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<document id="25E7EF157B91E091003C4A7A28C81755" ID-CLB-Dataset="298728" ID-DOI="10.11646/zootaxa.5471.1.3" ID-GBIF-Dataset="8c04b5de-4e97-4e7e-a0a0-9658beefb935" ID-ISSN="1175-5326" ID-Zenodo-Dep="12189573" ID-ZooBank="008B0723-9443-4D81-BE49-1C3C5D80B031" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="julia" IM.treatments_approvedBy="felipe" checkinTime="1718892294504" checkinUser="plazi" docAuthor="Chamorro, William, Lopera-Toro, Alejandro &amp; Génier, François" docDate="2024" docId="181587E6C24DFFCE7CEC82DAE46DD89F" docLanguage="en" docName="zootaxa.5471.1.3.pdf" docOrigin="Zootaxa 5471 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5471.1.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ontherus (Ontherus) araujoi Chamorro, Lopera, &amp; Genier 2024, new species" docType="treatment" docUuid="AE55C382-83D0-4168-9049-F3D0A343B989" docUuidSource="ZooBank" docVersion="6" lastPageNumber="60" masterDocId="E42CFF9EC24CFFCA7C7B8719E02CDE22" masterDocTitle="A new species of Ontherus Erichson, 1847 (Coleoptera: Scarabaeidae: Scarabaeinae: Coprini) and new distributional records of Cryptocanthon humidus Howden, 1973 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini) for Ecuador" masterLastPageNumber="66" masterPageNumber="56" pageNumber="57" updateTime="1719001740724" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="619EE2BDEBF8EB69ACF95D2427F64EB8">
<mods:title id="03FA80C89F85D07D7E344A2AB214476F">A new species of Ontherus Erichson, 1847 (Coleoptera: Scarabaeidae: Scarabaeinae: Coprini) and new distributional records of Cryptocanthon humidus Howden, 1973 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini) for Ecuador</mods:title>
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<mods:namePart id="5CCDA0E33899A80E6ED2064C19BB2B89">Chamorro, William</mods:namePart>
<mods:affiliation id="8AB4CCCFB88D8C859B28D85D2B1B1279">Universidad de la Amazonia, Research Group in Entomology, Entomology Laboratory, Florencia, Caquetá, Colombia.</mods:affiliation>
</mods:name>
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<mods:namePart id="38B0CCE5EE4AD31D820187561F9092AC">Lopera-Toro, Alejandro</mods:namePart>
<mods:affiliation id="AE24EFB95B32D65358B047A697935F8F">Manu Biological Station, Kosnipata, Cusco, Peru.</mods:affiliation>
</mods:name>
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<mods:namePart id="F6553D6F3F2DFB4D1620A3354D8089CC">Génier, François</mods:namePart>
<mods:nameIdentifier id="16909841217B26CBEFC66EC2F31041E3" type="ORCID">0000-0002-5399-8412</mods:nameIdentifier>
<mods:nameIdentifier id="D82850B2D42AD922C7FDA7C97FF9F2D2" type="email">fgenier@nature.ca</mods:nameIdentifier>
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<mods:title id="825F9B14CCE4E6AE87F6DE8A2FFBC745">Zootaxa</mods:title>
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<mods:date id="594FDC826D0081A00606DA37990331D1">2024</mods:date>
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<mods:number id="A7CBA7F779E905BB0D471D3BEA623119">2024-06-17</mods:number>
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<mods:number id="8D0537E6822F4DE8229706B7D8CB597C">5471</mods:number>
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<treatment id="181587E6C24DFFCE7CEC82DAE46DD89F" ID-DOI="http://doi.org/10.5281/zenodo.12190614" ID-GBIF-Taxon="233317628" ID-Zenodo-Dep="12190614" LSID="urn:lsid:zoobank.org:act:AE55C382-83D0-4168-9049-F3D0A343B989" httpUri="http://treatment.plazi.org/id/181587E6C24DFFCE7CEC82DAE46DD89F" lastPageId="4" lastPageNumber="60" pageId="1" pageNumber="57">
<subSubSection id="D8A6657BC24DFFCB7CEC82DAE427DBFF" box="[151,1035,1474,1502]" pageId="1" pageNumber="57" type="nomenclature">
<paragraph id="900336F0C24DFFCB7CEC82DAE427DBFF" blockId="1.[151,1035,1474,1537]" box="[151,1035,1474,1502]" pageId="1" pageNumber="57">
<heading id="CB4B819CC24DFFCB7CEC82DAE427DBFF" bold="true" box="[151,1035,1474,1502]" fontSize="11" level="1" pageId="1" pageNumber="57" reason="1">
<emphasis id="A2C8EAE2C24DFFCB7CEC82DAE427DBFF" bold="true" box="[151,1035,1474,1502]" pageId="1" pageNumber="57">
<taxonomicName id="57BC4D73C24DFFCB7CEC82DAE343DBFF" authority="Chamorro, Lopera, &amp; Genier" authorityName="Chamorro, Lopera, &amp; Genier" authorityYear="2024" box="[151,879,1474,1502]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="57" phylum="Arthropoda" rank="species" species="araujoi" status="sp. nov." subGenus="Ontherus">
<emphasis id="A2C8EAE2C24DFFCB7CEC82DAE125DBFC" bold="true" box="[151,265,1475,1502]" italics="true" pageId="1" pageNumber="57">Ontherus</emphasis>
(
<emphasis id="A2C8EAE2C24DFFCB7D6182DAE1A0DBFC" bold="true" box="[282,396,1475,1502]" italics="true" pageId="1" pageNumber="57">Ontherus</emphasis>
)
<emphasis id="A2C8EAE2C24DFFCB7DE582DAE1DADBFF" bold="true" box="[414,502,1475,1501]" italics="true" pageId="1" pageNumber="57">araujoi</emphasis>
Chamorro, Lopera, &amp; Génier
</taxonomicName>
,
<taxonomicNameLabel id="B9FB5799C24DFFCB7F0082DAE427DBFF" box="[891,1035,1475,1501]" pageId="1" pageNumber="57" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D8A6657BC24DFFCB7CEC82FEE1B5D823" box="[151,409,1511,1537]" pageId="1" pageNumber="57" type="description">
<paragraph id="900336F0C24DFFCB7CEC82FEE1B5D823" blockId="1.[151,1035,1474,1537]" box="[151,409,1511,1537]" pageId="1" pageNumber="57">
(
<figureCitation id="08872A75C24DFFCB7CE482FEE109D823" box="[159,293,1511,1537]" captionStart="FIGURE 1" captionStartId="2.[151,250,1846,1870]" captionTargetBox="[162,1425,190,1808]" captionTargetId="figure-17@2.[151,1436,179,1820]" captionTargetPageId="2" captionText="FIGURE 1. Ontherus araujoi new species. A, Ontherus araujoi (male, dorsal view); B, Ontherus araujoi (female, dorsal view); C, Ontherus araujoi (male, ventral view); D. Ontherus araujoi (male, lateral view)." figureDoi="http://doi.org/10.5281/zenodo.12189575" httpUri="https://zenodo.org/record/12189575/files/figure.png" pageId="1" pageNumber="57">Figs. 1A–D</figureCitation>
,
