434 lines
86 KiB
XML
434 lines
86 KiB
XML
<document ID-DOI="10.3897/zookeys.19.221" ID-GBIF-Dataset="e6ea8eea-7156-4785-8313-f4bd88dadad1" ID-ISSN="1313–2970" ID-Zenodo-Dep="576503" ID-ZooBank="C473F9F6-1AE7-4B3F-B17F-CA1C2709010C" checkinTime="1587956031210" checkinUser="plazi" docAuthor="Wesener, Thomas, Enghoff, Henrik & Sierwald, Petra" docDate="2009" docId="1772122E7846FF83FF013C12AB1BE880" docLanguage="en" docName="ZK_article_2037.pdf" docOrigin="ZooKeys 19 (19)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@11b73717" docStyleName="zookeys.2008.journal_article" docTitle="Granitobolus Wesener & Enghoff & Sierwald 2009, gen. n." docType="treatment" docUuid="81CDE7E7-992A-42FD-9FCB-16D88FA8E7B9" docUuidSource="ZooBank" docVersion="7" lastPageNumber="70" masterDocId="EB4B6A567807FFC6FF8D3F74AF59EA51" masterDocTitle="Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)" masterLastPageNumber="128" masterPageNumber="1" pageNumber="66" updateTime="1636630017160" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wesener, Thomas</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A. & Zoologisches Forschungsmuseum Alexander Koenig, Museumsmeile Bonn, Adenauerallee 160, D- 53113 Bonn, Germany. & B 54 D 3623309</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Enghoff, Henrik</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Natural History Museum of Denmark, Universitetsparken 15, DK- 2100 Copenhagen, Denmark</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sierwald, Petra</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A.</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2009</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2009-09-04</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>19</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>19</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>128</mods:end>
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</mods:extent>
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</mods:part>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.3897/zookeys.19.221</mods:identifier>
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<mods:identifier type="GBIF-Dataset">e6ea8eea-7156-4785-8313-f4bd88dadad1</mods:identifier>
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<mods:identifier type="ISSN">1313–2970</mods:identifier>
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<mods:identifier type="Zenodo-Dep">576503</mods:identifier>
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<mods:identifier type="ZooBank">C473F9F6-1AE7-4B3F-B17F-CA1C2709010C</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3791435" ID-GBIF-Taxon="164184220" ID-Zenodo-Dep="3791435" LSID="urn:lsid:zoobank.org:act:81CDE7E7-992A-42FD-9FCB-16D88FA8E7B9" httpUri="http://treatment.plazi.org/id/1772122E7846FF83FF013C12AB1BE880" lastPageId="69" lastPageNumber="70" pageId="65" pageNumber="66">
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<subSubSection box="[140,499,870,897]" pageId="65" pageNumber="66" type="nomenclature">
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<paragraph blockId="65.[140,987,870,932]" box="[140,499,870,897]" pageId="65" pageNumber="66">
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<heading box="[140,499,870,897]" fontSize="11" level="2" pageId="65" pageNumber="66" reason="3">
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<emphasis bold="true" box="[140,499,870,897]" pageId="65" pageNumber="66">
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<taxonomicName authority="Wesener" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[140,406,870,897]" class="Diplopoda" family="Pachybolidae" genus="Granitobolus" kingdom="Animalia" order="Spirobolida" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus" status="gen. n.">
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<emphasis bold="true" box="[140,298,870,897]" italics="true" pageId="65" pageNumber="66">Granitobolus</emphasis>
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Wesener
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</taxonomicName>
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,
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<taxonomicNameLabel box="[418,499,871,897]" pageId="65" pageNumber="66" rank="genus">gen. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection box="[140,987,906,932]" pageId="65" pageNumber="66" type="description">
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<paragraph blockId="65.[140,987,870,932]" box="[140,987,906,932]" pageId="65" pageNumber="66">
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<uri box="[140,987,906,932]" pageId="65" pageNumber="66">
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urn:lsid:zoobank.org:act:
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<uuid box="[421,987,906,932]" pageId="65" pageNumber="66">81CDE7E7-992A-42FD-9FCB-16D88FA8E7B9</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSection lastPageId="69" lastPageNumber="70" pageId="65" pageNumber="66" type="multiple">
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<subSubSection box="[140,628,976,1003]" pageId="65" pageNumber="66" type="type_taxon">
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<paragraph blockId="65.[140,628,976,1003]" box="[140,628,976,1003]" pageId="65" pageNumber="66">
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<emphasis bold="true" box="[140,287,976,1002]" pageId="65" pageNumber="66">
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<typeStatus box="[140,198,976,1002]" pageId="65" pageNumber="66">Type</typeStatus>
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species
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</emphasis>
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:
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<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[302,560,976,1002]" class="Diplopoda" family="Pachybolidae" genus="Granitobolus" kingdom="Animalia" order="Spirobolida" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" status="sp. n.">
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<emphasis box="[302,560,976,1002]" italics="true" pageId="65" pageNumber="66">Granitobolus endemicus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[567,628,977,1003]" pageId="65" pageNumber="66" rank="species">sp. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="65" pageNumber="66" type="reference_group">
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<paragraph blockId="65.[140,419,1046,1108]" box="[140,419,1046,1073]" pageId="65" pageNumber="66">
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<emphasis bold="true" box="[140,413,1046,1072]" pageId="65" pageNumber="66">Other species included</emphasis>
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:
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</paragraph>
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<paragraph blockId="65.[140,419,1046,1108]" box="[140,404,1082,1108]" pageId="65" pageNumber="66">
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<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[140,335,1082,1108]" class="Diplopoda" family="Pachybolidae" genus="Granitobolus" kingdom="Animalia" order="Spirobolida" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" status="sp. n.">
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<emphasis box="[140,335,1082,1108]" italics="true" pageId="65" pageNumber="66">G. andohahelensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[343,404,1082,1108]" pageId="65" pageNumber="66" rank="species">sp. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="65" pageNumber="66" type="diagnosis">
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<paragraph blockId="65.[140,1108,1152,1672]" pageId="65" pageNumber="66">
