557 lines
66 KiB
XML
557 lines
66 KiB
XML
<document id="594BE8C635965B668D46E5C4E5738D53" ID-DOI="10.5281/zenodo.213061" ID-GBIF-Dataset="948bdc05-d503-4677-8f6a-a3cea7b3193a" ID-ISSN="1175-5326" ID-Zenodo-Dep="213061" ID-ZooBank="DC681CEB-48B5-42E0-8C7C-D7A152DAC53A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460535248693" checkinUser="plazi" docAuthor="Kwun, Hyuck Joon & Kim, Jin Koo" docDate="2012" docId="182887C6FF9B9B79C994FF62F90EE7DF" docLanguage="en" docName="zt03493p034.pdf" docOrigin="Zootaxa 3493" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Eulophias koreanus Kwun & Kim, 2012, new species" docType="treatment" docVersion="10" lastPageNumber="32" masterDocId="E411FFBEFF999B7CC903FFF5FB41E045" masterDocTitle="A New Species of the Genus Eulophias (Zoarcoidei: Stichaeidae) from Korea" masterLastPageNumber="34" masterPageNumber="27" pageNumber="29" updateTime="1698316225394" updateUser="plazi">
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<mods:titleInfo id="CE3BCB3D1B2B85DFCD9BB4854C25FCA5">
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<mods:title id="6606E74512502A68F45330B0732B1E97">A New Species of the Genus Eulophias (Zoarcoidei: Stichaeidae) from Korea</mods:title>
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<mods:name id="DEEF96ED699FF45E96214C8F9AE0B8A5" type="personal">
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<mods:roleTerm id="1BD948C48574F7905EADFB46967A2F7C">Author</mods:roleTerm>
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<mods:namePart id="C6677D2C66CFFE90472C9D77AC068674">Kwun, Hyuck Joon</mods:namePart>
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<mods:name id="1E6BE926E2D94B1E07D6A6AB5769133E" type="personal">
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<mods:roleTerm id="949853330415933D6611F879C1984580">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="581A7CC529BFEC61ABC6B9B53930C82D">Kim, Jin Koo</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="7BB79AD32965A0A81D6CD049D0EC0B8E">text</mods:typeOfResource>
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<mods:title id="CEBC256CA8FC39E188ECAA61145708D4">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="F35EF1EE987B79503818C3A6CA645881">
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<mods:date id="795FBBDD11E80B461AB26F78F83E82FD">2012</mods:date>
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<mods:detail id="37B5BE361FC755A4D7120538DAA3BF67" type="volume">
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<mods:number id="B1D9BC0DF1077617512DF74A30C3F542">3493</mods:number>
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<mods:start id="516770E4BDF19672486D87A39DA10090">27</mods:start>
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<mods:end id="20896BD62E83A2F8DCE0F7443F2082BC">34</mods:end>
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<mods:classification id="FBC709EE786DB128F554E50B33621A56">journal article</mods:classification>
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<mods:identifier id="7667D2B983B8770D75DA66F1855EE075" type="DOI">10.5281/zenodo.213061</mods:identifier>
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<mods:identifier id="11A0EE98B23D3C4040D15127F01BC529" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="2D7DAC8AC8326D7F1BE6AFFEF7CF8637" type="Zenodo-Dep">213061</mods:identifier>
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<treatment id="182887C6FF9B9B79C994FF62F90EE7DF" ID-DOI="http://doi.org/10.5281/zenodo.5674279" ID-GBIF-Taxon="119527543" ID-Zenodo-Dep="5674279" LSID="urn:lsid:plazi:treatment:182887C6FF9B9B79C994FF62F90EE7DF" httpUri="http://treatment.plazi.org/id/182887C6FF9B9B79C994FF62F90EE7DF" lastPageId="5" lastPageNumber="32" pageId="2" pageNumber="29">
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<subSubSection id="D89B655BFF9B9B7EC994FF62FA6DE0BA" pageId="2" pageNumber="29" type="nomenclature">
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<paragraph id="903E36D0FF9B9B7EC994FF62F969E0F4" blockId="2.[151,1437,151,255]" box="[151,552,151,177]" pageId="2" pageNumber="29">
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<heading id="CB7681BCFF9B9B7EC994FF62F969E0F4" bold="true" box="[151,552,151,177]" fontSize="11" level="1" pageId="2" pageNumber="29" reason="1">
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<emphasis id="A2F5EAC2FF9B9B7EC994FF62F969E0F4" bold="true" box="[151,552,151,177]" pageId="2" pageNumber="29">
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<taxonomicName id="57814D53FF9B9B7EC994FF62FACBE0F4" ID-CoL="3CC4Z" box="[151,394,151,177]" class="Actinopterygii" family="Zoarcidae" genus="Eulophias" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="29" phylum="Chordata" rank="species" species="koreanus" status="sp. nov.">
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<emphasis id="A2F5EAC2FF9B9B7EC994FF62FACBE0F4" bold="true" box="[151,394,151,177]" italics="true" pageId="2" pageNumber="29">Eulophias koreanus</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="B9C657B9FF9B9B7EC89AFF62F969E0F4" box="[409,552,151,177]" pageId="2" pageNumber="29" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="903E36D0FF9B9B7EC994FF4AFA6DE0BA" blockId="2.[151,1437,151,255]" pageId="2" pageNumber="29">(New English name: Korean prickleback; new Korean genus name: Jul-ga-si-chi-sok; new Korean species name: Jul-ga-si-chi)</paragraph>
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</subSubSection>
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<subSubSection id="D89B655BFF9B9B7EC994FEC2FAB3E182" pageId="2" pageNumber="29" type="materials_examined">
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<paragraph id="903E36D0FF9B9B7EC994FEC2F8A2E132" blockId="2.[151,1436,311,615]" pageId="2" pageNumber="29">
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<emphasis id="A2F5EAC2FF9B9B7EC994FEC2FA47E115" bold="true" box="[151,262,311,336]" pageId="2" pageNumber="29">
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<emphasis id="A2F5EAC2FF9B9B7EC994FEC2FBBEE115" bold="true" box="[151,255,311,336]" italics="true" pageId="2" pageNumber="29">
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<typeStatus id="4F3A8872FF9B9B7EC994FEC2FBBEE115" box="[151,255,311,336]" pageId="2" pageNumber="29" type="holotype">Holotype</typeStatus>
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</emphasis>
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.
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</emphasis>
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PKU
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<quantity id="57799B35FF9B9B7EC855FEC2F959E10A" box="[342,536,311,335]" metricMagnitude="3" metricUnit="m" metricValue="1.4381606999999998" pageId="2" pageNumber="29" unit="mm" value="1438160.7">1438, 160.7 mm</quantity>
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in standard length (
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<collectionCode id="F690AE15FF9B9B7ECA0FFEC2F870E10A" box="[780,817,311,335]" pageId="2" pageNumber="29">SL</collectionCode>
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), near Yokji Island, southern sea of
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<collectingCountry id="E8967640FF9B9B7ECDE4FECDFE6EE10A" box="[1255,1327,312,335]" name="South Korea" pageId="2" pageNumber="29">Korea</collectingCountry>
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, 34°15ʹN 128°34ʹE,
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<quantity id="57799B35FF9B9B7EC80EFEAAFA07E132" box="[269,326,351,375]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="29" unit="m" value="80.0">80 m</quantity>
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depth,
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<date id="E43F1010FF9B9B7EC89AFEAAF969E132" box="[409,552,351,375]" pageId="2" pageNumber="29" value="2008-09-26">26 Sep 2008</date>
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, by bottom trawl, H.J. Kwun (
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<figureCitation id="08BA2A55FF9B9B7ECA7FFEAAF8FEE132" box="[892,959,351,375]" captionStart="FIGURE 2" captionStartId="4.[151,250,1979,2002]" captionTargetBox="[496,1112,280,1867]" captionTargetId="figure@4.[495,1113,279,1868]" captionTargetPageId="4" captionText="FIGURE 2. Eulophias koreanus sp. nov. (A) holotype, PKU 1438, 160.7 mm SL. (B) paratype, PKU 1440, 139.2 mm SL, from southern sea of Korea. Arrow indicates the location of anus." httpUri="https://zenodo.org/record/213063/files/figure.png" pageId="2" pageNumber="29">Fig. 2</figureCitation>
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A).
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</paragraph>
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<paragraph id="903E36D0FF9B9B7EC9C4FE7DFAB3E182" blockId="2.[151,1436,311,615]" pageId="2" pageNumber="29">
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<emphasis id="A2F5EAC2FF9B9B7EC9C4FE7DFA7EE1E5" bold="true" box="[199,319,392,416]" pageId="2" pageNumber="29">
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||
<emphasis id="A2F5EAC2FF9B9B7EC9C4FE7DFA78E1DA" bold="true" box="[199,313,392,415]" italics="true" pageId="2" pageNumber="29">
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<typeStatus id="4F3A8872FF9B9B7EC9C4FE7DFA78E1DA" box="[199,313,392,415]" pageId="2" pageNumber="29" type="paratype">Paratypes</typeStatus>
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</emphasis>
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.
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</emphasis>
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Four specimens: PKU
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<date id="E43F1010FF9B9B7ECB5CFE72F9A0E1DA" box="[607,737,391,415]" pageId="2" pageNumber="29" value="1439" valueMax="1441">1439–1441</date>
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, 125.0–
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<quantity id="57799B35FF9B9B7ECA43FE72F8F9E1DA" box="[832,952,391,415]" metricMagnitude="-1" metricUnit="m" metricValue="1.5459999999999998" pageId="2" pageNumber="29" unit="mm" value="154.6">154.6 mm</quantity>
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<collectionCode id="F690AE15FF9B9B7ECAC6FE72F8A8E1DA" box="[965,1001,391,415]" pageId="2" pageNumber="29">SL</collectionCode>
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;
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<collectionCode id="F690AE15FF9B9B7ECAF8FE72FF7FE1DA" box="[1019,1086,391,415]" httpUri="http://grbio.org/cool/r4iq-k5jx" name="National Institute of Biological Resources" pageId="2" pageNumber="29">NIBR</collectionCode>
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–P0000016593,
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<quantity id="57799B35FF9B9B7ECDFFFE72FE26E1DA" box="[1276,1383,391,415]" metricMagnitude="-2" metricUnit="m" metricValue="7.07" pageId="2" pageNumber="29" unit="mm" value="70.7">70.7 mm</quantity>
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<collectionCode id="F690AE15FF9B9B7ECC77FE72FED4E1DA" box="[1396,1429,391,415]" pageId="2" pageNumber="29">SL</collectionCode>
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, collecting data as for
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<typeStatus id="4F3A8872FF9B9B7EC88BFE5AFAAFE182" box="[392,494,431,455]" pageId="2" pageNumber="29" type="holotype">holotype</typeStatus>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="D89B655BFF9B9B7EC9C4FE22FF75E222" pageId="2" pageNumber="29" type="diagnosis">
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<paragraph id="903E36D0FF9B9B7EC9C4FE22FF75E222" blockId="2.[151,1436,311,615]" pageId="2" pageNumber="29">
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||
<emphasis id="A2F5EAC2FF9B9B7EC9C4FE22FA00E1B5" bold="true" box="[199,321,471,496]" pageId="2" pageNumber="29">
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||
<emphasis id="A2F5EAC2FF9B9B7EC9C4FE22FA7BE1B5" bold="true" box="[199,314,471,496]" italics="true" pageId="2" pageNumber="29">Diagnosis</emphasis>
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.