<figureCitation id="08872A75C24DFFCB7D4A82FEE15BD823" box="[305,375,1511,1537]" captionStart="FIGURE 2" captionStartId="3.[151,250,1732,1756]" captionTargetBox="[151,1436,192,1708]" captionTargetId="figure-26@3.[151,1436,181,1708]" captionTargetPageId="3" captionText="FIGURE 2. Ontherus diagnostic characters. A, Ontherus araujoi new species, aedeagus; B, Ontherus araujoi, endophallites. C, Ontherus araujoi, mesometasternal suture; D, Ontherus compressicornis Luederwaldt, 1931, mesometasternal suture." figureDoi="http://doi.org/10.5281/zenodo.12189577" httpUri="https://zenodo.org/record/12189577/files/figure.png" pageId="1" pageNumber="57">2A–C</figureCitation>
,
<figureCitation id="08872A75C24DFFCB7DF982FEE1BDD823" box="[386,401,1511,1537]" captionStart="FIGURE 4" captionStartId="8.[151,250,1889,1913]" captionTargetBox="[161,1426,191,1856]" captionTargetId="figure-17@8.[151,1436,181,1866]" captionTargetPageId="8" captionText="FIGURE 4. Distribution of Ontherus araujoi new species (red pentagon); Ontherus azteca Harold, 1869 (purple square); Ontherus edentulus Génier, 1996 (blue triangle); Ontherus pubens Génier, 1996 (green circle); and Cryptocanthon humidus Howden, 1973 (yellow octagon)." figureDoi="http://doi.org/10.5281/zenodo.12189581" httpUri="https://zenodo.org/record/12189581/files/figure.png" pageId="1" pageNumber="57">4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D8A6657BC24DFFCB7CEC8136E179D8FB" pageId="1" pageNumber="57" type="materials_examined">
<paragraph id="900336F0C24DFFCB7CEC8136E208D8B3" blockId="1.[151,1437,1583,2042]" pageId="1" pageNumber="57">
<materialsCitation id="20D43CADC24DFFCB7CEC8136E37DD84F" ID-GBIF-Occurrence="4898004321" collectingDate="2020-02-08" collectionCode="MEPN" collectorName="W. Chamorro" county="Male" elevation="1150" latitude="-4.0378723" location="Paltas" longLatPrecision="1" longitude="-79.80049" municipality="Catacocha" pageId="1" pageNumber="57" specimenCount="1" stateProvince="Loja" typeStatus="holotype">
<emphasis id="A2C8EAE2C24DFFCB7CEC8136E127D86B" bold="true" box="[151,267,1583,1609]" pageId="1" pageNumber="57">
<typeStatus id="4F078852C24DFFCB7CEC8136E12AD86B" box="[151,262,1583,1609]" pageId="1" pageNumber="57" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="79624E7CC24DFFCB7D6A8136E167D86B" box="[273,331,1583,1609]" pageId="1" pageNumber="57">Male</collectingCounty>
deposited in
<collectionCode id="F6ADAE35C24DFFCB7DA48129E21CD868" box="[479,560,1584,1610]" pageId="1" pageNumber="57">MEPN</collectionCode>
; labeled as: || ECU:
<collectingRegion id="5278F812C24DFFCB7F748136E374D86B" box="[783,856,1583,1609]" country="Ecuador" name="Loja" pageId="1" pageNumber="57">LOJA</collectingRegion>
:
<collectingMunicipality id="7067AC8AC24DFFCB7F198136E3FBD86B" box="[866,983,1583,1609]" pageId="1" pageNumber="57">Catacocha</collectingMunicipality>
|
<location id="9563602BC24DFFCB7F928136E403D86B" LSID="urn:lsid:plazi:treatment:181587E6C24DFFCE7CEC82DAE46DD89F:9563602BC24DFFCB7F928136E403D86B" box="[1001,1071,1583,1609]" county="Male" latitude="-4.0378723" longLatPrecision="1" longitude="-79.80049" municipality="Catacocha" name="Paltas" pageId="1" pageNumber="57" stateProvince="Loja">Paltas</location>
,
<geoCoordinate id="F5885037C24DFFCB78428136E4CAD86B" box="[1081,1254,1583,1609]" degrees="4" direction="south" minutes="02" orientation="latitude" pageId="1" pageNumber="57" precision="1" seconds="16.34" value="-4.0378723">4° 02’ 16.34’’ S</geoCoordinate>
|
<geoCoordinate id="F5885037C24DFFCB78838136E09ED84C" degrees="79" direction="west" minutes="48" orientation="longitude" pageId="1" pageNumber="57" precision="1" seconds="01.76" value="-79.80049">79° 48’ 01.76’’ W</geoCoordinate>
,
<quantity id="57449B15C24DFFCB7CC7814AE123D84C" box="[188,271,1619,1646]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="1" pageNumber="57" unit="m" value="1150.0">
<elevation id="1B91D1C3C24DFFCB7CC7814AE123D84C" box="[188,271,1619,1646]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="1" pageNumber="57" unit="m" value="1150.0">1150 m</elevation>
</quantity>
|
<date id="E4021030C24DFFCB7D5A814AE1A6D84F" box="[289,394,1619,1645]" pageId="1" pageNumber="57" value="2020-02-08">
<collectingDate id="F446E9D8C24DFFCB7D5A814AE1A6D84F" box="[289,394,1619,1645]" pageId="1" pageNumber="57" value="2020-02-08">8.II.2020</collectingDate>
</date>
,
<collectorName id="3D495326C24DFFCB7DEE814DE200D84F" box="[405,556,1619,1646]" pageId="1" pageNumber="57">W. Chamorro</collectorName>
|
<collectingMethod id="49FD4EE7C24DFFCB7E46814AE2ADD84F" box="[573,641,1619,1645]" pageId="1" pageNumber="57">Pitfall</collectingMethod>
human faeces ||; ||
</materialsCitation>
<typeStatus id="4F078852C24DFFCB7F23814AE3C0D84F" box="[856,1004,1619,1645]" pageId="1" pageNumber="57" type="holotype">HOLOTYPE</typeStatus>
<specimenCount id="86BAFD79C24DFFCB7F89814DE42BD84C" box="[1010,1031,1620,1646]" count="1" pageId="1" pageNumber="57" type="male">♂</specimenCount>
|
<taxonomicName id="57BC4D73C24DFFCB7862814AE4ACD84F" authorityName="Erichson" authorityYear="1847" box="[1049,1152,1619,1645]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="57" phylum="Arthropoda" rank="genus">Ontherus</taxonomicName>
| araujoi | dés. Chamorro, Lopera &amp; | Génier, 2024 || red card.
</paragraph>
<paragraph id="900336F0C24DFFCB7CBC8182E179D8FB" blockId="1.[151,1437,1583,2042]" pageId="1" pageNumber="57">
<materialsCitation id="20D43CADC24DFFCB7CBC8182E434D897" ID-GBIF-Occurrence="4898004315" box="[199,1048,1691,1718]" collectingDate="2020-02-08" collectionCode="MEPN" collectorName="W. Chamorro" county="Male" elevation="1150" latitude="-4.0378723" location="Paltas" longLatPrecision="1" longitude="-79.80049" municipality="Catacocha" pageId="1" pageNumber="57" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Loja" typeStatus="paratype">
<emphasis id="A2C8EAE2C24DFFCB7CBC8182E16AD897" bold="true" box="[199,326,1691,1717]" pageId="1" pageNumber="57">
<typeStatus id="4F078852C24DFFCB7CBC8182E16DD897" box="[199,321,1691,1717]" pageId="1" pageNumber="57" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="86BAFD79C24DFFCB7D368182E1BBD897" box="[333,407,1691,1717]" count="1" pageId="1" pageNumber="57" type="male">1 male</specimenCount>
and
<specimenCount id="86BAFD79C24DFFCB7DB68182E202D897" box="[461,558,1691,1717]" count="1" pageId="1" pageNumber="57" type="female">1 female</specimenCount>
deposited in
<collectionCode id="F6ADAE35C24DFFCB7EB88185E338D894" box="[707,788,1692,1718]" pageId="1" pageNumber="57">MEPN</collectionCode>
; same data as holotype
</materialsCitation>
.
