pageId="13" pageNumber="14">Fig. 5d</figureCitation>
). Intercoxal sclerite small, trapezoidal, unornamented, with slightly concave distal margin. Coxa bare. Basis robust, with long, slender, smooth outer seta and transverse spinule row above, and distal pore. Enp reduced to short seta. Exp-1 of characteristic shape: inner margin with large knob at ½ length and basal thin and pointed tubercle fused to exp; outer margin with proximal and distal rows of respectively one and four spinules. Exp-2 fused with exp-1, without ornamentation, prolonged into long apophysis slightly bent inwards, with pointed tip. Distal thumb represented by thin and pointed sigmoid segment, reaching past apophysis.
</paragraph>
<paragraph id="902836A9CD6EFFA9FF407765FF79C22B" blockId="13.[189,1399,1374,1825]" pageId="13" pageNumber="14">
P4 (
<figureCitation id="08AC2A2CCD6EFFA9FF117765FEB3C073" box="[236,302,1374,1401]" captionStart="Fig" captionStartId="14.[189,232,1862,1888]" captionTargetBox="[223,1358,296,1822]" captionTargetId="figure@14.[189,1398,294,1829]" captionTargetPageId="14" captionText="Fig. 5. Cottarellicaris sanctiangeli Bruno &amp; Cottarelli sp. nov., ♂ (NHMUK). a. P1, coxa, basis and enp, inner view. b. P1, basis and exp, outer view. c. P2, posterior view. d. P3, anterior view. e. P3, posterior view (variability). f. P4, anterior view. g. P4, intercoxal sclerite, coxa, basis and enp, outer view. h. P4, posterior view. a, c, e, g: Grotta Vucco Ucciardo; b, h: Grotta del Banco di ferro; d, f: Grotta superiore di SantAngelo. Scale bar: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3961748" httpUri="https://zenodo.org/record/3961748/files/figure.png" pageId="13" pageNumber="14">Fig. 5</figureCitation>
fh). Intercoxal sclerite smaller than in P1P3, with concave, smooth distal margin. Coxa bare. Basis armed with single slender seta on outer margin; ornamented with row of spinules at base of outer seta; row of three spiniform processes aligned along inner margin, of increasing size from outermost to innermost, all slightly curved inwards. Exp three-segmented, slender, all segments approximately of the same length; exp-1 slightly curved inwards, with distolateral pinnate spine, transversal spinular row at midlength of outer margin, spinular row at distolateral spine insertion, spinular row along distal margin; exp-2 unarmed, with spinular row along distal margin; exp-3 armed with outer unipinnate spine with spinules near insertion and very long apical pinnate seta, spine length less than ¼ of seta length; ornamentation represented by row of apical spinules and hyaline frill on inner distal corner. Enp onesegmented, as long as first two segments of corresponding exp, represented by curved plate with pointed inner tip carrying at outer border two outgrowths, distal one being elongate lamella, with undulating (crenulate) margins, reaching past end of enp, proximal outgrowth short curved lamella ending in thin tip.
</paragraph>
<paragraph id="902836A9CD6EFFA9FF407573FBC6C2A3" blockId="13.[189,1398,1864,1962]" pageId="13" pageNumber="14">
P5 (
<figureCitation id="08AC2A2CCD6EFFA9FF127573FEABC269" box="[239,310,1864,1891]" captionStart="Fig" captionStartId="15.[189,232,1825,1851]" captionTargetBox="[200,1378,266,1782]" captionTargetId="figure@15.[189,1398,237,1792]" captionTargetPageId="15" captionText="Fig. 6. Cottarellicaris sanctiangeli Bruno &amp; Cottarelli sp. nov. (NHMUK). a. ♂, P5, P6, first and second urosomites, ventral view. b. ♂, P5. c. ♂, P5 and first urosomite, ventral view. d. ♂, P5. e. ♀, first urosomite, P5, P6, genital double-somite and genital field, ventral view. f. ♀, anal somite, anal operculum and caudal rami, dorsal view. g. ♀, anal somite, anal operculum and caudal rami, lateral view. ab, e, g: Grotta Vucco Ucciardo; c: Grotta del Banco di ferro; d, f: Grotta superiore di SantAngelo. Scale bar: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3961750" httpUri="https://zenodo.org/record/3961750/files/figure.png" pageId="13" pageNumber="14">Fig. 6</figureCitation>
ad). Fused to intercoxal sclerite; represented by two trapezoidal cuticular plates with long basipodal seta and small pore near insertion. Armature on free distal margin, from inner to outer: one spiniform seta, three bare setae, outermost very short, remaining two subequal.
</paragraph>
<paragraph id="902836A9CD6EFFB4FF4075EAFE11C42F" blockId="13.[189,1186,2001,2028]" lastBlockId="16.[189,1399,267,658]" lastPageId="16" lastPageNumber="17" pageId="13" pageNumber="14">
P6 (
<figureCitation id="08AC2A2CCD6EFFA9FF1075EAFEDFC2E6" box="[237,322,2001,2028]" captionStart="Fig" captionStartId="15.[189,232,1825,1851]" captionTargetBox="[200,1378,266,1782]" captionTargetId="figure@15.[189,1398,237,1792]" captionTargetPageId="15" captionText="Fig. 6. Cottarellicaris sanctiangeli Bruno &amp; Cottarelli sp. nov. (NHMUK). a. ♂, P5, P6, first and second urosomites, ventral view. b. ♂, P5. c. ♂, P5 and first urosomite, ventral view. d. ♂, P5. e. ♀, first urosomite, P5, P6, genital double-somite and genital field, ventral view. f. ♀, anal somite, anal operculum and caudal rami, dorsal view. g. ♀, anal somite, anal operculum and caudal rami, lateral view. ab, e, g: Grotta Vucco Ucciardo; c: Grotta del Banco di ferro; d, f: Grotta superiore di SantAngelo. Scale bar: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3961750" httpUri="https://zenodo.org/record/3961750/files/figure.png" pageId="13" pageNumber="14">Fig. 6a</figureCitation>
). Vestigial, fused into simple cuticular plate, unornamented and unarmed.