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<emphasis bold="true" box="[140,259,1152,1178]" inLineHeading="true" pageId="65" pageNumber="66" reason="1">Diagnosis</emphasis>
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: telopodite tip of posterior gonopods in
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<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[727,867,1152,1178]" class="Diplopoda" family="Pachybolidae" genus="Granitobolus" kingdom="Animalia" order="Spirobolida" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus">
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<emphasis box="[727,867,1152,1178]" italics="true" pageId="65" pageNumber="66">Granitobolus</emphasis>
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</taxonomicName>
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uniquely shaped, resembling a duck’s head with an acutely projecting beak (
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<figureCitation box="[775,887,1187,1214]" captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="65" pageNumber="66">Fig. 37M</figureCitation>
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). Adults with 37 or 38 body rings, lowest number of any Malagasy genus of
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<taxonomicName authorityName="Cook" authorityYear="1895" box="[793,927,1223,1249]" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="65" pageNumber="66" phylum="Arthropoda" rank="order">Spirobolida</taxonomicName>
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. Natural colour usually contrasting, with red appendages, dark mesozonites and greyish or light brown metazonites (
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<figureCitation box="[289,373,1293,1319]" captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="65" pageNumber="66">Figs 37</figureCitation>
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A–D, 38A). Shares numerous characters only with species of the genus
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<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[211,404,1328,1354]" class="Diplopoda" family="Pachybolidae" genus="Pseudocentrobolus" kingdom="Animalia" order="Spirobolida" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus">
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<emphasis box="[211,404,1328,1354]" italics="true" pageId="65" pageNumber="66">Pseudocentrobolus</emphasis>
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</taxonomicName>
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: a stout preanal process which extends beyond anal valves (
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<figureCitation captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="65" pageNumber="66">Fig. 37F</figureCitation>
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), well-developed lips on the anal valves, thick moniliform body rings which widely overlap each other reaching the metazonite (
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<figureCitation box="[660,767,1399,1425]" captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="65" pageNumber="66">Fig. 37C</figureCitation>
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), rectangular extensions (
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<figureCitation captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="65" pageNumber="66">Fig. 37F</figureCitation>
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) on the posterior margins of the rings and a greatly enlarged collum, which is ventrally longer than body ring 2. Gnathochilarium with a subdivided mentum and a single sclerotized ledge on each stipites (
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<figureCitation box="[594,702,1504,1530]" captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="65" pageNumber="66">Fig. 37H</figureCitation>
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), like in
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<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[803,941,1504,1530]" class="Diplopoda" family="Pachybolidae" genus="Flagellobolus" kingdom="Animalia" order="Spirobolida" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus">
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<emphasis box="[803,941,1504,1530]" italics="true" pageId="65" pageNumber="66">Flagellobolus</emphasis>
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</taxonomicName>
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,
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<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[956,1099,1504,1530]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus">
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<emphasis box="[956,1099,1504,1530]" italics="true" pageId="65" pageNumber="66">Riotintobolus</emphasis>
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</taxonomicName>
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,
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<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[140,333,1539,1565]" class="Diplopoda" family="Pachybolidae" genus="Pseudocentrobolus" kingdom="Animalia" order="Spirobolida" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus">
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<emphasis box="[140,333,1539,1565]" italics="true" pageId="65" pageNumber="66">Pseudocentrobolus</emphasis>
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</taxonomicName>
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,
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<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[350,472,1539,1565]" class="Diplopoda" family="Pachybolidae" genus="Caprobolus" kingdom="Animalia" order="Spirobolida" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus" status="gen. n.">
|
||
<emphasis box="[350,472,1539,1565]" italics="true" pageId="65" pageNumber="66">Caprobolus</emphasis>
|
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</taxonomicName>
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||
<emphasis bold="true" box="[481,572,1539,1565]" pageId="65" pageNumber="66">
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||
<taxonomicNameLabel box="[481,565,1539,1565]" pageId="65" pageNumber="66" rank="genus">gen. n.</taxonomicNameLabel>
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||
,
|
||
</emphasis>
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||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[580,714,1539,1565]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus" status="gen. n.">
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||
<emphasis box="[580,714,1539,1565]" italics="true" pageId="65" pageNumber="66">Alluviobolus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[723,814,1539,1565]" pageId="65" pageNumber="66">
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<taxonomicNameLabel box="[723,807,1539,1565]" pageId="65" pageNumber="66" rank="genus">gen. n.</taxonomicNameLabel>
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||
,
|
||
</emphasis>
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[823,950,1539,1565]" class="Diplopoda" family="Pachybolidae" genus="Ostinobolus" kingdom="Animalia" order="Spirobolida" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus" status="gen. n.">
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||
<emphasis box="[823,950,1539,1565]" italics="true" pageId="65" pageNumber="66">Ostinobolus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[959,1042,1539,1566]" pageId="65" pageNumber="66" rank="genus">
|
||
<emphasis bold="true" box="[959,1034,1539,1565]" pageId="65" pageNumber="66">gen. n</emphasis>
|
||
.