<materialsCitation id="20D43CADC24DFFCB78598182E17DD8FB" ID-GBIF-Occurrence="4898004308" collectingDate="2020-02-08" collectionCode="CMNC" collectorName="W. Chamorro" county="Male" elevation="1150" latitude="-4.0378723" location="Paltas" longLatPrecision="1" longitude="-79.80049" municipality="Catacocha" pageId="1" pageNumber="57" specimenCount="1" specimenCount-male="1" stateProvince="Loja" typeStatus="paratype">
<specimenCount id="86BAFD79C24DFFCB78598182E441D897" box="[1058,1133,1691,1717]" count="1" pageId="1" pageNumber="57" type="male">1 male</specimenCount>
deposited in
<collectionCode id="F6ADAE35C24DFFCB79798182E574D897" box="[1282,1368,1691,1717]" pageId="1" pageNumber="57">CMNC</collectionCode>
; same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D8A6657BC24DFFCB7CBC81FAE0C1D967" pageId="1" pageNumber="57" type="etymology">
<paragraph id="900336F0C24DFFCB7CBC81FAE0C1D967" blockId="1.[151,1437,1583,2042]" pageId="1" pageNumber="57">
<emphasis id="A2C8EAE2C24DFFCB7CBC81FAE164D8DF" bold="true" box="[199,328,1763,1789]" pageId="1" pageNumber="57">Etymology</emphasis>
. The name
<taxonomicName id="57BC4D73C24DFFCB7DA581FAE211D8DF" authorityName="Chamorro, Lopera, &amp; Genier" authorityYear="2024" box="[478,573,1763,1789]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="57" phylum="Arthropoda" rank="species" species="araujoi">
“
<emphasis id="A2C8EAE2C24DFFCB7D9081FDE211D8DF" box="[491,573,1764,1789]" italics="true" pageId="1" pageNumber="57">araujoi</emphasis>
</taxonomicName>
” is an eponym for Pablo Araujo, biologist-entomologist, founder of the “Entomological section Escuela Politécnica Nacional” (Quito
<collectingCountry id="E8AB7660C24DFFCB7F02801EE3F4D903" box="[889,984,1799,1825]" name="Ecuador" pageId="1" pageNumber="57">Ecuador</collectingCountry>
) and a great researcher of
<taxonomicName id="57BC4D73C24DFFCB7950801EE5B1D903" authorityName="Latreille" authorityYear="1802" box="[1323,1437,1799,1825]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="57" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
beetles.
</paragraph>
</subSubSection>
<subSubSection id="D8A6657BC24DFFCB7CBC8056E5BCD9DB" pageId="1" pageNumber="57" type="diagnosis">
<paragraph id="900336F0C24DFFCB7CBC8056E5BCD9DB" blockId="1.[151,1437,1583,2042]" pageId="1" pageNumber="57">
<emphasis id="A2C8EAE2C24DFFCB7CBC8056E113D94B" bold="true" box="[199,319,1871,1897]" pageId="1" pageNumber="57">Diagnosis.</emphasis>
The male of
<taxonomicName id="57BC4D73C24DFFCB7DAE8049E264D94B" authorityName="Chamorro, Lopera, &amp; Genier" authorityYear="2024" box="[469,584,1872,1898]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="57" phylum="Arthropoda" rank="species" species="araujoi">
<emphasis id="A2C8EAE2C24DFFCB7DAE8049E264D94B" box="[469,584,1872,1898]" italics="true" pageId="1" pageNumber="57">O. araujoi</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="57BC4D73C24DFFCB7EA08056E404D94B" authority="Harold, 1869" authorityName="Harold" authorityYear="1869" box="[731,1064,1871,1897]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="57" phylum="Arthropoda" rank="species" species="azteca">
<emphasis id="A2C8EAE2C24DFFCB7EA08056E3A3D94B" box="[731,911,1871,1897]" italics="true" pageId="1" pageNumber="57">Ontherus azteca</emphasis>
Harold, 1869
</taxonomicName>
and
<taxonomicName id="57BC4D73C24DFFCB78248056E0FCD9AF" authority="Genier, 1996" authorityName="Genier" authorityYear="1996" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="57" phylum="Arthropoda" rank="species" species="cambeforti">
<emphasis id="A2C8EAE2C24DFFCB78248056E568D94B" box="[1119,1348,1871,1897]" italics="true" pageId="1" pageNumber="57">Ontherus cambeforti</emphasis>
<bibRefCitation id="F42D4B01C24DFFCB79378056E0FCD9AF" author="Genier, F." pageId="1" pageNumber="57" pagination="1 - 169" refId="ref6799" refString="Genier, F. (1996) A revision of the Neotropical genus Ontherus Erichson (Coleoptera: Scarabaeidae, Scarabaeinae). Memoirs of the Entomological Society of Canada, 170, 1 - 169. https: // doi. org / 10.4039 / entm 128170 fv" type="journal article" year="1996">Génier, 1996</bibRefCitation>
</taxonomicName>
, but it differs by the following characteristics: (1) apical tooth directed forward (anteriorly) and with the apex weakly flexed downward (
<figureCitation id="08872A75C24DFFCB7DBB808EE26AD993" box="[448,582,1943,1969]" captionStart="FIGURE 1" captionStartId="2.[151,250,1846,1870]" captionTargetBox="[162,1425,190,1808]" captionTargetId="figure-17@2.[151,1436,179,1820]" captionTargetPageId="2" captionText="FIGURE 1. Ontherus araujoi new species. A, Ontherus araujoi (male, dorsal view); B, Ontherus araujoi (female, dorsal view); C, Ontherus araujoi (male, ventral view); D. Ontherus araujoi (male, lateral view)." figureDoi="http://doi.org/10.5281/zenodo.12189575" httpUri="https://zenodo.org/record/12189575/files/figure.png" pageId="1" pageNumber="57">Figs. 1A, C</figureCitation>
); (2) protibial spur absent (
<figureCitation id="08872A75C24DFFCB7F02808EE3D2D993" box="[889,1022,1943,1969]" captionStart="FIGURE 1" captionStartId="2.[151,250,1846,1870]" captionTargetBox="[162,1425,190,1808]" captionTargetId="figure-17@2.[151,1436,179,1820]" captionTargetPageId="2" captionText="FIGURE 1. Ontherus araujoi new species. A, Ontherus araujoi (male, dorsal view); B, Ontherus araujoi (female, dorsal view); C, Ontherus araujoi (male, ventral view); D. Ontherus araujoi (male, lateral view)." figureDoi="http://doi.org/10.5281/zenodo.12189575" httpUri="https://zenodo.org/record/12189575/files/figure.png" pageId="1" pageNumber="57">Figs. 1A, C</figureCitation>
); (3) metafemurs with posteromedial triangular process (
<figureCitation id="08872A75C24DFFCB7D1D80A2E228D9F7" box="[358,516,1979,2005]" captionStart="FIGURE 1" captionStartId="2.[151,250,1846,1870]" captionTargetBox="[162,1425,190,1808]" captionTargetId="figure-17@2.[151,1436,179,1820]" captionTargetPageId="2" captionText="FIGURE 1. Ontherus araujoi new species. A, Ontherus araujoi (male, dorsal view); B, Ontherus araujoi (female, dorsal view); C, Ontherus araujoi (male, ventral view); D. Ontherus araujoi (male, lateral view)." figureDoi="http://doi.org/10.5281/zenodo.12189575" httpUri="https://zenodo.org/record/12189575/files/figure.png" pageId="1" pageNumber="57">Figs. 1A, C, D</figureCitation>
), never on posterodorsal edge as
<taxonomicName id="57BC4D73C24DFFCB7F1F80A5E3E6D9F7" authorityName="Harold" authorityYear="1869" box="[868,970,1980,2006]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="57" phylum="Arthropoda" rank="species" species="azteca">
<emphasis id="A2C8EAE2C24DFFCB7F1F80A5E3E6D9F7" box="[868,970,1980,2006]" italics="true" pageId="1" pageNumber="57">O. azteca</emphasis>
</taxonomicName>
(