<emphasis id="A2E3EABBCD73FFB4FF177330FE11C42F" bold="true" box="[234,396,267,293]" pageId="16" pageNumber="17">Adult female</emphasis>
</paragraph>
<caption id="C4E86621CD6DFFAAFF40757DFDD3C2E7" ID-DOI="http://doi.org/10.5281/zenodo.3961748" ID-Zenodo-Dep="3961748" httpUri="https://zenodo.org/record/3961748/files/figure.png" pageId="14" pageNumber="15" startId="14.[189,232,1862,1888]" targetBox="[223,1358,296,1822]" targetPageId="14">
<paragraph id="902836A9CD6DFFAAFF40757DFDD3C2E7" blockId="14.[189,1399,1862,2029]" pageId="14" pageNumber="15">
<emphasis id="A2E3EABBCD6DFFAAFF40757DFE94C26A" bold="true" box="[189,265,1862,1888]" pageId="14" pageNumber="15">Fig. 5.</emphasis>
<taxonomicName id="57974D2ACD6DFFAAFEF2757DFCADC26A" authority="Bruno &amp; Cottarelli" authorityName="Bruno &amp; Cottarelli &amp; Marrone &amp; Grasso &amp; Stefani &amp; Vecchioni &amp; Spena" authorityYear="2020" box="[271,816,1862,1888]" class="Maxillopoda" family="Parastenocarididae" genus="Cottarellicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="sanctiangeli" status="sp. nov.">
<emphasis id="A2E3EABBCD6DFFAAFEF2757DFDD2C26A" box="[271,591,1862,1888]" italics="true" pageId="14" pageNumber="15">Cottarellicaris sanctiangeli</emphasis>
Bruno &amp; Cottarelli
</taxonomicName>
<taxonomicNameLabel id="B9D057C0CD6DFFAAFCCB757DFC0FC26A" box="[822,914,1862,1888]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
, ♂ (NHMUK).
<emphasis id="A2E3EABBCD6DFFAAFBBA757DFBCBC26A" bold="true" box="[1095,1110,1862,1888]" pageId="14" pageNumber="15">a</emphasis>
. P1, coxa, basis and enp, inner view.
<emphasis id="A2E3EABBCD6DFFAAFEBA7552FECAC289" bold="true" box="[327,343,1897,1923]" pageId="14" pageNumber="15">b</emphasis>
. P1, basis and exp, outer view.
<emphasis id="A2E3EABBCD6DFFAAFD307552FD47C289" bold="true" box="[717,730,1897,1923]" pageId="14" pageNumber="15">c</emphasis>
. P2, posterior view.
<emphasis id="A2E3EABBCD6DFFAAFC307552FC40C289" bold="true" box="[973,989,1897,1923]" pageId="14" pageNumber="15">d</emphasis>
. P3, anterior view.
<emphasis id="A2E3EABBCD6DFFAAFB397552FB4CC289" bold="true" box="[1220,1233,1897,1923]" pageId="14" pageNumber="15">e</emphasis>
. P3, posterior view (variability).
<emphasis id="A2E3EABBCD6DFFAAFE6475B7FE3EC2AC" bold="true" box="[409,419,1932,1958]" pageId="14" pageNumber="15">f</emphasis>
. P4, anterior view.
<emphasis id="A2E3EABBCD6DFFAAFD7575B7FD0AC2AC" bold="true" box="[648,663,1932,1958]" pageId="14" pageNumber="15">g</emphasis>
. P4, intercoxal sclerite, coxa, basis and enp, outer view.
<emphasis id="A2E3EABBCD6DFFAAFACC75B7FADCC2AC" bold="true" box="[1329,1345,1932,1958]" pageId="14" pageNumber="15">h</emphasis>
. P4, posterior view. a, c, e, g: Grotta Vucco Ucciardo; b, h: Grotta del Banco di ferro; d, f: Grotta superiore di SantAngelo. Scale bar: 50 µm.
</paragraph>
</caption>
<caption id="C4E86621CD6CFFABFF40751AFE10C2E1" ID-DOI="http://doi.org/10.5281/zenodo.3961750" ID-Zenodo-Dep="3961750" httpUri="https://zenodo.org/record/3961750/files/figure.png" pageId="15" pageNumber="16" startId="15.[189,232,1825,1851]" targetBox="[200,1378,266,1782]" targetPageId="15">
<paragraph id="902836A9CD6CFFABFF40751AFE10C2E1" blockId="15.[189,1399,1824,2027]" pageId="15" pageNumber="16">
<emphasis id="A2E3EABBCD6CFFABFF40751AFE92C231" bold="true" box="[189,271,1825,1851]" pageId="15" pageNumber="16">Fig. 6.</emphasis>
<taxonomicName id="57974D2ACD6CFFABFEE6751AFCCBC231" authority="Bruno &amp; Cottarelli" authorityName="Bruno &amp; Cottarelli &amp; Marrone &amp; Grasso &amp; Stefani &amp; Vecchioni &amp; Spena" authorityYear="2020" box="[283,854,1825,1851]" class="Maxillopoda" family="Parastenocarididae" genus="Cottarellicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="sanctiangeli" status="sp. nov.">
<emphasis id="A2E3EABBCD6CFFABFEE6751AFDFFC231" box="[283,610,1825,1851]" italics="true" pageId="15" pageNumber="16">Cottarellicaris sanctiangeli</emphasis>
Bruno &amp; Cottarelli
</taxonomicName>
<taxonomicNameLabel id="B9D057C0CD6CFFABFC9C751AFC5CC231" box="[865,961,1825,1851]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
(NHMUK).
<emphasis id="A2E3EABBCD6CFFABFB9F751AFBECC231" bold="true" box="[1122,1137,1825,1851]" pageId="15" pageNumber="16">a</emphasis>
. ♂, P5, P6, first and second urosomites, ventral view.
<emphasis id="A2E3EABBCD6CFFABFDB1757FFDC1C254" bold="true" box="[588,604,1860,1886]" pageId="15" pageNumber="16">b</emphasis>
. ♂, P5.
<emphasis id="A2E3EABBCD6CFFABFD3E757FFD4DC254" bold="true" box="[707,720,1860,1886]" pageId="15" pageNumber="16">c</emphasis>
. ♂, P5 and first urosomite, ventral view.