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||
</taxonomicNameLabel>
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Both genera possess, however, completely different posterior gonopods.
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||
</paragraph>
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||
</subSubSection>
|
||
<subSubSection lastPageId="67" lastPageNumber="68" pageId="65" pageNumber="66" type="distribution">
|
||
<paragraph blockId="65.[140,1108,1152,1672]" lastBlockId="67.[140,1108,167,1672]" lastPageId="67" lastPageNumber="68" pageId="65" pageNumber="66">
|
||
<emphasis bold="true" box="[185,490,1609,1636]" pageId="65" pageNumber="66">
|
||
<emphasis box="[185,336,1609,1635]" inLineHeading="true" pageId="65" pageNumber="66" reason="1">Distribution</emphasis>
|
||
and ecology
|
||
</emphasis>
|
||
: species of
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[624,764,1610,1636]" class="Diplopoda" family="Pachybolidae" genus="Granitobolus" kingdom="Animalia" order="Spirobolida" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[624,764,1610,1636]" italics="true" pageId="65" pageNumber="66">Granitobolus</emphasis>
|
||
</taxonomicName>
|
||
have been recorded from numerous rainforest and littoral rainforest sites in Southeast
|
||
<collectingCountry box="[796,931,1645,1671]" name="Madagascar" pageId="65" pageNumber="66">Madagascar</collectingCountry>
|
||
. Unfortunately, mature males could be collected only from three sites (Andohahela-Vasiha, Manantantely and the Ambatotsirongorongo Mountain,
|
||
<figureCitation box="[663,746,202,228]" captionStart="Figure 30" captionStartId="56.[140,210,1654,1676]" captionTargetBox="[129,1120,1092,1644]" captionTargetId="figure@56.[140,1108,1098,1642]" captionTargetPageId="56" captionText="Figure 30. Distribution of the genera Pseudocentrobolus gen. n., Granitobolus gen. n. and Caprobolus gen.n." figureDoi="http://doi.org/10.5281/zenodo.3820818" httpUri="https://zenodo.org/record/3820818/files/figure.png" pageId="67" pageNumber="68">Fig. 30</figureCitation>
|
||
). At least single immature specimens found in the littoral forest of Mandena and the Sainte Luce fragments S8 and S9 might represent another undescribed species because of their uniformly dark, almost black coloration. The specimens from Isaka-Ivondro (Andohahela), Ebosika (Andohahela) and Ivorona (Anosyenne) closely resemble
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[716,912,343,369]" class="Diplopoda" family="Pachybolidae" genus="Granitobolus" kingdom="Animalia" order="Spirobolida" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis box="[716,912,343,369]" italics="true" pageId="67" pageNumber="68">G. andohahelensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[920,982,343,369]" pageId="67" pageNumber="68" rank="species">sp. n.</taxonomicNameLabel>
|
||
Except for two or three isolated specimens, which were collected on trails, all other specimens of
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[140,280,413,439]" class="Diplopoda" family="Pachybolidae" genus="Granitobolus" kingdom="Animalia" order="Spirobolida" pageId="67" pageNumber="68" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[140,280,413,439]" italics="true" pageId="67" pageNumber="68">Granitobolus</emphasis>
|
||
</taxonomicName>
|
||
were taken from large, massive, partly overgrown granitic outcrops. Specimens were collected in large numbers from thin layers of roots covering the stones. On a single rock, specimens of all sizes and both sexes were found; however, mature males remained rare. This might be related to the beginning of the dry season (June), when the collections were conducted. Both
|
||
<typeStatus box="[645,754,554,580]" pageId="67" pageNumber="68">holotypes</typeStatus>
|
||
were infested with rather long nematodes (P,
|
||
<figureCitation box="[305,408,589,615]" captionStart="Figure 38" captionStartId="68.[140,212,1494,1516]" captionTargetBox="[140,1108,821,1473]" captionTargetId="figure@68.[140,1108,1087,1476]" captionTargetPageId="68" captionText="Figure 38. Granitobolus andohahelensis sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view; E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; Md = mandible; ms = marginal setae; P = parasitic nematode?; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767324" httpUri="https://zenodo.org/record/3767324/files/figure.png" pageId="67" pageNumber="68">Fig. 38B</figureCitation>
|
||
). The legs and body rings of
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[741,888,589,616]" class="Diplopoda" family="Pachybolidae" genus="Granitobolus" kingdom="Animalia" order="Spirobolida" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis box="[741,888,589,616]" italics="true" pageId="67" pageNumber="68">G. endemicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[896,958,590,616]" pageId="67" pageNumber="68" rank="species">sp. n.</taxonomicNameLabel>
|
||
<typeStatus box="[967,1066,589,615]" pageId="67" pageNumber="68">holotype</typeStatus>
|
||
are also covered with fungi.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3767322" ID-Zenodo-Dep="3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="66" pageNumber="67" startId="66.[142,214,1430,1452]" targetBox="[140,1109,166,1342]" targetPageId="66">
|
||
<paragraph blockId="66.[142,1110,1430,1676]" pageId="66" pageNumber="67">
|
||
<emphasis bold="true" box="[142,251,1430,1452]" pageId="66" pageNumber="67">Figure 37.</emphasis>
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[258,469,1430,1451]" class="Diplopoda" family="Pachybolidae" genus="Granitobolus" kingdom="Animalia" order="Spirobolida" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis box="[258,469,1430,1451]" italics="true" pageId="66" pageNumber="67">Granitobolus endemicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[476,528,1431,1452]" pageId="66" pageNumber="67" rank="species">sp. n.</taxonomicNameLabel>
|
||
, A–C, G–J male paratype, K female paratype, D–F, L–O male holotype.