<figureCitation id="08872A75C24DFFCB7FAC80A2E406D9F7" box="[983,1066,1979,2005]" captionStart="FIGURE 3" captionStartId="7.[151,250,1909,1933]" captionTargetBox="[164,1424,193,1874]" captionTargetId="figure-26@7.[151,1436,181,1886]" captionTargetPageId="7" captionText="FIGURE 3. Ontherus and Cryptocanthon species. A, Ontherus azteca Harold, 1869, male in dorsal view; B, Ontherus edentulus Génier, 1996, female in dorsal view; C. Ontherus pubens Génier, 1996, male in dorsal view; D, Cryptocanthon humidus Howden, 1973, male in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12189579" httpUri="https://zenodo.org/record/12189579/files/figure.png" pageId="1" pageNumber="57">Fig. 3A</figureCitation>
) and
<taxonomicName id="57BC4D73C24DFFCB781F80A5E4D6D9F7" authorityName="Genier" authorityYear="1996" box="[1124,1274,1979,2006]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="57" phylum="Arthropoda" rank="species" species="cambeforti">
<emphasis id="A2C8EAE2C24DFFCB781F80A5E4D6D9F7" box="[1124,1274,1979,2006]" italics="true" pageId="1" pageNumber="57">O. cambeforti</emphasis>
</taxonomicName>
; (4) parameres (in lateral view) with rounded apices (
<figureCitation id="08872A75C24DFFCB7E4280C6E2BED9DB" box="[569,658,2015,2041]" captionStart="FIGURE 2" captionStartId="3.[151,250,1732,1756]" captionTargetBox="[151,1436,192,1708]" captionTargetId="figure-26@3.[151,1436,181,1708]" captionTargetPageId="3" captionText="FIGURE 2. Ontherus diagnostic characters. A, Ontherus araujoi new species, aedeagus; B, Ontherus araujoi, endophallites. C, Ontherus araujoi, mesometasternal suture; D, Ontherus compressicornis Luederwaldt, 1931, mesometasternal suture." figureDoi="http://doi.org/10.5281/zenodo.12189577" httpUri="https://zenodo.org/record/12189577/files/figure.png" pageId="1" pageNumber="57">Fig. 2A</figureCitation>
); (5) frontolateral peripheral endophallite triangular shaped (
<figureCitation id="08872A75C24DFFCB795780C6E5AFD9DB" box="[1324,1411,2015,2041]" captionStart="FIGURE 2" captionStartId="3.[151,250,1732,1756]" captionTargetBox="[151,1436,192,1708]" captionTargetId="figure-26@3.[151,1436,181,1708]" captionTargetPageId="3" captionText="FIGURE 2. Ontherus diagnostic characters. A, Ontherus araujoi new species, aedeagus; B, Ontherus araujoi, endophallites. C, Ontherus araujoi, mesometasternal suture; D, Ontherus compressicornis Luederwaldt, 1931, mesometasternal suture." figureDoi="http://doi.org/10.5281/zenodo.12189577" httpUri="https://zenodo.org/record/12189577/files/figure.png" pageId="1" pageNumber="57">Fig. 2B</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="C4C36678C24EFFC87CEC802FE3CBD950" ID-DOI="http://doi.org/10.5281/zenodo.12189575" ID-Zenodo-Dep="12189575" httpUri="https://zenodo.org/record/12189575/files/figure.png" pageId="2" pageNumber="58" startId="2.[151,250,1846,1870]" targetBox="[162,1425,190,1808]" targetPageId="2" targetType="figure">
<paragraph id="900336F0C24EFFC87CEC802FE3CBD950" blockId="2.[151,1436,1846,1906]" pageId="2" pageNumber="58">
<emphasis id="A2C8EAE2C24EFFC87CEC802FE13ED96C" bold="true" box="[151,274,1846,1870]" pageId="2" pageNumber="58">FIGURE 1.</emphasis>
<taxonomicName id="57BC4D73C24EFFC87D6C802FE1E9D96C" authority="Chamorro &amp; Lopera-Toro &amp; Génier, 2024" authorityName="Chamorro &amp; Lopera-Toro &amp; Génier" authorityYear="2024" box="[279,453,1846,1870]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="58" phylum="Arthropoda" rank="species" species="araujoi" status="sp. nov.">
<emphasis id="A2C8EAE2C24EFFC87D6C802FE1E9D96C" box="[279,453,1846,1870]" italics="true" pageId="2" pageNumber="58">Ontherus araujoi</emphasis>
</taxonomicName>
<emphasis id="A2C8EAE2C24EFFC87DB1802FE26AD96C" bold="true" box="[458,582,1846,1870]" pageId="2" pageNumber="58">
<taxonomicNameLabel id="B9FB5799C24EFFC87DB1802FE26AD96C" box="[458,582,1846,1870]" pageId="2" pageNumber="58" rank="species">new species</taxonomicNameLabel>
</emphasis>
.
<emphasis id="A2C8EAE2C24EFFC87E2A802FE24FD96C" bold="true" box="[593,611,1846,1870]" pageId="2" pageNumber="58">A</emphasis>
,
<taxonomicName id="57BC4D73C24EFFC87E14802FE331D96C" authorityName="Chamorro &amp; Lopera-Toro &amp; Génier" authorityYear="2024" box="[623,797,1846,1870]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="58" phylum="Arthropoda" rank="species" species="araujoi">
<emphasis id="A2C8EAE2C24EFFC87E14802FE331D96C" box="[623,797,1846,1870]" italics="true" pageId="2" pageNumber="58">Ontherus araujoi</emphasis>
</taxonomicName>
(male, dorsal view);
<emphasis id="A2C8EAE2C24EFFC87F8B802FE42DD96C" bold="true" box="[1008,1025,1846,1870]" pageId="2" pageNumber="58">B</emphasis>
,
<taxonomicName id="57BC4D73C24EFFC87877802FE497D96C" authorityName="Chamorro &amp; Lopera-Toro &amp; Génier" authorityYear="2024" box="[1036,1211,1846,1870]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="58" phylum="Arthropoda" rank="species" species="araujoi">
<emphasis id="A2C8EAE2C24EFFC87877802FE497D96C" box="[1036,1211,1846,1870]" italics="true" pageId="2" pageNumber="58">Ontherus araujoi</emphasis>
</taxonomicName>
(female, dorsal view);
<emphasis id="A2C8EAE2C24EFFC87CEC8043E085D950" bold="true" box="[151,169,1882,1906]" pageId="2" pageNumber="58">C</emphasis>
,
<taxonomicName id="57BC4D73C24EFFC87CCD8043E14AD950" authorityName="Chamorro &amp; Lopera-Toro &amp; Génier" authorityYear="2024" box="[182,358,1882,1906]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="58" phylum="Arthropoda" rank="species" species="araujoi">
<emphasis id="A2C8EAE2C24EFFC87CCD8043E14AD950" box="[182,358,1882,1906]" italics="true" pageId="2" pageNumber="58">Ontherus araujoi</emphasis>
</taxonomicName>
(male, ventral view);
<emphasis id="A2C8EAE2C24EFFC87E3D8043E274D950" bold="true" box="[582,600,1882,1906]" pageId="2" pageNumber="58">D</emphasis>
.
<taxonomicName id="57BC4D73C24EFFC87E1E8043E338D950" authorityName="Chamorro &amp; Lopera-Toro &amp; Génier" authorityYear="2024" box="[613,788,1882,1906]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="58" phylum="Arthropoda" rank="species" species="araujoi">
<emphasis id="A2C8EAE2C24EFFC87E1E8043E338D950" box="[613,788,1882,1906]" italics="true" pageId="2" pageNumber="58">Ontherus araujoi</emphasis>
</taxonomicName>
(male, lateral view).
</paragraph>
</caption>
<caption id="C4C36678C24FFFC97CEC81DDE56CD922" ID-DOI="http://doi.org/10.5281/zenodo.12189577" ID-Zenodo-Dep="12189577" httpUri="https://zenodo.org/record/12189577/files/figure.png" pageId="3" pageNumber="59" startId="3.[151,250,1732,1756]" targetBox="[151,1436,192,1708]" targetPageId="3" targetType="figure">
<paragraph id="900336F0C24FFFC97CEC81DDE56CD922" blockId="3.[151,1436,1732,1793]" pageId="3" pageNumber="59">
<emphasis id="A2C8EAE2C24FFFC97CEC81DDE13ED8FE" bold="true" box="[151,274,1732,1756]" pageId="3" pageNumber="59">FIGURE 2.</emphasis>
<taxonomicName id="57BC4D73C24FFFC97D6C81DDE159D8FE" authorityName="Erichson" authorityYear="1847" box="[279,373,1732,1756]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="A2C8EAE2C24FFFC97D6C81DDE159D8FE" box="[279,373,1732,1756]" italics="true" pageId="3" pageNumber="59">Ontherus</emphasis>
</taxonomicName>
diagnostic characters.