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</emphasis>
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A new species of
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<taxonomicName id="57814D53FF9B9B7ECB21FE2DF9D2E1B5" box="[546,659,472,496]" class="Actinopterygii" family="Zoarcidae" genus="Eulophias" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="29" phylum="Chordata" rank="genus">
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<emphasis id="A2F5EAC2FF9B9B7ECB21FE2DF9D2E1B5" box="[546,659,472,496]" italics="true" pageId="2" pageNumber="29">Eulophias</emphasis>
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</taxonomicName>
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with the following combination of characters: Vertebrae 141–143; dorsal-fin spiny rays 124–128, dorsal-fin soft rays 11–14; anal-fin spiny ray 1, anal-fin soft rays 102–103; caudalfin rays 9–10; pelvic-fin absent; a dark band present on posterior of orbit (
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<figureCitation id="08BA2A55FF9B9B7ECADCFDD2FF65E27A" box="[991,1060,551,575]" captionStart="FIGURE 3" captionStartId="5.[151,250,843,866]" captionTargetBox="[478,1109,284,789]" captionTargetId="figure@5.[448,1138,251,823]" captionTargetPageId="5" captionText="FIGURE 3. Heads of Eulophias koreanus sp. nov. (A) PKU 1438, holotype. (B) PKU 1440, paratype." httpUri="https://zenodo.org/record/213064/files/figure.png" pageId="2" pageNumber="29">Fig. 3</figureCitation>
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). Pectoral-fin length 36.1–39.5% of head length (
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<tableCitation id="DD03036BFF9B9B7EC846FDBAFAD8E222" box="[325,409,591,615]" captionStart="TABLE 1" captionStartId="2.[151,239,667,691]" captionTargetBox="[165,1417,756,1790]" captionTargetPageId="2" captionText="TABLE 1. Comparison of counts and measurements of the three species of Eulophias. Proportional data for Eulophias owashii is converted from Okada and Suzuki (1954). * Counted by Miki (1985); ** Present study." httpUri="http://table.plazi.org/id/C4FE6658FF9B9B7EC994FD6EFFE1E296" pageId="2" pageNumber="29" tableUuid="C4FE6658FF9B9B7EC994FD6EFFE1E296">Table 1</tableCitation>
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). The parhypural without a small spine ventrally (
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<figureCitation id="08BA2A55FF9B9B7ECACEFDBAFF51E222" box="[973,1040,591,615]" captionStart="FIGURE 4" captionStartId="5.[151,250,1361,1384]" captionTargetBox="[314,1279,916,1346]" captionTargetId="figure@5.[314,1279,916,1347]" captionTargetPageId="5" captionText="FIGURE 4. Caudal skeletons of (A) Eulophias koreanus (PKU 1438, holotype) and (B) Eulophias owashii (FRLM 7151, holotype, phothgraphed by Kimura). C, caudal fin ray; EP, epural; HY, hypural; LDP, last dorsal pterygiophore; NPU 2, neural spine of preural centrum 2 (last neural spine); PC, procurrent caudal ray; PH, parhypural; US, urostyle." httpUri="https://zenodo.org/record/213065/files/figure.png" pageId="2" pageNumber="29">Fig. 4</figureCitation>
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A).
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</paragraph>
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</subSubSection>
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<caption id="C4FE6658FF9B9B7EC994FD6EFFE1E296" ID-Table-UUID="C4FE6658FF9B9B7EC994FD6EFFE1E296" httpUri="http://table.plazi.org/id/C4FE6658FF9B9B7EC994FD6EFFE1E296" pageId="2" pageNumber="29" targetBox="[165,1417,756,1790]" targetIsTable="true" targetPageId="2">
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<paragraph id="903E36D0FF9B9B7EC994FD6EFFE1E296" blockId="2.[151,1436,667,723]" pageId="2" pageNumber="29">
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<emphasis id="A2F5EAC2FF9B9B7EC994FD6EFA4AE2F6" bold="true" box="[151,267,667,691]" pageId="2" pageNumber="29">TABLE 1.</emphasis>
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Comparison of counts and measurements of the three species of
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<taxonomicName id="57814D53FF9B9B7ECACEFD69FF78E2F6" box="[973,1081,668,691]" class="Actinopterygii" family="Zoarcidae" genus="Eulophias" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="29" phylum="Chordata" rank="genus">
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<emphasis id="A2F5EAC2FF9B9B7ECACEFD69FF78E2F6" box="[973,1081,668,691]" italics="true" pageId="2" pageNumber="29">Eulophias</emphasis>
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</taxonomicName>
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. Proportional data for
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<taxonomicName id="57814D53FF9B9B7ECC33FD69FBABE296" class="Actinopterygii" family="Zoarcidae" genus="Eulophias" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="29" phylum="Chordata" rank="species" species="owashii">
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<emphasis id="A2F5EAC2FF9B9B7ECC33FD69FBABE296" italics="true" pageId="2" pageNumber="29">Eulophias owashii</emphasis>
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</taxonomicName>
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is converted from Okada and Suzuki (1954). *Counted by Miki (1985); **Present study.
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</paragraph>
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</caption>
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||
<paragraph id="903E36D0FF9B9B7EC8A7FD01FFBEE6BB" pageId="2" pageNumber="29">
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||
<table id="E281C470FF9B6483C9A6FD01FEC8E6BB" box="[165,1417,756,1790]" gridcols="4" gridrows="24" pageId="2" pageNumber="29">
|
||
<tr id="2EB13492FF9B6483C9A6FD01FEC8E34F" box="[165,1417,756,778]" gridrow="0" pageId="2" pageNumber="29" rowspan-0="1">
|
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<th id="6D605DEEFF9B6483C8A7FD01F816E34F" box="[420,855,756,778]" gridcol="1" gridrow="0" pageId="2" pageNumber="29">
|
||
<taxonomicName id="57814D53FF9B9B7EC8A7FD01F92CE34F" box="[420,621,756,778]" class="Actinopterygii" family="Zoarcidae" genus="Eulophias" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="29" phylum="Chordata" rank="species" species="koreanus" status="sp. nov.">
|
||
<emphasis id="A2F5EAC2FF9B9B7EC8A7FD01F92CE34F" box="[420,621,756,778]" italics="true" pageId="2" pageNumber="29">Eulophias koreanus</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="B9C657B9FF9B9B7ECB70FD00F983E34F" box="[627,706,757,778]" pageId="2" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
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||
</th>
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||
<th id="6D605DEEFF9B6483CA74FD01FF3BE34F" box="[887,1146,756,778]" gridcol="2" gridrow="0" pageId="2" pageNumber="29">
|
||
<taxonomicName id="57814D53FF9B9B7ECA74FD01FF72E34F" box="[887,1075,756,778]" class="Actinopterygii" family="Zoarcidae" genus="Eulophias" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="29" phylum="Chordata" rank="species" species="owashii">
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||
<emphasis id="A2F5EAC2FF9B9B7ECA74FD01FF72E34F" box="[887,1075,756,778]" italics="true" pageId="2" pageNumber="29">Eulophias owashii</emphasis>
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</taxonomicName>
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||
</th>
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||
<th id="6D605DEEFF9B6483CDD1FD01FEC8E34F" box="[1234,1417,756,778]" gridcol="3" gridrow="0" pageId="2" pageNumber="29">
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||