|
||
<emphasis bold="true" box="[236,255,1462,1484]" pageId="66" pageNumber="67">A</emphasis>
|
||
living specimen, uncurling
|
||
<emphasis bold="true" box="[520,537,1462,1484]" pageId="66" pageNumber="67">B</emphasis>
|
||
living specimen, curled
|
||
<emphasis bold="true" box="[769,788,1462,1484]" pageId="66" pageNumber="67">C</emphasis>
|
||
dead male, curled
|
||
<emphasis bold="true" box="[969,988,1462,1484]" pageId="66" pageNumber="67">D</emphasis>
|
||
head, lateral view
|
||
<emphasis bold="true" box="[190,205,1494,1516]" pageId="66" pageNumber="67">E</emphasis>
|
||
legs 1–7, ventral view
|
||
<emphasis bold="true" box="[417,431,1494,1516]" pageId="66" pageNumber="67">F</emphasis>
|
||
telson
|
||
<emphasis bold="true" box="[498,518,1494,1516]" pageId="66" pageNumber="67">G</emphasis>
|
||
head, frontal view
|
||
<emphasis bold="true" box="[696,716,1494,1516]" pageId="66" pageNumber="67">H</emphasis>
|
||
gnathochilarium, ventral view;
|
||
<emphasis bold="true" box="[1011,1019,1494,1516]" pageId="66" pageNumber="67">I</emphasis>
|
||
midbody ring with legs
|
||
<emphasis bold="true" box="[271,306,1526,1548]" pageId="66" pageNumber="67">Į, K</emphasis>
|
||
left midbody legs
|
||
<emphasis bold="true" box="[475,490,1526,1548]" pageId="66" pageNumber="67">L</emphasis>
|
||
anterior gonopod, anterior view
|
||
<emphasis bold="true" box="[793,814,1526,1548]" pageId="66" pageNumber="67">M</emphasis>
|
||
left posterior gonopod, anterior view
|
||
<emphasis bold="true" box="[189,209,1558,1580]" pageId="66" pageNumber="67">N</emphasis>
|
||
anterior gonopod, posterior view
|
||
<emphasis bold="true" box="[525,546,1558,1580]" pageId="66" pageNumber="67">O</emphasis>
|
||
left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection lastPageId="69" lastPageNumber="70" pageId="67" pageNumber="68" type="description">
|
||
<paragraph blockId="67.[140,1108,167,1672]" pageId="67" pageNumber="68">
|
||
<emphasis bold="true" box="[185,335,659,685]" pageId="67" pageNumber="68">
|
||
<emphasis box="[185,330,659,685]" inLineHeading="true" pageId="67" pageNumber="68" reason="1">Description</emphasis>
|
||
.
|
||
</emphasis>
|
||
<emphasis box="[346,408,659,685]" italics="true" pageId="67" pageNumber="68">Males</emphasis>
|
||
: length up to
|
||
<quantity box="[579,664,659,686]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="67" pageNumber="68" unit="mm" value="35.0">35 mm</quantity>
|
||
(broken), diameter up to
|
||
<quantity box="[969,1064,660,686]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="67" pageNumber="68" unit="mm" value="3.3">3.3 mm</quantity>
|
||
. 37 or 38 podous rings. Females: length up to
|
||
<quantity box="[617,703,695,721]" metricMagnitude="-2" metricUnit="m" metricValue="4.8" pageId="67" pageNumber="68" unit="mm" value="48.0">48 mm</quantity>
|
||
, diameter up to
|
||
<quantity box="[890,997,694,721]" metricMagnitude="-3" metricUnit="m" metricValue="4.85" pageId="67" pageNumber="68" unit="mm" value="4.85">4.85 mm</quantity>
|
||
. 37 or 38 podous rings. Length-width ratio in both sexes 1:10. Apodous rings absent in all adult specimens.
|
||
</paragraph>
|
||
<paragraph blockId="67.[140,1108,167,1672]" pageId="67" pageNumber="68">
|
||
<emphasis box="[185,261,800,826]" italics="true" pageId="67" pageNumber="68">Colour</emphasis>
|
||
: mesozona dark olive green or blackish (
|
||
<figureCitation box="[728,834,800,826]" captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="67" pageNumber="68">Fig. 37A</figureCitation>
|
||
), metazona much light- er, brown to yellow (
|
||
<figureCitation box="[384,493,835,862]" captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="67" pageNumber="68">Fig. 37C</figureCitation>
|
||
). Head, antennae and legs red (faded to light brown in some specimens).