<emphasis id="A2E3EABBCD6CFFABFB39757FFB49C254" bold="true" box="[1220,1236,1860,1886]" pageId="15" pageNumber="16">d</emphasis>
. ♂, P5.
<emphasis id="A2E3EABBCD6CFFABFAC6757FFAD5C254" bold="true" box="[1339,1352,1860,1886]" pageId="15" pageNumber="16">e</emphasis>
. ♀, first urosomite, P5, P6, genital double-somite and genital field, ventral view.
<emphasis id="A2E3EABBCD6CFFABFB9B755CFBEDC28B" bold="true" box="[1126,1136,1895,1921]" pageId="15" pageNumber="16">f</emphasis>
. ♀, anal somite, anal operculum and caudal rami, dorsal view.
<emphasis id="A2E3EABBCD6CFFABFD6575B0FD3AC2AF" bold="true" box="[664,679,1931,1957]" pageId="15" pageNumber="16">g</emphasis>
. ♀, anal somite, anal operculum and caudal rami, lateral view. ab, e, g: Grotta Vucco Ucciardo; c: Grotta del Banco di ferro; d, f: Grotta superiore di SantAngelo. Scale bar: 50 µm.
</paragraph>
</caption>
<paragraph id="902836A9CD73FFB4FF40730BFAF8C798" blockId="16.[189,1399,267,658]" pageId="16" pageNumber="17">
HABITUS (
<figureCitation id="08AC2A2CCD73FFB4FEC97314FE10C440" box="[308,397,303,330]" captionStart="Fig" captionStartId="10.[189,232,1966,1992]" captionTargetBox="[222,1398,508,1920]" captionTargetId="figure@10.[189,1398,496,1928]" captionTargetPageId="10" captionText="Fig. 2. Cottarellicaris sanctiangeli Bruno &amp; Cottarelli sp. nov. from Grotta superiore di SantAngelo (NHMUK). a. ♂, habitus. b. ♀, habitus. Scale bar: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3961742" httpUri="https://zenodo.org/record/3961742/files/figure.png" pageId="16" pageNumber="17">Fig. 2b</figureCitation>
). Cylindrical and slender, without any demarcation between prosome and urosome. Free pedigerous somites without any lateral or dorsal expansions, all connected by well-developed arthrodial membranes. Integument weakly sclerotized, without cuticular pits, ornamented with sensilla on all somites except preanal one. Cuticular windows on urosomites and cephalothorax not present. Body length, excluding caudal setae, from 280 to 312 μm, mean 304 μm (n = 6), ornamentation of cephalothorax, somites, pigmentation and absence of nauplius eye as in male, except genital and first urosomite fused into double-somite. Cephalotorax representing about 19% of the total body length. Genital double-somite (
<figureCitation id="08AC2A2CCD73FFB4FE2C7015FDB7C742" box="[465,554,558,584]" captionStart="Fig" captionStartId="15.[189,232,1825,1851]" captionTargetBox="[200,1378,266,1782]" captionTargetId="figure@15.[189,1398,237,1792]" captionTargetPageId="15" captionText="Fig. 6. Cottarellicaris sanctiangeli Bruno &amp; Cottarelli sp. nov. (NHMUK). a. ♂, P5, P6, first and second urosomites, ventral view. b. ♂, P5. c. ♂, P5 and first urosomite, ventral view. d. ♂, P5. e. ♀, first urosomite, P5, P6, genital double-somite and genital field, ventral view. f. ♀, anal somite, anal operculum and caudal rami, dorsal view. g. ♀, anal somite, anal operculum and caudal rami, lateral view. ab, e, g: Grotta Vucco Ucciardo; c: Grotta del Banco di ferro; d, f: Grotta superiore di SantAngelo. Scale bar: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3961750" httpUri="https://zenodo.org/record/3961750/files/figure.png" pageId="16" pageNumber="17">Figs 6e</figureCitation>
,
<figureCitation id="08AC2A2CCD73FFB4FDCA7015FDCEC742" box="[567,595,558,584]" captionStart="Fig" captionStartId="18.[188,231,1033,1059]" captionTargetBox="[342,1246,265,991]" captionTargetId="figure@18.[342,1246,265,991]" captionTargetPageId="18" captionText="Fig. 8. Cottarellicaris sanctiangeli Bruno &amp; Cottarelli sp. nov. (CIME). a. ♂, anal somite, anal operculum and caudal rami, ventro-lateral view. b. ♀, anal somite, anal operculum and caudal rami, lateral view. c. ♀, P5, P6, genital double-somite and genital field, ventral view (outermost seta, which is folded behind the P5 arrowed). d. ♀, enp P4. Grotta superiore di SantAngelo. Scale bar: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3961754" httpUri="https://zenodo.org/record/3961754/files/figure.png" pageId="16" pageNumber="17">8c</figureCitation>
) without any trace of subdivision. Genital field (
<figureCitation id="08AC2A2CCD73FFB4FB767015FB79C742" box="[1163,1252,558,584]" captionStart="Fig" captionStartId="15.[189,232,1825,1851]" captionTargetBox="[200,1378,266,1782]" captionTargetId="figure@15.[189,1398,237,1792]" captionTargetPageId="15" captionText="Fig. 6. Cottarellicaris sanctiangeli Bruno &amp; Cottarelli sp. nov. (NHMUK). a. ♂, P5, P6, first and second urosomites, ventral view. b. ♂, P5. c. ♂, P5 and first urosomite, ventral view. d. ♂, P5. e. ♀, first urosomite, P5, P6, genital double-somite and genital field, ventral view. f. ♀, anal somite, anal operculum and caudal rami, dorsal view. g. ♀, anal somite, anal operculum and caudal rami, lateral view. ab, e, g: Grotta Vucco Ucciardo; c: Grotta del Banco di ferro; d, f: Grotta superiore di SantAngelo. Scale bar: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3961750" httpUri="https://zenodo.org/record/3961750/files/figure.png" pageId="16" pageNumber="17">Figs 6e</figureCitation>
,
<figureCitation id="08AC2A2CCD73FFB4FB0D7015FA90C742" box="[1264,1293,558,584]" captionStart="Fig" captionStartId="18.[188,231,1033,1059]" captionTargetBox="[342,1246,265,991]" captionTargetId="figure@18.[342,1246,265,991]" captionTargetPageId="18" captionText="Fig. 8. Cottarellicaris sanctiangeli Bruno &amp; Cottarelli sp. nov. (CIME). a. ♂, anal somite, anal operculum and caudal rami, ventro-lateral view. b. ♀, anal somite, anal operculum and caudal rami, lateral view. c. ♀, P5, P6, genital double-somite and genital field, ventral view (outermost seta, which is folded behind the P5 arrowed). d. ♀, enp P4. Grotta superiore di SantAngelo. Scale bar: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3961754" httpUri="https://zenodo.org/record/3961754/files/figure.png" pageId="16" pageNumber="17">8c</figureCitation>
) broader than tall, occupying anterior ventral ¼ of genital double-somite; single genital aperture covered by fused vestigial sixth legs; median copulatory pore located medially at ¼ of somite. Anal operculum convex.