<taxonomicName id="57814D53FF9B9B7ECDD0FD01FEC8E34F" box="[1235,1417,756,778]" class="Actinopterygii" family="Zoarcidae" genus="Eulophias" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="29" phylum="Chordata" rank="species" species="tanneri">
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||
<emphasis id="A2F5EAC2FF9B9B7ECDD0FD01FEC8E34F" box="[1235,1417,756,778]" italics="true" pageId="2" pageNumber="29">Eulophias tanneri</emphasis>
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</taxonomicName>
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||
</th>
|
||
</tr>
|
||
<tr id="2EB13492FF9B6483C9A6FCD5FEC8E372" box="[165,1417,800,823]" gridrow="1" pageId="2" pageNumber="29" rowspan-0="1">
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||
<td id="6D605DEEFF9B6483C8A7FCD5F816E372" box="[420,855,800,823]" gridcol="1" gridrow="1" pageId="2" pageNumber="29">Present study</td>
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||
<td id="6D605DEEFF9B6483CA74FCD5FF3BE372" box="[887,1146,800,823]" gridcol="2" gridrow="1" pageId="2" pageNumber="29">Okada and Suzuki (1954)</td>
|
||
<td id="6D605DEEFF9B6483CDD1FCD5FEC8E372" box="[1234,1417,800,823]" gridcol="3" gridrow="1" pageId="2" pageNumber="29">Smith (1902)</td>
|
||
</tr>
|
||
<tr id="2EB13492FF9B6483C9A6FCB9FEC8E327" box="[165,1417,844,866]" gridrow="2" pageId="2" pageNumber="29">
|
||
<th id="6D605DEEFF9B6483C9A6FCB9FAC6E327" box="[165,391,844,866]" gridcol="0" gridrow="2" pageId="2" pageNumber="29">Type status</th>
|
||
<td id="6D605DEEFF9B6483C8A7FCB9F816E327" box="[420,855,844,866]" gridcol="1" gridrow="2" pageId="2" pageNumber="29">Holotype Paratypes (n=4)</td>
|
||
<td id="6D605DEEFF9B6483CA74FCB9FF3BE327" box="[887,1146,844,866]" gridcol="2" gridrow="2" pageId="2" pageNumber="29">Holotype</td>
|
||
<td id="6D605DEEFF9B6483CDD1FCB9FEC8E327" box="[1234,1417,844,866]" gridcol="3" gridrow="2" pageId="2" pageNumber="29">Holotype</td>
|
||
</tr>
|
||
<tr id="2EB13492FF9B6483C9A6FC8DFEC8E3CA" box="[165,1417,888,911]" gridrow="3" pageId="2" pageNumber="29">
|
||
<th id="6D605DEEFF9B6483C9A6FC8DFAC6E3CA" box="[165,391,888,911]" gridcol="0" gridrow="3" pageId="2" pageNumber="29">Standard length (mm)</th>
|
||
<td id="6D605DEEFF9B6483C8A7FC8DF816E3CA" box="[420,855,888,911]" gridcol="1" gridrow="3" pageId="2" pageNumber="29">160.7 70.7–154.6</td>
|
||
<td id="6D605DEEFF9B6483CA74FC8DFF3BE3CA" box="[887,1146,888,911]" gridcol="2" gridrow="3" pageId="2" pageNumber="29">173</td>
|
||
<td id="6D605DEEFF9B6483CDD1FC8DFEC8E3CA" box="[1234,1417,888,911]" gridcol="3" gridrow="3" pageId="2" pageNumber="29">45</td>
|
||
</tr>
|
||
<tr id="2EB13492FF9B6483C9A6FC50FEC8E3FF" box="[165,1417,933,954]" gridrow="4" pageId="2" pageNumber="29" rowspan-1="1" rowspan-2="1" rowspan-3="1">
|
||
<th id="6D605DEEFF9B6483C9A6FC50FAC6E3FF" box="[165,391,933,954]" gridcol="0" gridrow="4" pageId="2" pageNumber="29">Counts</th>
|
||
</tr>
|
||
<tr id="2EB13492FF9B6483C9A6FC25FEC8E3A3" box="[165,1417,976,998]" gridrow="5" pageId="2" pageNumber="29">
|
||
<th id="6D605DEEFF9B6483C9A6FC25FAC6E3A3" box="[165,391,976,998]" gridcol="0" gridrow="5" pageId="2" pageNumber="29">Dorsal fin</th>
|
||
<td id="6D605DEEFF9B6483C8A7FC25F816E3A3" box="[420,855,976,998]" gridcol="1" gridrow="5" pageId="2" pageNumber="29">CXXVIII, 12 CXXIV–CXXVIII, 11–14</td>
|
||
<td id="6D605DEEFF9B6483CA74FC25FF3BE3A3" box="[887,1146,976,998]" gridcol="2" gridrow="5" pageId="2" pageNumber="29">CXXVII, 12</td>
|
||
<td id="6D605DEEFF9B6483CDD1FC25FEC8E3A3" box="[1234,1417,976,998]" gridcol="3" gridrow="5" pageId="2" pageNumber="29">CXXI, 13</td>
|
||
</tr>
|
||
<tr id="2EB13492FF9B6483C9A6FC09FEC8E456" box="[165,1417,1020,1043]" gridrow="6" pageId="2" pageNumber="29">
|
||
<th id="6D605DEEFF9B6483C9A6FC09FAC6E456" box="[165,391,1020,1043]" gridcol="0" gridrow="6" pageId="2" pageNumber="29">Anal fin</th>
|
||
<td id="6D605DEEFF9B6483C8A7FC09F816E456" box="[420,855,1020,1043]" gridcol="1" gridrow="6" pageId="2" pageNumber="29">I, 103 I, 102–103</td>
|
||
<td id="6D605DEEFF9B6483CA74FC09FF3BE456" box="[887,1146,1020,1043]" gridcol="2" gridrow="6" pageId="2" pageNumber="29">I, 95 (102*)</td>
|
||
<td id="6D605DEEFF9B6483CDD1FC09FEC8E456" box="[1234,1417,1020,1043]" gridcol="3" gridrow="6" pageId="2" pageNumber="29">I, 75</td>
|
||
</tr>
|
||
<tr id="2EB13492FF9B6483C9A6FBDDFEC8E47B" box="[165,1417,1064,1086]" gridrow="7" pageId="2" pageNumber="29">
|
||
<th id="6D605DEEFF9B6483C9A6FBDDFAC6E47B" box="[165,391,1064,1086]" gridcol="0" gridrow="7" pageId="2" pageNumber="29">Pectoral fin</th>
|
||
<td id="6D605DEEFF9B6483C8A7FBDDF816E47B" box="[420,855,1064,1086]" gridcol="1" gridrow="7" pageId="2" pageNumber="29">7 7</td>
|
||
<td id="6D605DEEFF9B6483CA74FBDDFF3BE47B" box="[887,1146,1064,1086]" gridcol="2" gridrow="7" pageId="2" pageNumber="29">7</td>
|
||
<td id="6D605DEEFF9B6483CDD1FBDDFEC8E47B" box="[1234,1417,1064,1086]" gridcol="3" gridrow="7" pageId="2" pageNumber="29">-</td>
|
||
</tr>
|
||
<tr id="2EB13492FF9B6483C9A6FBA1FEC8E42F" box="[165,1417,1108,1130]" gridrow="8" pageId="2" pageNumber="29">
|
||
<th id="6D605DEEFF9B6483C9A6FBA1FAC6E42F" box="[165,391,1108,1130]" gridcol="0" gridrow="8" pageId="2" pageNumber="29">Caudal fin</th>
|
||
<td id="6D605DEEFF9B6483C8A7FBA1F816E42F" box="[420,855,1108,1130]" gridcol="1" gridrow="8" pageId="2" pageNumber="29">4+10+1 3+9–10+1</td>
|
||
<td id="6D605DEEFF9B6483CA74FBA1FF3BE42F" box="[887,1146,1108,1130]" gridcol="2" gridrow="8" pageId="2" pageNumber="29">4+10+1**</td>
|
||
<td id="6D605DEEFF9B6483CDD1FBA1FEC8E42F" box="[1234,1417,1108,1130]" gridcol="3" gridrow="8" pageId="2" pageNumber="29">7</td>
|
||
</tr>
|
||
<tr id="2EB13492FF9B6483C9A6FB75FEC8E4D3" box="[165,1417,1152,1174]" gridrow="9" pageId="2" pageNumber="29">
|
||
<th id="6D605DEEFF9B6483C9A6FB75FAC6E4D3" box="[165,391,1152,1174]" gridcol="0" gridrow="9" pageId="2" pageNumber="29">Vertebrae</th>
|
||
<td id="6D605DEEFF9B6483C8A7FB75F816E4D3" box="[420,855,1152,1174]" gridcol="1" gridrow="9" pageId="2" pageNumber="29">141 141–143</td>
|
||
<td id="6D605DEEFF9B6483CA74FB75FF3BE4D3" box="[887,1146,1152,1174]" gridcol="2" gridrow="9" pageId="2" pageNumber="29">133*</td>
|
||
<td id="6D605DEEFF9B6483CDD1FB75FEC8E4D3" box="[1234,1417,1152,1174]" gridcol="3" gridrow="9" pageId="2" pageNumber="29">-</td>
|
||
</tr>
|
||
<tr id="2EB13492FF9B6483C9A6FB58FEC8E486" box="[165,1417,1197,1219]" gridrow="10" pageId="2" pageNumber="29" rowspan-1="1" rowspan-2="1" rowspan-3="1">
|
||
<th id="6D605DEEFF9B6483C9A6FB58FAC6E486" box="[165,391,1197,1219]" gridcol="0" gridrow="10" pageId="2" pageNumber="29">Measurements (%SL)</th>
|
||
</tr>
|
||
<tr id="2EB13492FF9B6483C9A6FB2DFEC8E4AB" box="[165,1417,1240,1262]" gridrow="11" pageId="2" pageNumber="29">
|
||
<th id="6D605DEEFF9B6483C9A6FB2DFAC6E4AB" box="[165,391,1240,1262]" gridcol="0" gridrow="11" pageId="2" pageNumber="29">Head length</th>
|
||
<td id="6D605DEEFF9B6483C8A7FB2DF816E4AB" box="[420,855,1240,1262]" gridcol="1" gridrow="11" pageId="2" pageNumber="29">10.3 9.3–12.2</td>
|
||
<td id="6D605DEEFF9B6483CA74FB2DFF3BE4AB" box="[887,1146,1240,1262]" gridcol="2" gridrow="11" pageId="2" pageNumber="29">10.4</td>
|
||
<td id="6D605DEEFF9B6483CDD1FB2DFEC8E4AB" box="[1234,1417,1240,1262]" gridcol="3" gridrow="11" pageId="2" pageNumber="29">12</td>
|
||
</tr>
|
||
<tr id="2EB13492FF9B6483C9A6FAF1FEC8E55F" box="[165,1417,1284,1306]" gridrow="12" pageId="2" pageNumber="29">
|
||
<th id="6D605DEEFF9B6483C9A6FAF1FAC6E55F" box="[165,391,1284,1306]" gridcol="0" gridrow="12" pageId="2" pageNumber="29">Body depth</th>
|
||
<td id="6D605DEEFF9B6483C8A7FAF1F816E55F" box="[420,855,1284,1306]" gridcol="1" gridrow="12" pageId="2" pageNumber="29">4.4 3.8–4.4</td>
|
||
<td id="6D605DEEFF9B6483CA74FAF1FF3BE55F" box="[887,1146,1284,1306]" gridcol="2" gridrow="12" pageId="2" pageNumber="29">4.6</td>
|
||