|
||
</paragraph>
|
||
<paragraph blockId="67.[140,1108,167,1672]" pageId="67" pageNumber="68">
|
||
<emphasis box="[185,244,906,932]" italics="true" pageId="67" pageNumber="68">Head</emphasis>
|
||
: each eye large, with circa 35–40 ocelli arranged in 6 or 7 vertical rows (
|
||
<figureCitation captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="67" pageNumber="68">Figs 37D, G</figureCitation>
|
||
). Labrum with standard three irregular teeth and one row of 10–12 stout marginal setae (
|
||
<figureCitation box="[275,380,976,1002]" captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="67" pageNumber="68">Fig. 37E</figureCitation>
|
||
). Clypeus with two setiferous foveolae on each side (
|
||
<figureCitation box="[990,1095,976,1002]" captionStart="Figure 38" captionStartId="68.[140,212,1494,1516]" captionTargetBox="[140,1108,821,1473]" captionTargetId="figure@68.[140,1108,1087,1476]" captionTargetPageId="68" captionText="Figure 38. Granitobolus andohahelensis sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view; E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; Md = mandible; ms = marginal setae; P = parasitic nematode?; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767324" httpUri="https://zenodo.org/record/3767324/files/figure.png" pageId="67" pageNumber="68">Fig. 38B</figureCitation>
|
||
). Length of antennae sexually dimorph, protruding back to ring 2 (females) or 4 (males,
|
||
<figureCitation box="[140,281,1047,1073]" captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="67" pageNumber="68">Figs 37A, D</figureCitation>
|
||
,
|
||
<figureCitation box="[294,346,1047,1073]" captionStart="Figure 38" captionStartId="68.[140,212,1494,1516]" captionTargetBox="[140,1108,821,1473]" captionTargetId="figure@68.[140,1108,1087,1476]" captionTargetPageId="68" captionText="Figure 38. Granitobolus andohahelensis sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view; E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; Md = mandible; ms = marginal setae; P = parasitic nematode?; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767324" httpUri="https://zenodo.org/record/3767324/files/figure.png" pageId="67" pageNumber="68">38A</figureCitation>
|
||
). Relative lengths of antennomeres: 1<2=3=4=5<6 (
|
||
<figureCitation box="[934,1042,1047,1073]" captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="67" pageNumber="68">Fig. 37G</figureCitation>
|
||
). Terminal antennomere with four large sensory cones located together inside a membranous area. Antennomere 5 latero-apically with three or four rows, antennomere 6 with two rows of sensilla basiconica.
|
||
</paragraph>
|
||
<paragraph blockId="67.[140,1108,167,1672]" pageId="67" pageNumber="68">
|
||
<emphasis box="[185,373,1187,1213]" italics="true" pageId="67" pageNumber="68">Gnathochilarium</emphasis>
|
||
unusual (
|
||
<figureCitation box="[489,599,1187,1214]" captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="67" pageNumber="68">Fig. 37H</figureCitation>
|
||
). Lamellae linguales each with two standard setae located behind one another. Stipites each with three apical setae. Mentum basally subdivided by a well-visible suture (
|
||
<figureCitation box="[560,671,1258,1284]" captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="67" pageNumber="68">Fig. 37H</figureCitation>
|
||
). Stipites each towards mentum with a wide sclerotized ledge (
|
||
<figureCitation box="[430,540,1293,1319]" captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="67" pageNumber="68">Fig. 37H</figureCitation>
|
||
). Hypopharyngeal crest with a field of spine-like structures. Central pads of endochilarium separated by a step into two levels. Medial palps with a large group of sensilla each, located towards hypopharyngeal crest.
|
||
</paragraph>
|
||
<paragraph blockId="67.[140,1108,167,1672]" pageId="67" pageNumber="68">
|
||
<emphasis box="[185,288,1399,1425]" italics="true" pageId="67" pageNumber="68">Mandible</emphasis>
|
||
: external tooth simple, rounded; mesal tooth with three cusps. Four pectinate lamellae. Molar plate with numerous (10 or 11) small and shallow transverse furrows.