<emphasis id="A2C8EAE2C24FFFC97E2181DDE240D8FE" bold="true" box="[602,620,1732,1756]" pageId="3" pageNumber="59">A</emphasis>
,
<taxonomicName id="57BC4D73C24FFFC97E0381DDE30AD8FE" authority="Chamorro &amp; Lopera-Toro &amp; Génier, 2024" authorityName="Chamorro &amp; Lopera-Toro &amp; Génier" authorityYear="2024" box="[632,806,1732,1756]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="59" phylum="Arthropoda" rank="species" species="araujoi" status="sp. nov.">
<emphasis id="A2C8EAE2C24FFFC97E0381DDE30AD8FE" box="[632,806,1732,1756]" italics="true" pageId="3" pageNumber="59">Ontherus araujoi</emphasis>
</taxonomicName>
<emphasis id="A2C8EAE2C24FFFC97F5081DCE38BD8FE" bold="true" box="[811,935,1732,1757]" pageId="3" pageNumber="59">
<taxonomicNameLabel id="B9FB5799C24FFFC97F5081DCE38BD8FE" box="[811,935,1732,1757]" pageId="3" pageNumber="59" rank="species">new species</taxonomicNameLabel>
</emphasis>
, aedeagus;
<emphasis id="A2C8EAE2C24FFFC9786081DCE400D8FF" bold="true" box="[1051,1068,1733,1757]" pageId="3" pageNumber="59">B</emphasis>
,
<taxonomicName id="57BC4D73C24FFFC9784C81DDE4CAD8FE" authorityName="Chamorro &amp; Lopera-Toro &amp; Génier" authorityYear="2024" box="[1079,1254,1732,1756]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="59" phylum="Arthropoda" rank="species" species="araujoi">
<emphasis id="A2C8EAE2C24FFFC9784C81DDE4CAD8FE" box="[1079,1254,1732,1756]" italics="true" pageId="3" pageNumber="59">Ontherus araujoi</emphasis>
</taxonomicName>
, endophallites.
<emphasis id="A2C8EAE2C24FFFC979FF81DDE5BAD8FE" bold="true" box="[1412,1430,1732,1756]" pageId="3" pageNumber="59">C</emphasis>
,
<taxonomicName id="57BC4D73C24FFFC97CEC81F1E16BD922" authorityName="Chamorro &amp; Lopera-Toro &amp; Génier" authorityYear="2024" box="[151,327,1768,1792]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="59" phylum="Arthropoda" rank="species" species="araujoi">
<emphasis id="A2C8EAE2C24FFFC97CEC81F1E16BD922" box="[151,327,1768,1792]" italics="true" pageId="3" pageNumber="59">Ontherus araujoi</emphasis>
</taxonomicName>
, mesometasternal suture;
<emphasis id="A2C8EAE2C24FFFC97E3481F0E24DD923" bold="true" box="[591,609,1769,1793]" pageId="3" pageNumber="59">D</emphasis>
,
<taxonomicName id="57BC4D73C24FFFC97E1581F1E46CD923" authority="Luederwaldt, 1931" authorityName="Luederwaldt" authorityYear="1931" box="[622,1088,1768,1793]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="59" phylum="Arthropoda" rank="species" species="compressicornis">
<emphasis id="A2C8EAE2C24FFFC97E1581F1E354D922" box="[622,888,1768,1792]" italics="true" pageId="3" pageNumber="59">Ontherus compressicornis</emphasis>
Luederwaldt, 1931
</taxonomicName>
, mesometasternal suture.
</paragraph>
</caption>
<subSubSection id="D8A6657BC24FFFCE7CBC8053E14BDBE3" lastPageId="4" lastPageNumber="60" pageId="3" pageNumber="59" type="description">
<paragraph id="900336F0C24FFFCE7CBC8053E18CDB77" blockId="3.[151,1437,1866,2037]" lastBlockId="4.[151,1437,151,1725]" lastPageId="4" lastPageNumber="60" pageId="3" pageNumber="59">
<emphasis id="A2C8EAE2C24FFFC97CBC8053E175D946" bold="true" box="[199,345,1866,1892]" pageId="3" pageNumber="59">Description.</emphasis>
Male (
<figureCitation id="08872A75C24FFFC97DD78052E22BD947" box="[428,519,1867,1893]" captionStart="FIGURE 1" captionStartId="2.[151,250,1846,1870]" captionTargetBox="[162,1425,190,1808]" captionTargetId="figure-17@2.[151,1436,179,1820]" captionTargetPageId="2" captionText="FIGURE 1. Ontherus araujoi new species. A, Ontherus araujoi (male, dorsal view); B, Ontherus araujoi (female, dorsal view); C, Ontherus araujoi (male, ventral view); D. Ontherus araujoi (male, lateral view)." figureDoi="http://doi.org/10.5281/zenodo.12189575" httpUri="https://zenodo.org/record/12189575/files/figure.png" pageId="3" pageNumber="59">Fig. 1A</figureCitation>
): length:
<quantity id="57449B15C24FFFC97E0E8052E337D947" box="[629,795,1867,1893]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" metricValueMax="1.22" metricValueMin="1.1199999999999999" pageId="3" pageNumber="59" unit="mm" value="11.7" valueMax="12.2" valueMin="11.2">11.2–12.2 mm</quantity>
, width 5.7–6.0 mm. Female (
<figureCitation id="08872A75C24FFFC978168052E4E9D947" box="[1133,1221,1867,1893]" captionStart="FIGURE 1" captionStartId="2.[151,250,1846,1870]" captionTargetBox="[162,1425,190,1808]" captionTargetId="figure-17@2.[151,1436,179,1820]" captionTargetPageId="2" captionText="FIGURE 1. Ontherus araujoi new species. A, Ontherus araujoi (male, dorsal view); B, Ontherus araujoi (female, dorsal view); C, Ontherus araujoi (male, ventral view); D. Ontherus araujoi (male, lateral view)." figureDoi="http://doi.org/10.5281/zenodo.12189575" httpUri="https://zenodo.org/record/12189575/files/figure.png" pageId="3" pageNumber="59">Fig. 1B</figureCitation>
): length
<quantity id="57449B15C24FFFC979578053E0E9D9AB" metricMagnitude="-2" metricUnit="m" metricValue="1.185" metricValueMax="1.3199999999999998" metricValueMin="1.05" pageId="3" pageNumber="59" unit="mm" value="11.85" valueMax="13.2" valueMin="10.5">10.5–13.2 mm</quantity>
, width: 5.0–6.0 mm.
<emphasis id="A2C8EAE2C24FFFC97DB88077E23BD9AA" bold="true" box="[451,535,1902,1928]" pageId="3" pageNumber="59">Colour</emphasis>
. Body dark reddish brown, pubescence fulvous, length
<quantity id="57449B15C24FFFC978DE8077E562D9AB" box="[1189,1358,1902,1929]" metricMagnitude="-2" metricUnit="m" metricValue="1.185" metricValueMax="1.3199999999999998" metricValueMin="1.05" pageId="3" pageNumber="59" unit="mm" value="11.85" valueMax="13.2" valueMin="10.5">10.5–13.2 mm</quantity>
.