<td id="6D605DEEFF9B6483CDD1FAF1FEC8E55F" box="[1234,1417,1284,1306]" gridcol="3" gridrow="12" pageId="2" pageNumber="29">5</td>
|
||
</tr>
|
||
<tr id="2EB13492FF9B6483C9A6FAC5FEC8E503" box="[165,1417,1328,1350]" gridrow="13" pageId="2" pageNumber="29">
|
||
<th id="6D605DEEFF9B6483C9A6FAC5FAC6E503" box="[165,391,1328,1350]" gridcol="0" gridrow="13" pageId="2" pageNumber="29">Body width</th>
|
||
<td id="6D605DEEFF9B6483C8A7FAC5F816E503" box="[420,855,1328,1350]" gridcol="1" gridrow="13" pageId="2" pageNumber="29">3.2 3.0–4.2</td>
|
||
<td id="6D605DEEFF9B6483CA74FAC5FF3BE503" box="[887,1146,1328,1350]" gridcol="2" gridrow="13" pageId="2" pageNumber="29">-</td>
|
||
<td id="6D605DEEFF9B6483CDD1FAC5FEC8E503" box="[1234,1417,1328,1350]" gridcol="3" gridrow="13" pageId="2" pageNumber="29">-</td>
|
||
</tr>
|
||
<tr id="2EB13492FF9B6483C9A6FAA9FEC8E537" box="[165,1417,1372,1394]" gridrow="14" pageId="2" pageNumber="29">
|
||
<th id="6D605DEEFF9B6483C9A6FAA9FAC6E537" box="[165,391,1372,1394]" gridcol="0" gridrow="14" pageId="2" pageNumber="29">Predorsal length</th>
|
||
<td id="6D605DEEFF9B6483C8A7FAA9F816E537" box="[420,855,1372,1394]" gridcol="1" gridrow="14" pageId="2" pageNumber="29">9.6 9.0–11.0</td>
|
||
<td id="6D605DEEFF9B6483CA74FAA9FF3BE537" box="[887,1146,1372,1394]" gridcol="2" gridrow="14" pageId="2" pageNumber="29">10.4</td>
|
||
<td id="6D605DEEFF9B6483CDD1FAA9FEC8E537" box="[1234,1417,1372,1394]" gridcol="3" gridrow="14" pageId="2" pageNumber="29">-</td>
|
||
</tr>
|
||
<tr id="2EB13492FF9B6483C9A6FA7DFEC8E5DB" box="[165,1417,1416,1438]" gridrow="15" pageId="2" pageNumber="29">
|
||
<th id="6D605DEEFF9B6483C9A6FA7DFAC6E5DB" box="[165,391,1416,1438]" gridcol="0" gridrow="15" pageId="2" pageNumber="29">Preanal fin length</th>
|
||
<td id="6D605DEEFF9B6483C8A7FA7DF816E5DB" box="[420,855,1416,1438]" gridcol="1" gridrow="15" pageId="2" pageNumber="29">35.0 34.8–36.1</td>
|
||
<td id="6D605DEEFF9B6483CA74FA7DFF3BE5DB" box="[887,1146,1416,1438]" gridcol="2" gridrow="15" pageId="2" pageNumber="29">35.8</td>
|
||
<td id="6D605DEEFF9B6483CDD1FA7DFEC8E5DB" box="[1234,1417,1416,1438]" gridcol="3" gridrow="15" pageId="2" pageNumber="29">40</td>
|
||
</tr>
|
||
<tr id="2EB13492FF9B6483C9A6FA41FEC8E58F" box="[165,1417,1460,1482]" gridrow="16" pageId="2" pageNumber="29">
|
||
<th id="6D605DEEFF9B6483C9A6FA41FAC6E58F" box="[165,391,1460,1482]" gridcol="0" gridrow="16" pageId="2" pageNumber="29">Preanus length</th>
|
||
<td id="6D605DEEFF9B6483C8A7FA41F816E58F" box="[420,855,1460,1482]" gridcol="1" gridrow="16" pageId="2" pageNumber="29">33.6 33.1–35.1</td>
|
||
<td id="6D605DEEFF9B6483CA74FA41FF3BE58F" box="[887,1146,1460,1482]" gridcol="2" gridrow="16" pageId="2" pageNumber="29">33.6</td>
|
||
<td id="6D605DEEFF9B6483CDD1FA41FEC8E58F" box="[1234,1417,1460,1482]" gridcol="3" gridrow="16" pageId="2" pageNumber="29">-</td>
|
||
</tr>
|
||
<tr id="2EB13492FF9B6483C9A6FA15FEC8E5B3" box="[165,1417,1504,1526]" gridrow="17" pageId="2" pageNumber="29">
|
||
<th id="6D605DEEFF9B6483C9A6FA15FAC6E5B3" box="[165,391,1504,1526]" gridcol="0" gridrow="17" pageId="2" pageNumber="29">Postorbital length</th>
|
||
<td id="6D605DEEFF9B6483C8A7FA15F816E5B3" box="[420,855,1504,1526]" gridcol="1" gridrow="17" pageId="2" pageNumber="29">6.3 5.2–7.2</td>
|
||
<td id="6D605DEEFF9B6483CA74FA15FF3BE5B3" box="[887,1146,1504,1526]" gridcol="2" gridrow="17" pageId="2" pageNumber="29">-</td>
|
||
<td id="6D605DEEFF9B6483CDD1FA15FEC8E5B3" box="[1234,1417,1504,1526]" gridcol="3" gridrow="17" pageId="2" pageNumber="29">-</td>
|
||
</tr>
|
||
<tr id="2EB13492FF9B6483C9A6F9F8FEC8E666" box="[165,1417,1549,1571]" gridrow="18" pageId="2" pageNumber="29" rowspan-1="1" rowspan-2="1" rowspan-3="1">
|
||
<th id="6D605DEEFF9B6483C9A6F9F8FAC6E666" box="[165,391,1549,1571]" gridcol="0" gridrow="18" pageId="2" pageNumber="29">Measurements (%HL)</th>
|
||
</tr>
|
||
<tr id="2EB13492FF9B6483C9A6F9CDFEC8E60B" box="[165,1417,1592,1614]" gridrow="19" pageId="2" pageNumber="29">
|
||
<th id="6D605DEEFF9B6483C9A6F9CDFAC6E60B" box="[165,391,1592,1614]" gridcol="0" gridrow="19" pageId="2" pageNumber="29">Snout length</th>
|
||
<td id="6D605DEEFF9B6483C8A7F9CDF816E60B" box="[420,855,1592,1614]" gridcol="1" gridrow="19" pageId="2" pageNumber="29">14.5 12.9–16.0</td>
|
||
<td id="6D605DEEFF9B6483CA74F9CDFF3BE60B" box="[887,1146,1592,1614]" gridcol="2" gridrow="19" pageId="2" pageNumber="29">16.7</td>
|
||
<td id="6D605DEEFF9B6483CDD1F9CDFEC8E60B" box="[1234,1417,1592,1614]" gridcol="3" gridrow="19" pageId="2" pageNumber="29">-</td>
|
||
</tr>
|
||
<tr id="2EB13492FF9B6483C9A6F991FEC8E63F" box="[165,1417,1636,1658]" gridrow="20" pageId="2" pageNumber="29">
|
||
<th id="6D605DEEFF9B6483C9A6F991FAC6E63F" box="[165,391,1636,1658]" gridcol="0" gridrow="20" pageId="2" pageNumber="29">Interorbiral width</th>
|
||
<td id="6D605DEEFF9B6483C8A7F991F816E63F" box="[420,855,1636,1658]" gridcol="1" gridrow="20" pageId="2" pageNumber="29">5.4 4.7–7.6</td>
|
||
<td id="6D605DEEFF9B6483CA74F991FF3BE63F" box="[887,1146,1636,1658]" gridcol="2" gridrow="20" pageId="2" pageNumber="29">22.2</td>
|
||
<td id="6D605DEEFF9B6483CDD1F991FEC8E63F" box="[1234,1417,1636,1658]" gridcol="3" gridrow="20" pageId="2" pageNumber="29">-</td>
|
||
</tr>
|
||
<tr id="2EB13492FF9B6483C9A6F965FEC8E6E3" box="[165,1417,1680,1702]" gridrow="21" pageId="2" pageNumber="29">
|
||
<th id="6D605DEEFF9B6483C9A6F965FAC6E6E3" box="[165,391,1680,1702]" gridcol="0" gridrow="21" pageId="2" pageNumber="29">Eye length</th>
|
||
<td id="6D605DEEFF9B6483C8A7F965F816E6E3" box="[420,855,1680,1702]" gridcol="1" gridrow="21" pageId="2" pageNumber="29">22.9 21.5–26.4</td>
|
||
<td id="6D605DEEFF9B6483CA74F965FF3BE6E3" box="[887,1146,1680,1702]" gridcol="2" gridrow="21" pageId="2" pageNumber="29">27.8</td>
|
||
<td id="6D605DEEFF9B6483CDD1F965FEC8E6E3" box="[1234,1417,1680,1702]" gridcol="3" gridrow="21" pageId="2" pageNumber="29">-</td>
|
||
</tr>
|
||
<tr id="2EB13492FF9B6483C9A6F949FEC8E697" box="[165,1417,1724,1746]" gridrow="22" pageId="2" pageNumber="29">
|
||
<th id="6D605DEEFF9B6483C9A6F949FAC6E697" box="[165,391,1724,1746]" gridcol="0" gridrow="22" pageId="2" pageNumber="29">Upper jaw length</th>
|
||
<td id="6D605DEEFF9B6483C8A7F949F816E697" box="[420,855,1724,1746]" gridcol="1" gridrow="22" pageId="2" pageNumber="29">25.9 18.8–33.3</td>
|
||
<td id="6D605DEEFF9B6483CA74F949FF3BE697" box="[887,1146,1724,1746]" gridcol="2" gridrow="22" pageId="2" pageNumber="29">27.8</td>
|
||
<td id="6D605DEEFF9B6483CDD1F949FEC8E697" box="[1234,1417,1724,1746]" gridcol="3" gridrow="22" pageId="2" pageNumber="29">-</td>
|
||
</tr>
|
||
<tr id="2EB13492FF9B6483C9A6F91DFEC8E6BB" box="[165,1417,1768,1790]" gridrow="23" pageId="2" pageNumber="29">
|
||
<th id="6D605DEEFF9B6483C9A6F91DFAC6E6BB" box="[165,391,1768,1790]" gridcol="0" gridrow="23" pageId="2" pageNumber="29">Pectoral fin length</th>
|
||
<td id="6D605DEEFF9B6483C8A7F91DF816E6BB" box="[420,855,1768,1790]" gridcol="1" gridrow="23" pageId="2" pageNumber="29">38.0 36.1–39.5</td>
|
||
<td id="6D605DEEFF9B6483CA74F91DFF3BE6BB" box="[887,1146,1768,1790]" gridcol="2" gridrow="23" pageId="2" pageNumber="29">27.8</td>
|
||
<td id="6D605DEEFF9B6483CDD1F91DFEC8E6BB" box="[1234,1417,1768,1790]" gridcol="3" gridrow="23" pageId="2" pageNumber="29">35.7</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<subSubSection id="D89B655BFF9B9B7FC9C4F891F9D5E42A" lastPageId="3" lastPageNumber="30" pageId="2" pageNumber="29" type="description">
|
||
<paragraph id="903E36D0FF9B9B7EC9C4F891FF6BE739" blockId="2.[151,1436,1892,2037]" box="[199,1066,1892,1917]" pageId="2" pageNumber="29">
|
||
<emphasis id="A2F5EAC2FF9B9B7EC9C4F891FA12E738" bold="true" box="[199,339,1892,1917]" pageId="2" pageNumber="29">
|
||
<emphasis id="A2F5EAC2FF9B9B7EC9C4F891FA0CE738" bold="true" box="[199,333,1892,1917]" italics="true" pageId="2" pageNumber="29">Description</emphasis>
|
||
.