|
||
</paragraph>
|
||
<paragraph blockId="67.[140,1108,167,1672]" pageId="67" pageNumber="68">
|
||
<emphasis box="[185,274,1469,1495]" italics="true" pageId="67" pageNumber="68">Collum:</emphasis>
|
||
smooth, laterally longer than body ring 2. Lobe enlarged, covering antenna and part of head (
|
||
<figureCitation box="[411,520,1504,1530]" captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="67" pageNumber="68">Figs 37D</figureCitation>
|
||
,
|
||
<figureCitation box="[532,585,1504,1530]" captionStart="Figure 38" captionStartId="68.[140,212,1494,1516]" captionTargetBox="[140,1108,821,1473]" captionTargetId="figure@68.[140,1108,1087,1476]" captionTargetPageId="68" captionText="Figure 38. Granitobolus andohahelensis sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view; E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; Md = mandible; ms = marginal setae; P = parasitic nematode?; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767324" httpUri="https://zenodo.org/record/3767324/files/figure.png" pageId="67" pageNumber="68">38A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="67.[140,1108,167,1672]" pageId="67" pageNumber="68">
|
||
<emphasis box="[185,305,1539,1565]" italics="true" pageId="67" pageNumber="68">Body rings:</emphasis>
|
||
moniliform, overlapping up to anterior half of mesozona. Metazona of swollen appearance, width larger than pro- or mesozona (
|
||
<figureCitation box="[785,868,1575,1601]" captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="67" pageNumber="68">Figs 37</figureCitation>
|
||
A–C). Posterior margin of rings slightly excavated into rectangular extensions (
|
||
<figureCitation box="[790,889,1610,1636]" captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="67" pageNumber="68">Fig. 37F</figureCitation>
|
||
). Ozopores starting at ring 6, touching suture between mesozona and metazona (
|
||
<figureCitation box="[827,935,1645,1671]" captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="67" pageNumber="68">Fig. 37D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="68.[140,1108,167,756]" pageId="68" pageNumber="69">
|
||
<emphasis box="[185,251,167,193]" italics="true" pageId="68" pageNumber="69">Telson</emphasis>
|
||
: anal valves with well-developed lips and micropunctation, but with neither grooves, nor setae. Preanal process well-rounded or sharp, extending beyond anal valves (
|
||
<figureCitation box="[220,318,237,263]" captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="68" pageNumber="69">Figs 37F</figureCitation>
|
||
,
|
||
<figureCitation box="[330,383,237,263]" captionStart="Figure 38" captionStartId="68.[140,212,1494,1516]" captionTargetBox="[140,1108,821,1473]" captionTargetId="figure@68.[140,1108,1087,1476]" captionTargetPageId="68" captionText="Figure 38. Granitobolus andohahelensis sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view; E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; Md = mandible; ms = marginal setae; P = parasitic nematode?; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767324" httpUri="https://zenodo.org/record/3767324/files/figure.png" pageId="68" pageNumber="69">38C</figureCitation>
|
||
). Subanal scale inconspicuous.
|
||
</paragraph>
|
||
<paragraph blockId="68.[140,1108,167,756]" pageId="68" pageNumber="69">
|
||
<emphasis box="[185,230,273,298]" italics="true" pageId="68" pageNumber="69">Legs</emphasis>
|
||
: coxae 1 and 2 elongated, fused with sternum, podomeres from prefemur to tarsus in both sexes each with 4–10 ventral/mesal setae. Length of midbody legs circa 1.0 times body diameter in males (
|
||
<figureCitation box="[534,628,343,369]" captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="68" pageNumber="69">Fig. 37J</figureCitation>
|
||
), 0.6 times body diameter in females (
|
||
<figureCitation captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="68" pageNumber="69">Fig. 37K</figureCitation>
|
||
). Each podomere with an apical, ventral seta (
|
||
<figureCitation box="[717,822,378,404]" captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="68" pageNumber="69">Fig. 37K</figureCitation>
|
||
). Coxae 3 and beyond of rectangular shape (
|
||
<figureCitation box="[355,451,413,439]" captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="68" pageNumber="69">Fig. 37I</figureCitation>
|
||
). Tarsus with a stout dorso-apical seta and a pair of stout ventro-apical ones in males, three pairs of setae in females. In females apical setae pair stout, others successively much smaller.
|
||
</paragraph>
|
||
<paragraph blockId="68.[140,1108,167,756]" pageId="68" pageNumber="69">
|
||
<emphasis box="[185,429,519,545]" italics="true" pageId="68" pageNumber="69">Male sexual characters</emphasis>
|
||
: tarsi starting at leg 3 up to midbody legs wide and with a large tarsal pad not protruding past base of claw (
|
||
<figureCitation box="[712,808,554,580]" captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="68" pageNumber="69">Fig. 37J</figureCitation>
|
||
). Tibia shorter than in females (
|
||
<figureCitation box="[218,350,589,616]" captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="68" pageNumber="69">Figs 37J, K</figureCitation>
|
||
). Coxae 3–7 unmodified (
|
||
<figureCitation box="[646,749,589,615]" captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="68" pageNumber="69">Figs 37E</figureCitation>
|
||
,
|
||
<figureCitation box="[761,813,589,615]" captionStart="Figure 38" captionStartId="68.[140,212,1494,1516]" captionTargetBox="[140,1108,821,1473]" captionTargetId="figure@68.[140,1108,1087,1476]" captionTargetPageId="68" captionText="Figure 38. Granitobolus andohahelensis sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view; E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; Md = mandible; ms = marginal setae; P = parasitic nematode?; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767324" httpUri="https://zenodo.org/record/3767324/files/figure.png" pageId="68" pageNumber="69">38B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="68.[140,1108,167,756]" lastBlockId="69.[140,1108,167,721]" lastPageId="69" lastPageNumber="70" pageId="68" pageNumber="69">