<emphasis id="A2C8EAE2C24FFFC979278076E5B0D9AB" bold="true" box="[1372,1436,1903,1929]" pageId="3" pageNumber="59">Head</emphasis>
(
<figureCitation id="08872A75C24FFFC97CE4808AE130D98F" box="[159,284,1939,1965]" captionStart="FIGURE 1" captionStartId="2.[151,250,1846,1870]" captionTargetBox="[162,1425,190,1808]" captionTargetId="figure-17@2.[151,1436,179,1820]" captionTargetPageId="2" captionText="FIGURE 1. Ontherus araujoi new species. A, Ontherus araujoi (male, dorsal view); B, Ontherus araujoi (female, dorsal view); C, Ontherus araujoi (male, ventral view); D. Ontherus araujoi (male, lateral view)." figureDoi="http://doi.org/10.5281/zenodo.12189575" httpUri="https://zenodo.org/record/12189575/files/figure.png" pageId="3" pageNumber="59">Fig. 1A, C</figureCitation>
). Anterior edge arcuate, weakly upturned, slightly emarginate medially, clypeogenal junction straight. Clypeus transversely wrinkled on anterior half, either with confluent and feeble punctures on posterior half. Clypeofrontal suture arcuate, feebly carinate, with small and rounded conical process medially. Genal surface with punctures confluent throughout. Vertex finely and shallowly punctate on anterior two-thirds, posterior third smooth.
<emphasis id="A2C8EAE2C248FFCE7CEC87A2E13CDEF7" bold="true" box="[151,272,187,213]" pageId="4" pageNumber="60">Pronotum</emphasis>
(
<figureCitation id="08872A75C248FFCE7D5E87A2E15BDEF7" box="[293,375,187,213]" captionStart="FIGURE 1" captionStartId="2.[151,250,1846,1870]" captionTargetBox="[162,1425,190,1808]" captionTargetId="figure-17@2.[151,1436,179,1820]" captionTargetPageId="2" captionText="FIGURE 1. Ontherus araujoi new species. A, Ontherus araujoi (male, dorsal view); B, Ontherus araujoi (female, dorsal view); C, Ontherus araujoi (male, ventral view); D. Ontherus araujoi (male, lateral view)." figureDoi="http://doi.org/10.5281/zenodo.12189575" httpUri="https://zenodo.org/record/12189575/files/figure.png" pageId="4" pageNumber="60">Fig.1A</figureCitation>
). Transverse wider than long W/L = 2.0; anterior edge narrowly membranous and almost straight behind head insertion. Disc slightly declivous on anterior fifth behind head insertion, and almost convex in lateral view; punctures rounded, moderate in size on disc and lateral swellings, slightly larger and denser along anterior and lateral declivities, not confluent on anterior angles.
<emphasis id="A2C8EAE2C248FFCE7F15863EE396DF63" bold="true" box="[878,954,295,321]" pageId="4" pageNumber="60">Elytra</emphasis>
(
<figureCitation id="08872A75C248FFCE7FB0863EE409DF63" box="[971,1061,295,321]" captionStart="FIGURE 1" captionStartId="2.[151,250,1846,1870]" captionTargetBox="[162,1425,190,1808]" captionTargetId="figure-17@2.[151,1436,179,1820]" captionTargetPageId="2" captionText="FIGURE 1. Ontherus araujoi new species. A, Ontherus araujoi (male, dorsal view); B, Ontherus araujoi (female, dorsal view); C, Ontherus araujoi (male, ventral view); D. Ontherus araujoi (male, lateral view)." figureDoi="http://doi.org/10.5281/zenodo.12189575" httpUri="https://zenodo.org/record/12189575/files/figure.png" pageId="4" pageNumber="60">Fig. 1A</figureCitation>
). Subquadrate, L/W = 1.2. Striae moderately wide and impressed on disc and apical declivity, punctures rounded, deeper than stria, approximately 1.2 times as wide as stria and separated by 1–2 diameters on disc and apical declivity, slightly encroaching on intervals. Interstriae slightly convex, surface feebly punctate, with a trace of alutaceous microsculpture.
<emphasis id="A2C8EAE2C248FFCE794A868BE0CEDFF3" bold="true" pageId="4" pageNumber="60">Thoracic sterna</emphasis>
(
<figureCitation id="08872A75C248FFCE7C8886AEE166DFF3" box="[243,330,439,465]" captionStart="FIGURE 1" captionStartId="2.[151,250,1846,1870]" captionTargetBox="[162,1425,190,1808]" captionTargetId="figure-17@2.[151,1436,179,1820]" captionTargetPageId="2" captionText="FIGURE 1. Ontherus araujoi new species. A, Ontherus araujoi (male, dorsal view); B, Ontherus araujoi (female, dorsal view); C, Ontherus araujoi (male, ventral view); D. Ontherus araujoi (male, lateral view)." figureDoi="http://doi.org/10.5281/zenodo.12189575" httpUri="https://zenodo.org/record/12189575/files/figure.png" pageId="4" pageNumber="60">Fig. 1C</figureCitation>
). Proepisternum with posterolateral portion densely pubescent and finely alutaceous throughout. Prosternum behind procoxa finely pubescent, setae apposed, with a brush of longer setae medially on posterior edge. Mesosternum (
<figureCitation id="08872A75C248FFCE7DF386E6E1C1DC3B" box="[392,493,511,537]" captionStart="FIGURE 1" captionStartId="2.[151,250,1846,1870]" captionTargetBox="[162,1425,190,1808]" captionTargetId="figure-17@2.[151,1436,179,1820]" captionTargetPageId="2" captionText="FIGURE 1. Ontherus araujoi new species. A, Ontherus araujoi (male, dorsal view); B, Ontherus araujoi (female, dorsal view); C, Ontherus araujoi (male, ventral view); D. Ontherus araujoi (male, lateral view)." figureDoi="http://doi.org/10.5281/zenodo.12189575" httpUri="https://zenodo.org/record/12189575/files/figure.png" pageId="4" pageNumber="60">Figs. 1C</figureCitation>
,
<figureCitation id="08872A75C248FFCE7D8686E6E232DC3B" box="[509,542,511,537]" captionStart="FIGURE 2" captionStartId="3.[151,250,1732,1756]" captionTargetBox="[151,1436,192,1708]" captionTargetId="figure-26@3.[151,1436,181,1708]" captionTargetPageId="3" captionText="FIGURE 2. Ontherus diagnostic characters. A, Ontherus araujoi new species, aedeagus; B, Ontherus araujoi, endophallites. C, Ontherus araujoi, mesometasternal suture; D, Ontherus compressicornis Luederwaldt, 1931, mesometasternal suture." figureDoi="http://doi.org/10.5281/zenodo.12189577" httpUri="https://zenodo.org/record/12189577/files/figure.png" pageId="4" pageNumber="60">2C</figureCitation>
) with feeble and alutaceous microsculpture, surface slightly and transversely impressed on disc, with rounded punctures moderate in size, coarse and dense throughout, pubescence fine and short throughout. Mesepisternum regularly punctate, punctures of different sizes, rounded and umbilicate, surface finely alutaceous, strongly and sharply carinate parallel to posterior edge. Metasternal disc with fine and rounded punctures, surface widely and somewhat deeply sulcate; lateral lobes on anterior half with punctures rounded and umbilicate, posterior half with punctures rounded, surface weakly alutaceous on anterior half, pubescence long and obliquely oriented; median lobe of mesosternon proportion (length/medial width) = 1.6, lateral margins almost parallel on anterior half, slightly convergent on posterior half, lateral margins narrow throughout and anterior angles of lateral margins rounded internally; surface finely punctate, anterior-most punctures slightly larger and finely setiferous; surface between punctures finely alutaceous anteriorly; junction with mesosternum angulate medially, finely carinate throughout, not produced into short carina longitudinally on mesosternum.