</paragraph>
<paragraph id="902836A9CD73FFB4FF407081FEE4C7F3" blockId="16.[189,1399,698,761]" pageId="16" pageNumber="17">
CAUDAL RAMI (
<figureCitation id="08AC2A2CCD73FFB4FE897081FE22C7DE" box="[372,447,698,724]" captionStart="Fig" captionStartId="15.[189,232,1825,1851]" captionTargetBox="[200,1378,266,1782]" captionTargetId="figure@15.[189,1398,237,1792]" captionTargetPageId="15" captionText="Fig. 6. Cottarellicaris sanctiangeli Bruno &amp; Cottarelli sp. nov. (NHMUK). a. ♂, P5, P6, first and second urosomites, ventral view. b. ♂, P5. c. ♂, P5 and first urosomite, ventral view. d. ♂, P5. e. ♀, first urosomite, P5, P6, genital double-somite and genital field, ventral view. f. ♀, anal somite, anal operculum and caudal rami, dorsal view. g. ♀, anal somite, anal operculum and caudal rami, lateral view. ab, e, g: Grotta Vucco Ucciardo; c: Grotta del Banco di ferro; d, f: Grotta superiore di SantAngelo. Scale bar: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3961750" httpUri="https://zenodo.org/record/3961750/files/figure.png" pageId="16" pageNumber="17">Figs 6</figureCitation>
fg, 8b). Shape, ornamentation and armature similar to those of male, length to width ratio: 3.5.
</paragraph>
<paragraph id="902836A9CD73FFB4FF40711AFC9BC631" blockId="16.[189,774,801,827]" box="[189,774,801,827]" pageId="16" pageNumber="17">ROSTRUM, A2 AND ORAL APPENDAGES. As in male.</paragraph>
<paragraph id="902836A9CD73FFB4FF40715FFDABC6E1" blockId="16.[189,1399,867,1003]" pageId="16" pageNumber="17">
A1 (
<figureCitation id="08AC2A2CCD73FFB4FF0F7158FEDBC674" box="[242,326,867,894]" captionStart="Fig" captionStartId="17.[189,232,1859,1885]" captionTargetBox="[214,1388,351,1791]" captionTargetId="figure@17.[189,1398,338,1818]" captionTargetPageId="17" captionText="Fig. 7. Cottarellicaris sanctiangeli Bruno &amp; Cottarelli sp. nov., ♀ (NHMUK). a. Anal somite, anal operculum and caudal rami, dorsal view (variability). b. Anal somite, anal operculum and caudal rami, lateral view (variability). c. Rostrum, A1. d. P1, anterior view. e. P2, inner view. f. P3, posterior view. g. Coxa, basis, enp P3, inner view. h. P4, posterior view. i. Coxa, basis, enp P4, inner view (variability). a, ci: Grotta Grotta Vucco Ucciardo; b: Grotta del Banco di ferro. Scale bar: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3961752" httpUri="https://zenodo.org/record/3961752/files/figure.png" pageId="16" pageNumber="17">Fig. 7c</figureCitation>
). Seven-segmented, aesthetasc on fourth segment longer than in male, reaching beyond end of seventh segment. First segment bare. Apical acrothek represented by two setae of different length and slender aesthetasc. Armature formula: 1-[0], 2-[1 unipinnate + 4 bare], 3-[4 bare], 4-[2 bare + ae], 5-[0], 6-[0], 7-[7 bare + (2 bare + ae)].
</paragraph>
<paragraph id="902836A9CD73FFB4FF40762FFCDEC158" blockId="16.[189,1398,1043,1106]" pageId="16" pageNumber="17">
P1 (
<figureCitation id="08AC2A2CCD73FFB4FF137628FED8C124" box="[238,325,1043,1070]" captionStart="Fig" captionStartId="17.[189,232,1859,1885]" captionTargetBox="[214,1388,351,1791]" captionTargetId="figure@17.[189,1398,338,1818]" captionTargetPageId="17" captionText="Fig. 7. Cottarellicaris sanctiangeli Bruno &amp; Cottarelli sp. nov., ♀ (NHMUK). a. Anal somite, anal operculum and caudal rami, dorsal view (variability). b. Anal somite, anal operculum and caudal rami, lateral view (variability). c. Rostrum, A1. d. P1, anterior view. e. P2, inner view. f. P3, posterior view. g. Coxa, basis, enp P3, inner view. h. P4, posterior view. i. Coxa, basis, enp P4, inner view (variability). a, ci: Grotta Grotta Vucco Ucciardo; b: Grotta del Banco di ferro. Scale bar: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3961752" httpUri="https://zenodo.org/record/3961752/files/figure.png" pageId="16" pageNumber="17">Fig. 7d</figureCitation>
). Intercoxal sclerite, coxa, basis as in male but lamellar hook missing; exp and enp similar to those of male in shape, ornamentation and armature.