|
||
<emphasis box="[185,375,624,651]" italics="true" pageId="68" pageNumber="69">Anterior gonopods</emphasis>
|
||
: median sternal projection triangular, with a wide, rectangular tip (
|
||
<figureCitation box="[147,251,659,686]" captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="68" pageNumber="69">Figs 37L</figureCitation>
|
||
,
|
||
<figureCitation box="[266,321,660,686]" captionStart="Figure 38" captionStartId="68.[140,212,1494,1516]" captionTargetBox="[140,1108,821,1473]" captionTargetId="figure@68.[140,1108,1087,1476]" captionTargetPageId="68" captionText="Figure 38. Granitobolus andohahelensis sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view; E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; Md = mandible; ms = marginal setae; P = parasitic nematode?; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767324" httpUri="https://zenodo.org/record/3767324/files/figure.png" pageId="68" pageNumber="69">38D</figureCitation>
|
||
). Sternite never protruding as high as coxite. Coxite wide, smoothly rounded, laterally curving around telopodite, but only extending slightly at posterior side (
|
||
<figureCitation box="[199,307,730,756]" captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="68" pageNumber="69">Figs 37N</figureCitation>
|
||
,
|
||
<figureCitation box="[322,370,730,756]" captionStart="Figure 38" captionStartId="68.[140,212,1494,1516]" captionTargetBox="[140,1108,821,1473]" captionTargetId="figure@68.[140,1108,1087,1476]" captionTargetPageId="68" captionText="Figure 38. Granitobolus andohahelensis sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view; E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; Md = mandible; ms = marginal setae; P = parasitic nematode?; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767324" httpUri="https://zenodo.org/record/3767324/files/figure.png" pageId="68" pageNumber="69">38F</figureCitation>
|
||
). Coxite anteriorly almost completely covering telopodite. Coxite processes absent, but mesal margin towards sternite sometimes projecting (
|
||
<figureCitation box="[956,1054,167,193]" captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="69" pageNumber="70">Fig. 37L</figureCitation>
|
||
). Telopodite with a short, well-rounded, laterally curved process, which only slightly projects above telopodite margin and never protrudes above coxite margin (
|
||
<figureCitation box="[877,983,237,263]" captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="69" pageNumber="70">Figs 37N</figureCitation>
|
||
,
|
||
<figureCitation box="[995,1044,237,263]" captionStart="Figure 38" captionStartId="68.[140,212,1494,1516]" captionTargetBox="[140,1108,821,1473]" captionTargetId="figure@68.[140,1108,1087,1476]" captionTargetPageId="68" captionText="Figure 38. Granitobolus andohahelensis sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view; E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; Md = mandible; ms = marginal setae; P = parasitic nematode?; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767324" httpUri="https://zenodo.org/record/3767324/files/figure.png" pageId="69" pageNumber="70">38F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3767324" ID-Zenodo-Dep="3767324" httpUri="https://zenodo.org/record/3767324/files/figure.png" pageId="68" pageNumber="69" startId="68.[140,212,1494,1516]" targetBox="[140,1108,821,1473]" targetPageId="68">
|
||
<paragraph blockId="68.[140,1108,1494,1676]" pageId="68" pageNumber="69">
|
||
<emphasis bold="true" box="[140,253,1494,1516]" pageId="68" pageNumber="69">Figure 38.</emphasis>
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[260,514,1494,1515]" class="Diplopoda" family="Pachybolidae" genus="Granitobolus" kingdom="Animalia" order="Spirobolida" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis box="[260,514,1494,1515]" italics="true" pageId="68" pageNumber="69">Granitobolus andohahelensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[521,574,1495,1516]" pageId="68" pageNumber="69" rank="species">sp. n.</taxonomicNameLabel>
|
||
, male holotype.
|
||
<emphasis bold="true" box="[732,751,1494,1516]" pageId="68" pageNumber="69">A</emphasis>
|
||
head, lateral view
|
||
<emphasis bold="true" box="[929,946,1494,1516]" pageId="68" pageNumber="69">B</emphasis>
|
||
legs 1–7, ventral view
|
||
<emphasis bold="true" box="[191,210,1526,1548]" pageId="68" pageNumber="69">C</emphasis>
|
||
telson
|
||
<emphasis bold="true" box="[281,300,1526,1548]" pageId="68" pageNumber="69">D</emphasis>
|
||
anterior gonopod, anterior view;
|
||
<emphasis bold="true" box="[627,642,1526,1548]" pageId="68" pageNumber="69">E</emphasis>
|
||
left posterior gonopod, anterior view
|
||
<emphasis bold="true" box="[1011,1025,1526,1548]" pageId="68" pageNumber="69">F</emphasis>
|
||
anterior gonopod, posterior view
|
||
<emphasis bold="true" box="[375,395,1558,1580]" pageId="68" pageNumber="69">G</emphasis>
|
||
left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; Md = mandible; ms = marginal setae; P = parasitic nematode?; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; T = telopodite. Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="69.[140,1108,167,721]" pageId="69" pageNumber="70">
|
||
<emphasis box="[185,460,272,298]" italics="true" pageId="69" pageNumber="70">Posterior gonopods in situ</emphasis>
|
||
almost completely covered by anterior gonopods. Suture between coxite and telopodite well-visible (
|
||