|
||
</emphasis>
|
||
The count and proportional measurements are shown in
|
||
<tableCitation id="DD03036BFF9B9B7ECAD3F891FF65E739" box="[976,1060,1892,1916]" captionStart="TABLE 1" captionStartId="2.[151,239,667,691]" captionTargetBox="[165,1417,756,1790]" captionTargetPageId="2" captionText="TABLE 1. Comparison of counts and measurements of the three species of Eulophias. Proportional data for Eulophias owashii is converted from Okada and Suzuki (1954). * Counted by Miki (1985); ** Present study." httpUri="http://table.plazi.org/id/C4FE6658FF9B9B7EC994FD6EFFE1E296" pageId="2" pageNumber="29" tableUuid="C4FE6658FF9B9B7EC994FD6EFFE1E296">Table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="903E36D0FF9B9B7FC9C4F879F837E3B2" blockId="2.[151,1436,1892,2037]" lastBlockId="3.[151,1437,151,1776]" lastPageId="3" lastPageNumber="30" pageId="2" pageNumber="29">
|
||
Body slender and very elongate, its depth 3.8–4.4% of standard length (SL) and its width 3.0–4.2% of SL; dorsal and ventral profiles of body nearly a straight form, but gently curved in front of eye (snout region). Head very small and slightly compressed, its length 9.3–12.2% of SL; head without cirri or fleshy protrusions; snout short, blunt (
|
||
<figureCitation id="08BA2A55FF9A9B7FC82FFF62FA31E0EA" box="[300,368,151,175]" captionStart="FIGURE 3" captionStartId="5.[151,250,843,866]" captionTargetBox="[478,1109,284,789]" captionTargetId="figure@5.[448,1138,251,823]" captionTargetPageId="5" captionText="FIGURE 3. Heads of Eulophias koreanus sp. nov. (A) PKU 1438, holotype. (B) PKU 1440, paratype." httpUri="https://zenodo.org/record/213064/files/figure.png" pageId="3" pageNumber="30">Fig. 3</figureCitation>
|
||
A) or slightly pointed anteriorly (
|
||
<figureCitation id="08BA2A55FF9A9B7FCBF6FF62F87BE0EA" box="[757,826,151,175]" captionStart="FIGURE 3" captionStartId="5.[151,250,843,866]" captionTargetBox="[478,1109,284,789]" captionTargetId="figure@5.[448,1138,251,823]" captionTargetPageId="5" captionText="FIGURE 3. Heads of Eulophias koreanus sp. nov. (A) PKU 1438, holotype. (B) PKU 1440, paratype." httpUri="https://zenodo.org/record/213064/files/figure.png" pageId="3" pageNumber="30">Fig. 3</figureCitation>
|
||
B), showing ontogenetic variation in snout shape, its length shorter than eye diameter; eye moderate size, its diameter 21.5–26.4% of head length (HL), eye little elevated and advanced in head; interorbital space narrow, somewhat convex transversely, its width 4.7–7.6% of HL; single nostril tubular on each side, its length similar to half of pupil; right and left gill membranes continuous across isthmus. Mouth small, terminal and oblique; upper jaw length 18.8–33.3% of HL; lower jaw slightly projecting or upper and lower jaws equally protruding (
|
||
<figureCitation id="08BA2A55FF9A9B7FCA24FEAAF82EE132" box="[807,879,351,375]" captionStart="FIGURE 3" captionStartId="5.[151,250,843,866]" captionTargetBox="[478,1109,284,789]" captionTargetId="figure@5.[448,1138,251,823]" captionTargetPageId="5" captionText="FIGURE 3. Heads of Eulophias koreanus sp. nov. (A) PKU 1438, holotype. (B) PKU 1440, paratype." httpUri="https://zenodo.org/record/213064/files/figure.png" pageId="3" pageNumber="30">Fig. 3</figureCitation>
|
||
); maxilla straight or curved, reaching below to anterior border or center of pupil; two to three rows of conical teeth on both jaws (
|
||
<typeStatus id="4F3A8872FF9A9B7FCD2FFE72FFD1E1DA" box="[1068,1168,391,415]" pageId="3" pageNumber="30" type="holotype">holotype</typeStatus>
|
||
with two and four large conical teeth on upper and lower jaws respectively); no teeth on vomer or palatine; the tip of snout located to the horizontal line from the lower border of eye or from the center of eye (
|
||
<figureCitation id="08BA2A55FF9A9B7FCAD6FE22FF5DE1AA" box="[981,1052,471,495]" captionStart="FIGURE 3" captionStartId="5.[151,250,843,866]" captionTargetBox="[478,1109,284,789]" captionTargetId="figure@5.[448,1138,251,823]" captionTargetPageId="5" captionText="FIGURE 3. Heads of Eulophias koreanus sp. nov. (A) PKU 1438, holotype. (B) PKU 1440, paratype." httpUri="https://zenodo.org/record/213064/files/figure.png" pageId="3" pageNumber="30">Fig. 3</figureCitation>
|
||
). Body covered by small cycloid scales, but head naked; lateral-line scales pit-shaped, a single lateral line extending from behind operculum to base of caudal-fin. Dorsal-fin composed of spiny rayed and soft rayed portions, those connected smoothly, of which the former very long (85.7–86.2% of SL), but the latter very short (5.2–5.5% of SL); origin of dorsal-fin located above posterior margin of operculum; dorsal-fin height very low, but the height of spiny ray tended to increase posteriorly. Pectoral-fin short and rounded, its length 36.1–39.5% of HL; pelvic-fin absent; anal-fin composed of single spiny ray and numerous soft rays. Dorsal- and anal-fin soft rays unbranched, but 1-2 of the longest pectoral- and 8-9 of caudal-fin rays branched. Anus located below between 35th and 36th dorsal-fin spiny rays. Caudal-fin rounded, connected with dorsal- and anal-fins without notch. Procurrent caudal rays present: 3–4 rays dorsally, 1 ray ventrally (
|
||
<figureCitation id="08BA2A55FF9A9B7FC837FCCAFA37E312" box="[308,374,831,855]" captionStart="FIGURE 4" captionStartId="5.[151,250,1361,1384]" captionTargetBox="[314,1279,916,1346]" captionTargetId="figure@5.[314,1279,916,1347]" captionTargetPageId="5" captionText="FIGURE 4. Caudal skeletons of (A) Eulophias koreanus (PKU 1438, holotype) and (B) Eulophias owashii (FRLM 7151, holotype, phothgraphed by Kimura). C, caudal fin ray; EP, epural; HY, hypural; LDP, last dorsal pterygiophore; NPU 2, neural spine of preural centrum 2 (last neural spine); PC, procurrent caudal ray; PH, parhypural; US, urostyle." httpUri="https://zenodo.org/record/213065/files/figure.png" pageId="3" pageNumber="30">Fig. 4</figureCitation>
|
||
A). The last dorsal pterygiophore inserted between the last 1st and 2nd (including the
|
||
<typeStatus id="4F3A8872FF9A9B7FCC32FCCAFED4E312" box="[1329,1429,831,855]" pageId="3" pageNumber="30" type="holotype">holotype</typeStatus>
|
||
) or 2nd and 3rd neural spines (
|
||
<figureCitation id="08BA2A55FF9A9B7FC8EAFC92F96DE33A" box="[489,556,871,895]" captionStart="FIGURE 4" captionStartId="5.[151,250,1361,1384]" captionTargetBox="[314,1279,916,1346]" captionTargetId="figure@5.[314,1279,916,1347]" captionTargetPageId="5" captionText="FIGURE 4. Caudal skeletons of (A) Eulophias koreanus (PKU 1438, holotype) and (B) Eulophias owashii (FRLM 7151, holotype, phothgraphed by Kimura). C, caudal fin ray; EP, epural; HY, hypural; LDP, last dorsal pterygiophore; NPU 2, neural spine of preural centrum 2 (last neural spine); PC, procurrent caudal ray; PH, parhypural; US, urostyle." httpUri="https://zenodo.org/record/213065/files/figure.png" pageId="3" pageNumber="30">Fig. 4</figureCitation>
|
||
A). The number of epurals of the
|
||
<typeStatus id="4F3A8872FF9A9B7FCAAAFC92FF4AE33A" box="[937,1035,871,895]" pageId="3" pageNumber="30" type="holotype">holotype</typeStatus>
|
||
was three (
|
||
<figureCitation id="08BA2A55FF9A9B7FCD8DFC92FF92E33A" box="[1166,1235,871,895]" captionStart="FIGURE 4" captionStartId="5.[151,250,1361,1384]" captionTargetBox="[314,1279,916,1346]" captionTargetId="figure@5.[314,1279,916,1347]" captionTargetPageId="5" captionText="FIGURE 4. Caudal skeletons of (A) Eulophias koreanus (PKU 1438, holotype) and (B) Eulophias owashii (FRLM 7151, holotype, phothgraphed by Kimura). C, caudal fin ray; EP, epural; HY, hypural; LDP, last dorsal pterygiophore; NPU 2, neural spine of preural centrum 2 (last neural spine); PC, procurrent caudal ray; PH, parhypural; US, urostyle." httpUri="https://zenodo.org/record/213065/files/figure.png" pageId="3" pageNumber="30">Fig. 4</figureCitation>
|
||
A), but that of the
|
||
<typeStatus id="4F3A8872FF9A9B7FC994FC64FA43E3EC" box="[151,258,913,937]" pageId="3" pageNumber="30" type="paratype">paratypes</typeStatus>
|
||
was uncertain. Parhypural of all specimens had no spines on its ventral portion (
|
||
<figureCitation id="08BA2A55FF9A9B7FCDC1FC7AFE4BE3E2" box="[1218,1290,911,935]" captionStart="FIGURE 4" captionStartId="5.[151,250,1361,1384]" captionTargetBox="[314,1279,916,1346]" captionTargetId="figure@5.[314,1279,916,1347]" captionTargetPageId="5" captionText="FIGURE 4. Caudal skeletons of (A) Eulophias koreanus (PKU 1438, holotype) and (B) Eulophias owashii (FRLM 7151, holotype, phothgraphed by Kimura). C, caudal fin ray; EP, epural; HY, hypural; LDP, last dorsal pterygiophore; NPU 2, neural spine of preural centrum 2 (last neural spine); PC, procurrent caudal ray; PH, parhypural; US, urostyle." httpUri="https://zenodo.org/record/213065/files/figure.png" pageId="3" pageNumber="30">Fig. 4</figureCitation>
|
||
A). Cephalic sensory pores well-developed: nasal pores 2; interorbital pore 1; mandibular pores 3; preopercular pores 4; infraorbital pores 7; postorbital pores 4; occipital pores 3 (
|
||
<figureCitation id="08BA2A55FF9A9B7FCA27FC2AF829E3B2" box="[804,872,991,1015]" captionStart="FIGURE 5" captionStartId="6.[151,250,910,933]" captionTargetBox="[570,1025,219,859]" captionTargetId="figure@6.[550,1040,193,893]" captionTargetPageId="6" captionText="FIGURE 5. Sensory system of Eulophias koreanus sp. nov. IFOP, infraorbital pores; IOP, interorbital pore; MP, mandibular pores; NP, nasal pores; OP, occipital pores; POP, postorbital pores; POPP, preopercular pores." httpUri="https://zenodo.org/record/213066/files/figure.png" pageId="3" pageNumber="30">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="903E36D0FF9A9B7FC9C4FBF2F9D5E42A" blockId="3.[151,1437,151,1776]" pageId="3" pageNumber="30">
|
||
<emphasis id="A2F5EAC2FF9A9B7FC9C4FBF2FA0BE465" bold="true" box="[199,330,1031,1056]" pageId="3" pageNumber="30">
|
||
<emphasis id="A2F5EAC2FF9A9B7FC9C4FBF2FA02E465" bold="true" box="[199,323,1031,1056]" italics="true" pageId="3" pageNumber="30">Coloration</emphasis>
|
||
.