</paragraph>
<paragraph id="902836A9CD73FFB4FF407640FCE5C1D7" blockId="16.[189,1399,1146,1246]" pageId="16" pageNumber="17">
P2 (
<figureCitation id="08AC2A2CCD73FFB4FF117641FEA2C19F" box="[236,319,1146,1173]" captionStart="Fig" captionStartId="17.[189,232,1859,1885]" captionTargetBox="[214,1388,351,1791]" captionTargetId="figure@17.[189,1398,338,1818]" captionTargetPageId="17" captionText="Fig. 7. Cottarellicaris sanctiangeli Bruno &amp; Cottarelli sp. nov., ♀ (NHMUK). a. Anal somite, anal operculum and caudal rami, dorsal view (variability). b. Anal somite, anal operculum and caudal rami, lateral view (variability). c. Rostrum, A1. d. P1, anterior view. e. P2, inner view. f. P3, posterior view. g. Coxa, basis, enp P3, inner view. h. P4, posterior view. i. Coxa, basis, enp P4, inner view (variability). a, ci: Grotta Grotta Vucco Ucciardo; b: Grotta del Banco di ferro. Scale bar: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3961752" httpUri="https://zenodo.org/record/3961752/files/figure.png" pageId="16" pageNumber="17">Fig. 7e</figureCitation>
). Intercoxal sclerite longer and narrower than in male, coxa narrower than in male, basis as in male, but with spinular row below enp insertion. Exp as in male. Enp similar in shape and ornamentation to that of male, but with two apical spinule instead of three.
</paragraph>
<paragraph id="902836A9CD73FFB4FF40773DFD1DC0DC" blockId="16.[189,1399,1286,1494]" pageId="16" pageNumber="17">
P3 (
<figureCitation id="08AC2A2CCD73FFB4FF11773DFEB3C02B" box="[236,302,1286,1313]" captionStart="Fig" captionStartId="17.[189,232,1859,1885]" captionTargetBox="[214,1388,351,1791]" captionTargetId="figure@17.[189,1398,338,1818]" captionTargetPageId="17" captionText="Fig. 7. Cottarellicaris sanctiangeli Bruno &amp; Cottarelli sp. nov., ♀ (NHMUK). a. Anal somite, anal operculum and caudal rami, dorsal view (variability). b. Anal somite, anal operculum and caudal rami, lateral view (variability). c. Rostrum, A1. d. P1, anterior view. e. P2, inner view. f. P3, posterior view. g. Coxa, basis, enp P3, inner view. h. P4, posterior view. i. Coxa, basis, enp P4, inner view (variability). a, ci: Grotta Grotta Vucco Ucciardo; b: Grotta del Banco di ferro. Scale bar: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3961752" httpUri="https://zenodo.org/record/3961752/files/figure.png" pageId="16" pageNumber="17">Fig. 7</figureCitation>
fg). Intercoxal sclerite small, with concave margin, bare. Coxa bare. Basis with outer seta and spinular row near seta insertion. Exp two-segmented: exp-1 as long as exp-2, with distolateral curved unipinnate spine, transversal row of spinules at ¼ and ¼ of outer margin, hyaline frill on inner distal corner; exp-2 with outer unipinnate spine and apical pinnate seta, spine length about ¼ of seta, hyaline frill on inner distal corner. Enp represented by thin and pointed segment, slightly shorter than half of corresponding exp-1, apically pinnate.
</paragraph>
<paragraph id="902836A9CD73FFB4FF4077C4FB77C334" blockId="16.[189,1399,1535,1598]" pageId="16" pageNumber="17">
P4 (
<figureCitation id="08AC2A2CCD73FFB4FF0C77C4FED0C313" box="[241,333,1535,1561]" captionStart="Fig" captionStartId="17.[189,232,1859,1885]" captionTargetBox="[214,1388,351,1791]" captionTargetId="figure@17.[189,1398,338,1818]" captionTargetPageId="17" captionText="Fig. 7. Cottarellicaris sanctiangeli Bruno &amp; Cottarelli sp. nov., ♀ (NHMUK). a. Anal somite, anal operculum and caudal rami, dorsal view (variability). b. Anal somite, anal operculum and caudal rami, lateral view (variability). c. Rostrum, A1. d. P1, anterior view. e. P2, inner view. f. P3, posterior view. g. Coxa, basis, enp P3, inner view. h. P4, posterior view. i. Coxa, basis, enp P4, inner view (variability). a, ci: Grotta Grotta Vucco Ucciardo; b: Grotta del Banco di ferro. Scale bar: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3961752" httpUri="https://zenodo.org/record/3961752/files/figure.png" pageId="16" pageNumber="17">Figs 7h</figureCitation>
,
<figureCitation id="08AC2A2CCD73FFB4FEA277C4FEE0C313" box="[351,381,1535,1561]" captionStart="Fig" captionStartId="18.[188,231,1033,1059]" captionTargetBox="[342,1246,265,991]" captionTargetId="figure@18.[342,1246,265,991]" captionTargetPageId="18" captionText="Fig. 8. Cottarellicaris sanctiangeli Bruno &amp; Cottarelli sp. nov. (CIME). a. ♂, anal somite, anal operculum and caudal rami, ventro-lateral view. b. ♀, anal somite, anal operculum and caudal rami, lateral view. c. ♀, P5, P6, genital double-somite and genital field, ventral view (outermost seta, which is folded behind the P5 arrowed). d. ♀, enp P4. Grotta superiore di SantAngelo. Scale bar: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3961754" httpUri="https://zenodo.org/record/3961754/files/figure.png" pageId="16" pageNumber="17">8d</figureCitation>
). Intercoxal sclerite, coxa and exp as in male. Basis bare. Enp represented by thin cylindrical segment, about ¼ length of corresponding exp-1, ending in short bipinnate seta.
</paragraph>
<paragraph id="902836A9CD73FFB4FF40745DFD1BC3AF" blockId="16.[189,1398,1638,1701]" pageId="16" pageNumber="17">
P5 (
<figureCitation id="08AC2A2CCD73FFB4FF13745DFEDEC38B" box="[238,323,1638,1665]" captionStart="Fig" captionStartId="15.[189,232,1825,1851]" captionTargetBox="[200,1378,266,1782]" captionTargetId="figure@15.[189,1398,237,1792]" captionTargetPageId="15" captionText="Fig. 6. Cottarellicaris sanctiangeli Bruno &amp; Cottarelli sp. nov. (NHMUK). a. ♂, P5, P6, first and second urosomites, ventral view. b. ♂, P5. c. ♂, P5 and first urosomite, ventral view. d. ♂, P5. e. ♀, first urosomite, P5, P6, genital double-somite and genital field, ventral view. f. ♀, anal somite, anal operculum and caudal rami, dorsal view. g. ♀, anal somite, anal operculum and caudal rami, lateral view. ab, e, g: Grotta Vucco Ucciardo; c: Grotta del Banco di ferro; d, f: Grotta superiore di SantAngelo. Scale bar: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3961750" httpUri="https://zenodo.org/record/3961750/files/figure.png" pageId="16" pageNumber="17">Fig. 6e</figureCitation>
). Fused to intercoxal sclerite, represented by two cuticular plates shorter than in male, with same armature and without distal pore.