<figureCitation box="[615,719,307,334]" captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="69" pageNumber="70">Fig. 37O</figureCitation>
|
||
), sternite triangular and sclerotized but often hidden by a large, finger-shaped coxite process (
|
||
<figureCitation box="[815,925,343,369]" captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="69" pageNumber="70">Fig. 37O</figureCitation>
|
||
). Finger-shaped process not curved, running parallel to basal margin (
|
||
<figureCitation box="[770,883,378,404]" captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="69" pageNumber="70">Figs 37O</figureCitation>
|
||
,
|
||
<figureCitation box="[899,955,378,404]" captionStart="Figure 38" captionStartId="68.[140,212,1494,1516]" captionTargetBox="[140,1108,821,1473]" captionTargetId="figure@68.[140,1108,1087,1476]" captionTargetPageId="68" captionText="Figure 38. Granitobolus andohahelensis sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view; E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; Md = mandible; ms = marginal setae; P = parasitic nematode?; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767324" httpUri="https://zenodo.org/record/3767324/files/figure.png" pageId="69" pageNumber="70">38G</figureCitation>
|
||
). Telopodites arranged face-to-face with one another (
|
||
<figureCitation box="[606,720,413,439]" captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="69" pageNumber="70">Fig. 37M</figureCitation>
|
||
). Shape of apical part resembling ‘beak’ (
|
||
<figureCitation box="[219,332,448,474]" captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="69" pageNumber="70">Figs 37M</figureCitation>
|
||
,
|
||
<figureCitation box="[344,395,448,474]" captionStart="Figure 38" captionStartId="68.[140,212,1494,1516]" captionTargetBox="[140,1108,821,1473]" captionTargetId="figure@68.[140,1108,1087,1476]" captionTargetPageId="68" captionText="Figure 38. Granitobolus andohahelensis sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view; E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; Md = mandible; ms = marginal setae; P = parasitic nematode?; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767324" httpUri="https://zenodo.org/record/3767324/files/figure.png" pageId="69" pageNumber="70">38E</figureCitation>
|
||
). Beak acutely projecting, partly sclerotized and carrying sperm canal (
|
||
<figureCitation box="[216,325,483,510]" captionStart="Figure 38" captionStartId="68.[140,212,1494,1516]" captionTargetBox="[140,1108,821,1473]" captionTargetId="figure@68.[140,1108,1087,1476]" captionTargetPageId="68" captionText="Figure 38. Granitobolus andohahelensis sp. n., male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view; E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; Md = mandible; ms = marginal setae; P = parasitic nematode?; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767324" httpUri="https://zenodo.org/record/3767324/files/figure.png" pageId="69" pageNumber="70">Fig. 38G</figureCitation>
|
||
). Telopodite between apex and beak with a small membranous area (
|
||
<figureCitation box="[148,259,519,545]" captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="69" pageNumber="70">Fig. 37M</figureCitation>
|
||
), in basal half mesally of sperm canal with a lower elevated area (
|
||
<figureCitation box="[984,1095,519,545]" captionStart="Figure 37" captionStartId="66.[142,214,1430,1452]" captionTargetBox="[140,1109,166,1342]" captionTargetId="figure@66.[140,1110,165,965]" captionTargetPageId="66" captionText="Figure 37. Granitobolus endemicus sp. n., A–C, G–J male paratype, K female paratype, D–F, L–O male holotype. A living specimen, uncurling B living specimen, curled C dead male, curled D head, lateral view E legs 1–7, ventral view F telson G head, frontal view H gnathochilarium, ventral view; I midbody ring with legs Į, K left midbody legs L anterior gonopod, anterior view M left posterior gonopod, anterior view N anterior gonopod, posterior view O left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; fs-p = finger-shaped process; Gn = gnathochilarium; L = ledge; Md = mandible; Me = mentum; ms = marginal setae; Pre = preanal ring; re = rectangular extension; sf = setiferous foveolae; St = sternite; ST = stipites; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767322" httpUri="https://zenodo.org/record/3767322/files/figure.png" pageId="69" pageNumber="70">Fig. 37M</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="69.[140,1108,167,721]" pageId="69" pageNumber="70">
|
||
<emphasis box="[185,448,554,580]" italics="true" pageId="69" pageNumber="70">Female sexual characters:</emphasis>
|
||
vulva simple, with a small, poorly sclerotized operculum at base, bivalve-like. Posterior valve apically slightly overlapping anterior one. Both valves smooth, sensory cones absent. Each valve basally towards opening with two rows of setae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="69" pageNumber="70" type="etymology">
|
||
<paragraph blockId="69.[140,1108,167,721]" pageId="69" pageNumber="70">
|
||
<emphasis bold="true" box="[185,314,659,685]" inLineHeading="true" pageId="69" pageNumber="70" reason="1">Etymology</emphasis>
|
||
:
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[328,468,659,685]" class="Diplopoda" family="Pachybolidae" genus="Granitobolus" kingdom="Animalia" order="Spirobolida" pageId="69" pageNumber="70" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[328,468,659,685]" italics="true" pageId="69" pageNumber="70">Granitobolus</emphasis>
|
||
</taxonomicName>
|
||
, masculine, after Granit (German for granite) for the special habitat, granitic outcrops, on which species of this genus were found, and
|
||
<emphasis box="[1019,1083,695,721]" italics="true" pageId="69" pageNumber="70">-bolus</emphasis>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</subSection>
|
||
</treatment>
|
||
</document> |