<emphasis id="A2C8EAE2C248FFCE780F847EE480DDA3" bold="true" box="[1140,1196,871,897]" pageId="4" pageNumber="60">Legs</emphasis>
(
<figureCitation id="08872A75C248FFCE78C2847EE51EDDA3" box="[1209,1330,871,897]" captionStart="FIGURE 1" captionStartId="2.[151,250,1846,1870]" captionTargetBox="[162,1425,190,1808]" captionTargetId="figure-17@2.[151,1436,179,1820]" captionTargetPageId="2" captionText="FIGURE 1. Ontherus araujoi new species. A, Ontherus araujoi (male, dorsal view); B, Ontherus araujoi (female, dorsal view); C, Ontherus araujoi (male, ventral view); D. Ontherus araujoi (male, lateral view)." figureDoi="http://doi.org/10.5281/zenodo.12189575" httpUri="https://zenodo.org/record/12189575/files/figure.png" pageId="4" pageNumber="60">Fig. 1A–C</figureCitation>
). Protibia quadridentate, apical tooth projecting anteriorly and with the apex weakly flexed down, unmodified in females; ventromedial carina lacking intervening setae on basal third in both sexes; apical spur absents in males, in females recurved and weakly bent down apically; metafemurs (
<figureCitation id="08872A75C248FFCE7E8684CAE3ACDDCF" box="[765,896,979,1005]" captionStart="FIGURE 1" captionStartId="2.[151,250,1846,1870]" captionTargetBox="[162,1425,190,1808]" captionTargetId="figure-17@2.[151,1436,179,1820]" captionTargetPageId="2" captionText="FIGURE 1. Ontherus araujoi new species. A, Ontherus araujoi (male, dorsal view); B, Ontherus araujoi (female, dorsal view); C, Ontherus araujoi (male, ventral view); D. Ontherus araujoi (male, lateral view)." figureDoi="http://doi.org/10.5281/zenodo.12189575" httpUri="https://zenodo.org/record/12189575/files/figure.png" pageId="4" pageNumber="60">Figs. 1A, C</figureCitation>
) with triangular projection on posterodorsal and distal edge, posteroventral surface sulcate, sulcus extending anteriorly at the apex and apicoventral surface with shallow deep impression in males, absent in females. Metatibiae (
<figureCitation id="08872A75C248FFCE7F068302E436DA17" box="[893,1050,1051,1077]" captionStart="FIGURE 1" captionStartId="2.[151,250,1846,1870]" captionTargetBox="[162,1425,190,1808]" captionTargetId="figure-17@2.[151,1436,179,1820]" captionTargetPageId="2" captionText="FIGURE 1. Ontherus araujoi new species. A, Ontherus araujoi (male, dorsal view); B, Ontherus araujoi (female, dorsal view); C, Ontherus araujoi (male, ventral view); D. Ontherus araujoi (male, lateral view)." figureDoi="http://doi.org/10.5281/zenodo.12189575" httpUri="https://zenodo.org/record/12189575/files/figure.png" pageId="4" pageNumber="60">Fig. 1A, C–D</figureCitation>
) with internal edge slightly lobate on basal half in males, simple in females.
<emphasis id="A2C8EAE2C248FFCE7E128327E2F2DA7A" bold="true" box="[617,734,1086,1112]" pageId="4" pageNumber="60">Abdomen</emphasis>
(
<figureCitation id="08872A75C248FFCE7E968326E368DA7B" box="[749,836,1087,1113]" captionStart="FIGURE 1" captionStartId="2.[151,250,1846,1870]" captionTargetBox="[162,1425,190,1808]" captionTargetId="figure-17@2.[151,1436,179,1820]" captionTargetPageId="2" captionText="FIGURE 1. Ontherus araujoi new species. A, Ontherus araujoi (male, dorsal view); B, Ontherus araujoi (female, dorsal view); C, Ontherus araujoi (male, ventral view); D. Ontherus araujoi (male, lateral view)." figureDoi="http://doi.org/10.5281/zenodo.12189575" httpUri="https://zenodo.org/record/12189575/files/figure.png" pageId="4" pageNumber="60">Fig. 1C</figureCitation>
). Sternites 2–5 punctate laterally, anterior edge with a row of setiferous punctures laterally, the suture between sternites 5–6 deeply and sharply sulcate laterally.
<emphasis id="A2C8EAE2C248FFCE7952837BE5B0DA5E" bold="true" box="[1321,1436,1122,1148]" pageId="4" pageNumber="60">Pygidium</emphasis>
(
<figureCitation id="08872A75C248FFCE7CE4839EE131DA83" box="[159,285,1159,1185]" captionStart="FIGURE 1" captionStartId="2.[151,250,1846,1870]" captionTargetBox="[162,1425,190,1808]" captionTargetId="figure-17@2.[151,1436,179,1820]" captionTargetPageId="2" captionText="FIGURE 1. Ontherus araujoi new species. A, Ontherus araujoi (male, dorsal view); B, Ontherus araujoi (female, dorsal view); C, Ontherus araujoi (male, ventral view); D. Ontherus araujoi (male, lateral view)." figureDoi="http://doi.org/10.5281/zenodo.12189575" httpUri="https://zenodo.org/record/12189575/files/figure.png" pageId="4" pageNumber="60">Fig. 1C–D</figureCitation>
). width/length = 2.0 in males or
<quantity id="57449B15C248FFCE7EE6839EE2CCDA83" box="[669,736,1159,1185]" metricMagnitude="-2" metricUnit="m" metricValue="5.334" pageId="4" pageNumber="60" unit="in" value="2.1">2.1 in</quantity>
females, punctures small, evenly distributed, surface glossy; apical margin clearly delimited in males, basal portion slightly sulcate along margin medially.
<emphasis id="A2C8EAE2C248FFCE789583B2E5B1DAE6" bold="true" box="[1262,1437,1194,1221]" pageId="4" pageNumber="60">Male genitalia</emphasis>
(
<figureCitation id="08872A75C248FFCE7CE483D6E0D6DACB" box="[159,250,1231,1257]" captionStart="FIGURE 2" captionStartId="3.[151,250,1732,1756]" captionTargetBox="[151,1436,192,1708]" captionTargetId="figure-26@3.[151,1436,181,1708]" captionTargetPageId="3" captionText="FIGURE 2. Ontherus diagnostic characters. A, Ontherus araujoi new species, aedeagus; B, Ontherus araujoi, endophallites. C, Ontherus araujoi, mesometasternal suture; D, Ontherus compressicornis Luederwaldt, 1931, mesometasternal suture." figureDoi="http://doi.org/10.5281/zenodo.12189577" httpUri="https://zenodo.org/record/12189577/files/figure.png" pageId="4" pageNumber="60">Fig. 2A</figureCitation>
). Phallobase approximately 2.0 times as long as parameres, which is convex apically. Parameres in lateral view elongate and with apex rounded, ventral portion slightly concave, deeply notched basally, and posteroventral portion long.
<emphasis id="A2C8EAE2C248FFCE7D4B820EE1EBDB13" bold="true" box="[304,455,1303,1329]" pageId="4" pageNumber="60">Endophallus</emphasis>
. Fronto-lateral peripheral endophallite, axial endophallite, and subaxial endophallite as in
<figureCitation id="08872A75C248FFCE7CCF8222E124DB77" box="[180,264,1339,1365]" captionStart="FIGURE 2" captionStartId="3.[151,250,1732,1756]" captionTargetBox="[151,1436,192,1708]" captionTargetId="figure-26@3.[151,1436,181,1708]" captionTargetPageId="3" captionText="FIGURE 2. Ontherus diagnostic characters. A, Ontherus araujoi new species, aedeagus; B, Ontherus araujoi, endophallites. C, Ontherus araujoi, mesometasternal suture; D, Ontherus compressicornis Luederwaldt, 1931, mesometasternal suture." figureDoi="http://doi.org/10.5281/zenodo.12189577" httpUri="https://zenodo.org/record/12189577/files/figure.png" pageId="4" pageNumber="60">Fig. 2B</figureCitation>
(see arrows).