</paragraph>
<paragraph id="902836A9CD73FFB4FF4074F6FAEAC3ED" blockId="16.[189,1399,1741,1768]" box="[189,1399,1741,1768]" pageId="16" pageNumber="17">
P6 (
<figureCitation id="08AC2A2CCD73FFB4FF1674F6FEA2C3E2" box="[235,319,1741,1768]" captionStart="Fig" captionStartId="15.[189,232,1825,1851]" captionTargetBox="[200,1378,266,1782]" captionTargetId="figure@15.[189,1398,237,1792]" captionTargetPageId="15" captionText="Fig. 6. Cottarellicaris sanctiangeli Bruno &amp; Cottarelli sp. nov. (NHMUK). a. ♂, P5, P6, first and second urosomites, ventral view. b. ♂, P5. c. ♂, P5 and first urosomite, ventral view. d. ♂, P5. e. ♀, first urosomite, P5, P6, genital double-somite and genital field, ventral view. f. ♀, anal somite, anal operculum and caudal rami, dorsal view. g. ♀, anal somite, anal operculum and caudal rami, lateral view. ab, e, g: Grotta Vucco Ucciardo; c: Grotta del Banco di ferro; d, f: Grotta superiore di SantAngelo. Scale bar: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3961750" httpUri="https://zenodo.org/record/3961750/files/figure.png" pageId="16" pageNumber="17">Fig. 6e</figureCitation>
). Vestigial, fused into simple cuticular plate, covering gonopore, unornamented and unarmed.
</paragraph>
<paragraph id="902836A9CD73FFB4FF407534FEDEC223" blockId="16.[189,323,1807,1833]" box="[189,323,1807,1833]" pageId="16" pageNumber="17">
<heading id="CB6081C5CD73FFB4FF407534FEDEC223" bold="true" box="[189,323,1807,1833]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="A2E3EABBCD73FFB4FF407534FEDEC223" bold="true" box="[189,323,1807,1833]" pageId="16" pageNumber="17">Variability</emphasis>
</heading>
</paragraph>
<paragraph id="902836A9CD73FFB4FF407504FD4AC2E6" blockId="16.[189,1399,1855,2028]" pageId="16" pageNumber="17">
The ornamentation of male P3 exp-1 can vary, as one specimen from Vucco Ucciardo has the proximal and distal rows of respectively two and three spinules (
<figureCitation id="08AC2A2CCD73FFB4FCB9755FFC07C274" box="[836,922,1891,1918]" captionStart="Fig" captionStartId="14.[189,232,1862,1888]" captionTargetBox="[223,1358,296,1822]" captionTargetId="figure@14.[189,1398,294,1829]" captionTargetPageId="14" captionText="Fig. 5. Cottarellicaris sanctiangeli Bruno &amp; Cottarelli sp. nov., ♂ (NHMUK). a. P1, coxa, basis and enp, inner view. b. P1, basis and exp, outer view. c. P2, posterior view. d. P3, anterior view. e. P3, posterior view (variability). f. P4, anterior view. g. P4, intercoxal sclerite, coxa, basis and enp, outer view. h. P4, posterior view. a, c, e, g: Grotta Vucco Ucciardo; b, h: Grotta del Banco di ferro; d, f: Grotta superiore di SantAngelo. Scale bar: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3961748" httpUri="https://zenodo.org/record/3961748/files/figure.png" pageId="16" pageNumber="17">Fig. 5e</figureCitation>
). Two males (
<figureCitation id="08AC2A2CCD73FFB4FBC2755FFB2EC275" box="[1087,1203,1892,1919]" captionStart="Fig" captionStartId="11.[189,232,1825,1851]" captionTargetBox="[195,1384,351,1760]" captionTargetId="figure@11.[189,1398,340,1777]" captionTargetPageId="11" captionText="Fig. 3. Cottarellicaris sanctiangeli Bruno &amp; Cottarelli sp. nov., ♂ (NHMUK). a. Anal somite, anal operculum and caudal rami, dorsal view. b. Anal somite, anal operculum and caudal rami, dorsal view (variability). c. Anal somite, anal operculum and caudal rami, dorsal view. d. Anal somite, anal operculum and caudal rami, lateral view (variability). e. Anal somite, anal operculum and caudal rami, lateral view (variability). a: Grotta superiore di SantAngelo; b, d: Grotta del Banco di ferro; c, e: Grotta Vucco Ucciardo. Scale bar: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3961744" httpUri="https://zenodo.org/record/3961744/files/figure.png" pageId="16" pageNumber="17">Fig. 3b, d</figureCitation>
) and one female (
<figureCitation id="08AC2A2CCD73FFB4FF3B75B3FE82C2A9" box="[198,287,1928,1955]" captionStart="Fig" captionStartId="17.[189,232,1859,1885]" captionTargetBox="[214,1388,351,1791]" captionTargetId="figure@17.[189,1398,338,1818]" captionTargetPageId="17" captionText="Fig. 7. Cottarellicaris sanctiangeli Bruno &amp; Cottarelli sp. nov., ♀ (NHMUK). a. Anal somite, anal operculum and caudal rami, dorsal view (variability). b. Anal somite, anal operculum and caudal rami, lateral view (variability). c. Rostrum, A1. d. P1, anterior view. e. P2, inner view. f. P3, posterior view. g. Coxa, basis, enp P3, inner view. h. P4, posterior view. i. Coxa, basis, enp P4, inner view (variability). a, ci: Grotta Grotta Vucco Ucciardo; b: Grotta del Banco di ferro. Scale bar: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3961752" httpUri="https://zenodo.org/record/3961752/files/figure.png" pageId="16" pageNumber="17">Fig. 7b</figureCitation>
) from Grotta del Banco di ferro, one male (