|
||
</emphasis>
|
||
When fresh, head and body whitish brown. Dark band present on posterior of orbit (
|
||
<figureCitation id="08BA2A55FF9A9B7FCC46FBF2FECFE45A" box="[1349,1422,1031,1055]" captionStart="FIGURE 3" captionStartId="5.[151,250,843,866]" captionTargetBox="[478,1109,284,789]" captionTargetId="figure@5.[448,1138,251,823]" captionTargetPageId="5" captionText="FIGURE 3. Heads of Eulophias koreanus sp. nov. (A) PKU 1438, holotype. (B) PKU 1440, paratype." httpUri="https://zenodo.org/record/213064/files/figure.png" pageId="3" pageNumber="30">Fig. 3</figureCitation>
|
||
). Approximately 19–23 dark brown blotches present midlaterally on body (
|
||
<figureCitation id="08BA2A55FF9A9B7FCAD3FBDAFF54E402" box="[976,1045,1071,1095]" captionStart="FIGURE 2" captionStartId="4.[151,250,1979,2002]" captionTargetBox="[496,1112,280,1867]" captionTargetId="figure@4.[495,1113,279,1868]" captionTargetPageId="4" captionText="FIGURE 2. Eulophias koreanus sp. nov. (A) holotype, PKU 1438, 160.7 mm SL. (B) paratype, PKU 1440, 139.2 mm SL, from southern sea of Korea. Arrow indicates the location of anus." httpUri="https://zenodo.org/record/213063/files/figure.png" pageId="3" pageNumber="30">Fig. 2</figureCitation>
|
||
). All fins whitish. After fixation in formalin, head and body became dark brown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D89B655BFF9A9B7FC9C4FB8AFAB7E4FA" pageId="3" pageNumber="30" type="distribution">
|
||
<paragraph id="903E36D0FF9A9B7FC9C4FB8AFAB7E4FA" blockId="3.[151,1437,151,1776]" pageId="3" pageNumber="30">
|
||
<emphasis id="A2F5EAC2FF9A9B7FC9C4FB8AFA1BE4DD" bold="true" box="[199,346,1151,1176]" pageId="3" pageNumber="30">
|
||
<emphasis id="A2F5EAC2FF9A9B7FC9C4FB8AFA12E4DD" bold="true" box="[199,339,1151,1176]" italics="true" pageId="3" pageNumber="30">Distribution</emphasis>
|
||
.
|
||
</emphasis>
|
||
<taxonomicName id="57814D53FF9A9B7FC862FB75F901E4DD" box="[353,576,1152,1176]" class="Actinopterygii" family="Zoarcidae" genus="Eulophias" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="30" phylum="Chordata" rank="species" species="koreanus">
|
||
<emphasis id="A2F5EAC2FF9A9B7FC862FB75F901E4DD" box="[353,576,1152,1176]" italics="true" pageId="3" pageNumber="30">Eulophias koreanus</emphasis>
|
||
</taxonomicName>
|
||
occurs in the southern sea of
|
||
<collectingCountry id="E8967640FF9A9B7FCA8EFB75F893E4D2" box="[909,978,1152,1175]" name="South Korea" pageId="3" pageNumber="30">Korea</collectingCountry>
|
||
(near Yokji Island, the bottom at a depth of approximately
|
||
<quantity id="57799B35FF9A9B7FC85DFB52FAD8E4FA" box="[350,409,1191,1215]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="30" unit="m" value="80.0">80 m</quantity>
|
||
,
|
||
<figureCitation id="08BA2A55FF9A9B7FC8A6FB52FAA4E4FA" box="[421,485,1191,1215]" captionStart="FIGURE 1" captionStartId="1.[151,250,1349,1372]" captionTargetBox="[438,1136,848,1295]" captionTargetId="figure@1.[434,1147,842,1326]" captionTargetPageId="1" captionText="FIGURE 1. Distribution of the three species of Eulophias. Eulophias koreanus (black squares), Eulophias owashii (black circle), and Eulophias tanneri (black triangle)." httpUri="https://zenodo.org/record/213062/files/figure.png" pageId="3" pageNumber="30">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D89B655BFF9A9B7FC9C4FB3AF97BE6AA" pageId="3" pageNumber="30" type="etymology">
|
||
<paragraph id="903E36D0FF9A9B7FC9C4FB3AFFC8E4A2" blockId="3.[151,1437,151,1776]" box="[199,1161,1231,1256]" pageId="3" pageNumber="30">
|
||
<emphasis id="A2F5EAC2FF9A9B7FC9C4FB3AFA06E4AD" bold="true" box="[199,327,1231,1256]" pageId="3" pageNumber="30">
|
||
<emphasis id="A2F5EAC2FF9A9B7FC9C4FB3AFA01E4AD" bold="true" box="[199,320,1231,1256]" italics="true" pageId="3" pageNumber="30">Etymology</emphasis>
|
||
.
|
||
</emphasis>
|
||
The specific epithet “
|
||
<taxonomicName id="57814D53FF9A9B7FCB3DFB25F9E2E4AD" box="[574,675,1232,1256]" class="Actinopterygii" family="Zoarcidae" genus="Eulophias" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="30" phylum="Chordata" rank="species" species="koreanus">
|
||
<emphasis id="A2F5EAC2FF9A9B7FCB3DFB25F9E2E4AD" box="[574,675,1232,1256]" italics="true" pageId="3" pageNumber="30">koreanus</emphasis>
|
||
</taxonomicName>
|
||
” is derived from ‘Korea’, the
|
||
<typeStatus id="4F3A8872FF9A9B7FCAFBFB25FF69E4AD" box="[1016,1064,1232,1256]" pageId="3" pageNumber="30">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
<paragraph id="903E36D0FF9A9B7FC9C4FB02F97BE6AA" blockId="3.[151,1437,151,1776]" pageId="3" pageNumber="30">
|
||
<emphasis id="A2F5EAC2FF9A9B7FC9C4FB02FA1AE555" bold="true" box="[199,347,1271,1296]" pageId="3" pageNumber="30">
|
||
<emphasis id="A2F5EAC2FF9A9B7FC9C4FB02FA15E555" bold="true" box="[199,340,1271,1296]" italics="true" pageId="3" pageNumber="30">Comparison</emphasis>
|
||
.
|
||
</emphasis>
|
||
When comparing
|
||
<taxonomicName id="57814D53FF9A9B7FCB4BFB0DF871E555" box="[584,816,1272,1296]" class="Actinopterygii" family="Zoarcidae" genus="Eulophias" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="30" phylum="Chordata" rank="species" species="koreanus">
|
||
<emphasis id="A2F5EAC2FF9A9B7FCB4BFB0DF871E555" box="[584,816,1272,1296]" italics="true" pageId="3" pageNumber="30">Eulophias koreanus</emphasis>
|
||
</taxonomicName>
|
||
with two congeneric species in external features,
|
||
<taxonomicName id="57814D53FF9A9B7FC994FAD5FA34E57D" box="[151,373,1312,1336]" class="Actinopterygii" family="Zoarcidae" genus="Eulophias" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="30" phylum="Chordata" rank="species" species="koreanus">
|
||
<emphasis id="A2F5EAC2FF9A9B7FC994FAD5FA34E57D" box="[151,373,1312,1336]" italics="true" pageId="3" pageNumber="30">Eulophias koreanus</emphasis>
|
||
</taxonomicName>
|
||
is more similar to
|
||
<taxonomicName id="57814D53FF9A9B7FCB4BFAD4F9F8E57D" box="[584,697,1313,1336]" class="Actinopterygii" family="Zoarcidae" genus="Eulophias" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="30" phylum="Chordata" rank="species" species="tanneri">
|
||
<emphasis id="A2F5EAC2FF9A9B7FCB4BFAD4F9F8E57D" box="[584,697,1313,1336]" italics="true" pageId="3" pageNumber="30">E. tanneri</emphasis>
|
||
</taxonomicName>
|
||
in having a dark band behind the eye (absent in
|
||
<taxonomicName id="57814D53FF9A9B7FCDDAFAD4FE0EE57D" box="[1241,1359,1312,1336]" class="Actinopterygii" family="Zoarcidae" genus="Eulophias" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="30" phylum="Chordata" rank="species" species="owashii">
|
||
<emphasis id="A2F5EAC2FF9A9B7FCDDAFAD4FE0EE57D" box="[1241,1359,1312,1336]" italics="true" pageId="3" pageNumber="30">E. owashii</emphasis>
|
||
</taxonomicName>
|
||
) and a long pectoral-fin (short in
|
||
<taxonomicName id="57814D53FF9A9B7FC8BFFABCF973E525" box="[444,562,1352,1376]" class="Actinopterygii" family="Zoarcidae" genus="Eulophias" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="30" phylum="Chordata" rank="species" species="owashii">
|
||
<emphasis id="A2F5EAC2FF9A9B7FC8BFFABCF973E525" box="[444,562,1352,1376]" italics="true" pageId="3" pageNumber="30">E. owashii</emphasis>
|
||
</taxonomicName>
|
||
), but is more similar to
|
||
<taxonomicName id="57814D53FF9A9B7FCA3FFABCF8F3E525" box="[828,946,1352,1376]" class="Actinopterygii" family="Zoarcidae" genus="Eulophias" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="30" phylum="Chordata" rank="species" species="owashii">
|
||
<emphasis id="A2F5EAC2FF9A9B7FCA3FFABCF8F3E525" box="[828,946,1352,1376]" italics="true" pageId="3" pageNumber="30">E. owashii</emphasis>
|
||
</taxonomicName>
|
||
in having more numerous anal- and caudalfin rays (only a few in
|
||
<taxonomicName id="57814D53FF9A9B7FC89AFA84F94BE5CD" box="[409,522,1393,1416]" class="Actinopterygii" family="Zoarcidae" genus="Eulophias" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="30" phylum="Chordata" rank="species" species="tanneri">
|
||
<emphasis id="A2F5EAC2FF9A9B7FC89AFA84F94BE5CD" box="[409,522,1393,1416]" italics="true" pageId="3" pageNumber="30">E. tanneri</emphasis>
|
||
</taxonomicName>
|
||
). However,
|
||
<taxonomicName id="57814D53FF9A9B7FCB96FA85F835E5CD" box="[661,884,1392,1416]" class="Actinopterygii" family="Zoarcidae" genus="Eulophias" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="30" phylum="Chordata" rank="species" species="koreanus">
|
||
<emphasis id="A2F5EAC2FF9A9B7FCB96FA85F835E5CD" box="[661,884,1392,1416]" italics="true" pageId="3" pageNumber="30">Eulophias koreanus</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="57814D53FF9A9B7FCD09FA84FF3BE5CD" box="[1034,1146,1393,1416]" class="Actinopterygii" family="Zoarcidae" genus="Eulophias" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="30" phylum="Chordata" rank="species" species="tanneri">
|
||
<emphasis id="A2F5EAC2FF9A9B7FCD09FA84FF3BE5CD" box="[1034,1146,1393,1416]" italics="true" pageId="3" pageNumber="30">E. tanneri</emphasis>
|
||
</taxonomicName>
|
||
in the number of anal-fin rays (102–103 vs.