</paragraph>
<paragraph id="900336F0C248FFCE7CBC8247E14BDBE3" blockId="4.[151,1437,151,1725]" pageId="4" pageNumber="60">
<emphasis id="A2C8EAE2C248FFCE7CBC8247E194DB5A" bold="true" box="[199,440,1374,1400]" pageId="4" pageNumber="60">Sexual dimorphism.</emphasis>
Females (
<figureCitation id="08872A75C248FFCE7E548246E2A9DB5B" box="[559,645,1375,1401]" captionStart="FIGURE 1" captionStartId="2.[151,250,1846,1870]" captionTargetBox="[162,1425,190,1808]" captionTargetId="figure-17@2.[151,1436,179,1820]" captionTargetPageId="2" captionText="FIGURE 1. Ontherus araujoi new species. A, Ontherus araujoi (male, dorsal view); B, Ontherus araujoi (female, dorsal view); C, Ontherus araujoi (male, ventral view); D. Ontherus araujoi (male, lateral view)." figureDoi="http://doi.org/10.5281/zenodo.12189575" httpUri="https://zenodo.org/record/12189575/files/figure.png" pageId="4" pageNumber="60">Fig. 1B</figureCitation>
) can be distinguished from males by the following characteristics: 1) length less than
<quantity id="57449B15C248FFCE7D37829AE1B0DBBF" box="[332,412,1411,1437]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="4" pageNumber="60" unit="mm" value="11.0">11 mm</quantity>
; 2) apical tooth not forwardly directed; 3) protibial spur present; and 4). metafemur without triangular process.
</paragraph>
</subSubSection>
<subSubSection id="D8A6657BC248FFCE7CBC82D2E46DD89F" pageId="4" pageNumber="60" type="discussion">
<paragraph id="900336F0C248FFCE7CBC82D2E5B6D80F" blockId="4.[151,1437,151,1725]" pageId="4" pageNumber="60">
<emphasis id="A2C8EAE2C248FFCE7CBC82D2E11ADBC7" bold="true" box="[199,310,1483,1509]" pageId="4" pageNumber="60">Remarks.</emphasis>
<taxonomicName id="57BC4D73C248FFCE7D4682D2E1DADBC6" authorityName="Chamorro &amp; Lopera-Toro &amp; Génier" authorityYear="2024" box="[317,502,1483,1509]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="60" phylum="Arthropoda" rank="species" species="araujoi">
<emphasis id="A2C8EAE2C248FFCE7D4682D2E1DADBC6" box="[317,502,1483,1509]" italics="true" pageId="4" pageNumber="60">Ontherus araujoi</emphasis>
</taxonomicName>
is distributed in
<collectingCountry id="E8AB7660C248FFCE7ED582D2E32BDBC7" box="[686,775,1483,1509]" name="Ecuador" pageId="4" pageNumber="60">Ecuador</collectingCountry>
(with the
<typeStatus id="4F078852C248FFCE7F0C82D2E389DBC7" box="[887,933,1483,1509]" pageId="4" pageNumber="60">type</typeStatus>
locality
<quantity id="57449B15C248FFCE787E82D3E465DBC7" box="[1029,1097,1482,1509]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="60" unit="km" value="50.0">50 km</quantity>
north of the border with
<collectingCountry id="E8AB7660C248FFCE792782D2E5BEDBC7" box="[1372,1426,1483,1509]" name="Peru" pageId="4" pageNumber="60">Peru</collectingCountry>
). It inhabits dry montane forest, in the southern Ecuadorian Andes (
<bibRefCitation id="F42D4B01C248FFCE7F1982F6E3C4D82B" author="Sierra, R." box="[866,1000,1519,1545]" pageId="4" pageNumber="60" refId="ref7502" refString="Sierra, R. (1999) Propuesta Preliminar de un Sistema de Clasificacion de Vegetacion para el Ecuador Continental. Proyecto INEFAN / GEF-BIRF y EcoCiencia, Quito, 193 pp." type="book" year="1999">Sierra, 1999</bibRefCitation>
) over
<quantity id="57449B15C248FFCE785482F6E4ADD82B" box="[1071,1153,1519,1545]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="60" unit="m" value="1000.0">1000 m</quantity>
(
<figureCitation id="08872A75C248FFCE78EB82F6E4FFD82B" box="[1168,1235,1519,1545]" captionStart="FIGURE 4" captionStartId="8.[151,250,1889,1913]" captionTargetBox="[161,1426,191,1856]" captionTargetId="figure-17@8.[151,1436,181,1866]" captionTargetPageId="8" captionText="FIGURE 4. Distribution of Ontherus araujoi new species (red pentagon); Ontherus azteca Harold, 1869 (purple square); Ontherus edentulus Génier, 1996 (blue triangle); Ontherus pubens Génier, 1996 (green circle); and Cryptocanthon humidus Howden, 1973 (yellow octagon)." figureDoi="http://doi.org/10.5281/zenodo.12189581" httpUri="https://zenodo.org/record/12189581/files/figure.png" pageId="4" pageNumber="60">Fig. 4</figureCitation>
). According to the biogeographic regionalization proposed by
<bibRefCitation id="F42D4B01C248FFCE7E1D810AE33FD80F" author="Morrone, J. J." box="[614,787,1555,1581]" pageId="4" pageNumber="60" pagination="1 - 110" refId="ref7338" refString="Morrone, J. J. (2014) Biogeographical regionalisation of the Neotropical region. Zootaxa, 3782 (1), 1 - 110. https: // doi. org / 10.11646 / zootaxa. 3782.1.1" type="journal article" year="2014">Morrone (2014)</bibRefCitation>
, this new species occurs in the “Western
<collectingCountry id="E8AB7660C248FFCE78B2810AE50ED80F" box="[1225,1314,1555,1581]" name="Ecuador" pageId="4" pageNumber="60">Ecuador</collectingCountry>
Province”.
</paragraph>
<paragraph id="900336F0C248FFCE7CB5812EE46DD89F" blockId="4.[151,1437,151,1725]" pageId="4" pageNumber="60">
Females similar to the
<taxonomicName id="57BC4D73C248FFCE7DB7812EE2B4D873" baseAuthorityName="Mannerheim" baseAuthorityYear="1828" box="[460,664,1591,1617]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="60" phylum="Arthropoda" rank="species" species="appendiculatus">
<emphasis id="A2C8EAE2C248FFCE7DB7812EE1CCD873" box="[460,480,1591,1617]" italics="true" pageId="4" pageNumber="60">O</emphasis>
.
<emphasis id="A2C8EAE2C248FFCE7D96812EE2B4D873" box="[493,664,1591,1617]" italics="true" pageId="4" pageNumber="60">appendiculatus</emphasis>
</taxonomicName>
species group (see
<bibRefCitation id="F42D4B01C248FFCE7F09812EE42BD873" author="Genier, F." box="[882,1031,1591,1617]" pageId="4" pageNumber="60" pagination="1 - 169" refId="ref6799" refString="Genier, F. (1996) A revision of the Neotropical genus Ontherus Erichson (Coleoptera: Scarabaeidae, Scarabaeinae). Memoirs of the Entomological Society of Canada, 170, 1 - 169. https: // doi. org / 10.4039 / entm 128170 fv" type="journal article" year="1996">Génier, 1996</bibRefCitation>
), but differ in the vertex punctate on all area of the head (with punctures transverse along the posterior surface). However, the shape of the parameres looks similar to the species included in the
<taxonomicName id="57BC4D73C248FFCE7E038166E2F7D8BB" baseAuthorityName="Blanchard" baseAuthorityYear="1846" box="[632,731,1663,1689]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="60" phylum="Arthropoda" rank="species" species="alexis">
<emphasis id="A2C8EAE2C248FFCE7E038166E2A0D8BB" box="[632,652,1663,1689]" italics="true" pageId="4" pageNumber="60">O</emphasis>
.
<emphasis id="A2C8EAE2C248FFCE7EE18166E2F7D8BB" box="[666,731,1663,1689]" italics="true" pageId="4" pageNumber="60">alexis</emphasis>
</taxonomicName>
species group of the subgenus
<taxonomicName id="57BC4D73C248FFCE784C8166E4E2D8BB" box="[1079,1230,1663,1689]" pageId="4" pageNumber="60" rank="subGenus" subGenus="Caelontherus">
<emphasis id="A2C8EAE2C248FFCE784C8166E4E2D8BB" box="[1079,1230,1663,1689]" italics="true" pageId="4" pageNumber="60">Caelontherus</emphasis>
</taxonomicName>
. We will keep this question open for now and resolve the issue when additional specimens are studied.
</paragraph>
</subSubSection>
</treatment>
</document>
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