<figureCitation id="08AC2A2CCD73FFB4FCBF75B3FC04C2A8" box="[834,921,1928,1954]" captionStart="Fig" captionStartId="11.[189,232,1825,1851]" captionTargetBox="[195,1384,351,1760]" captionTargetId="figure@11.[189,1398,340,1777]" captionTargetPageId="11" captionText="Fig. 3. Cottarellicaris sanctiangeli Bruno &amp; Cottarelli sp. nov., ♂ (NHMUK). a. Anal somite, anal operculum and caudal rami, dorsal view. b. Anal somite, anal operculum and caudal rami, dorsal view (variability). c. Anal somite, anal operculum and caudal rami, dorsal view. d. Anal somite, anal operculum and caudal rami, lateral view (variability). e. Anal somite, anal operculum and caudal rami, lateral view (variability). a: Grotta superiore di SantAngelo; b, d: Grotta del Banco di ferro; c, e: Grotta Vucco Ucciardo. Scale bar: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3961744" httpUri="https://zenodo.org/record/3961744/files/figure.png" pageId="16" pageNumber="17">Fig. 3e</figureCitation>
) and one female (
<figureCitation id="08AC2A2CCD73FFB4FB8075B3FB49C2A9" box="[1149,1236,1928,1955]" captionStart="Fig" captionStartId="17.[189,232,1859,1885]" captionTargetBox="[214,1388,351,1791]" captionTargetId="figure@17.[189,1398,338,1818]" captionTargetPageId="17" captionText="Fig. 7. Cottarellicaris sanctiangeli Bruno &amp; Cottarelli sp. nov., ♀ (NHMUK). a. Anal somite, anal operculum and caudal rami, dorsal view (variability). b. Anal somite, anal operculum and caudal rami, lateral view (variability). c. Rostrum, A1. d. P1, anterior view. e. P2, inner view. f. P3, posterior view. g. Coxa, basis, enp P3, inner view. h. P4, posterior view. i. Coxa, basis, enp P4, inner view (variability). a, ci: Grotta Grotta Vucco Ucciardo; b: Grotta del Banco di ferro. Scale bar: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3961752" httpUri="https://zenodo.org/record/3961752/files/figure.png" pageId="16" pageNumber="17">Fig. 7a</figureCitation>
) from Vucco Ucciardo carry seven setae on the caudal rami instead then 6. In one female from Vucco Ucciardo the enp P4 is fused with the apical spine (
<figureCitation id="08AC2A2CCD73FFB4FD8475EAFD54C2E6" box="[633,713,2001,2028]" captionStart="Fig" captionStartId="17.[189,232,1859,1885]" captionTargetBox="[214,1388,351,1791]" captionTargetId="figure@17.[189,1398,338,1818]" captionTargetPageId="17" captionText="Fig. 7. Cottarellicaris sanctiangeli Bruno &amp; Cottarelli sp. nov., ♀ (NHMUK). a. Anal somite, anal operculum and caudal rami, dorsal view (variability). b. Anal somite, anal operculum and caudal rami, lateral view (variability). c. Rostrum, A1. d. P1, anterior view. e. P2, inner view. f. P3, posterior view. g. Coxa, basis, enp P3, inner view. h. P4, posterior view. i. Coxa, basis, enp P4, inner view (variability). a, ci: Grotta Grotta Vucco Ucciardo; b: Grotta del Banco di ferro. Scale bar: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3961752" httpUri="https://zenodo.org/record/3961752/files/figure.png" pageId="16" pageNumber="17">Fig. 7i</figureCitation>
).
</paragraph>
<caption id="C4E86621CD72FFB5FF407578FB18C2E1" ID-DOI="http://doi.org/10.5281/zenodo.3961752" ID-Zenodo-Dep="3961752" httpUri="https://zenodo.org/record/3961752/files/figure.png" pageId="17" pageNumber="18" startId="17.[189,232,1859,1885]" targetBox="[214,1388,351,1791]" targetPageId="17">
<paragraph id="902836A9CD72FFB5FF407578FB18C2E1" blockId="17.[189,1399,1859,2027]" pageId="17" pageNumber="18">
<emphasis id="A2E3EABBCD72FFB5FF407578FE93C254" bold="true" box="[189,270,1859,1886]" pageId="17" pageNumber="18">Fig. 7.</emphasis>
<taxonomicName id="57974D2ACD72FFB5FEE47578FCCDC257" authority="Bruno &amp; Cottarelli" authorityName="Bruno &amp; Cottarelli &amp; Marrone &amp; Grasso &amp; Stefani &amp; Vecchioni &amp; Spena" authorityYear="2020" box="[281,848,1859,1886]" class="Maxillopoda" family="Parastenocarididae" genus="Cottarellicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="sanctiangeli" status="sp. nov.">
<emphasis id="A2E3EABBCD72FFB5FEE47578FDC2C257" box="[281,607,1859,1885]" italics="true" pageId="17" pageNumber="18">Cottarellicaris sanctiangeli</emphasis>
Bruno &amp; Cottarelli
</taxonomicName>
<taxonomicNameLabel id="B9D057C0CD72FFB5FCA6757FFC21C254" box="[859,956,1860,1886]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
, ♀ (NHMUK).
<emphasis id="A2E3EABBCD72FFB5FB7C757FFB0DC254" bold="true" box="[1153,1168,1860,1886]" pageId="17" pageNumber="18">a</emphasis>
. Anal somite, anal operculum and caudal rami, dorsal view (variability).
<emphasis id="A2E3EABBCD72FFB5FCC4755CFCD4C28B" bold="true" box="[825,841,1895,1921]" pageId="17" pageNumber="18">b</emphasis>
. Anal somite, anal operculum and caudal rami, lateral view (variability).
<emphasis id="A2E3EABBCD72FFB5FE1A75B1FE69C2AE" bold="true" box="[487,500,1930,1956]" pageId="17" pageNumber="18">c</emphasis>
. Rostrum, A1.