|
||
<quantity id="57799B35FF9A9B7FC877FA62FAF3E5EA" box="[372,434,1431,1455]" metricMagnitude="0" metricUnit="m" metricValue="1.905" pageId="3" pageNumber="30" unit="in" value="75.0">75 in</quantity>
|
||
<taxonomicName id="57814D53FF9A9B7FC8BEFA6CF973E5F5" box="[445,562,1433,1456]" class="Actinopterygii" family="Zoarcidae" genus="Eulophias" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="30" phylum="Chordata" rank="species" species="tanneri">
|
||
<emphasis id="A2F5EAC2FF9A9B7FC8BEFA6CF973E5F5" box="[445,562,1433,1456]" italics="true" pageId="3" pageNumber="30">E. tanneri</emphasis>
|
||
</taxonomicName>
|
||
) and caudal-fin rays (9–10 vs. 7), and from
|
||
<taxonomicName id="57814D53FF9A9B7FCD4EFA6CFF89E5F5" box="[1101,1224,1432,1456]" class="Actinopterygii" family="Zoarcidae" genus="Eulophias" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="30" phylum="Chordata" rank="species" species="owashii">
|
||
<emphasis id="A2F5EAC2FF9A9B7FCD4EFA6CFF89E5F5" box="[1101,1224,1432,1456]" italics="true" pageId="3" pageNumber="30">E. owashii</emphasis>
|
||
</taxonomicName>
|
||
in the number of vertebrae (141–143 vs.
|
||
<quantity id="57799B35FF9A9B7FC89DFA4AFAA5E592" box="[414,484,1471,1495]" metricMagnitude="0" metricUnit="m" metricValue="3.3781999999999996" pageId="3" pageNumber="30" unit="in" value="133.0">133 in</quantity>
|
||
<taxonomicName id="57814D53FF9A9B7FC8E9FA34F921E59D" box="[490,608,1472,1496]" class="Actinopterygii" family="Zoarcidae" genus="Eulophias" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="30" phylum="Chordata" rank="species" species="owashii">
|
||
<emphasis id="A2F5EAC2FF9A9B7FC8E9FA34F921E59D" box="[490,608,1472,1496]" italics="true" pageId="3" pageNumber="30">E. owashii</emphasis>
|
||
</taxonomicName>
|
||
) and pectoral-fin length (36.1–39.5% of head length vs. 27.8%) (
|
||
<tableCitation id="DD03036BFF9A9B7FCC34FA4AFECBE592" box="[1335,1418,1471,1495]" captionStart="TABLE 1" captionStartId="2.[151,239,667,691]" captionTargetBox="[165,1417,756,1790]" captionTargetPageId="2" captionText="TABLE 1. Comparison of counts and measurements of the three species of Eulophias. Proportional data for Eulophias owashii is converted from Okada and Suzuki (1954). * Counted by Miki (1985); ** Present study." httpUri="http://table.plazi.org/id/C4FE6658FF9B9B7EC994FD6EFFE1E296" pageId="3" pageNumber="30" tableUuid="C4FE6658FF9B9B7EC994FD6EFFE1E296">Table 1</tableCitation>
|
||
).
|
||
<taxonomicName id="57814D53FF9A9B7FC994FA1DFA3AE645" box="[151,379,1512,1536]" class="Actinopterygii" family="Zoarcidae" genus="Eulophias" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="30" phylum="Chordata" rank="species" species="koreanus">
|
||
<emphasis id="A2F5EAC2FF9A9B7FC994FA1DFA3AE645" box="[151,379,1512,1536]" italics="true" pageId="3" pageNumber="30">Eulophias koreanus</emphasis>
|
||
</taxonomicName>
|
||
also differs from
|
||
<taxonomicName id="57814D53FF9A9B7FCB5AFA1CF994E645" box="[601,725,1512,1536]" class="Actinopterygii" family="Zoarcidae" genus="Eulophias" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="30" phylum="Chordata" rank="species" species="owashii">
|
||
<emphasis id="A2F5EAC2FF9A9B7FCB5AFA1CF994E645" box="[601,725,1512,1536]" italics="true" pageId="3" pageNumber="30">E. owashii</emphasis>
|
||
</taxonomicName>
|
||
in the following osteological characteristics: the last dorsal pterygiophore inserted between the last 1st and 2nd or 2nd and 3rd neural spines in
|
||
<taxonomicName id="57814D53FF9A9B7FCD4BF9E4FF8DE66D" box="[1096,1228,1552,1576]" class="Actinopterygii" family="Zoarcidae" genus="Eulophias" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="30" phylum="Chordata" rank="species" species="koreanus">
|
||
<emphasis id="A2F5EAC2FF9A9B7FCD4BF9E4FF8DE66D" box="[1096,1228,1552,1576]" italics="true" pageId="3" pageNumber="30">E. koreanus</emphasis>
|
||
</taxonomicName>
|
||
, whereas between the last 3rd and 4th neural spines in the
|
||
<typeStatus id="4F3A8872FF9A9B7FCB50F9C2F9F4E60A" box="[595,693,1591,1615]" pageId="3" pageNumber="30" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="57814D53FF9A9B7FCBD9F9CCF811E615" box="[730,848,1592,1616]" class="Actinopterygii" family="Zoarcidae" genus="Eulophias" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="30" phylum="Chordata" rank="species" species="owashii">
|
||
<emphasis id="A2F5EAC2FF9A9B7FCBD9F9CCF811E615" box="[730,848,1592,1616]" italics="true" pageId="3" pageNumber="30">E. owashii</emphasis>
|
||
</taxonomicName>
|
||
; the parhypural without a small spine ventrally in
|
||
<taxonomicName id="57814D53FF9A9B7FCC87F9CCFBBCE63D" class="Actinopterygii" family="Zoarcidae" genus="Eulophias" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="30" phylum="Chordata" rank="species" species="koreanus">
|
||
<emphasis id="A2F5EAC2FF9A9B7FCC87F9CCFBBCE63D" italics="true" pageId="3" pageNumber="30">E. koreanus</emphasis>
|
||
</taxonomicName>
|
||
, whereas with a small spine in
|
||
<taxonomicName id="57814D53FF9A9B7FCB60F994F99BE63D" box="[611,730,1632,1656]" class="Actinopterygii" family="Zoarcidae" genus="Eulophias" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="30" phylum="Chordata" rank="species" species="owashii">
|
||
<emphasis id="A2F5EAC2FF9A9B7FCB60F994F99BE63D" box="[611,730,1632,1656]" italics="true" pageId="3" pageNumber="30">E. owashii</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="08BA2A55FF9A9B7FCBE8F9AAF873E632" box="[747,818,1631,1655]" captionStart="FIGURE 4" captionStartId="5.[151,250,1361,1384]" captionTargetBox="[314,1279,916,1346]" captionTargetId="figure@5.[314,1279,916,1347]" captionTargetPageId="5" captionText="FIGURE 4. Caudal skeletons of (A) Eulophias koreanus (PKU 1438, holotype) and (B) Eulophias owashii (FRLM 7151, holotype, phothgraphed by Kimura). C, caudal fin ray; EP, epural; HY, hypural; LDP, last dorsal pterygiophore; NPU 2, neural spine of preural centrum 2 (last neural spine); PC, procurrent caudal ray; PH, parhypural; US, urostyle." httpUri="https://zenodo.org/record/213065/files/figure.png" pageId="3" pageNumber="30">Fig. 4</figureCitation>
|
||
). In addition, we newly found ontogenetic variation in head shape, especially in the snout region: the
|
||
<typeStatus id="4F3A8872FF9A9B7FCBB5F972F859E6DA" box="[694,792,1671,1695]" pageId="3" pageNumber="30" type="holotype">holotype</typeStatus>
|
||
and the largest
|
||
<typeStatus id="4F3A8872FF9A9B7FCAD0F97CFF72E6E4" box="[979,1075,1673,1697]" pageId="3" pageNumber="30" type="paratype">paratype</typeStatus>
|
||
(
|
||
<quantity id="57799B35FF9A9B7FCD46F972FFFAE6E5" box="[1093,1211,1671,1696]" metricMagnitude="-1" metricUnit="m" metricValue="1.5459999999999998" pageId="3" pageNumber="30" unit="mm" value="154.6">154.6 mm</quantity>
|
||
and
|
||
<quantity id="57799B35FF9A9B7FCDF4F972FE2AE6E5" box="[1271,1387,1671,1696]" metricMagnitude="-1" metricUnit="m" metricValue="1.607" pageId="3" pageNumber="30" unit="mm" value="160.7">160.7 mm</quantity>
|
||
SL, respectively) have a blunt snout, whereas the rest three smaller
|
||
<typeStatus id="4F3A8872FF9A9B7FCA5EF944F889E68C" box="[861,968,1713,1737]" pageId="3" pageNumber="30" type="paratype">paratypes</typeStatus>
|
||
(
|
||
<quantity id="57799B35FF9A9B7FCAD4F95AFF7EE68D" box="[983,1087,1711,1736]" metricMagnitude="-2" metricUnit="m" metricValue="7.07" pageId="3" pageNumber="30" unit="mm" value="70.7">70.7 mm</quantity>
|
||
, 125.0 mm, and
|
||
<quantity id="57799B35FF9A9B7FCDFAF95AFE2AE68D" box="[1273,1387,1711,1736]" metricMagnitude="-1" metricUnit="m" metricValue="1.392" pageId="3" pageNumber="30" unit="mm" value="139.2">139.2 mm</quantity>
|
||
SL) have a slightly pointed snout (
|
||
<figureCitation id="08BA2A55FF9A9B7FC8EBF922F96DE6AA" box="[488,556,1751,1775]" captionStart="FIGURE 3" captionStartId="5.[151,250,843,866]" captionTargetBox="[478,1109,284,789]" captionTargetId="figure@5.[448,1138,251,823]" captionTargetPageId="5" captionText="FIGURE 3. Heads of Eulophias koreanus sp. nov. (A) PKU 1438, holotype. (B) PKU 1440, paratype." httpUri="https://zenodo.org/record/213064/files/figure.png" pageId="3" pageNumber="30">Fig. 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="C4FE6658FF9D9B78C994F84EF9B9E7B5" httpUri="https://zenodo.org/record/213063/files/figure.png" pageId="4" pageNumber="31" targetBox="[496,1112,280,1867]" targetPageId="4">
|
||
<paragraph id="903E36D0FF9D9B78C994F84EF9B9E7B5" blockId="4.[151,1436,1979,2032]" pageId="4" pageNumber="31">
|
||
<emphasis id="A2F5EAC2FF9D9B78C994F84EFA53E794" bold="true" box="[151,274,1979,2002]" pageId="4" pageNumber="31">FIGURE 2.</emphasis>
|
||
<taxonomicName id="57814D53FF9D9B78C81BF849FAA1E797" box="[280,480,1980,2002]" class="Actinopterygii" family="Zoarcidae" genus="Eulophias" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="31" phylum="Chordata" rank="species" species="koreanus" status="sp. nov.">
|
||
<emphasis id="A2F5EAC2FF9D9B78C81BF849FAA1E797" box="[280,480,1980,2002]" italics="true" pageId="4" pageNumber="31">Eulophias koreanus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B9C657B9FF9D9B78C8E5F848F975E797" box="[486,564,1981,2002]" pageId="4" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(A) holotype, PKU 1438, 160.7 mm SL. (B) paratype, PKU 1440, 139.2 mm SL, from southern sea of Korea. Arrow indicates the location of anus.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="C4FE6658FF9C9B79C994FCBEFFE2E327" box="[151,1187,843,866]" httpUri="https://zenodo.org/record/213064/files/figure.png" pageId="5" pageNumber="32" targetBox="[478,1109,284,789]" targetPageId="5">