<emphasis id="A2E3EABBCD72FFB5FD5875B1FD28C2AE" bold="true" box="[677,693,1930,1956]" pageId="17" pageNumber="18">d</emphasis>
. P1, anterior view.
<emphasis id="A2E3EABBCD72FFB5FC6A75B1FC39C2AE" bold="true" box="[919,932,1930,1956]" pageId="17" pageNumber="18">e</emphasis>
. P2, inner view.
<emphasis id="A2E3EABBCD72FFB5FB9575B1FBEFC2AE" bold="true" box="[1128,1138,1930,1956]" pageId="17" pageNumber="18">f</emphasis>
. P3, posterior view.
<emphasis id="A2E3EABBCD72FFB5FA9D75B1FAF2C2AE" bold="true" box="[1376,1391,1930,1956]" pageId="17" pageNumber="18">g</emphasis>
. Coxa, basis, enp P3, inner view.
<emphasis id="A2E3EABBCD72FFB5FDC77596FDD7C2CD" bold="true" box="[570,586,1965,1991]" pageId="17" pageNumber="18">h</emphasis>
. P4, posterior view.
<emphasis id="A2E3EABBCD72FFB5FCC77596FCDFC2CD" bold="true" box="[826,834,1965,1991]" pageId="17" pageNumber="18">i</emphasis>
. Coxa, basis, enp P4, inner view (variability). a, ci: Grotta Grotta Vucco Ucciardo; b: Grotta del Banco di ferro. Scale bar: 50 µm.
</paragraph>
</caption>
<caption id="C4E86621CD71FFB6FF417632FB45C184" ID-DOI="http://doi.org/10.5281/zenodo.3961754" ID-Zenodo-Dep="3961754" httpUri="https://zenodo.org/record/3961754/files/figure.png" pageId="18" pageNumber="19" startId="18.[188,231,1033,1059]" targetBox="[342,1246,265,991]" targetPageId="18">
<paragraph id="902836A9CD71FFB6FF417632FB45C184" blockId="18.[188,1398,1033,1166]" pageId="18" pageNumber="19">
<emphasis id="A2E3EABBCD71FFB6FF417632FE9AC129" bold="true" box="[188,263,1033,1059]" pageId="18" pageNumber="19">Fig. 8.</emphasis>
<taxonomicName id="57974D2ACD71FFB6FEF17632FCB7C129" authority="Bruno &amp; Cottarelli" authorityName="Bruno &amp; Cottarelli &amp; Marrone &amp; Grasso &amp; Stefani &amp; Vecchioni &amp; Spena" authorityYear="2020" box="[268,810,1033,1060]" class="Maxillopoda" family="Parastenocarididae" genus="Cottarellicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="sanctiangeli" status="sp. nov.">
<emphasis id="A2E3EABBCD71FFB6FEF17632FDD1C129" box="[268,588,1033,1059]" italics="true" pageId="18" pageNumber="19">Cottarellicaris sanctiangeli</emphasis>
Bruno &amp; Cottarelli
</taxonomicName>
<taxonomicNameLabel id="B9D057C0CD71FFB6FCD27631FC1AC12E" box="[815,903,1034,1060]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
(CIME).
<emphasis id="A2E3EABBCD71FFB6FC087631FB99C12E" bold="true" box="[1013,1028,1034,1060]" pageId="18" pageNumber="19">a</emphasis>
. ♂, anal somite, anal operculum and caudal rami, ventro-lateral view.
<emphasis id="A2E3EABBCD71FFB6FD847616FD14C14D" bold="true" box="[633,649,1069,1095]" pageId="18" pageNumber="19">b</emphasis>
. ♀, anal somite, anal operculum and caudal rami, lateral view.
<emphasis id="A2E3EABBCD71FFB6FF41766BFF54C160" bold="true" box="[188,201,1104,1130]" pageId="18" pageNumber="19">c</emphasis>
. ♀, P5, P6, genital double-somite and genital field, ventral view (outermost seta, which is folded behind the P5 arrowed).
<emphasis id="A2E3EABBCD71FFB6FE237648FE73C187" bold="true" box="[478,494,1139,1165]" pageId="18" pageNumber="19">d</emphasis>
. ♀, enp P4. Grotta superiore di SantAngelo. Scale bar: 50 µm.
</paragraph>
</caption>
<paragraph id="902836A9CD71FFB6FF4076D1FE55C00E" blockId="18.[189,456,1258,1284]" box="[189,456,1258,1284]" pageId="18" pageNumber="19">
<heading id="CB6081C5CD71FFB6FF4076D1FE55C00E" bold="true" box="[189,456,1258,1284]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="A2E3EABBCD71FFB6FF4076D1FE55C00E" bold="true" box="[189,456,1258,1284]" pageId="18" pageNumber="19">Accompanying fauna</emphasis>
</heading>
</paragraph>
<paragraph id="902836A9CD71FFB6FF407722FA99C039" blockId="18.[189,1284,1305,1332]" box="[189,1284,1305,1332]" pageId="18" pageNumber="19">
Grotta del Banco di ferro,
<date id="E4291069CD71FFB6FE0E7722FD0EC039" box="[499,659,1305,1332]" pageId="18" pageNumber="19" value="2017-05-10">10 May 2017</date>
:
<taxonomicName id="57974D2ACD71FFB6FD637722FCCCC039" box="[670,849,1305,1331]" class="Maxillopoda" family="Canthocamptidae" genus="Maraenobiotus" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="A2E3EABBCD71FFB6FD637722FCCCC039" box="[670,849,1305,1331]" italics="true" pageId="18" pageNumber="19">Maraenobiotus</emphasis>
</taxonomicName>
sp.: pool 4;
<taxonomicName id="57974D2ACD71FFB6FC197722FBE1C039" box="[996,1148,1305,1331]" class="Maxillopoda" family="Canthocamptidae" genus="Elaphoidella" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="A2E3EABBCD71FFB6FC197722FBE1C039" box="[996,1148,1305,1331]" italics="true" pageId="18" pageNumber="19">Elaphoidella</emphasis>
</taxonomicName>
sp.: pool 2.
</paragraph>
</subSubSection>
</treatment>
</document>