|
||
<paragraph id="903E36D0FF9C9B79C994FCBEFFE2E327" blockId="5.[151,1187,843,866]" box="[151,1187,843,866]" pageId="5" pageNumber="32">
|
||
<emphasis id="A2F5EAC2FF9C9B79C994FCBEFA53E324" bold="true" box="[151,274,843,866]" pageId="5" pageNumber="32">FIGURE 3.</emphasis>
|
||
Heads of
|
||
<taxonomicName id="57814D53FF9C9B79C879FCB9F902E327" box="[378,579,844,866]" class="Actinopterygii" family="Zoarcidae" genus="Eulophias" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="32" phylum="Chordata" rank="species" species="koreanus" status="sp. nov.">
|
||
<emphasis id="A2F5EAC2FF9C9B79C879FCB9F902E327" box="[378,579,844,866]" italics="true" pageId="5" pageNumber="32">Eulophias koreanus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B9C657B9FF9C9B79CB4AFCB8F9D9E327" box="[585,664,845,866]" pageId="5" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(A) PKU 1438, holotype. (B) PKU 1440, paratype.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="C4FE6658FF9C9B79C994FAA4FFEBE5E0" httpUri="https://zenodo.org/record/213065/files/figure.png" pageId="5" pageNumber="32" targetBox="[314,1279,916,1346]" targetPageId="5">
|
||
<paragraph id="903E36D0FF9C9B79C994FAA4FFEBE5E0" blockId="5.[151,1436,1361,1445]" pageId="5" pageNumber="32">
|
||
<emphasis id="A2F5EAC2FF9C9B79C994FAA4FA57E522" bold="true" box="[151,278,1361,1384]" pageId="5" pageNumber="32">FIGURE 4.</emphasis>
|
||
Caudal skeletons of (A)
|
||
<taxonomicName id="57814D53FF9C9B79CB20FAA7F9B1E52D" box="[547,752,1362,1384]" class="Actinopterygii" family="Zoarcidae" genus="Eulophias" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="32" phylum="Chordata" rank="species" species="koreanus">
|
||
<emphasis id="A2F5EAC2FF9C9B79CB20FAA7F9B1E52D" box="[547,752,1362,1384]" italics="true" pageId="5" pageNumber="32">Eulophias koreanus</emphasis>
|
||
</taxonomicName>
|
||
(PKU 1438, holotype) and (B)
|
||
<taxonomicName id="57814D53FF9C9B79CD47FAA7FE42E52D" box="[1092,1283,1362,1384]" class="Actinopterygii" family="Zoarcidae" genus="Eulophias" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="32" phylum="Chordata" rank="species" species="owashii">
|
||
<emphasis id="A2F5EAC2FF9C9B79CD47FAA7FE42E52D" box="[1092,1283,1362,1384]" italics="true" pageId="5" pageNumber="32">Eulophias owashii</emphasis>
|
||
</taxonomicName>
|
||
(FRLM 7151, holotype, phothgraphed by Kimura). C, caudal fin ray; EP, epural; HY, hypural; LDP, last dorsal pterygiophore; NPU2, neural spine of preural centrum 2 (last neural spine); PC, procurrent caudal ray; PH, parhypural; US, urostyle.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="D89B655BFF9C9B79C9C4FA03F90EE7DF" pageId="5" pageNumber="32" type="discussion">
|
||
<paragraph id="903E36D0FF9C9B79C9C4FA03F81CE6DB" blockId="5.[151,1436,1526,1946]" pageId="5" pageNumber="32">
|
||
<emphasis id="A2F5EAC2FF9C9B79C9C4FA03FA75E64A" bold="true" box="[199,308,1526,1551]" pageId="5" pageNumber="32">
|
||
<emphasis id="A2F5EAC2FF9C9B79C9C4FA03FA6FE64A" bold="true" box="[199,302,1526,1551]" italics="true" pageId="5" pageNumber="32">Remarks</emphasis>
|
||
.
|
||
</emphasis>
|
||
We found a new species belonging to the genus
|
||
<taxonomicName id="57814D53FF9C9B79CA61FA03F895E64B" box="[866,980,1526,1550]" class="Actinopterygii" family="Zoarcidae" genus="Eulophias" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="32" phylum="Chordata" rank="genus">
|
||
<emphasis id="A2F5EAC2FF9C9B79CA61FA03F895E64B" box="[866,980,1526,1550]" italics="true" pageId="5" pageNumber="32">Eulophias</emphasis>
|
||
</taxonomicName>
|
||
based on five specimens collected from the southern sea of
|
||
<collectingCountry id="E8967640FF9C9B79C87BF9EEFA81E677" box="[376,448,1563,1586]" name="South Korea" pageId="5" pageNumber="32">Korea</collectingCountry>
|
||
. Although they are remarkably consistent in all meristic and morphometric characters, they showed slightly variation in head shape with growth. However, analyses of 500-bp sequences of mtDNA COI showed identical sequences among five specimens as 0.000 Kimura 2-parameter distances (
|
||
<bibRefCitation id="F4104B21FF9C9B79CDA5F997FE08E63F" author="Kimura" box="[1190,1353,1634,1658]" pageId="5" pageNumber="32" refString="Kimura, M. (1980) A simple method for estimating evolutionary rates of base substitutions through comparative studies of nucleotide sequences. Journal of Molecular Evolution, 16, 111 - 120." type="journal article" year="1980">Kimura, 1980</bibRefCitation>
|
||
). Thus, such difference in head shape must be an ontogenetic variation.
|
||
</paragraph>
|
||
<paragraph id="903E36D0FF9C9B79C9C4F95FF90EE7DF" blockId="5.[151,1436,1526,1946]" pageId="5" pageNumber="32">
|
||
The new species revealed here brings the worldwide total species of
|
||
<taxonomicName id="57814D53FF9C9B79CAD1F95FFF05E687" box="[978,1092,1706,1730]" class="Actinopterygii" family="Zoarcidae" genus="Eulophias" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="32" phylum="Chordata" rank="genus">
|
||
<emphasis id="A2F5EAC2FF9C9B79CAD1F95FFF05E687" box="[978,1092,1706,1730]" italics="true" pageId="5" pageNumber="32">Eulophias</emphasis>
|
||
</taxonomicName>
|
||
to three:
|
||
<taxonomicName id="57814D53FF9C9B79CDB3F95EFE77E687" box="[1200,1334,1706,1730]" class="Actinopterygii" family="Zoarcidae" genus="Eulophias" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="32" phylum="Chordata" rank="species" species="koreanus" status="sp. nov.">
|
||
<emphasis id="A2F5EAC2FF9C9B79CDB3F95EFE77E687" box="[1200,1334,1706,1730]" italics="true" pageId="5" pageNumber="32">E. koreanus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B9C657B9FF9C9B79CC3DF95EFED9E687" box="[1342,1432,1707,1730]" pageId="5" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="57814D53FF9C9B79C994F925FA57E6A3" box="[151,278,1742,1767]" class="Actinopterygii" family="Zoarcidae" genus="Eulophias" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="32" phylum="Chordata" rank="species" species="owashii">
|
||
<emphasis id="A2F5EAC2FF9C9B79C994F925FA57E6A3" box="[151,278,1742,1767]" italics="true" pageId="5" pageNumber="32">E. owashii</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="57814D53FF9C9B79C865F925FA9EE6A3" box="[358,479,1743,1767]" class="Actinopterygii" family="Zoarcidae" genus="Eulophias" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="32" phylum="Chordata" rank="species" species="tanneri">
|
||
<emphasis id="A2F5EAC2FF9C9B79C865F925FA9EE6A3" box="[358,479,1743,1767]" italics="true" pageId="5" pageNumber="32">E. tanneri</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="08BA2A55FF9C9B79C8FBF93BF903E6A3" box="[504,578,1742,1766]" captionStart="FIGURE 1" captionStartId="1.[151,250,1349,1372]" captionTargetBox="[438,1136,848,1295]" captionTargetId="figure@1.[434,1147,842,1326]" captionTargetPageId="1" captionText="FIGURE 1. Distribution of the three species of Eulophias. Eulophias koreanus (black squares), Eulophias owashii (black circle), and Eulophias tanneri (black triangle)." httpUri="https://zenodo.org/record/213062/files/figure.png" pageId="5" pageNumber="32">Fig. 1</figureCitation>
|
||
). Together with two known
|
||
<taxonomicName id="57814D53FF9C9B79CAACF93BFF60E6A3" box="[943,1057,1742,1766]" class="Actinopterygii" family="Zoarcidae" genus="Eulophias" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="32" phylum="Chordata" rank="genus">
|
||
<emphasis id="A2F5EAC2FF9C9B79CAACF93BFF60E6A3" box="[943,1057,1742,1766]" italics="true" pageId="5" pageNumber="32">Eulophias</emphasis>
|
||
</taxonomicName>
|
||
species, several morphological combinations are likely to be diagnostic characters of the genus
|
||
<taxonomicName id="57814D53FF9C9B79CA62F907F893E74F" box="[865,978,1778,1802]" class="Actinopterygii" family="Zoarcidae" genus="Eulophias" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="32" phylum="Chordata" rank="genus">
|
||
<emphasis id="A2F5EAC2FF9C9B79CA62F907F893E74F" box="[865,978,1778,1802]" italics="true" pageId="5" pageNumber="32">Eulophias</emphasis>
|
||
</taxonomicName>
|
||
: a slender and very elongated body, dark blotches on the sides of the body, head without cirri and fleshy protrusions, absence of a pelvic-fin, and dorsal-fin with numerous spiny rays (i.e., more than approx. 120) and only a few soft rays (i.e., less than approx. 15). Also, the first two hypurals are well fused with parhypural, which seems to be a diagnostic character of the subfamily
|
||
<taxonomicName id="57814D53FF9C9B79C994F877FA70E7DF" box="[151,305,1922,1946]" class="Actinopterygii" family="Zoarcidae" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="32" phylum="Chordata" rank="subFamily" subFamily="Neozoarcinae">Neozoarcinae</taxonomicName>
|
||
(Sensu
|
||
<bibRefCitation id="F4104B21FF9C9B79C888F877F903E7DF" author="Anderson" box="[395,578,1922,1946]" pageId="5" pageNumber="32" refString="Anderson, M. E. (1994) Systematics and osteology of the Zoarcidae (Teleostei: Perciformes). Ichthyological Bulletin, 60, 1 - 120." type="journal article" year="1994">Anderson, 1994</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |